347 lines
39 KiB
XML
347 lines
39 KiB
XML
<document id="089663455DF03E1AD4B5127293F41E2B" ID-DOI="10.11646/zootaxa.5026.4.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="5300792" ID-ZooBank="E712BC8E-3E75-422C-8587-69ED17B6CDD5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630041845836" checkinUser="plazi" docAuthor="Fomichev, Alexander A." docDate="2021" docId="D220E4309B17A402BFED8932FB13FF55" docLanguage="en" docName="zootaxa.5026.4.7.pdf" docOrigin="Zootaxa 5026 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acantholycosa sidorovi Fomichev 2021, sp. n." docType="treatment" docVersion="4" lastPageNumber="576" masterDocId="2E199C489B12A40BBF7A8D7DFF9FFFAC" masterDocTitle="New data on the wolf spiders from theAcantholycosa-complex (Araneae: Lycosidae) from the South Siberia" masterLastPageNumber="585" masterPageNumber="567" pageNumber="572" updateTime="1699139093203" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="5EA08E312746C752A3B6A6A02D89EABE">New data on the wolf spiders from theAcantholycosa-complex (Araneae: Lycosidae) from the South Siberia</mods:title>
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<mods:name id="B4D430A3B3B25985F875F932A0D681E7" type="personal">
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<mods:roleTerm id="13423D3AF9F573310F5B7DBA8EF3E313">Author</mods:roleTerm>
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<mods:namePart id="45126E0C33CC3720F3063ABB46A03CDF">Fomichev, Alexander A.</mods:namePart>
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<mods:affiliation id="A830A438413CEA985B9240927A742AF3">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. & Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia.</mods:affiliation>
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<mods:title id="12152A23E4674DDB3753B7ABA8C1079B">Zootaxa</mods:title>
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<mods:part id="138C798E3822D44C887600DD9AA8317A">
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<mods:date id="91196D8741552C932EABDECD7AE85B11">2021</mods:date>
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<mods:number id="23ECEDB5CC8DC89ED28CF29431D03567">2021-08-27</mods:number>
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<emphasis id="68FD89349B17A40EBFED8932FE65FBC6" bold="true" box="[151,506,1103,1130]" pageId="5" pageNumber="572">
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<taxonomicName id="9D892EA59B17A40EBFED8932FE2FFBC6" ID-CoL="999ML" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[151,432,1103,1130]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="sidorovi" status="sp. nov.">
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<emphasis id="68FD89349B17A40EBFED8932FE2FFBC6" bold="true" box="[151,432,1103,1130]" italics="true" pageId="5" pageNumber="572">Acantholycosa sidorovi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="73CE344F9B17A40EBEC2892DFE65FBC6" box="[440,506,1104,1130]" pageId="5" pageNumber="572" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="129306AD9B17A40EBFED890EFBB2FA95" pageId="5" pageNumber="572" type="description">
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<paragraph id="5A3655269B17A40EBFED890EFDFBFB22" blockId="5.[151,612,1103,1166]" box="[151,612,1139,1166]" pageId="5" pageNumber="572">
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<figureCitation id="C2B249A39B17A40EBFED890EFE88FB21" box="[151,279,1139,1165]" captionStart="FIGURES 10–19" captionStartId="3.[151,264,1849,1873]" captionTargetBox="[155,1425,188,1810]" captionTargetId="figure-15@3.[151,1436,181,1825]" captionTargetPageId="3" captionText="FIGURES 10–19. Male palp (10–13) and bulb (14–19) of Acantholycosa sergeevi sp. n. (10–11), A. sayanensis (14–15), A. sidorovi sp. n. (16–17), A. levinae (18-19) and Mongolicosa lizae sp. n. (12–13). 10, 12—retrolateral; 11, 13, 14, 16, 18— ventral; 15, 17, 19—anterior. Scale bars=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300796" httpUri="https://zenodo.org/record/5300796/files/figure.png" pageId="5" pageNumber="572">Figs 16–17</figureCitation>
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, 24–27, 34–35, 60–62, 75–76
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</paragraph>
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<paragraph id="5A3655269B17A40EBFED89C9FBB2FA95" blockId="5.[151,1437,1204,2041]" pageId="5" pageNumber="572">
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<emphasis id="68FD89349B17A40EBFED89C9FF7BFB62" bold="true" box="[151,228,1204,1230]" pageId="5" pageNumber="572">Types.</emphasis>
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<collectingCountry id="229E15B69B17A40EBF9689C9FECBFB62" box="[236,340,1204,1230]" name="Russia" pageId="5" pageNumber="572">RUSSIA</collectingCountry>
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:
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<collectingRegion id="984D9BC49B17A40EBE2589C9FE45FB62" box="[351,474,1204,1230]" country="Russia" name="Altay" pageId="5" pageNumber="572">Altai Krai</collectingRegion>
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:
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(
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, 001.8653),
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, without precise locality and date, collector unknown (donated by
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.
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. Sidorov)
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;
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:
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(
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, 001.8654),
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(
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, 001.8724), together with the
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;
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(
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, 001.8655),
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, the upper reaches of
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,
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,
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,
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,
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<collectingDate id="3E738A0E9B17A40EBED68860FD80FA9B" box="[428,543,1309,1335]" pageId="5" pageNumber="572" value="2016-07-09">9.07.2016</collectingDate>
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</date>
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, collector unknown (donated by
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<collectionCode id="3C98CDE39B17A40EBCF78863FC3EFA94" box="[909,929,1310,1336]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="572" type="Museum">V</collectionCode>
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.
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<collectionCode id="3C98CDE39B17A40EBCDF8863FC26FA94" box="[933,953,1310,1336]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="572" type="Museum">V</collectionCode>
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. Sidorov)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="129306AD9B17A40EBFBD883CFD4EFAD2" pageId="5" pageNumber="572" type="etymology">
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<paragraph id="5A3655269B17A40EBFBD883CFD4EFAD2" blockId="5.[151,1437,1204,2041]" pageId="5" pageNumber="572">
|
||
<emphasis id="68FD89349B17A40EBFBD883CFED1FAF7" bold="true" box="[199,334,1345,1371]" pageId="5" pageNumber="572">Etymology.</emphasis>
|
||
The specific name is a patronym given in honor of my friend, Vladislav V. Sidorov (Barnaul,
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||
<collectingCountry id="229E15B69B17A40EBFED8819FF79FAD3" box="[151,230,1380,1407]" name="Russia" pageId="5" pageNumber="572">Russia</collectingCountry>
|
||
), who donated the
|
||
<typeStatus id="8532EB849B17A40EBEC38818FE6BFAD3" box="[441,500,1381,1407]" pageId="5" pageNumber="572">types</typeStatus>
|
||
of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="129306AD9B17A40EBFBD88FAFD29F934" pageId="5" pageNumber="572" type="diagnosis">
|
||
<paragraph id="5A3655269B17A40EBFBD88FAFD29F934" blockId="5.[151,1437,1204,2041]" pageId="5" pageNumber="572">
|
||
<emphasis id="68FD89349B17A40EBFBD88FAFEDDFA0D" bold="true" box="[199,322,1415,1441]" pageId="5" pageNumber="572">Diagnosis.</emphasis>
|
||
The new species is closely related to
|
||
<taxonomicName id="9D892EA59B17A40EBDA588F5FCF7FA0D" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[735,872,1415,1442]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="altaiensis">
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||
<emphasis id="68FD89349B17A40EBDA588F5FCF7FA0D" box="[735,872,1415,1442]" italics="true" pageId="5" pageNumber="572">A. altaiensis</emphasis>
|
||
</taxonomicName>
|
||
, which is also known from the Western
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||
<collectingRegion id="984D9BC49B17A40EBA5B88FAFAC2FA0D" box="[1313,1373,1415,1441]" country="Russia" name="Altay" pageId="5" pageNumber="572">Altai</collectingRegion>
|
||
. Both species have a spine (
|
||
<emphasis id="68FD89349B17A40EBEF388D6FE3AFA69" box="[393,421,1451,1477]" italics="true" pageId="5" pageNumber="572">Sp</emphasis>
|
||
) near base of the embolus (
|
||
<emphasis id="68FD89349B17A40EBDA688D6FC9EFA68" box="[732,769,1451,1476]" italics="true" pageId="5" pageNumber="572">Em</emphasis>
|
||
), laminar apical arm (
|
||
<emphasis id="68FD89349B17A40EBC8088D6FB86FA69" box="[1018,1049,1451,1477]" italics="true" pageId="5" pageNumber="572">Aa</emphasis>
|
||
) of the tegular apophysis (
|
||
<emphasis id="68FD89349B17A40EBA3C88D6FAFBFA68" box="[1350,1380,1451,1476]" italics="true" pageId="5" pageNumber="572">Tg</emphasis>
|
||
) and diamond-shaped epigynal fovea (
|
||
<emphasis id="68FD89349B17A40EBD7788B3FDB3FA4B" box="[525,556,1486,1511]" italics="true" pageId="5" pageNumber="572">Fo</emphasis>
|
||
) with pentagonal septal base (
|
||
<emphasis id="68FD89349B17A40EBCFB88B3FC02FA44" box="[897,925,1486,1512]" italics="true" pageId="5" pageNumber="572">Sb</emphasis>
|
||
). The male of
|
||
<taxonomicName id="9D892EA59B17A40EBB3B88B3FB24FA44" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[1089,1211,1486,1512]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="sidorovi" status="sp. nov.">
|
||
<emphasis id="68FD89349B17A40EBB3B88B3FB24FA44" box="[1089,1211,1486,1512]" italics="true" pageId="5" pageNumber="572">A. sidorovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="68FD89349B17A40EBBB888B3FA9EFA44" bold="true" box="[1218,1281,1486,1512]" pageId="5" pageNumber="572">
|
||
<taxonomicNameLabel id="73CE344F9B17A40EBBB888B3FA9EFA44" box="[1218,1281,1486,1512]" pageId="5" pageNumber="572" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs by the enlarged and more curved paleal outgrowth (
|
||
<emphasis id="68FD89349B17A40EBDE3888FFD27F9A7" box="[665,696,1522,1547]" italics="true" pageId="5" pageNumber="572">Po</emphasis>
|
||
), more prominent spine near base of the embolus (in retrolateral view) and by underdeveloped apical arm of the tegular apophysis (cf. Figs 25, 27, 34 and figs 2, 4,
|
||
<quantity id="9D71F8C39B17A40EBB8B8B69FA83F982" box="[1265,1308,1556,1582]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="5" pageNumber="572" unit="in" value="6.0">6 in</quantity>
|
||
<bibRefCitation id="3E1828D79B17A40EBA598B69FF68F9FD" author="Marusik, Y. M. & Azarkina, G. N. & Koponen, S." pageId="5" pageNumber="572" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. & Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
|
||
Marusik
|
||
<emphasis id="68FD89349B17A40EBAF28B68FF32F9FD" italics="true" pageId="5" pageNumber="572">et al</emphasis>
|
||
. 2004
|
||
</bibRefCitation>
|
||
). The female of the new species can be separated from the female of
|
||
<taxonomicName id="9D892EA59B17A40EBC8F8B45FBE1F9FD" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[1013,1150,1591,1618]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="572" phylum="Arthropoda" rank="species" species="altaiensis">
|
||
<emphasis id="68FD89349B17A40EBC8F8B45FBE1F9FD" box="[1013,1150,1591,1618]" italics="true" pageId="5" pageNumber="572">A. altaiensis</emphasis>
|
||
</taxonomicName>
|
||
by the apical pocket (
|
||
<emphasis id="68FD89349B17A40EBA0E8B45FA0CF9FE" box="[1396,1427,1592,1618]" italics="true" pageId="5" pageNumber="572">Ap</emphasis>
|
||
) almost as wide as the epigynal fovea (vs. significantly narrower) and by the longer and thinner receptacles (
|
||
<emphasis id="68FD89349B17A40EBA2E8B26FAEDF9D8" box="[1364,1394,1627,1652]" italics="true" pageId="5" pageNumber="572">Re</emphasis>
|
||
, cf.
|
||
<figureCitation id="C2B249A39B17A40EBFED8B03FE8BF93B" box="[151,276,1661,1688]" captionStart="FIGURES 54–62" captionStartId="8.[151,264,1578,1602]" captionTargetBox="[151,1436,181,1554]" captionTargetId="figure-33@8.[151,1436,181,1554]" captionTargetPageId="8" captionText="FIGURES 54–62. Epigyne and female internal genitalia of Acantholycosa sayanensis (54–56), A. azyuzini (57–59) and A. sidorovi sp. n. (60–62). 54, 57, 60—intact, ventral; 55, 58, 61—macerated, ventral; 56, 59, 62—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300802" httpUri="https://zenodo.org/record/5300802/files/figure.png" pageId="5" pageNumber="572">Figs 60, 62</figureCitation>
|
||
and figs
|
||
<quantity id="9D71F8C39B17A40EBE068B03FE5CF934" box="[380,451,1662,1688]" metricMagnitude="-1" metricUnit="m" metricValue="2.159" metricValueMax="2.286" metricValueMin="2.032" pageId="5" pageNumber="572" unit="in" value="8.5" valueMax="9.0" valueMin="8.0">8–9 in</quantity>
|
||
<bibRefCitation id="3E1828D79B17A40EBEB08B03FD35F934" author="Marusik, Y. M. & Azarkina, G. N. & Koponen, S." box="[458,682,1662,1688]" pageId="5" pageNumber="572" pagination="101 - 148" refId="ref10349" refString="Marusik, Y. M., Azarkina, G. N. & Koponen, S. (2004) A survey of east Palearctic Lycosidae (Aranei). II. Genus Acantholycosa F. Dahl, 1908 and related new genera. Arthropoda Selecta, 12 (2), 101 - 148." type="journal article" year="2004">
|
||
Marusik
|
||
<emphasis id="68FD89349B17A40EBD548B03FDC0F934" box="[558,607,1662,1688]" italics="true" pageId="5" pageNumber="572">et al</emphasis>
|
||
. 2004
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="129306AD9B17A402BFBD8BDCFB18FF79" lastPageId="9" lastPageNumber="576" pageId="5" pageNumber="572" type="description">
|
||
<paragraph id="5A3655269B17A40EBFBD8BDCFF6FF823" blockId="5.[151,1437,1204,2041]" pageId="5" pageNumber="572">
|
||
<emphasis id="68FD89349B17A40EBFBD8BDCFEC6F917" bold="true" box="[199,345,1697,1723]" pageId="5" pageNumber="572">Description.</emphasis>
|
||
Male (
|
||
<typeStatus id="8532EB849B17A40EBEDC8BDCFD90F910" box="[422,527,1697,1724]" pageId="5" pageNumber="572" type="holotype">holotype</typeStatus>
|
||
). Total length 7.5. Carapace: 3.8 long, 3.2 wide. Coloration. Carapace dark brown. Eye field almost black. Chelicerae, sternum and palps brown. Labium, endites and coxae yellow-gray. Legs yellowbrown. Femora of legs with brown transverse stripes and longitudinal yellow stripes. Abdomen yellow-brown dorsally. Cardiac mark almost indistinct. Venter of the abdomen yellow-gray. Spinnerets dark gray. Spination of leg I: Fe d1-1-1 p0-0-2 r0-1-1; Ti p1-0-0 r1-0-0 v2-2-2-2-1-2; Mt p1-1-0 r0-1-0 v2-2-0. Leg measurements: I 4.4, 1.7, 4.2, 4.6, 1.8 (16.7); II 4.3, 1.7, 4.0, 4.7, 1.85 (16.55); III 4.2, 1.55, 3.6, 5.45, 1.85 (16.65); IV 4.9, 1.6, 4.3, 6.3, 1.55 (18.65).
|
||
</paragraph>
|
||
<paragraph id="5A3655269B17A40EBFBD8AEAFDB4F855" blockId="5.[151,1437,1204,2041]" pageId="5" pageNumber="572">
|
||
Palp as in
|
||
<figureCitation id="C2B249A39B17A40EBE428AEAFE28F81D" box="[312,439,1943,1969]" captionStart="FIGURES 10–19" captionStartId="3.[151,264,1849,1873]" captionTargetBox="[155,1425,188,1810]" captionTargetId="figure-15@3.[151,1436,181,1825]" captionTargetPageId="3" captionText="FIGURES 10–19. Male palp (10–13) and bulb (14–19) of Acantholycosa sergeevi sp. n. (10–11), A. sayanensis (14–15), A. sidorovi sp. n. (16–17), A. levinae (18-19) and Mongolicosa lizae sp. n. (12–13). 10, 12—retrolateral; 11, 13, 14, 16, 18— ventral; 15, 17, 19—anterior. Scale bars=0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5300796" httpUri="https://zenodo.org/record/5300796/files/figure.png" pageId="5" pageNumber="572">Figs 16–17</figureCitation>
|
||
, 24–27, 34–35. Tegular apophysis almost square, with a poorly developed basal arm (
|
||
<emphasis id="68FD89349B17A40EBA178AE5FA13F81D" box="[1389,1420,1944,1969]" italics="true" pageId="5" pageNumber="572">Ba</emphasis>
|
||
). Palea (
|
||
<emphasis id="68FD89349B17A40EBF9C8AC7FF60F878" box="[230,255,1978,2004]" italics="true" pageId="5" pageNumber="572">Pl</emphasis>
|
||
) rounded. Paleal outgrowth strongly bent. Terminal apophysis (
|
||
<emphasis id="68FD89349B17A40EBCA08AC6FC6AF878" box="[986,1013,1979,2004]" italics="true" pageId="5" pageNumber="572">Ta</emphasis>
|
||
) straight, spine-like. Embolus with a semicircular notch in the distal part.
|
||
</paragraph>
|
||
<caption id="0EF605AE9B14A40DBFED8BB3FDBBF8DB" ID-DOI="http://doi.org/10.5281/zenodo.5300798" ID-Zenodo-Dep="5300798" httpUri="https://zenodo.org/record/5300798/files/figure.png" pageId="6" pageNumber="573" startId="6.[151,264,1742,1766]" targetBox="[154,1435,189,1719]" targetPageId="6">
|
||
<paragraph id="5A3655269B14A40DBFED8BB3FDBBF8DB" blockId="6.[151,1437,1742,1911]" pageId="6" pageNumber="573">
|
||
<emphasis id="68FD89349B14A40DBFED8BB3FECBF94B" bold="true" box="[151,340,1742,1767]" pageId="6" pageNumber="573">FIGURES 32–43.</emphasis>
|
||
Tegulum and subtegulum (32–37), bulb (38–40) and embolic division (41–43) of
|
||
<taxonomicName id="9D892EA59B14A40DBBE98BB2FA03F94A" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[1171,1436,1743,1767]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="sayanensis">
|
||
<emphasis id="68FD89349B14A40DBBE98BB2FA03F94A" box="[1171,1436,1743,1767]" italics="true" pageId="6" pageNumber="573">Acantholycosa sayanensis</emphasis>
|
||
</taxonomicName>
|
||
(32–33),
|
||
<taxonomicName id="9D892EA59B14A40DBF8F8B8EFEFCF8A7" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[245,355,1779,1803]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="sidorovi" status="sp. nov.">
|
||
<emphasis id="68FD89349B14A40DBF8F8B8EFEFCF8A7" box="[245,355,1779,1803]" italics="true" pageId="6" pageNumber="573">A. sidorovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="68FD89349B14A40DBE118B8EFE3BF8A7" bold="true" box="[363,420,1779,1803]" pageId="6" pageNumber="573">
|
||
<taxonomicNameLabel id="73CE344F9B14A40DBE118B8EFE3BF8A7" box="[363,420,1779,1803]" pageId="6" pageNumber="573" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(34–35),
|
||
<taxonomicName id="9D892EA59B14A40DBD738B8EFDF1F8A7" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[521,622,1779,1803]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="levinae">
|
||
<emphasis id="68FD89349B14A40DBD738B8EFDF1F8A7" box="[521,622,1779,1803]" italics="true" pageId="6" pageNumber="573">A. levinae</emphasis>
|
||
</taxonomicName>
|
||
(36–37) and
|
||
<taxonomicName id="9D892EA59B14A40DBD828B8EFCF7F8A6" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[760,872,1779,1802]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="573" phylum="Arthropoda" rank="species" species="sergeevi" status="sp. nov.">
|
||
<emphasis id="68FD89349B14A40DBD828B8EFCF7F8A6" box="[760,872,1779,1802]" italics="true" pageId="6" pageNumber="573">A. sergeevi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="68FD89349B14A40DBC0A8B8EFC36F8A7" bold="true" box="[880,937,1779,1803]" pageId="6" pageNumber="573">
|
||
<taxonomicNameLabel id="73CE344F9B14A40DBC0A8B8EFC36F8A7" box="[880,937,1779,1803]" pageId="6" pageNumber="573" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(38–43). 32, 34, 36, 38, 41—ventral; 33, 35, 37, 39, 42—anterior; 40, 43—retrolateral. Scale bars=0.2 mm. Abbreviations:
|
||
<emphasis id="68FD89349B14A40DBCE58A6AFC24F882" box="[927,955,1815,1838]" italics="true" pageId="6" pageNumber="573">Aa</emphasis>
|
||
apical arm of the tegular apophysis,
|
||
<emphasis id="68FD89349B14A40DBA398A6AFAC0F882" box="[1347,1375,1815,1838]" italics="true" pageId="6" pageNumber="573">Ba</emphasis>
|
||
basal arm of the tegular apophysis,
|
||
<emphasis id="68FD89349B14A40DBEBF8A46FE78F8FE" box="[453,487,1851,1874]" italics="true" pageId="6" pageNumber="573">Em</emphasis>
|
||
embolus,
|
||
<emphasis id="68FD89349B14A40DBD2A8A46FDF4F8FE" box="[592,619,1851,1874]" italics="true" pageId="6" pageNumber="573">Pe</emphasis>
|
||
extension of the palea,
|
||
<emphasis id="68FD89349B14A40DBC268A46FCECF8FF" box="[860,883,1851,1875]" italics="true" pageId="6" pageNumber="573">Pl</emphasis>
|
||
palea,
|
||
<emphasis id="68FD89349B14A40DBCC18A46FC4CF8FE" box="[955,979,1851,1874]" italics="true" pageId="6" pageNumber="573">Ta</emphasis>
|
||
terminal apophysis,
|
||
<emphasis id="68FD89349B14A40DBBDD8A46FB5DF8FE" box="[1191,1218,1851,1874]" italics="true" pageId="6" pageNumber="573">Tg</emphasis>
|
||
tegular apophysis,
|
||
<emphasis id="68FD89349B14A40DBAFD8A46FA02F8FE" box="[1415,1437,1851,1874]" italics="true" pageId="6" pageNumber="573">Ts</emphasis>
|
||
triangle-shaped projection of the palea.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0EF605AE9B15A40CBFED8A7FFC7CF82A" ID-DOI="http://doi.org/10.5281/zenodo.5300800" ID-Zenodo-Dep="5300800" httpUri="https://zenodo.org/record/5300800/files/figure.png" pageId="7" pageNumber="574" startId="7.[151,264,1794,1818]" targetBox="[163,1427,188,1770]" targetPageId="7">
|
||
<paragraph id="5A3655269B15A40CBFED8A7FFC7CF82A" blockId="7.[151,1436,1794,1926]" pageId="7" pageNumber="574">
|
||
<emphasis id="68FD89349B15A40CBFED8A7FFEC8F8B6" bold="true" box="[151,343,1794,1818]" pageId="7" pageNumber="574">FIGURES 44–53.</emphasis>
|
||
Tegulum and subtegulum (44–45, 52–53), bulb (46–48) and embolic division (49–51) of
|
||
<taxonomicName id="9D892EA59B15A40CBA7D8A7FFF75F892" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="sergeevi" status="sp. nov.">
|
||
<emphasis id="68FD89349B15A40CBA7D8A7FFF75F892" italics="true" pageId="7" pageNumber="574">Acantholycosa sergeevi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="68FD89349B15A40CBF8E8A5BFEAFF892" bold="true" box="[244,304,1830,1854]" pageId="7" pageNumber="574">
|
||
<taxonomicNameLabel id="73CE344F9B15A40CBF8E8A5BFEAFF892" box="[244,304,1830,1854]" pageId="7" pageNumber="574" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(44–45) and
|
||
<taxonomicName id="9D892EA59B15A40CBEBB8A5BFDE0F892" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" box="[449,639,1830,1854]" class="Arachnida" family="Lycosidae" genus="Mongolicosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="574" phylum="Arthropoda" rank="species" species="lizae" status="sp. nov.">
|
||
<emphasis id="68FD89349B15A40CBEBB8A5BFDE0F892" box="[449,639,1830,1854]" italics="true" pageId="7" pageNumber="574">Mongolicosa lizae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="68FD89349B15A40CBDF38A5BFD5AF892" bold="true" box="[649,709,1830,1854]" pageId="7" pageNumber="574">
|
||
<taxonomicNameLabel id="73CE344F9B15A40CBDF38A5BFD5AF892" box="[649,709,1830,1854]" pageId="7" pageNumber="574" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(46–53). 44, 46, 49, 52—ventral; 45, 47, 50, 53—anterior; 48, 51— retrolateral. Scale bars=0.2 mm. Abbreviations:
|
||
<emphasis id="68FD89349B15A40CBDFA8A37FD03F8CD" box="[640,668,1866,1889]" italics="true" pageId="7" pageNumber="574">Aa</emphasis>
|
||
apical arm of the tegular apophysis,
|
||
<emphasis id="68FD89349B15A40CBB6C8A36FBADF8CE" box="[1046,1074,1867,1890]" italics="true" pageId="7" pageNumber="574">Ba</emphasis>
|
||
basal arm of the tegular apophysis,
|
||
<emphasis id="68FD89349B15A40CBFED8A12FF26F82A" box="[151,185,1903,1926]" italics="true" pageId="7" pageNumber="574">Em</emphasis>
|
||
embolus,
|
||
<emphasis id="68FD89349B15A40CBE588A13FEA6F82A" box="[290,313,1902,1926]" italics="true" pageId="7" pageNumber="574">Pl</emphasis>
|
||
palea,
|
||
<emphasis id="68FD89349B15A40CBEFB8A13FE05F82A" box="[385,410,1902,1926]" italics="true" pageId="7" pageNumber="574">Sd</emphasis>
|
||
sperm duct,
|
||
<emphasis id="68FD89349B15A40CBD678A12FDAAF82A" box="[541,565,1903,1926]" italics="true" pageId="7" pageNumber="574">Ta</emphasis>
|
||
terminal apophysis,
|
||
<emphasis id="68FD89349B15A40CBC728A12FCBCF82A" box="[776,803,1903,1926]" italics="true" pageId="7" pageNumber="574">Tg</emphasis>
|
||
tegular apophysis.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0EF605AE9B1AA403BFED8B57FE37F902" ID-DOI="http://doi.org/10.5281/zenodo.5300802" ID-Zenodo-Dep="5300802" httpUri="https://zenodo.org/record/5300802/files/figure.png" pageId="8" pageNumber="575" startId="8.[151,264,1578,1602]" targetBox="[151,1436,181,1554]" targetPageId="8">
|
||
<paragraph id="5A3655269B1AA403BFED8B57FE37F902" blockId="8.[151,1437,1578,1711]" pageId="8" pageNumber="575">
|
||
<emphasis id="68FD89349B1AA403BFED8B57FEC8F9EF" bold="true" box="[151,343,1578,1603]" pageId="8" pageNumber="575">FIGURES 54–62.</emphasis>
|
||
Epigyne and female internal genitalia of
|
||
<taxonomicName id="9D892EA59B1AA403BC758B56FB84F9EE" authorityName="Marusik, Azarkina & Koponen" authorityYear="2004" box="[783,1051,1579,1603]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="575" phylum="Arthropoda" rank="species" species="sayanensis">
|
||
<emphasis id="68FD89349B1AA403BC758B56FB84F9EE" box="[783,1051,1579,1603]" italics="true" pageId="8" pageNumber="575">Acantholycosa sayanensis</emphasis>
|
||
</taxonomicName>
|
||
(54–56),
|
||
<taxonomicName id="9D892EA59B1AA403BBFE8B56FB69F9EE" authorityName="Marusik, Hippa & Koponen" authorityYear="1996" box="[1156,1270,1579,1602]" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="575" phylum="Arthropoda" rank="species" species="azyuzini">
|
||
<emphasis id="68FD89349B1AA403BBFE8B56FB69F9EE" box="[1156,1270,1579,1602]" italics="true" pageId="8" pageNumber="575">A. azyuzini</emphasis>
|
||
</taxonomicName>
|
||
(57–59) and
|
||
<taxonomicName id="9D892EA59B1AA403BAFC8B56FF76F9CB" authority="Fomichev, 2021" authorityName="Fomichev" authorityYear="2021" class="Arachnida" family="Lycosidae" genus="Acantholycosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="575" phylum="Arthropoda" rank="species" species="sidorovi" status="sp. nov.">
|
||
<emphasis id="68FD89349B1AA403BAFC8B56FF76F9CB" italics="true" pageId="8" pageNumber="575">A. sidorovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="68FD89349B1AA403BF898B32FEAFF9CB" bold="true" box="[243,304,1615,1639]" pageId="8" pageNumber="575">
|
||
<taxonomicNameLabel id="73CE344F9B1AA403BF898B32FEAFF9CB" box="[243,304,1615,1639]" pageId="8" pageNumber="575" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(60–62). 54, 57, 60—intact, ventral; 55, 58, 61—macerated, ventral; 56, 59, 62—macerated, dorsal. Scale bars=0.2 mm. Abbreviations:
|
||
<emphasis id="68FD89349B1AA403BEBF8B0EFE7EF926" box="[453,481,1651,1674]" italics="true" pageId="8" pageNumber="575">Ap</emphasis>
|
||
apical pocket,
|
||
<emphasis id="68FD89349B1AA403BD038B0EFD08F927" box="[633,663,1651,1675]" italics="true" pageId="8" pageNumber="575">Cd</emphasis>
|
||
copulatory ducts,
|
||
<emphasis id="68FD89349B1AA403BC288B0EFCF1F926" box="[850,878,1651,1674]" italics="true" pageId="8" pageNumber="575">Fo</emphasis>
|
||
epigynal fovea,
|
||
<emphasis id="68FD89349B1AA403BB6C8B0EFBB4F926" box="[1046,1067,1651,1674]" italics="true" pageId="8" pageNumber="575">Li</emphasis>
|
||
lip of the epigyne,
|
||
<emphasis id="68FD89349B1AA403BB958B0EFA95F926" box="[1263,1290,1651,1674]" italics="true" pageId="8" pageNumber="575">Re</emphasis>
|
||
receptacle,
|
||
<emphasis id="68FD89349B1AA403BAF98B0EFA03F927" box="[1411,1436,1651,1675]" italics="true" pageId="8" pageNumber="575">Sb</emphasis>
|
||
septal base,
|
||
<emphasis id="68FD89349B1AA403BE688BEAFEB6F902" box="[274,297,1687,1710]" italics="true" pageId="8" pageNumber="575">Ss</emphasis>
|
||
septal stem.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5A3655269B1AA403BFBD8A7CFCACF87C" blockId="8.[151,1437,1793,2035]" pageId="8" pageNumber="575">Female. Total length 8.7. Carapace: 3.4 long, 2.75 wide. Coloration. Carapace dark brown. Eye field almost black. Chelicerae, sternum and labium brown. Endites and coxae yellow-gray. Legs and palps yellow-gray with brown annulation. Abdomen brown-gray dorsally. Cardiac mark dark brown. Venter of the abdomen yellow. Spinnerets yellow-gray. Spination of leg I: Fe d1-1-1 p0-0-2 r0-0-1; Ti p1-0-0 r1-0-0 v2-2-2-2-1-2; Mt p0-1-0 r0- 1-0 v2-2-0. Leg measurements: I 3.15, 1.45, 2.8, 2.75, 1.35 (11.5); II 3.15, 1.4, 2.65, 2.85, 1.3 (11.35); III 2.9, 1.25, 2.45, 3.45, 1.4 (11.45); IV 3.75, 1.4, 3.25, 5.15, 1.95 (15.5).</paragraph>
|
||
<paragraph id="5A3655269B1AA402BFBD8AA4FB18FF79" blockId="8.[151,1437,1793,2035]" lastBlockId="9.[151,1436,151,249]" lastPageId="9" lastPageNumber="576" pageId="8" pageNumber="575">
|
||
Epigyne and internal genitalia as in
|
||
<figureCitation id="C2B249A39B1AA403BD258AA4FD7EF85F" box="[607,737,2009,2035]" captionStart="FIGURES 54–62" captionStartId="8.[151,264,1578,1602]" captionTargetBox="[151,1436,181,1554]" captionTargetId="figure-33@8.[151,1436,181,1554]" captionTargetPageId="8" captionText="FIGURES 54–62. Epigyne and female internal genitalia of Acantholycosa sayanensis (54–56), A. azyuzini (57–59) and A. sidorovi sp. n. (60–62). 54, 57, 60—intact, ventral; 55, 58, 61—macerated, ventral; 56, 59, 62—macerated, dorsal. Scale bars=0.2 mm. Abbreviations: Ap apical pocket, Cd copulatory ducts, Fo epigynal fovea, Li lip of the epigyne, Re receptacle, Sb septal base, Ss septal stem." figureDoi="http://doi.org/10.5281/zenodo.5300802" httpUri="https://zenodo.org/record/5300802/files/figure.png" pageId="8" pageNumber="575">Figs 60–62</figureCitation>
|
||
. Fovea slightly longer than wide. Septal base smooth, 2 times longer than wide. Septal stem undeveloped. Lips of the epigyne (
|
||
<emphasis id="68FD89349B1BA402BC1C8DEAFCE2FF1C" box="[870,893,151,176]" italics="true" pageId="9" pageNumber="576">Li</emphasis>
|
||
) not touching each other. Apical pocket with one hood. Receptacles fusiform. Copulatory ducts (
|
||
<emphasis id="68FD89349B1BA402BDD38DC6FD55FF79" box="[681,714,187,213]" italics="true" pageId="9" pageNumber="576">Cd</emphasis>
|
||
) almost straight, as long as receptacles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="129306AD9B1BA402BFBD8DA3FB13FF55" box="[199,1164,222,249]" pageId="9" pageNumber="576" type="distribution">
|
||
<paragraph id="5A3655269B1BA402BFBD8DA3FB13FF55" blockId="9.[151,1436,151,249]" box="[199,1164,222,249]" pageId="9" pageNumber="576">
|
||
<emphasis id="68FD89349B1BA402BFBD8DA3FEFEFF54" bold="true" box="[199,353,222,248]" pageId="9" pageNumber="576">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="8532EB849B1BA402BD1A8DA2FD0FFF55" box="[608,656,223,249]" pageId="9" pageNumber="576">type</typeStatus>
|
||
locality in the
|
||
<collectingRegion id="984D9BC49B1BA402BC4D8DA2FCF0FF55" box="[823,879,223,249]" country="Russia" name="Altay" pageId="9" pageNumber="576">Altai</collectingRegion>
|
||
Mountains (
|
||
<figureCitation id="C2B249A39B1BA402BC878DA2FB1FFF55" box="[1021,1152,222,249]" captionStart="FIGURES 72–77" captionStartId="13.[151,264,1676,1700]" captionTargetBox="[151,1436,309,1652]" captionTargetId="figure-76@13.[151,1436,309,1652]" captionTargetPageId="13" captionText="FIGURES 72–77. Male of Acantholycosa sayanensis from Borus Mt. Range (72), female of Acantholycosa sp. (A. azyuzini or A. levinae) with egg sac, from Katunskyi Mt. Range (73), habitat of A. sayanensis and A. sergeevi sp. n. in Borus Mt. Range (74), collecting localities of studied species from the Acantholycosa-complex (75–76) and map showing the main mountain systems of South Siberia and adjacent regions (77). Diamond—A. sayanensis, square—A. sidorovi sp. n., triangle—A. kulikovi sp. n., star—A. levinae, circle—A. sergeevi sp. n., inverted triangle—Mongolicosa lizae sp. n. The frame of Fig. 76 refers to the content of Fig. 75." figureDoi="http://doi.org/10.5281/zenodo.5300806" httpUri="https://zenodo.org/record/5300806/files/figure.png" pageId="9" pageNumber="576">Figs 75–76</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |