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<document id="0EE0063F10751C0E65C68C4B7FBF555C" ID-CLB-Dataset="3499" ID-DOI="10.11646/zootaxa.5068.1.4" ID-GBIF-Dataset="f4246026-f48b-4fe0-8555-4faf0639bd35" ID-ISSN="1175-5326" ID-Zenodo-Dep="5702167" ID-ZooBank="4DAB2CFA-B192-434D-A697-8B1067BBFDED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1636961835283" checkinUser="plazi" docAuthor="De Oliveira, Denize Wgliana G., Linzmeier, Adelita M. &amp; Konstantinov, Alexander S." docDate="2021" docId="D26D412EFFFF8F50FF14F16985E7FB4A" docLanguage="en" docName="zootaxa.5068.1.4.pdf" docOrigin="Zootaxa 5068 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tupiniquim confusa Linzmeier, Oliveira &amp; Konstantinov 2021, new species" docType="treatment" docVersion="8" lastPageNumber="109" masterDocId="2E543956FFF58F5AFF83F0228732FFA3" masterDocTitle="Discovery of the first leaf litter inhabiting flea beetles in Brazil (Coleoptera: Chrysomelidae: Galerucinae) with description of two new genera and three new species" masterLastPageNumber="114" masterPageNumber="99" pageNumber="109" updateTime="1699468582102" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="3B2BEB4E224426AC36B1FD846472D431">
<mods:title id="3E89A29BF41744F311AC7F1AC04B8622">Discovery of the first leaf litter inhabiting flea beetles in Brazil (Coleoptera: Chrysomelidae: Galerucinae) with description of two new genera and three new species</mods:title>
</mods:titleInfo>
<mods:name id="FA614298D9C4C9736CCC307398534E6B" type="personal">
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<mods:roleTerm id="6F28708D119D02F166881C047FAF2C42">Author</mods:roleTerm>
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<mods:namePart id="A0E49623D0464459C89C603A4C0AD0DE">De Oliveira, Denize Wgliana G.</mods:namePart>
</mods:name>
<mods:name id="4EDBC5025FBCA99CF062740BE95F6CD0" type="personal">
<mods:role id="2CC925805E4259D129B61194368EB35D">
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<mods:namePart id="AAB172B1154FC11712000E8CADA91FA7">Linzmeier, Adelita M.</mods:namePart>
</mods:name>
<mods:name id="4BBF42D7DEE07A8270241D8BCF71173C" type="personal">
<mods:role id="1D127C2F4F4E4D6DC16C7619386C11F3">
<mods:roleTerm id="4938D14AC4EED74CBDEA3885651EE6B7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="054B6E9AC94FD72A2C44AD03CBF933A5">Konstantinov, Alexander S.</mods:namePart>
<mods:nameIdentifier id="2F66344ECEF3F4BEB3E0A063D84AD984" type="ORCID">0000-0002-9289-8078</mods:nameIdentifier>
<mods:affiliation id="BEBE32E92F8A4C43F0213008982CB09A">Systematic Entomology Laboratory, USDA, c / o Smithsonian Institution, P. O. Box 37012, National Museum of Natural History, Washington, DC 20013 - 7012, USA. alex. konstantinov @ usda. gov; https: // orcid. org / 0000 - 0001 - 6578 - 6735</mods:affiliation>
<mods:nameIdentifier id="C2D3E6F738F14403986BAD163F202054" type="email">wgliana@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="1D93216BCF9523B9A2B9841F9ACF2A9F">text</mods:typeOfResource>
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<mods:title id="D34A3037920B14BCD59D5C510BB1B83B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D1F166C15B79547E99B8F865A8F15B09">
<mods:date id="BB9310D5369F0BD3E8D92E1B9788615E">2021</mods:date>
<mods:detail id="0775ED84190B65966609A6D4B19BBCE2" type="pubDate">
<mods:number id="C334DB63925FEDE425114F71B6FE540A">2021-11-15</mods:number>
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<mods:detail id="4954B624342CB434D3C745DA8F0BFD0F" type="volume">
<mods:number id="45B8AB82959CF30C503056404737F1DF">5068</mods:number>
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<mods:classification id="DEFDE376EF1E78FFE7FEEE7A140971BC">journal article</mods:classification>
<mods:identifier id="B09D9EE019E871D35FC559C237FE705F" type="CLB-Dataset">3499</mods:identifier>
<mods:identifier id="3064BC033940F77E4C4A32C59E7DEC9B" type="DOI">10.11646/zootaxa.5068.1.4</mods:identifier>
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<treatment id="D26D412EFFFF8F50FF14F16985E7FB4A" ID-DOI="http://doi.org/10.5281/zenodo.5702253" ID-GBIF-Taxon="190302924" ID-Zenodo-Dep="5702253" LSID="urn:lsid:plazi:treatment:D26D412EFFFF8F50FF14F16985E7FB4A" httpUri="http://treatment.plazi.org/id/D26D412EFFFF8F50FF14F16985E7FB4A" lastPageNumber="109" pageId="10" pageNumber="109">
<subSubSection id="12DEA3B3FFFF8F50FF14F16984C7FEC7" box="[151,1013,330,357]" pageId="10" pageNumber="109" type="nomenclature">
<paragraph id="5A7BF038FFFF8F50FF14F16984C7FEC7" blockId="10.[151,1013,330,393]" box="[151,1013,330,357]" pageId="10" pageNumber="109">
<heading id="01334754FFFF8F50FF14F16984C7FEC7" bold="true" box="[151,1013,330,357]" fontSize="11" level="1" pageId="10" pageNumber="109" reason="1">
<emphasis id="68B02C2AFFFF8F50FF14F16984C7FEC7" bold="true" box="[151,1013,330,357]" pageId="10" pageNumber="109">
<taxonomicName id="9DC48BBBFFFF8F50FF14F169846BFEC7" authority="Linzmeier, Oliveira &amp; Konstantinov" authorityName="Linzmeier, Oliveira &amp; Konstantinov" authorityYear="2021" box="[151,857,330,357]" class="Insecta" family="Chrysomelidae" genus="Tupiniquim" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="109" phylum="Arthropoda" rank="species" species="confusa" status="sp. nov.">
<emphasis id="68B02C2AFFFF8F50FF14F16986BFFEC7" bold="true" box="[151,397,330,357]" italics="true" pageId="10" pageNumber="109">Tupiniquim confusa</emphasis>
Linzmeier, Oliveira &amp; Konstantinov
</taxonomicName>
,
<taxonomicNameLabel id="73839151FFFF8F50FCE6F16984C7FEC7" box="[869,1013,330,357]" pageId="10" pageNumber="109" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12DEA3B3FFFF8F50FF14F14D8615FE2A" box="[151,295,367,393]" pageId="10" pageNumber="109" type="description">
<paragraph id="5A7BF038FFFF8F50FF14F14D8615FE2A" blockId="10.[151,1013,330,393]" box="[151,295,367,393]" pageId="10" pageNumber="109">
(
<figureCitation id="C2FFECBDFFFF8F50FF1CF14D862DFE2A" box="[159,287,367,393]" captionStart-0="FIGURES 14–16" captionStart-1="FIGURES 17–22" captionStartId-0="8.[151,264,1888,1912]" captionStartId-1="9.[151,264,1924,1948]" captionTargetBox-0="[182,1414,217,1844]" captionTargetBox-1="[151,1416,202,1885]" captionTargetId-0="figure-24@8.[151,1436,181,1864]" captionTargetId-1="figure-19@9.[151,1436,181,1901]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 14–16. Tupiniquim confusa Linzmeier, Oliveira &amp; Konstantinov, new species, holotype, male, habitus. 14, dorsal view. 15, lateral view. 16, frontal view." captionText-1="FIGURES 17–22. Tupiniquim confusa Linzmeier, Oliveira &amp; Konstantinov, new species. 17, female abdomen. 18, tignum and spermatheca. 19, right metafemur. 20, tignum. 21, median lobe of aedeagus (ventral, lateral and dorsal views). 22, spermatheca." figureDoi-0="http://doi.org/10.5281/zenodo.5702177" figureDoi-1="http://doi.org/10.5281/zenodo.5702179" httpUri-0="https://zenodo.org/record/5702177/files/figure.png" httpUri-1="https://zenodo.org/record/5702179/files/figure.png" pageId="10" pageNumber="109">Figs 14–22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="12DEA3B3FFFF8F50FF14F1948495FD25" pageId="10" pageNumber="109" type="diagnosis">
<paragraph id="5A7BF038FFFF8F50FF14F1948495FD25" blockId="10.[151,1437,438,1257]" pageId="10" pageNumber="109">
<emphasis id="68B02C2AFFFF8F50FF14F1948620FE73" bold="true" box="[151,274,438,464]" pageId="10" pageNumber="109">Diagnosis.</emphasis>
<taxonomicName id="9DC48BBBFFFF8F50FE98F19586CEFE72" authorityName="Linzmeier, Oliveira &amp; Konstantinov" authorityYear="2021" box="[283,508,439,465]" class="Insecta" family="Chrysomelidae" genus="Tupiniquim" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="109" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="68B02C2AFFFF8F50FE98F19586CEFE72" box="[283,508,439,465]" italics="true" pageId="10" pageNumber="109">Tupiniquim confusa</emphasis>
</taxonomicName>
can easily be differentiated from its congener (in parentheses) by the following features: setiferous pores in a small cluster of three on each side of vertex near eyes (four setiferous pores organized in two longitudinal lines); posterior margin of pronotum without depressions (two small depression on posterior margin of pronotum); pronotal surface finely textured (pronotal surface almost smooth, with fine punctures); elytra with punctures confusedly organized (elytra with punctures barely organized in rows); median lobe of aedeagus at apex bent ventrally in lateral view (median lobe of aedeagus straight).
</paragraph>
</subSubSection>
<subSubSection id="12DEA3B3FFFF8F50FF44F2AC8505FBB2" pageId="10" pageNumber="109" type="description">
<paragraph id="5A7BF038FFFF8F50FF44F2AC8681FC06" blockId="10.[151,1437,438,1257]" pageId="10" pageNumber="109">
<emphasis id="68B02C2AFFFF8F50FF44F2AC8663FD0B" bold="true" box="[199,337,654,680]" pageId="10" pageNumber="109">Description</emphasis>
. Body
<quantity id="9D3C5DDDFFFF8F50FE2AF2AC857DFD0A" box="[425,591,654,681]" metricMagnitude="-3" metricUnit="m" metricValue="1.89" metricValueMax="2.05" metricValueMin="1.73" pageId="10" pageNumber="109" unit="mm" value="1.89" valueMax="2.05" valueMin="1.73">1.73–2.05 mm</quantity>
long and
<quantity id="9D3C5DDDFFFF8F50FD45F2AD8459FD0A" box="[710,875,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="1.105" metricValueMax="1.13" metricValueMin="1.08" pageId="10" pageNumber="109" unit="mm" value="1.105" valueMax="1.13" valueMin="1.08">1.08–1.13 mm</quantity>
wide. Color entirely yellow or brown; in brown specimens pronotum and head sometimes darker and antennae and legs lighter. Frons and vertex forming about 135º angle, in lateral view. Supraorbital pore almost indistinguishable from two or three other pores closely placed in cluster (
<figureCitation id="C2FFECBDFFFF8F50FE92F2D98654FCB5" box="[273,358,763,790]" captionStart="FIGURES 14–16" captionStartId="8.[151,264,1888,1912]" captionTargetBox="[182,1414,217,1844]" captionTargetId="figure-24@8.[151,1436,181,1864]" captionTargetPageId="8" captionText="FIGURES 14–16. Tupiniquim confusa Linzmeier, Oliveira &amp; Konstantinov, new species, holotype, male, habitus. 14, dorsal view. 15, lateral view. 16, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5702177" httpUri="https://zenodo.org/record/5702177/files/figure.png" pageId="10" pageNumber="109">Fig. 16</figureCitation>
). Ridge from inner margin of antennal socket to eye not inclined towards insertion of antenna. Pronotum rectangular, width 1.5 times length. Surface of pronotum microreticulate. Elytra punctured, with punctures confused (
<figureCitation id="C2FFECBDFFFF8F50FEC9F36186ACFCFE" box="[330,414,835,861]" captionStart="FIGURES 14–16" captionStartId="8.[151,264,1888,1912]" captionTargetBox="[182,1414,217,1844]" captionTargetId="figure-24@8.[151,1436,181,1864]" captionTargetPageId="8" captionText="FIGURES 14–16. Tupiniquim confusa Linzmeier, Oliveira &amp; Konstantinov, new species, holotype, male, habitus. 14, dorsal view. 15, lateral view. 16, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5702177" httpUri="https://zenodo.org/record/5702177/files/figure.png" pageId="10" pageNumber="109">Fig. 14</figureCitation>
). Median lobe simple, tubular, in ventral view lateral expansion slightly longer and farther from apex, apex with long, wide denticle; in lateral view abruptly curved basally, apex bent ventrally, straightening further apically (
<figureCitation id="C2FFECBDFFFF8F50FED1F3A98694FC06" box="[338,422,907,933]" captionStart="FIGURES 17–22" captionStartId="9.[151,264,1924,1948]" captionTargetBox="[151,1416,202,1885]" captionTargetId="figure-19@9.[151,1436,181,1901]" captionTargetPageId="9" captionText="FIGURES 17–22. Tupiniquim confusa Linzmeier, Oliveira &amp; Konstantinov, new species. 17, female abdomen. 18, tignum and spermatheca. 19, right metafemur. 20, tignum. 21, median lobe of aedeagus (ventral, lateral and dorsal views). 22, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5702179" httpUri="https://zenodo.org/record/5702179/files/figure.png" pageId="10" pageNumber="109">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="5A7BF038FFFF8F50FF44F38D8505FBB2" blockId="10.[151,1437,438,1257]" pageId="10" pageNumber="109">
Spermatheca with receptacle having external side nearly straight and internal side strongly convex; pump with longitudinal part much shorter than transverse part (
<figureCitation id="C2FFECBDFFFF8F50FD57F3F18415FC4E" box="[724,807,979,1005]" captionStart="FIGURES 17–22" captionStartId="9.[151,264,1924,1948]" captionTargetBox="[151,1416,202,1885]" captionTargetId="figure-19@9.[151,1436,181,1901]" captionTargetPageId="9" captionText="FIGURES 17–22. Tupiniquim confusa Linzmeier, Oliveira &amp; Konstantinov, new species. 17, female abdomen. 18, tignum and spermatheca. 19, right metafemur. 20, tignum. 21, median lobe of aedeagus (ventral, lateral and dorsal views). 22, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5702179" httpUri="https://zenodo.org/record/5702179/files/figure.png" pageId="10" pageNumber="109">Fig. 22</figureCitation>
). Tignum with posterior sclerotization about half as wide as tignum entire length (
<figureCitation id="C2FFECBDFFFF8F50FE24F3D58518FBB2" box="[423,554,1015,1041]" captionStart="FIGURES 17–22" captionStartId="9.[151,264,1924,1948]" captionTargetBox="[151,1416,202,1885]" captionTargetId="figure-19@9.[151,1436,181,1901]" captionTargetPageId="9" captionText="FIGURES 17–22. Tupiniquim confusa Linzmeier, Oliveira &amp; Konstantinov, new species. 17, female abdomen. 18, tignum and spermatheca. 19, right metafemur. 20, tignum. 21, median lobe of aedeagus (ventral, lateral and dorsal views). 22, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.5702179" httpUri="https://zenodo.org/record/5702179/files/figure.png" pageId="10" pageNumber="109">Figs 18, 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="12DEA3B3FFFF8F50FF44F43984E8FB02" pageId="10" pageNumber="109" type="materials_examined">
<paragraph id="5A7BF038FFFF8F50FF44F43987ECFBDE" blockId="10.[151,1437,438,1257]" pageId="10" pageNumber="109">
<emphasis id="68B02C2AFFFF8F50FF44F4398640FB97" bold="true" box="[199,370,1050,1077]" pageId="10" pageNumber="109">Type material.</emphasis>
<materialsCitation id="EAACFA65FFFF8F50FEF5F439869BFBFA" ID-GBIF-Occurrence="3406813302" collectingDate="2012-11-16" collectingDateMax="2012-11-18" collectingDateMin="2012-11-16" collectionCode="DZUP, R" collectorName="S. &amp; Sv. Kurbatov" country="Brazil" elevation="900" location="Camacan" pageId="10" pageNumber="109" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
<typeStatus id="857F4E9AFFFF8F50FEF5F43986D1FB96" box="[374,483,1051,1077]" pageId="10" pageNumber="109" type="holotype">Holotype</typeStatus>
:male (
<collectionCode id="3CD568FDFFFF8F50FDAEF439854FFB96" box="[557,637,1051,1077]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="10" pageNumber="109" type="University or college">DZUP</collectionCode>
).
<collectingCountry id="22D3B0A8FFFF8F50FD0FF43985E7FB96" box="[652,725,1051,1077]" name="Brazil" pageId="10" pageNumber="109">Brazil</collectingCountry>
:
<collectingRegion id="98003EDAFFFF8F50FD5FF4398413FB96" box="[732,801,1051,1077]" country="Brazil" name="Bahia" pageId="10" pageNumber="109">Bahia</collectingRegion>
; W of
<location id="5F1BA6E3FFFF8F50FCE1F43984FFFB96" LSID="urn:lsid:plazi:treatment:D26D412EFFFF8F50FF14F16985E7FB4A:5F1BA6E3FFFF8F50FCE1F43984FFFB96" box="[866,973,1051,1077]" country="Brazil" name="Camacan" pageId="10" pageNumber="109" stateProvince="Bahia">Camacan</location>
,
<quantity id="9D3C5DDDFFFF8F50FC56F439832AFB96" box="[981,1048,1051,1077]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="109" unit="m" value="900.0">
<elevation id="D1E9170BFFFF8F50FC56F439832AFB96" box="[981,1048,1051,1077]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="109" unit="m" value="900.0">900m</elevation>
</quantity>
;
<collectorName id="F73195EEFFFF8F50FBA2F43983FEFB96" box="[1057,1228,1051,1077]" pageId="10" pageNumber="109">
Serra Bonita
<taxonomicNameLabel id="73839151FFFF8F50FB2CF43983FEFB96" box="[1199,1228,1051,1077]" pageId="10" pageNumber="109" rank="subSpecies">N.</taxonomicNameLabel>
</collectorName>
<collectionCode id="3CD568FDFFFF8F50FB4FF43983D1FB96" box="[1228,1251,1051,1077]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="109">R</collectionCode>
.;
<date id="2E7AD6F8FFFF8F50FB73F43982AAFB96" box="[1264,1432,1051,1077]" pageId="10" pageNumber="109" value="2012-11-16" valueMax="2012-11-18" valueMin="2012-11-16">
<collectingDate id="3E3E2F10FFFF8F50FB73F43982AAFB96" box="[1264,1432,1051,1077]" pageId="10" pageNumber="109" value="2012-11-16" valueMax="2012-11-18" valueMin="2012-11-16">16-18.XI.2012</collectingDate>
</date>
;
<collectorName id="F73195EEFFFF8F50FF14F41D8659FBFA" box="[151,363,1087,1113]" pageId="10" pageNumber="109">S. &amp; Sv. Kurbatov</collectorName>
, legs
</materialsCitation>
. /
<typeStatus id="857F4E9AFFFF8F50FE46F41D8502FBFA" box="[453,560,1087,1113]" pageId="10" pageNumber="109" type="holotype">Holotype</typeStatus>
,
<taxonomicName id="9DC48BBBFFFF8F50FDBEF41D83F3FBFA" authority="Linzmeier, Oliveira &amp; Konstantinov" authorityName="Linzmeier, Oliveira &amp; Konstantinov" authorityYear="2021" box="[573,1217,1087,1113]" class="Insecta" family="Chrysomelidae" genus="Tupiniquim" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="109" phylum="Arthropoda" rank="species" species="confusa" status="sp. nov.">
<emphasis id="68B02C2AFFFF8F50FDBEF41D842EFBFA" box="[573,796,1087,1113]" italics="true" pageId="10" pageNumber="109">Tupiniquim confusa</emphasis>
Linzmeier, Oliveira &amp; Konstantinov
</taxonomicName>
,
<taxonomicNameLabel id="73839151FFFF8F50FB4DF41D8214FBFA" box="[1230,1318,1087,1113]" pageId="10" pageNumber="109" rank="species">
sp.
<date id="2E7AD6F8FFFF8F50FB74F41D8214FBFA" box="[1271,1318,1087,1113]" pageId="10" pageNumber="109" value="2021-11">nov.</date>
</taxonomicNameLabel>
2021 [red label].
</paragraph>
<paragraph id="5A7BF038FFFF8F50FF44F4A584E8FB02" blockId="10.[151,1437,438,1257]" box="[199,986,1159,1185]" pageId="10" pageNumber="109">
<materialsCitation id="EAACFA65FFFF8F50FF44F4A584E4FB02" ID-GBIF-Occurrence="3406813305" box="[199,982,1159,1185]" collectionCode="USNM" pageId="10" pageNumber="109" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="857F4E9AFFFF8F50FF44F4A58606FB02" box="[199,308,1159,1185]" pageId="10" pageNumber="109" type="paratype">Paratypes</typeStatus>
(3):
<specimenCount id="4CC23BB1FFFF8F50FEE9F4A586F1FB02" box="[362,451,1159,1185]" pageId="10" pageNumber="109" type="male">2 males</specimenCount>
,
<specimenCount id="4CC23BB1FFFF8F50FE4CF4A58506FB02" box="[463,564,1159,1185]" pageId="10" pageNumber="109" type="female">1 female</specimenCount>
;
<collectionCode id="3CD568FDFFFF8F50FDBCF4A585A6FB02" box="[575,660,1159,1185]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="109" type="Museum">USNM</collectionCode>
: same data label as
<typeStatus id="857F4E9AFFFF8F50FCF3F4A584E4FB02" box="[880,982,1159,1185]" pageId="10" pageNumber="109" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="12DEA3B3FFFF8F50FF44F48985E7FB4A" pageId="10" pageNumber="109" type="etymology">
<paragraph id="5A7BF038FFFF8F50FF44F48985E7FB4A" blockId="10.[151,1437,438,1257]" pageId="10" pageNumber="109">
<emphasis id="68B02C2AFFFF8F50FF44F489867CFB66" bold="true" box="[199,334,1195,1221]" pageId="10" pageNumber="109">Etymology.</emphasis>
The specific epithet is a Latin adjective referring to one of the distinctive characteristics of this species, punctures on elytra confusedly distributed.
</paragraph>
</subSubSection>
</treatment>
</document>
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