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<document ID-CLB-Dataset="72036" ID-DOI="http://dx.doi.org/10.3897/dez.69.83840" ID-GBIF-Dataset="eafc7326-93e2-4d55-b708-2da4b77e2e5e" ID-Pensoft-Pub="1860-1324-1-71" ID-Pensoft-UUID="DF59A16543B05799A4FD7B66E721FC76" ID-ZooBank="7DEB0587A3734B06BAADCA6102EC6581" ModsDocID="1860-1324-69-1-71" checkinTime="1655205803791" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan &amp; Machard, Patrice" docDate="2022" docId="D3B37B2A59DE540E8A18075F47E7AB0A" docLanguage="en" docName="DeutEntomolZeit 69(1): 71-122" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-06-13" docSource="http://dx.doi.org/10.3897/dez.69.83840" docTitle="Platyderus (Eremoderus) iranicus Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard 2022, sp. nov." docType="treatment" docUuid="3C731C6C-346E-4E8A-B5F3-D2D0B3E27D97" docUuidSource="ZooBank" docVersion="6" id="DF59A16543B05799A4FD7B66E721FC76" lastPageNumber="71" masterDocId="DF59A16543B05799A4FD7B66E721FC76" masterDocTitle="Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)" masterLastPageNumber="122" masterPageNumber="71" pageNumber="71" updateTime="1732679132351" updateUser="ExternalLinkService">
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<mods:title id="2F45CC83ECA3D7140C8A7E27ABDFE57B">Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)</mods:title>
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<mods:namePart id="26CD3A846C315ACA509764C84C3D62AB">Gueorguiev, Borislav</mods:namePart>
<mods:nameIdentifier id="87F415C4D2600245FB9F9FE531BAFB2C" type="ORCID">https://orcid.org/0000-0002-8532-0657</mods:nameIdentifier>
<mods:affiliation id="69D98706FE53B9EF7CD2A3B1270B1B3F">National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
<mods:nameIdentifier id="FC55283349B7D013601687996EEB58BC" type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
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<mods:namePart id="B5D8F9146A3A99D1CA728ABF09A181E6">Wrase, David W.</mods:namePart>
<mods:nameIdentifier id="B5B9DE86FB04392B9F0724CF7AD6881C" type="ORCID">https://orcid.org/0000-0003-1779-2684</mods:nameIdentifier>
<mods:affiliation id="5539CB8B368E924678E7B6C1D569FD63">Oderstr. 2, 15306, Gusow-Platkow, Germany</mods:affiliation>
</mods:name>
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<mods:namePart id="401CBA9928FAB2E69A5A366133945086">Assmann, Thorsten</mods:namePart>
<mods:nameIdentifier id="3A71D9418C9DFB0742AA53FBE880742C" type="ORCID">https://orcid.org/0000-0002-9203-769X</mods:nameIdentifier>
<mods:affiliation id="41F2B7BFEC55B1DE3A5F1721EDC54103">Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
</mods:name>
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<mods:namePart id="B70805EC43849567E02741502C9BE844">Muilwijk, Jan</mods:namePart>
<mods:affiliation id="44588618FC80B8308D1C3F04B2F500EE">Department of Entomology, Naturalis Biodiversity Centre, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
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<mods:namePart id="2C0A189A1482C446EABEF9996B69B17B">Machard, Patrice</mods:namePart>
<mods:affiliation id="DE6EE475D79F60F79A50F59C95993FBB">29 Chemin de Champigny, Molineuf, F- 41190 Valencisse, France</mods:affiliation>
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<treatment id="D3B37B2A59DE540E8A18075F47E7AB0A" ID-GBIF-Taxon="195868742" LSID="urn:lsid:zoobank.org:act:3C731C6C-346E-4E8A-B5F3-D2D0B3E27D97" httpUri="http://treatment.plazi.org/id/D3B37B2A59DE540E8A18075F47E7AB0A" lastPageNumber="71" pageId="0" pageNumber="71" scope_family="Carabidae" scope_order="Coleoptera" scope_subGenus="Platyderus" scope_tribe="Sphodrini">
<subSubSection id="DB162637846135D0CCE89906E999B546" pageId="0" pageNumber="71" type="nomenclature">
<paragraph id="ADA718A83024B8E0BFAE6175AC91477B" pageId="0" pageNumber="71">
4.
<taxonomicName id="0233931A19EA45971E8FE1D552F55EDF" LSID="http://zoobank.org/3C731C6C-346E-4E8A-B5F3-D2D0B3E27D97" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) iranicus</taxonomicName>
<taxonomicNameLabel id="E1EA6D87C016ECFE8ADB71597A5CAEF5" pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="2633F1EFF0B1410DD140C0F4E6CA28B4" pageId="0" pageNumber="71" type="description">
<paragraph id="B55C46A5FD2E99750B3731210FA9E3D5" pageId="0" pageNumber="71">
<figureCitation id="06D2050E404806BEAC4A3A623494BD4A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">Figs 1D</figureCitation>
<figureCitation id="5AB27AE93E02DBD2B141C2DEDD8E6445" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">, 5C</figureCitation>
<figureCitation id="FE27E7B42E3DEDF29A50815039C3A597" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">, 8D</figureCitation>
<figureCitation id="EAA501A9224713751E2D16FDCEDF72BA" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">, 10D</figureCitation>
<figureCitation id="D1956C969863AFC2AF5F25C3EDE06629" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">, 12D</figureCitation>
<figureCitation id="B5A582E372E4EB45D7FA00C1507D2F44" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">, 13C</figureCitation>
<figureCitation id="6AE0B3F7646B0536E7FC6C626AE9D301" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">, 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BB775C2BE6894BC89590234051E2541E" pageId="0" pageNumber="71" type="type locality">
<paragraph id="2C108ECE6F629568FFDB6D4D682D479A" pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph id="FFE426D7E201EC233E32772E81F0D4AC" pageId="0" pageNumber="71">Iran, Chahar Mahal va Bakhtiari Province, 7 km NE Naghan Town, 31.97472, 50.77694, 2400 m.</paragraph>
</subSubSection>
<subSubSection id="2A30974A247E7A6646C63FD0CF92A197" pageId="0" pageNumber="71" type="type material">
<paragraph id="4F9D60705E36AEA98131E28BAD0A38F5" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="D7A7C3669F9BF9B4CA65BC5D65C8855D" pageId="0" pageNumber="71">
<emphasis id="533F49A49C6F819308F86B2418A31B90" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
1♂, 'IRAN, /
<normalizedToken id="0010CB94F2F601CA6C9A6A662587CDAB" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="D418D8A1DB8ED17C01C54817A05D0470" originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken id="0977CC27676AE17B806558B9995290CE" originalValue="vā">va</normalizedToken>
<normalizedToken id="BC262F18498745B82B0791F9629EB212" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
/ 7km NE Naghan Town /
<geoCoordinate id="9C041529D174B09765C13704FF835CA9" degrees="31" direction="north" minutes="58" orientation="latitude" precision="15" seconds="29" value="31.974722">31°58'29&quot;N</geoCoordinate>
,
<geoCoordinate id="509770EB677F655A6F52ED5D6FA06C28" degrees="050" direction="east" minutes="46" orientation="longitude" precision="15" seconds="37" value="50.776943">050°46'37&quot;E</geoCoordinate>
2400m / 03.04.2007, leg. Jan
<normalizedToken id="2474ADB2B8FE3C8CD9F6DBC913C5678F" originalValue="Muilwijk’">Muilwijk'</normalizedToken>
(HMIM). Paratypes: 1♀, labelled as holotype (cMUI); 1♀, 'IRAN (
<normalizedToken id="CD30CE6B3ED58C23B43AAA768D90EA9A" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="050CA976D6C145183E01B06879C35EF8" originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken id="AF95B8FD5C68D5F26FFAF587719AD7D5" originalValue="vā">va</normalizedToken>
<normalizedToken id="562FF6397AE36999AC5266369944BF21" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
) / Zagros Mts. / Boldaghi vill. (nr Choghaklor lake) / 3 km S Sibak, 2600-2700 m /
<geoCoordinate id="B252EA66EDEDD4D4110D053E462EE42F" degrees="31" direction="north" minutes="51" orientation="latitude" precision="15" seconds="47" value="31.863056">31°51'47&quot;N</geoCoordinate>
,
<geoCoordinate id="43F34B5180F5AD8A93C4279072E04741" degrees="50" direction="east" minutes="55" orientation="longitude" precision="15" seconds="24" value="50.923336">50°55'24&quot; E</geoCoordinate>
/ (subalpine slopes with snowfields / under stones) / 20.IV.2018 Wrase &amp; Laser [07]' (cWR); 1♂, 'IRAN (
<normalizedToken id="F0CE8E171F2DE187295407D9294F4954" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="9038CE8BFDE4DE446DCADD8ADEA17CA7" originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken id="FAD98227876FECB971045C40134704C9" originalValue="vā">va</normalizedToken>
<normalizedToken id="8050C7000E13F6CA06E52A0E4360EA09" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
) / Zagros Mts., Zard Koh Mt. / Cheri pass 20 km W Samsami / 2775 m,
<geoCoordinate id="830706224DA72CF0C0D3DB51E99C294C" degrees="32" direction="north" minutes="09" orientation="latitude" precision="15" seconds="55" value="32.16528">32°09'55&quot;N</geoCoordinate>
/
<geoCoordinate id="0708910A3E494856F2ACB0DD0E407F00" degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="37" value="50.176945">50°10'37&quot;E</geoCoordinate>
/ (subalpine slopes/under stones) / 21.IV.2018 Wrase &amp; Laser [08]' (cWR); 5♂♂, 1♀, 'IRAN, Zagros Mts., P: / Chahar Mahal va / Bachtiari, Zard Koh Mt. / pass, subalpine slope // 2770m, 21.IV.2018 /
<geoCoordinate id="105EAB20E4AF1D98DA479F4C85FB20CD" degrees="32" direction="north" minutes="09" orientation="latitude" precision="15" seconds="55" value="32.16528">32°09'55&quot;N</geoCoordinate>
/
<geoCoordinate id="2DAC0992AEB0C7D84D95BEEBF108DFDC" degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="37" value="50.176945">50°10'37&quot;E</geoCoordinate>
/ leg. M.
<normalizedToken id="68E3653CC52A1B5B623F31B8BE90EB8F" originalValue="Hartmann’">Hartmann'</normalizedToken>
(c NME); 10♂♂, 5♀♀,
<geoCoordinate id="3320F72027F03AC81C561C33FA8B35B0" degrees="32" direction="north" minutes="09" orientation="latitude" precision="15" seconds="55" value="32.16528">32°09'55&quot;N</geoCoordinate>
,
<geoCoordinate id="BCD6C44B6B8BD53479EAB6BE3FE24949" degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="37" value="50.176945">50°10'37&quot;E</geoCoordinate>
Iran / Zagros Mts., p.
<normalizedToken id="CBC2A82839DD4971677DDD9811D64D6A" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="8DAEC9EE492F9A41EC0E28E82153558D" originalValue="Mahāll">Mahall</normalizedToken>
/
<normalizedToken id="69D270E3EA03315FA628D1B7236A35FA" originalValue="vā">va</normalizedToken>
<normalizedToken id="7CA2D5267ABDA57AD1B5E537654FCC64" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
, Zard Koh Mts., pass / 20km W Samsami, subalpine / slopes 21.IV.2018 2.775m NN / leg.: Schnitter IR49' (NME); 12♂♂, 2♀♀, 'IRAN: Zagros Mts., p.
<normalizedToken id="61512F531FB60675378DE949941A3564" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="164A5A0EA61406D2F63683F20659C946" originalValue="Mahāll">Mahall</normalizedToken>
/
<normalizedToken id="B58A10B94E1788686CF451A36EB633F4" originalValue="vā">va</normalizedToken>
<normalizedToken id="06E8C8FED56B92E355EA0E2F26968F44" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
, Zard Koh Mts.,pass 20km / W Samsami,
<geoCoordinate id="DA6982F1EF78F1EA03F0E01DCBB6BEF3" degrees="32" direction="north" minutes="09" orientation="latitude" precision="15" seconds="55" value="32.16528">32°09'55&quot;N</geoCoordinate>
,
<geoCoordinate id="4F496C39829FDC25CED6169644878A17" degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="37" value="50.176945">50°10'37&quot;E</geoCoordinate>
/ 2775m, 21.IV.2018, subalp. slopes / leg. A.Weigel #49' (NME); 1♀, 'IR Chaharmahal and / Bakhtiari Cheri pass / 4.vi.2018 3000m /
<geoCoordinate id="08807C17ADF033152AB2281C799AC2C3" degrees="32" direction="north" minutes="09" orientation="latitude" precision="15" seconds="31" value="32.15861">32°09'31&quot;N</geoCoordinate>
,
<geoCoordinate id="05DC41C3A80CF63103F4E21EAC6752B4" degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="47" value="50.17972">50°10'47&quot;E</geoCoordinate>
/ Muilwijk J.' (cMUI); 1♂, 'IR Chaharmahal and / Bakhtiari Cheri pass / 5.vi.2018 2450m /
<geoCoordinate id="F9CBA0E8B2422A6EDCE1D2EAD00A2815" degrees="32" direction="north" minutes="10" orientation="latitude" precision="15" seconds="36" value="32.176666">32°10'36&quot;N</geoCoordinate>
,
<geoCoordinate id="504D437304ED467E5DD4A9D4FB417E52" degrees="50" direction="east" minutes="10" orientation="longitude" precision="15" seconds="06" value="50.168335">50°10'06&quot;E</geoCoordinate>
/ Muilwijk J.' (cMUI); 9♀♀, 'IR Chahar Mahaal Va / Bakhtiari Chery pass / 31.v.2019 Muilwijk J. / 2950-3260 m /
<geoCoordinate id="22CAC833687D5AB663BA93547501B799" degrees="32" direction="north" minutes="10" orientation="latitude" precision="15" seconds="51" value="32.180836">32°10'51&quot;N</geoCoordinate>
,
<geoCoordinate id="DBB14EC337B2374C455286BF16B033F5" degrees="50" direction="east" minutes="09" orientation="longitude" precision="15" seconds="59" value="50.16639">50°09'59&quot;E</geoCoordinate>
(cMUI, HMIM); 1♂, 2♀♀, 'IRAN (
<normalizedToken id="4D106C356D783996BF3A2A906FF13E2C" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="F8B1B86701249C44D3B139279F052913" originalValue="Mahāll">Mahall</normalizedToken>
<normalizedToken id="1401F40A059A8E150659D4BD2B6E6002" originalValue="vā">va</normalizedToken>
<normalizedToken id="D9EB7D999030834330778944FC4547FC" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
) / Zagros Mts., 5 km SW Asad Abad /
<geoCoordinate id="F231DD68CEE0A9F2C112DB6E3A878D1D" degrees="32" direction="north" minutes="20" orientation="latitude" precision="15" seconds="30" value="32.341667">32°20'30&quot;N</geoCoordinate>
/
<geoCoordinate id="337141E15B2C4A14A6C6A35A0FAFA060" degrees="50" direction="east" minutes="32" orientation="longitude" precision="15" seconds="59" value="50.54972">50°32'59&quot;E</geoCoordinate>
/ ca 2400 m (subalpine slopes/ / under stones) / 19.IV.2018 Wrase &amp; Laser [03]' (cWR); 3♂♂,
<geoCoordinate id="84107A4794293139D94D398AF911DA4A" degrees="32" direction="north" minutes="20" orientation="latitude" precision="15" seconds="30" value="32.341667">32°20'30&quot;N</geoCoordinate>
,
<geoCoordinate id="E1E06787294C6B48B6012AD88F0949E9" degrees="50" direction="east" minutes="32" orientation="longitude" precision="15" seconds="59" value="50.54972">50°32'59&quot;E</geoCoordinate>
Iran / Zagros Mts., p.
<normalizedToken id="24D902F8DA06E54DF5A3BDF9DEDD40B5" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="DEEF6F5029ADA4714A2701B8B30DF864" originalValue="Mahāll">Mahall</normalizedToken>
/
<normalizedToken id="D75BB33CFB93A9EB0B5F3968716F3A23" originalValue="vā">va</normalizedToken>
<normalizedToken id="1EC1043E6F922B3D216D382C441565D1" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
, Asad Abad 5 km SW / snow field, high pasture / 19.IV.2018 2.500-2.770m NN / leg.: Schnitter IR 44' (NME); 1♂ (genitalia lost after study), 3♀♀, 'IRAN Zagros Mts., P: Chahar / Mahall va Bachtiari, Asad / Abad 5 km SW,
<geoCoordinate id="B31387129408E0AF48AC1FF2F74B1BCC" degrees="32" direction="north" minutes="20" orientation="latitude" precision="15" seconds="30" value="32.341667">32°20'30&quot;N</geoCoordinate>
, /
<geoCoordinate id="1D61B5C88E99FABCF425B79789C83F2E" degrees="50" direction="east" minutes="32" orientation="longitude" precision="15" seconds="59" value="50.54972">50°32'59&quot;E</geoCoordinate>
snow field, high / pasture, 19.IV.2018, 2.500- / 2.770m, leg. A. Weigel #18-04' (NME); 3♂♂, 'IRAN: Zagros Mts., p.
<normalizedToken id="2AE34B65E6791DB093336B3C916E9279" originalValue="Chahār">Chahar</normalizedToken>
<normalizedToken id="17752E78145C52DA83DC5EECBDC0E835" originalValue="Mahāll">Mahall</normalizedToken>
/
<normalizedToken id="D1C63BFB9BC08991987DC20F4DD7E226" originalValue="vā">va</normalizedToken>
<normalizedToken id="AFD707C107DB73AEF24712A566D3C9DB" originalValue="Bachtīārī">Bachtīarī</normalizedToken>
, Asad Abad 5 km SW, / 19.IV.2018,
<geoCoordinate id="34CF8A40A6D0489760E95068245E21D8" degrees="32" direction="north" minutes="20" orientation="latitude" precision="15" seconds="30" value="32.341667">32°20'30&quot;N</geoCoordinate>
,
<geoCoordinate id="4F23A8EB4358F9AE944D99E72290A1B3" degrees="50" direction="east" minutes="32" orientation="longitude" precision="15" seconds="59" value="50.54972">50°32'59&quot;E</geoCoordinate>
/ 2500-2770m, snow field, high / pasture leg. A. Weigel #44' (NME); 1♂, 'IRAN, Zagros Mts., P: / Chahar Mahal va / Bachtiari, Asad Abad / 5 km SW, 2500-2770m // 19.IV.2018, snow field / high pasture, 32°20'30&quot;N, 50°32'59&quot;E / leg. M. Hartmann, #3/18' (NME); 4♂♂, 3♀♀, 'IR Esfahan Kamran Pass / 1.vi.2019 Muilwijl J. / 3000-3300m /
<geoCoordinate id="ECE5D799621A25F992492B2798E6A10D" degrees="32" direction="north" minutes="45" orientation="latitude" precision="15" seconds="49" value="32.76361">32°45'49&quot;N</geoCoordinate>
,
<geoCoordinate id="FBA0FB1D1DD155D15CB0C88387A59DE3" degrees="50" direction="east" minutes="02" orientation="longitude" precision="15" seconds="34" value="50.042778">50°02'34&quot;E</geoCoordinate>
' (cMUI); 1♂, 'IR Esfahan Pashadagan / 2.vi.2019 / Muilwijl J. 3000-3300m /
<geoCoordinate id="B301A05E686C0A707D6ED66EBBD38CA0" degrees="32" direction="north" minutes="45" orientation="latitude" precision="15" seconds="48" value="32.763332">32°45'48&quot;N</geoCoordinate>
,
<geoCoordinate id="AD6B9F84C50B1DE52283479982DA92F8" degrees="49" direction="east" minutes="59" orientation="longitude" precision="15" seconds="52" value="49.99778">49°59'52&quot;E</geoCoordinate>
' (cMUI).
</paragraph>
<caption id="BE3587610FBEE3D7C93C5DBDADB42081" doi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71" start="Figure 4" startId="F4">
<paragraph id="AA1D335EE6AE36D6ECF0FD510233238C" pageId="0" pageNumber="71">
<emphasis id="30923E4D5DC1EE2E5DEF00F35A437205" bold="true" pageId="0" pageNumber="71">Figure 4.</emphasis>
Dorsal habitus.
<emphasis id="AB4ABAFCB9D3160D474A0E872398D720" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="B3B0D1FC7DD5895399518DC28B7EDB32" authority="insignitus" authorityName="insignitus Bedel" authorityYear="1902" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subGenus="Eremoderus" subSpecies="insignitus">Platyderus (Eremoderus) insignitus insignitus</taxonomicName>
Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco;
<emphasis id="47F17E3DF73E5054F71BE39BBD64F0E5" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="865F06177EF9FC26B5B201D6266469EF" class="Insecta" kingdom="Animalia" lsidName="P. (E.) irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis" subGenus="E.">P. (E.) irakensis</taxonomicName>
, sp. nov., holotype;
<emphasis id="338A1524A011A145C3194C6D3CC5380A" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="6669B1FE4CA1B761FED1247B1DFBE493" class="Insecta" kingdom="Animalia" lsidName="P. (E.) jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis" subGenus="E.">P. (E.) jordanensis</taxonomicName>
, sp. nov., holotype;
<emphasis id="EC0913A171172910F4881A8229FCAECC" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="AE9D76FC751F7D793A961C29414EBB6F" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), lectotype and its labels;
<emphasis id="F62301DA9FDA90BFE454155DFCA984C7" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="26F0E1023703AAD76E499D9D040CB567" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), male specimen, &quot;Jerusalem Syria&quot;, Jerusalem District, Israel. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="FE6F8D76F8D0A043B639A781E8D70D7B" pageId="0" pageNumber="71">TME: 72 specimens. TGE: 16♂♂, 3♀♀.</paragraph>
</subSubSection>
<subSubSection id="6BA95D3737F085992C9572419E98BE7D" pageId="0" pageNumber="71" type="etymology">
<paragraph id="2EFDC422E77B4A1BA157DA13265ED319" pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph id="1483EDF18D2792E30C711FC7592B3A28" pageId="0" pageNumber="71">Adjective, derived from the name of the country where the species was collected.</paragraph>
</subSubSection>
<subSubSection id="D89110C98ADA47D97E8C9F75BEB9C568" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="FD140145B0305EB0CBF716F07FB043B0" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="666CF3298DE8876E16AA015497F53D05" pageId="0" pageNumber="71">
See
<normalizedToken id="201908CCCEAE13A9B8CD40BA41C8301B" originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
under
<taxonomicName id="7803A6891CEDB8D3D7C8577C9496F9EA" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="B340ADAA2C5047C9A8798BBDB09E2FF5" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection id="0137C4AB5C9DCCFB0A3B15D8A256F6A8" pageId="0" pageNumber="71" type="description">
<paragraph id="F37316CD05186C22A17079C3CC5370B0" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="DAE4CD739765378647CFF7014659586C" pageId="0" pageNumber="71">
<emphasis id="E6495CDD8D7F3EA9333AF92FDC41BACB" bold="true" pageId="0" pageNumber="71">
<emphasis id="7DDBECB89C63CC58FA09D8A9B4C3910F" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of small size for
<taxonomicName id="5FDBBF6B7C99ED644AFC861AA9044576" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="5C926A9F0037B335669FD22C99FB7940" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 4.30-6.65 mm; BW: 1.50-2.30 mm), with relatively short, moderately convex body (Fig.
<figureCitation id="291F1E8EB6B3DC37960600D165E675DE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype and its labels; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., male paratype, Cheri pass 20 km W Samsami, Chahar Mahal va Bakhtiari Province, Iran; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure1" httpUri="https://binary.pensoft.net/fig/695638" pageId="0" pageNumber="71">1D</figureCitation>
).
<emphasis id="6C06C1787F7DB7BA8FA1C8E55FEDE6D7" bold="true" pageId="0" pageNumber="71">
<emphasis id="A756914BB66C7148BCB4927FA130E5B2" italics="true" pageId="0" pageNumber="71">Measurements and ratio</emphasis>
s.
</emphasis>
See Table
<tableCitation id="97BF5D8525B1FC566E17AB731D822600" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric data for species of the ' Platyderus weiratheri ' and ' Platyderus Eremoderus iranicus - Platyderus Eremoderus vanensis ' groups of Platyderus." httpUri="http://table.plazi.org/id/ADBAD32C4643AAB3D1823CD894BA80BA" pageId="0" pageNumber="71" tableUuid="ADBAD32C4643AAB3D1823CD894BA80BA">1</tableCitation>
.
<emphasis id="B735B4103EA779E2C784A04FD1367052" bold="true" pageId="0" pageNumber="71">
<emphasis id="B26AC7AA82FC845C68918DFA6803A34E" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Integument uniformly yellow-brown to reddish-brown, appendages somewhat lighter than body. Surface moderately shiny.
<emphasis id="F199DDAED346ADBA6F72186E415C4BAF" bold="true" pageId="0" pageNumber="71">
<emphasis id="93D1EAEBF5D0EBD3AE50378747970D57" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Clypeus, vertex and disc of head with reduced microsculpture. Pronotum with regular isodiametric sculpticells posterolaterally and scarcely-visible to absent microsculpture in middle and anteriorly. Ventral surface with microsculpture and dorsal punctation as in
<taxonomicName id="CE464DD7813596897CED75E379BC5AF7" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="29428B7C77A86D206D5D92D6929179D5" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="4B1EAB9A2AB914B2BB4CF31B66220C9F" bold="true" pageId="0" pageNumber="71">
<emphasis id="27078162DB9A7934C836071ACD6BC185" italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
About two thirds as wide as pronotum (mean PW/HW= 1.37). Eyes moderately convex. Labrum subrectangular, as long as or slightly shorter than clypeus, with anterior margin straight. Frontoclypeal suture and frontal furrows as in
<taxonomicName id="8B94DD3B831D43C42AFDEE3ACDD22D15" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="93C2FCB4762445741CA702DDE5AFD41A" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov. Paraorbital sulci moderately deep, ending before level of posterior supraorbital pore.
<emphasis id="361A2FF17FCABCD8A9627B49F9163300" bold="true" pageId="0" pageNumber="71">
<emphasis id="5F2275EBB5D2CDDAE64A50D6DDD9D318" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum wider than long (mean PW/PL= 1.27), with widest point at second quarter. Anterior and posterior transverse impressions indistinct. Anterior bead lacking in medial 1/10 to 1/8. Sides convex anteriorly, straight posteriorly. Metepisternum somewhat longer than wide, MA/MI= 0.9-1.0.
<emphasis id="F86C0BFB39D818CB75D959B5C09179B6" bold="true" pageId="0" pageNumber="71">
<emphasis id="1E532F8357BDFEED182461C98DF75B3C" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Cylindrical, one and a half times or more as long as wide (mean EL/EW= 1.61), wider than pronotum and relatively long in relation to pronotum length (mean EW/PW= 1.31; mean EL/PL= 2.66). Parascutellar striole and striae as in
<taxonomicName id="13C287E8C4DC813B89B31167C0A0CD9B" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="9F910B4A7250820ACE5EE3BCA6CF79D8" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov. Interval 3 with two or three minute discal setiferous punctures, anterior puncture being mostly absent. Umbilicate setiferous series consisting mostly of 15 or 16 punctures.
<emphasis id="68AF99EA3EADDDA9E37F3FCE6717CE55" bold="true" pageId="0" pageNumber="71">
<emphasis id="2FABA926C876C7CF19A4FEB6D9E9D47C" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one in medial third. Anterior side of mesofemur ventrally with four setiferous punctures (Fig.
<figureCitation id="F83B54D2202EFA2368FFAF4B87A6A46D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures). A. Platyderus (Eremoderus) weiratheri Maran, 1940, female specimen, Boz dag koey env., Izmir District, Turkey; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; E. P. (E.) klapperichi, sp. nov., holotype; F. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; G. P. (E.) afghanistanicus, sp. nov., holotype; H. P. (E.) brunneus ferrantei Reitter, 1909, lectotype; I. P. (E.) languidus (Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71">5C</figureCitation>
). Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis id="01E37C0E5DA1AB3196364C8937AB81EB" bold="true" pageId="0" pageNumber="71">
<emphasis id="0453F6CA381E12FCDA824C70ACF72C30" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Median lobe of aedeagus in lateral view as in
<taxonomicName id="47B334FF391BF292CAC1D4E6E761D142" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="813FFABCA91FE8CA191E2CFDCB670A52" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov., but significantly narrower, with shaft less protruding dorsally and more constricted proximally (Fig.
<figureCitation id="E58846D34B9FC4F1FDD2C0FC1762FACD" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Median lobe of aedeagus, left lateral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure8" httpUri="https://binary.pensoft.net/fig/695645" pageId="0" pageNumber="71">8D</figureCitation>
); median lobe in ventral view straight, ca. 3.7 times longer than wide; apical lamella nearly symmetrical (Fig.
<figureCitation id="E52B42FDC609A4EBF32B3BC769E096F7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vanensis, sp. nov., holotype; F. P. (E.) vrabeci, sp. nov., holotype; G. P. (E.) lassallei, sp. nov., holotype; H. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; I. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; J. P. (Platyderus) sp., male specimen, vicinity of Izmir City, Turkey; K. P. (P.) sp., male specimen, Boz Daglar Mtn., Turkey. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure10" httpUri="https://binary.pensoft.net/fig/695647" pageId="0" pageNumber="71">10D</figureCitation>
). Internal sac in lateral view as in
<taxonomicName id="7EACB159C45A9A11DC69EBAFB51002D4" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="D03CA187AD51846A6EC4226638115D6A" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov., but with ventral sclerite truncate distally; same in ventral view as in
<taxonomicName id="22D3003C4D2052A5DE9F45348F08FB64" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="7626D3EC5D26ED70C0CA01FFD50A3DFF" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov. Right paramere as that of
<taxonomicName id="0FD6782A0986FB95600FF7BECDEB5222" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="EC0E2937E1E88A013FEA4D353175F836" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov. but somewhat finer (Fig.
<figureCitation id="C0714296BCE7DFCD2BB64432AC56BACC" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Right paramere, external face. A. Platyderus (Eremoderus) chatzakiae, sp. nov., holotype; B. P. (E.) weiratheri Maran, 1940, lectotype; C. P. (E.) felixi, sp. nov., holotype; D. P. (E.) iranicus, sp. nov., holotype; E. P. (E.) vrabeci, sp. nov., holotype; F. P. (E.) lassallei, sp. nov., holotype; G. P. (E.) ledouxi Morvan, 1974, male specimen, 10 km S Hasan Keif, Mazandaran Province, Iran; H. P. (E.) taghizadehi Morvan, 1974, male specimen, Tochal, Tehran Province, Iran; I. P. (E.) afghanistanicus, sp. nov., holotype; J. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; K. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; L. P. (E.) jordanensis, sp. nov., holotype; M. P. (E.) languidus (Reiche &amp; Saulcy, 1855), lectotype; N. P. (E.) languidus (Reiche &amp; Saulcy, 1855), male paralectotype, &quot; Syria &quot;. Scale bars: 0.5 mm." figureDoi="10.3897/dez.69.83840.figure12" httpUri="https://binary.pensoft.net/fig/695649" pageId="0" pageNumber="71">12D</figureCitation>
).
<emphasis id="5AA55889EC069378726D5D3125EE8AE8" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation id="040ABA6BCDD6221048C496612E07E88E" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) weiratheri Maran, 1940, topotype female specimen; B. P. (E.) felixi, sp. nov., topotype female paratype; C. P. (E.) iranicus, sp. nov., topotype female paratype; D. P. (E.) vanensis, sp. nov., topotype female paratype; E. P. (E.) vanensis, sp. nov., female paratype, col de Buglan 1500 m, border area of Bingoel and Mus provinces, Turkey; F. P. (E.) vrabeci, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations: ag - apical gonocoxite; at - atrium; bc - bursa copulatrix; bg - basal gonocoxite; co - common oviduct; rc - receptaculum; sg - spermathecal gland; smc - seminal canal; spc - spermathecal canal." figureDoi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71">13C</figureCitation>
). Apical gonocoxite with blunt apex and two dorsolateral ensiform setae. Spermathecal canal connected in medial third of receptaculum.
</paragraph>
</subSubSection>
<subSubSection id="2B77E96B32CF7C9BDCC4E7D390264BAE" pageId="0" pageNumber="71" type="habitat">
<paragraph id="B4DF59BA991DEE9ACD5DD8B45589FF61" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="770C9ECBEF054B7BB58BC981F38FA8FA" pageId="0" pageNumber="71">
The holotype and one female paratype were collected along an artificial lake situated in a valley northwest of village Aliabad. They were caught at grassland with lots of stones situated along this lake. The vegetation there consisted of grassland, fields and small orchards - as a whole a cultivated landscape. Other ground beetles collected at this place, together with the new species, were
<taxonomicName id="80EF32C0CF2EF40C1C4AA23A04FBDFDD" authorityName="Osculati" authorityYear="1844" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus maurus subsp. osculatii" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="maurus" subSpecies="osculatii">
<emphasis id="958448BAF260FF2C87BF9FD2BCB247D9" italics="true" pageId="0" pageNumber="71">Carabus maurus osculatii</emphasis>
</taxonomicName>
Villa, 1844,
<taxonomicName id="5DEAFC138CD0CD39936212B736511502" baseAuthorityName="Chaudoir" baseAuthorityYear="1868" class="Insecta" family="Carabidae" genus="Poecilus" kingdom="Animalia" lsidName="Poecilus festivus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="festivus">
<emphasis id="274343B574FFE65D50B2A0635FB68C51" italics="true" pageId="0" pageNumber="71">Poecilus festivus</emphasis>
</taxonomicName>
(Chaudoir, 1868) and
<taxonomicName id="097B0D19F89146CDEC290A2BC18E76D0" baseAuthorityName="Heer" baseAuthorityYear="1841" class="Insecta" family="Carabidae" genus="Amblystomus" kingdom="Animalia" lsidName="Amblystomus niger" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="937A36F67C359028FF3B5CB54E27CFDA" italics="true" pageId="0" pageNumber="71">Amblystomus niger</emphasis>
</taxonomicName>
(Heer, 1841).
</paragraph>
<paragraph id="CA2AA32A5C4163292290DA94603BB603" pageId="0" pageNumber="71">Cheri Pass environments were represented by overgrazed grasslands with rocks and stones at lower altitudes and stone slopes with sparse vegetation and snow fields at higher altitudes. Similarly, Kamran Pass habitats were consisted of overgrazed grasslands at lower altitude and of stony (sub)alpine slopes with sparse vegetation and snow fields at higher altitudes.</paragraph>
<paragraph id="256FA30DAFC7A4D19399703D645996E0" pageId="0" pageNumber="71">Habitats around Asad Abad represent grasslands on subalpine slopes; specimens there were collected under stones, sometimes near edges of snowfields.</paragraph>
</subSubSection>
<subSubSection id="35301B938720F4DE746565825BFDDF92" pageId="0" pageNumber="71" type="distribution">
<paragraph id="AB68C52710B59C0CAB4B07DEA7C5BB20" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="E579426763A327FCD66212EF6E2712F0" pageId="0" pageNumber="71">Southwest Iran: Central Zagros Mountain Chains. So far, four isolated populations are known: (1) Borujen County; (2) Kuhrang County (type locality); (3) Shahr-e Kord County (locality southwest Asad Abad); (4) Fereydunshahr County (Kamran Pass), the first three in Chahar Mahal va Bakhtiari Province, the fourth one in Isfahan Province (near border area with Chahar Mahal va Bakhtiari Province).</paragraph>
</subSubSection>
<subSubSection id="53420298D6D6A81D3FDBCEA6DDC2DFF9" pageId="0" pageNumber="71" type="notes">
<paragraph id="D896A0F75E918FCE236065D1CB306C88" pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph id="8AA8DDD4AF52910003932EF60FF78C04" pageId="0" pageNumber="71">
The two specimens collected 7 km northeast of Naghan Town represent the nearest population of
<taxonomicName id="C77E768AE7E28648CD246C4D9AA09681" class="Insecta" kingdom="Animalia" lsidName="P. iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus">
<emphasis id="20133AA4FC6C28E2D80111D95B8E5848" italics="true" pageId="0" pageNumber="71">P. iranicus</emphasis>
</taxonomicName>
sp. nov. to the type locality of
<taxonomicName id="9EF4B585EB9893A56C961C1970F2E09D" class="Insecta" kingdom="Animalia" lsidName="P. felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi">
<emphasis id="E5228728FDCFE5805BF74517B7AF830E" italics="true" pageId="0" pageNumber="71">P. felixi</emphasis>
</taxonomicName>
sp. nov., ca. 17 km in a straight line (Fig.
<figureCitation id="654F03018AA6CFA5B8A57BB1BA8CA288" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of species from subgenus Platyderus Eremoderus in Iran and Afghanistan: Platyderus felixi sp. nov. (red circle), P. iranicus sp. nov. (blue circles), P. lassallei sp. nov. (yellow circles), P. davatchii (green triangle), P. ledouxi (red triangle), P. taghizadehi (blue triangle), P. klapperichi sp. nov. (black triangle), and P. afghanistanicus sp. nov. (red question mark)." figureDoi="10.3897/dez.69.83840.figure17" httpUri="https://binary.pensoft.net/fig/695654" pageId="0" pageNumber="71">17</figureCitation>
). The specimen from the environments of the Choghaklor Lake was collected together with three specimens of
<taxonomicName id="58F908B9DEF916484810DCBF58518756" baseAuthorityName="Menetries" baseAuthorityYear="1832" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus umbratus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="umbratus">Platyderus (s. str.) umbratus</taxonomicName>
(Men.).
</paragraph>
<paragraph id="F699B38DC622C20A1B13C99A9F56F0B8" pageId="0" pageNumber="71">The distance between the population from the type locality (near Cheri Pass) and that one southwest of Asad Abad is ca. 45 km in a straight line. On the other hand, the distance between Cheri Pass and locality northeast of Naghan Town is ca. 62 km in a straight line. Respectively, the distance between the locality southwest of Asad Abad and that one northeast of Naghan Town is about 47 km in a straight line. The distance between the type locality (near Cheri Pass) and the locality of Kamran Pass is about 67 km in a straight line (the same distance is between the populations from the Kamran Pass and that of Asad Abad). The distance between the Kamran Pass (Fereydunshahr County) and the locality northeast of Naghan Town is about 112 km in a straight line.</paragraph>
<caption id="60C9768E5594E7C17DC142D58579ABB7" doi="10.3897/dez.69.83840.figure5" httpUri="https://binary.pensoft.net/fig/695642" pageId="0" pageNumber="71" start="Figure 5" startId="F5">
<paragraph id="6CD6FF3A75EF78699C9736420E8ED0EF" pageId="0" pageNumber="71">
<emphasis id="55E0F9645EA65A11D15FE9E25A53B04D" bold="true" pageId="0" pageNumber="71">Figure 5.</emphasis>
Mesofemora, anterior side (A-D: left mesofemur, E-I: right mesofemur; white arrows and dots indicate position of setiferous punctures).
<emphasis id="3ED81C56EC195D854092DC791E04D2F8" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="F46DB3F1BFDC9C1E5E4CD5A7DF8A8D92" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="Eremoderus">Platyderus (Eremoderus) weiratheri</taxonomicName>
<normalizedToken id="81C7FE8B61E63C8BCF7B059A93DEEAD3" originalValue="Mařan">Maran</normalizedToken>
, 1940, female specimen, Boz
<normalizedToken id="C696279DA373452A9FA3BEE0C1320393" originalValue="dağ">dag</normalizedToken>
<normalizedToken id="89A9621B2F642EEF72308B8BF4843807" originalValue="köy">koey</normalizedToken>
env., Izmir District, Turkey;
<emphasis id="8039EF565473717C542719E511C0F190" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="4447411521BEC0760FAE743F91F608AC" class="Insecta" kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
, sp. nov., topotype female paratype;
<emphasis id="BBE89BBAED171813DBA4F1694DBD169D" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="F0658CB01CB51CDFFB7AA1F34082907E" class="Insecta" kingdom="Animalia" lsidName="P. (E.) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="E.">P. (E.) iranicus</taxonomicName>
, sp. nov., topotype female paratype;
<emphasis id="42E3AF97AB1A2A508C67354DD58EAAB4" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="B7922B043E45CA18637C8C34FA5E2963" class="Insecta" kingdom="Animalia" lsidName="P. (E.) lassallei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="lassallei" subGenus="E.">P. (E.) lassallei</taxonomicName>
, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol;
<emphasis id="EC8A2D9E4C7694B0A69A47B2501E1393" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="AFEA46346CE200117E730F28F97EEBAC" class="Insecta" kingdom="Animalia" lsidName="P. (E.) klapperichi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="klapperichi" subGenus="E.">P. (E.) klapperichi</taxonomicName>
, sp. nov., holotype;
<emphasis id="EACE73164284DC0FB5A34BCE6D8D21D2" bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName id="98AC6949753FE8C36958513E67B01A93" class="Insecta" kingdom="Animalia" lsidName="P. (E.) taghizadehi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="taghizadehi" subGenus="E.">P. (E.) taghizadehi</taxonomicName>
Morvan, 1974, male specimen, Tochal, Tehran Province, Iran;
<emphasis id="F3002E08AF6240E11A90EE0925244BB2" bold="true" pageId="0" pageNumber="71">G.</emphasis>
<taxonomicName id="442899284229F9E714C80D6E32FCA7F8" class="Insecta" kingdom="Animalia" lsidName="P. (E.) afghanistanicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="afghanistanicus" subGenus="E.">P. (E.) afghanistanicus</taxonomicName>
, sp. nov., holotype;
<emphasis id="A57306EA78C25E287BD810BF33F2CE02" bold="true" pageId="0" pageNumber="71">H.</emphasis>
<taxonomicName id="5857BE09CF994448A231ED5A0C34A328" class="Insecta" kingdom="Animalia" lsidName="P. (E.) brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subGenus="E." subSpecies="ferrantei">P. (E.) brunneus ferrantei</taxonomicName>
Reitter, 1909, lectotype;
<emphasis id="81BC1435291B956D15820CA59DBF232C" bold="true" pageId="0" pageNumber="71">I.</emphasis>
<taxonomicName id="95C3CC3C5CE1AD3B9EA0E6B3ED264495" class="Insecta" kingdom="Animalia" lsidName="P. (E.) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="E.">P. (E.) languidus</taxonomicName>
(Reiche &amp; Saulcy, 1855), female specimen, Netanya 1.3.97, Central District, Israel. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="1990771B4AAC5EC239895F3EC41E7754" pageId="0" pageNumber="71">
&quot;
<emphasis id="818FC640FD26E1049A47E01A6C8EDAB5" bold="true" pageId="0" pageNumber="71">
<taxonomicName id="F3E74EF3BD38D5B34FB96765071B8A0D" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="Eremoderus">
<emphasis id="F22B550870FEB1CF4BEB935D06FABD33" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus vanensis</emphasis>
</taxonomicName>
&quot; subgroup
</emphasis>
</paragraph>
<paragraph id="265E4111A8B6629CED1D4E48E68B12FA" pageId="0" pageNumber="71">
<emphasis id="A498351687713657A6B55B737AFF3E51" bold="true" pageId="0" pageNumber="71">Notes.</emphasis>
By its parascutellar striole and striae 1-8 more coarsely and densely punctate and more deeply impressed, the species from Eastern Turkey differ from those of the &quot;
<taxonomicName id="CB734DA94A9515AE6169C1334AD8F3B3" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="Eremoderus">
<emphasis id="18D3A56F2EF324338854BE8C95DECBE0" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus iranicus</emphasis>
</taxonomicName>
&quot; subgroup.
</paragraph>
</subSubSection>
</treatment>
</document>
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