treatments-xml/data/D2/18/96/D218962A7F38FFDCC099FA1FFDF3521A.xml
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<document id="6804B3F039CE8F1F09657D6E97FC6891" ID-CLB-Dataset="4041" ID-DOI="10.11646/zootaxa.5051.1.12" ID-GBIF-Dataset="61c578ca-75a0-4623-a1e8-e1d64bdd9c4a" ID-ISSN="1175-5326" ID-Zenodo-Dep="5563569" ID-ZooBank="A99E653A-EBDF-48B1-BF24-0194136E03F9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634021289632" checkinUser="plazi" docAuthor="Gómez, Samuel" docDate="2021" docId="D218962A7F38FFDCC099FA1FFDF3521A" docLanguage="en" docName="zootaxa.5051.1.12.pdf" docOrigin="Zootaxa 5051 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Delavalia reducta Gómez 2021, sp. nov." docType="treatment" docUuid="4490AD3C-AB74-4025-B975-2D89DD7CB309" docUuidSource="ZooBank" docVersion="8" lastPageNumber="204" masterDocId="2E21EE527F21FFFEC00EFF8FFFD7535A" masterDocTitle="On some deep-sea Stenheliinae from the Gulf of California and the west coast of the Baja California Peninsula (Mexico): the genus Delavalia Brady, 1869 and proposal of Archaeohuysia gen. nov. and Diarthropodella gen. nov. (Copepoda Harpacticoida: Miraciidae)" masterLastPageNumber="235" masterPageNumber="171" pageNumber="196" updateTime="1699459531294" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5CA6EA4397B0C3E9850AAF10D08F58F8">On some deep-sea Stenheliinae from the Gulf of California and the west coast of the Baja California Peninsula (Mexico): the genus Delavalia Brady, 1869 and proposal of Archaeohuysia gen. nov. and Diarthropodella gen. nov. (Copepoda Harpacticoida: Miraciidae)</mods:title>
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<mods:namePart id="CAE8A49DDAEBFE9A7C113AEB1E720D83">Gómez, Samuel</mods:namePart>
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<mods:date id="611D3AF8122015DB03F2D023A6606516">2021</mods:date>
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<mods:number id="9BF4A80D2FFE4CB1679D74732F107BB4">2021-10-12</mods:number>
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<treatment id="D218962A7F38FFDCC099FA1FFDF3521A" ID-DOI="http://doi.org/10.5281/zenodo.5570777" ID-GBIF-Taxon="188640380" ID-Zenodo-Dep="5570777" LSID="urn:lsid:zoobank.org:act:4490AD3C-AB74-4025-B975-2D89DD7CB309" httpUri="http://treatment.plazi.org/id/D218962A7F38FFDCC099FA1FFDF3521A" lastPageId="34" lastPageNumber="204" pageId="25" pageNumber="196">
<subSubSection id="12AB74B77F38FFE7C099FA1FFE0556F1" box="[151,466,1424,1451]" pageId="25" pageNumber="196" type="nomenclature">
<paragraph id="5A0E273C7F38FFE7C099FA1FFE0556F1" blockId="25.[151,977,1424,1523]" box="[151,466,1424,1451]" pageId="25" pageNumber="196">
<heading id="014690507F38FFE7C099FA1FFE0556F1" bold="true" box="[151,466,1424,1451]" fontSize="11" level="1" pageId="25" pageNumber="196" reason="1">
<emphasis id="68C5FB2E7F38FFE7C099FA1FFE0556F1" bold="true" box="[151,466,1424,1451]" pageId="25" pageNumber="196">
<taxonomicName id="9DB15CBF7F38FFE7C099FA1FFEBA56F1" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[151,365,1424,1451]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="25" pageNumber="196" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">
<emphasis id="68C5FB2E7F38FFE7C099FA1FFEBA56F1" bold="true" box="[151,365,1424,1451]" italics="true" pageId="25" pageNumber="196">Delavalia reducta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73F646557F38FFE7C17BFA1EFE0556F1" box="[373,466,1425,1451]" pageId="25" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12AB74B77F38FFE7C099FA3AFC0656A9" pageId="25" pageNumber="196" type="description">
<paragraph id="5A0E273C7F38FFE7C099FA3AFEF95695" blockId="25.[151,977,1424,1523]" box="[151,302,1461,1487]" pageId="25" pageNumber="196">
(
<figureCitation id="C28A3BB97F38FFE7C091FA3AFEF15695" box="[159,294,1461,1487]" captionStart-0="FIGURE 19" captionStart-1="FIGURE 20" captionStart-2="FIGURE 21" captionStart-3="FIGURE 22" captionStart-4="FIGURE 23" captionStartId-0="26.[151,250,1911,1935]" captionStartId-1="28.[151,250,1875,1899]" captionStartId-2="29.[151,250,1813,1837]" captionStartId-3="31.[151,250,1933,1957]" captionStartId-4="32.[151,250,1873,1897]" captionTargetBox-0="[163,1428,319,1879]" captionTargetBox-1="[161,1425,193,1838]" captionTargetBox-2="[159,1431,191,1782]" captionTargetBox-3="[157,1427,532,1903]" captionTargetBox-4="[162,1426,191,1839]" captionTargetId-0="figure-56@26.[151,1436,309,1887]" captionTargetId-1="figure-20@28.[151,1436,181,1851]" captionTargetId-2="figure-15@29.[151,1436,181,1789]" captionTargetId-3="figure-205@31.[151,1436,525,1910]" captionTargetId-4="figure-20@32.[151,1436,181,1849]" captionTargetPageId-0="26" captionTargetPageId-1="28" captionTargetPageId-2="29" captionTargetPageId-3="31" captionTargetPageId-4="32" captionText-0="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." captionText-1="FIGURE 20. Delavalia reducta sp. nov., female: A, rostrum and antennule (microphotograph available in http://metadata. icmyl.unam.mx/handle/20.500.12201/10625); B, antenna (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10618)." captionText-2="FIGURE 21. Delavalia reducta sp. nov., female: A, mandible (microphotograph available in http://metadata.icmyl.unam. mx/handle/20.500.12201/10622); B, maxillule; C, maxilla; D, maxilliped (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10616)." captionText-3="FIGURE 22. Delavalia reducta sp. nov., female: A, P1, anterior (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10620); B, P2, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10621)." captionText-4="FIGURE 23. Delavalia reducta sp. nov., female: A, P3, anterior (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10617); B, P4, anterior (microphotograph available in http://metadata.icmyl.unam. mx/handle/20.500.12201/10623); C, P5, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10624)." figureDoi-0="http://doi.org/10.5281/zenodo.5563607" figureDoi-1="http://doi.org/10.5281/zenodo.5563609" figureDoi-2="http://doi.org/10.5281/zenodo.5563611" figureDoi-3="http://doi.org/10.5281/zenodo.5563613" figureDoi-4="http://doi.org/10.5281/zenodo.5563615" httpUri-0="https://zenodo.org/record/5563607/files/figure.png" httpUri-1="https://zenodo.org/record/5563609/files/figure.png" httpUri-2="https://zenodo.org/record/5563611/files/figure.png" httpUri-3="https://zenodo.org/record/5563613/files/figure.png" httpUri-4="https://zenodo.org/record/5563615/files/figure.png" pageId="25" pageNumber="196">Figs. 1923</figureCitation>
)
</paragraph>
<paragraph id="5A0E273C7F38FFE7C099FA57FC0656A9" blockId="25.[151,977,1424,1523]" box="[151,977,1496,1523]" pageId="25" pageNumber="196">
<uri id="2E202B3E7F38FFE7C099FA57FC0656A9" box="[151,977,1496,1523]" pageId="25" pageNumber="196">
urn:lsid:zoobank.org:act:
<uuid id="2E171DE97F38FFE7C1A7FA57FC0656A9" box="[425,977,1496,1523]" pageId="25" pageNumber="196">4490AD3C-AB74-4025-B975-2D89DD7CB309</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="12AB74B77F38FFE7C099F9AEFE275591" pageId="25" pageNumber="196" type="materials_examined">
<paragraph id="5A0E273C7F38FFE7C099F9AEFD7955D9" blockId="25.[151,1437,1568,2028]" pageId="25" pageNumber="196">
<materialsCitation id="EAD92D617F38FFE7C099F9AEFD4455D9" ID-GBIF-Occurrence="3383566309" country="Mexico" county="Talud" elevation="825" latitude="31.805" location="Off Todos Santos Bay" longLatPrecision="72" longitude="-116.925" municipality="Eastern Tropical Pacific" pageId="25" pageNumber="196" specimenCount="1" stateProvince="Baja California" typeStatus="holotype">
<emphasis id="68C5FB2E7F38FFE7C099F9AEFEED5560" bold="true" box="[151,314,1568,1595]" pageId="25" pageNumber="196">
<typeStatus id="850A999E7F38FFE7C099F9AEFF055561" box="[151,210,1569,1595]" pageId="25" pageNumber="196">Type</typeStatus>
locality.
</emphasis>
<location id="5F6E71E77F38FFE7C14BF9AEFD9D5561" LSID="urn:lsid:plazi:treatment:D218962A7F38FFDCC099FA1FFDF3521A:5F6E71E77F38FFE7C14BF9AEFD9D5561" box="[325,586,1569,1595]" country="Mexico" county="Talud" latitude="31.805" longLatPrecision="72" longitude="-116.925" municipality="Eastern Tropical Pacific" name="Off Todos Santos Bay" pageId="25" pageNumber="196" stateProvince="Baja California">Off Todos Santos Bay</location>
,
<collectingRegion id="9875E9DE7F38FFE7C257F9AEFCDD5561" box="[601,778,1569,1595]" country="Mexico" name="Baja California" pageId="25" pageNumber="196">Baja California</collectingRegion>
(
<collectingMunicipality id="BA6ABD467F38FFE7C312F9AEFBEF5566" box="[796,1080,1569,1596]" pageId="25" pageNumber="196">Eastern Tropical Pacific</collectingMunicipality>
),
<collectingCountry id="22A667AC7F38FFE7C45EF9AEFB7E5561" box="[1104,1193,1569,1595]" name="Mexico" pageId="25" pageNumber="196">Mexico</collectingCountry>
;
<collectingCounty id="B36F5FB07F38FFE7C4B6F9AEFB205561" box="[1208,1271,1569,1595]" pageId="25" pageNumber="196">Talud</collectingCounty>
XVIB cruise, sampling station 25 (
<geoCoordinate id="3F8541FB7F38FFE7C18DF9CBFE245505" box="[387,499,1604,1631]" degrees="31.805" direction="north" orientation="latitude" pageId="25" pageNumber="196" precision="55" value="31.805">31.805°N</geoCoordinate>
,
<geoCoordinate id="3F8541FB7F38FFE7C1F1F9CBFD545505" box="[511,643,1604,1631]" degrees="116.925" direction="west" orientation="longitude" pageId="25" pageNumber="196" precision="55" value="-116.925">116.925°W</geoCoordinate>
); depth
<quantity id="9D498AD97F38FFE7C2D2F9CBFCF05505" box="[732,807,1604,1631]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="25" pageNumber="196" unit="m" value="825.0">
<elevation id="D19CC00F7F38FFE7C2D2F9CBFCF05505" box="[732,807,1604,1631]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="25" pageNumber="196" unit="m" value="825.0">825 m</elevation>
</quantity>
; organic carbon content, 4.17%; organic matter content, 7.17%; sand 2.13%; clay, 10.36%; silt, 87.51
</materialsCitation>
%.
</paragraph>
<paragraph id="5A0E273C7F38FFE7C0C9F903FE275591" blockId="25.[151,1437,1568,2028]" pageId="25" pageNumber="196">
<emphasis id="68C5FB2E7F38FFE7C0C9F903FE1055FC" bold="true" box="[199,455,1676,1702]" pageId="25" pageNumber="196">Specimens examined.</emphasis>
Adult female
<typeStatus id="850A999E7F38FFE7C265F902FD1A55FD" box="[619,717,1677,1703]" pageId="25" pageNumber="196" type="holotype">holotype</typeStatus>
dissected and mounted onto ten slides (EMUCOP-260514-01);
<date id="2E0F01FC7F38FFE7C099F93EFEEF5591" box="[151,312,1713,1739]" pageId="25" pageNumber="196" value="2014-05-26">May 26, 2014</date>
; coll. S. Gómez.
</paragraph>
</subSubSection>
<subSubSection id="12AB74B77F38FFE7C0C9F95AFE8B546D" pageId="25" pageNumber="196" type="etymology">
<paragraph id="5A0E273C7F38FFE7C0C9F95AFE8B546D" blockId="25.[151,1437,1568,2028]" pageId="25" pageNumber="196">
<emphasis id="68C5FB2E7F38FFE7C0C9F95AFE9955B5" bold="true" box="[199,334,1749,1775]" pageId="25" pageNumber="196">Etymology.</emphasis>
The specific epithet from the Latin
<taxonomicName id="9DB15CBF7F38FFE7C2D6F95AFCFB55B5" authorityName="Gómez" authorityYear="2021" box="[728,812,1749,1775]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="25" pageNumber="196" phylum="Arthropoda" rank="species" species="reducta">
<emphasis id="68C5FB2E7F38FFE7C2D6F95AFCFB55B5" box="[728,812,1749,1775]" italics="true" pageId="25" pageNumber="196">reducta</emphasis>
</taxonomicName>
, reduced, makes reference to the loss of the inner seta of P1 ENP1, and to the reduced armature complement of P1 ENP2 from three to two. It is in the nominative singular. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="12AB74B77F38FFE0C0C9F8CFFB1A5449" lastPageId="30" lastPageNumber="201" pageId="25" pageNumber="196" type="description">
<paragraph id="5A0E273C7F38FFE7C0C9F8CFFE8954F9" blockId="25.[151,1437,1568,2028]" pageId="25" pageNumber="196">
<emphasis id="68C5FB2E7F38FFE7C0C9F8CFFE1B5400" bold="true" box="[199,460,1856,1882]" pageId="25" pageNumber="196">Description of female.</emphasis>
Total body length measured from tip of rostrum to posterior margin of caudal rami, 525 µm; habitus pyriform, widest at posterior end of cephalothorax, tapering posteriad (
<figureCitation id="C28A3BB97F38FFE7C44FF8EAFB7E5425" box="[1089,1193,1893,1919]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="25" pageNumber="196">Fig. 19A</figureCitation>
); cephalothorax/body length ratio, 0.35.
</paragraph>
<paragraph id="5A0E273C7F38FFE7C0C9F822FBDC549D" blockId="25.[151,1437,1568,2028]" box="[199,1035,1965,1991]" pageId="25" pageNumber="196">
Prosome and pedigerous somites (
<figureCitation id="C28A3BB97F38FFE7C248F822FD7B549D" box="[582,684,1965,1991]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="25" pageNumber="196">Fig. 19A</figureCitation>
) largely as in previous species.
</paragraph>
<paragraph id="5A0E273C7F38FFE4C0C9F85EFCA6538F" blockId="25.[151,1437,1568,2028]" lastBlockId="26.[151,1437,151,285]" lastPageId="26" lastPageNumber="197" pageId="25" pageNumber="196">
Urosome (
<figureCitation id="C28A3BB97F38FFE7C132F85EFE1554B1" box="[316,450,2001,2027]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="25" pageNumber="196">Fig. 19AC</figureCitation>
) consisting of fifth pedigerous somite (first urosomite), genital double-somite (genital— second urosomite—and third urosomites fused), two free urosomites, and anal somite. Urosomites without expansions laterally nor dorsally; integument weakly sclerotized.
</paragraph>
<paragraph id="5A0E273C7F3BFFE4C0C9FF50FF2C5247" blockId="26.[151,1437,151,285]" pageId="26" pageNumber="197">
Fifth pedigerous somite (
<figureCitation id="C28A3BB97F3BFFE4C1FEFF50FD8C53A3" box="[496,603,223,249]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="26" pageNumber="197">Fig. 19A</figureCitation>
) narrower than preceding somites; with two dorsolateral sets of small spinules.
</paragraph>
<caption id="0ECE77B47F3BFFE4C099F8F8FE6B54E9" ID-DOI="http://doi.org/10.5281/zenodo.5563607" ID-Zenodo-Dep="5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="26" pageNumber="197" startId="26.[151,250,1911,1935]" targetBox="[163,1428,319,1879]" targetPageId="26">
<paragraph id="5A0E273C7F3BFFE4C099F8F8FE6B54E9" blockId="26.[151,1436,1911,1971]" pageId="26" pageNumber="197">
<emphasis id="68C5FB2E7F3BFFE4C099F8F8FEC854D5" bold="true" box="[151,287,1911,1935]" pageId="26" pageNumber="197">FIGURE 19.</emphasis>
<taxonomicName id="9DB15CBF7F3BFFE4C12AF8F8FE0D54D5" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[292,474,1911,1935]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="26" pageNumber="197" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">
<emphasis id="68C5FB2E7F3BFFE4C12AF8F8FE0D54D5" box="[292,474,1911,1935]" italics="true" pageId="26" pageNumber="197">Delavalia reducta</emphasis>
</taxonomicName>
<emphasis id="68C5FB2E7F3BFFE4C1EEF8F8FDE754D5" bold="true" box="[480,560,1911,1935]" pageId="26" pageNumber="197">
<taxonomicNameLabel id="73F646557F3BFFE4C1EEF8F8FDE754D5" box="[480,560,1911,1935]" pageId="26" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted).
</paragraph>
</caption>
<paragraph id="5A0E273C7F3AFFE5C0C9FF18FBB452D3" blockId="27.[151,1437,151,2014]" pageId="27" pageNumber="198">
Second and third urosomites completely fused dorsally and ventrally forming genital double-somite (
<figureCitation id="C28A3BB97F3AFFE5C524FF18FA5853EB" box="[1322,1423,151,177]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19C</figureCitation>
), with dorsolateral trace of division (
<figureCitation id="C28A3BB97F3AFFE5C23AFF34FD77538F" box="[564,672,187,213]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19A</figureCitation>
); genital double-somite as long as wide, widest part measured in proximal third close to P6; proximal half of genital double-somite with sensilla and two sets of small spinules dorsally (
<figureCitation id="C28A3BB97F3AFFE5C10AFE8CFEB95247" box="[260,366,259,285]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19A</figureCitation>
), ventrally without sensilla nor spinules (
<figureCitation id="C28A3BB97F3AFFE5C343FE8CFC615247" box="[845,950,259,285]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19C</figureCitation>
); distal half of genital double-somite with posterior sensilla and dorsolateral spinular rows (
<figureCitation id="C28A3BB97F3AFFE5C2CAFEA8FCFA521B" box="[708,813,295,321]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19A</figureCitation>
), ventrally with few sensilla and without spinules (
<figureCitation id="C28A3BB97F3AFFE5C561FEA8FF1E523F" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19C</figureCitation>
); posterior hyaline fringe broad and smooth; genital complex hardly distinguishable, copulatory pores not exposed, paired genital apertures located ventrolaterally and covered by P6 (
<figureCitation id="C28A3BB97F3AFFE5C3FEFEE0FB8052D3" box="[1008,1111,367,393]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19C</figureCitation>
).
</paragraph>
<paragraph id="5A0E273C7F3AFFE5C0C9FE1CFAD452F7" blockId="27.[151,1437,151,2014]" box="[199,1283,403,429]" pageId="27" pageNumber="198">
Fourth urosomite (
<figureCitation id="C28A3BB97F3AFFE5C196FE1CFDC852F7" box="[408,543,403,429]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19A, C</figureCitation>
) as distal half of genital double-somite; no ventral pores detected.
</paragraph>
<paragraph id="5A0E273C7F3AFFE5C0C9FE38FC7D52AF" blockId="27.[151,1437,151,2014]" pageId="27" pageNumber="198">
Fifth urosomite with dorsolateral spinules and without dorsal sensilla (
<figureCitation id="C28A3BB97F3AFFE5C3D5FE38FB94528B" box="[987,1091,439,465]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19A</figureCitation>
); ventrally without sensilla but with two medial sets of spinules (
<figureCitation id="C28A3BB97F3AFFE5C201FE54FDA252AF" box="[527,629,475,501]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19C</figureCitation>
); no ventral pores detected.
</paragraph>
<paragraph id="5A0E273C7F3AFFE5C0C9FE70FE2851DF" blockId="27.[151,1437,151,2014]" pageId="27" pageNumber="198">
Anal somite about 2.2. times as wide as long (
<figureCitation id="C28A3BB97F3AFFE5C2E7FE70FCA15143" box="[745,886,511,537]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19AB</figureCitation>
); with dorsal, ventrolateral and ventral spinules around joint of caudal rami (
<figureCitation id="C28A3BB97F3AFFE5C1D0FDACFDBE5167" box="[478,617,547,573]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19AC</figureCitation>
); with spinules along medial cleft ventrally (
<figureCitation id="C28A3BB97F3AFFE5C468FDACFB195167" box="[1126,1230,547,573]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19C</figureCitation>
); with one ventral pore on each side (
<figureCitation id="C28A3BB97F3AFFE5C165FDC8FE05513B" box="[363,466,583,609]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19C</figureCitation>
); anal operculum with transverse row of minute spinules, semicircular, flanked by one sensilla on each side (
<figureCitation id="C28A3BB97F3AFFE5C182FDE4FE2551DF" box="[396,498,619,645]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19B</figureCitation>
).
</paragraph>
<paragraph id="5A0E273C7F3AFFE5C0C9FD00FB045007" blockId="27.[151,1437,151,2014]" pageId="27" pageNumber="198">
Caudal rami elongate, about 3.6 times as long as wide (
<figureCitation id="C28A3BB97F3AFFE5C321FD00FC6251F3" box="[815,949,655,681]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19AC</figureCitation>
) and about as long as fifth and anal somites combined; outer margin nearly straight, inner margin convex; with spinules at base of setae I and II, and VII, and with inner medial spinules (
<figureCitation id="C28A3BB97F3AFFE5C1C7FD58FD8751AB" box="[457,592,727,753]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19AC</figureCitation>
), with one medial pore ventrally (
<figureCitation id="C28A3BB97F3AFFE5C3CDFD58FBFF51AB" box="[963,1064,727,753]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19C</figureCitation>
); with seven elements (
<figureCitation id="C28A3BB97F3AFFE5C527FD58FA5951AB" box="[1321,1422,727,753]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19B</figureCitation>
); setae I and II subdistal, lateral, the former a small seta ventral to seta II, the latter long; seta III subdistal, arising ventrally (
<figureCitation id="C28A3BB97F3AFFE5C107FC90FE465063" box="[265,401,799,825]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="27" pageNumber="198">Fig. 19BC</figureCitation>
); setae IV and V distal, rat-tail like in distal half, with fracture plane; seta VI small, issuing at inner distal corner; dorsal seta VII triarticulate at base, situated subdistally close to inner margin.
</paragraph>
<paragraph id="5A0E273C7F3AFFE5C0C9FCE8FA5F50DB" blockId="27.[151,1437,151,2014]" box="[199,1416,871,897]" pageId="27" pageNumber="198">
Rostrum (
<figureCitation id="C28A3BB97F3AFFE5C13CFCE8FE4450DB" box="[306,403,871,897]" captionStart="FIGURE 20" captionStartId="28.[151,250,1875,1899]" captionTargetBox="[161,1425,193,1838]" captionTargetId="figure-20@28.[151,1436,181,1851]" captionTargetPageId="28" captionText="FIGURE 20. Delavalia reducta sp. nov., female: A, rostrum and antennule (microphotograph available in http://metadata. icmyl.unam.mx/handle/20.500.12201/10625); B, antenna (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10618)." figureDoi="http://doi.org/10.5281/zenodo.5563609" httpUri="https://zenodo.org/record/5563609/files/figure.png" pageId="27" pageNumber="198">Fig. 20A</figureCitation>
) trapezoidal, not fused to cephalothorax, bifid, with two subdistal sensilla, without dorsal pore.
</paragraph>
<paragraph id="5A0E273C7F3AFFE5C0C9FC04FCB4576F" blockId="27.[151,1437,151,2014]" pageId="27" pageNumber="198">
Antennule (
<figureCitation id="C28A3BB97F3AFFE5C142FC04FE6450FF" box="[332,435,907,933]" captionStart="FIGURE 20" captionStartId="28.[151,250,1875,1899]" captionTargetBox="[161,1425,193,1838]" captionTargetId="figure-20@28.[151,1436,181,1851]" captionTargetPageId="28" captionText="FIGURE 20. Delavalia reducta sp. nov., female: A, rostrum and antennule (microphotograph available in http://metadata. icmyl.unam.mx/handle/20.500.12201/10625); B, antenna (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10618)." figureDoi="http://doi.org/10.5281/zenodo.5563609" httpUri="https://zenodo.org/record/5563609/files/figure.png" pageId="27" pageNumber="198">Fig. 20A</figureCitation>
) seven-segmented; all segments smooth, except for first segment with proximal spinular row; first segment without pore. All setae smooth, except for one pinnate seta on second, third and last segments; second and third segments with one seta with fracture plane each; sixth segment with two, seventh segment with three articulated setae. Armature formula: 1(1); 2(10); 3(9); 4(5 + (1 + ae)), 5(3); 6(8); 7(5 + acro). Acrothek consisting of two setae and one minute aesthetasc fused basally.
</paragraph>
<paragraph id="5A0E273C7F3AFFE5C0C9FBB0FD925623" blockId="27.[151,1437,151,2014]" pageId="27" pageNumber="198">
Antenna (
<figureCitation id="C28A3BB97F3AFFE5C138FBB0FE495703" box="[310,414,1087,1113]" captionStart="FIGURE 20" captionStartId="28.[151,250,1875,1899]" captionTargetBox="[161,1425,193,1838]" captionTargetId="figure-20@28.[151,1436,181,1851]" captionTargetPageId="28" captionText="FIGURE 20. Delavalia reducta sp. nov., female: A, rostrum and antennule (microphotograph available in http://metadata. icmyl.unam.mx/handle/20.500.12201/10625); B, antenna (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10618)." figureDoi="http://doi.org/10.5281/zenodo.5563609" httpUri="https://zenodo.org/record/5563609/files/figure.png" pageId="27" pageNumber="198">Fig. 20B</figureCitation>
). Coxa short, with some outer spinules. Allobasis as long as free endopodal segment; with some slender spinules at base of exopod, and with few long slender inner spinules at proximal third; with one abexopodal seta arising slightly above the middle of segment. Free endopodal segment elongate; proximal half with longitudinal row of strong inner spinules, distal half with subdistal outer strong spinules, with two outer subdistal frills; armature composed of two lateral spines and two setae, distally with one inner apical element, three apical geniculate setae and one slender element, and one outer distal pinnate element fused basally to slender seta. Exopod three-segmented; first and third segments longest; first and middle segment without, third segment with spinules as shown; first and second segment with one distal seta each, third segment with one proximal and three apical setae, two of which seemingly fused basally.
</paragraph>
<paragraph id="5A0E273C7F3AFFE5C0C9FA0CFBFD552F" blockId="27.[151,1437,151,2014]" pageId="27" pageNumber="198">
Mandible (
<figureCitation id="C28A3BB97F3AFFE5C14DFA0CFE7B56C7" box="[323,428,1411,1437]" captionStart="FIGURE 21" captionStartId="29.[151,250,1813,1837]" captionTargetBox="[159,1431,191,1782]" captionTargetId="figure-15@29.[151,1436,181,1789]" captionTargetPageId="29" captionText="FIGURE 21. Delavalia reducta sp. nov., female: A, mandible (microphotograph available in http://metadata.icmyl.unam. mx/handle/20.500.12201/10622); B, maxillule; C, maxilla; D, maxilliped (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10616)." figureDoi="http://doi.org/10.5281/zenodo.5563611" httpUri="https://zenodo.org/record/5563611/files/figure.png" pageId="27" pageNumber="198">Fig. 21A</figureCitation>
). Coxa relatively short. Gnathobase wide; ventral distal corner produced into small sharp semi-hyaline process; with two strong and several smaller teeth, two spines and two setae, one of which pinnate. Basis elongate, spinular ornamentation difficult to see, not detected, with three subdistal outer setae. Exopod arising from short pedestal, one-segmented, elongate, about 2.2 times as long as wide, and 0.3 times as long as basis, with three lateral and three apical setae, none of which fused basally. Endopod recurved, twisted over exopod, with three lateral setae, and five distal elements (three slender setae, one of which spinulose and one strong element, and longest element fused to endopod basally and with hyaline flange in middle part).
</paragraph>
<paragraph id="5A0E273C7F3AFFE5C0C9F9F0FC415473" blockId="27.[151,1437,151,2014]" pageId="27" pageNumber="198">
Maxillule (
<figureCitation id="C28A3BB97F3AFFE5C148F9F0FE7855C3" box="[326,431,1663,1689]" captionStart="FIGURE 21" captionStartId="29.[151,250,1813,1837]" captionTargetBox="[159,1431,191,1782]" captionTargetId="figure-15@29.[151,1436,181,1789]" captionTargetPageId="29" captionText="FIGURE 21. Delavalia reducta sp. nov., female: A, mandible (microphotograph available in http://metadata.icmyl.unam. mx/handle/20.500.12201/10622); B, maxillule; C, maxilla; D, maxilliped (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10616)." figureDoi="http://doi.org/10.5281/zenodo.5563611" httpUri="https://zenodo.org/record/5563611/files/figure.png" pageId="27" pageNumber="198">Fig. 21B</figureCitation>
). Arthrite of praecoxa with two surface setae and some dorsal spinules; distal armature composed of seven strong spines as shown —no setiform elements detected—, and one lateral pinnate seta. Coxal endite with three setae, spinular ornamentation not detected. Basis with two endites; proximal endite with four, distal endite with three slender setae. Exopod and endopod fused basally, separated from basis, one-segmented; endopod larger than exopod, with four setae; exopod with two setae.
</paragraph>
<paragraph id="5A0E273C7F3AFFE5C0C9F8BCFED65484" blockId="27.[151,1437,151,2014]" pageId="27" pageNumber="198">
Maxilla (
<figureCitation id="C28A3BB97F3AFFE5C121F8BCFE405417" box="[303,407,1843,1869]" captionStart="FIGURE 21" captionStartId="29.[151,250,1813,1837]" captionTargetBox="[159,1431,191,1782]" captionTargetId="figure-15@29.[151,1436,181,1789]" captionTargetPageId="29" captionText="FIGURE 21. Delavalia reducta sp. nov., female: A, mandible (microphotograph available in http://metadata.icmyl.unam. mx/handle/20.500.12201/10622); B, maxillule; C, maxilla; D, maxilliped (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10616)." figureDoi="http://doi.org/10.5281/zenodo.5563611" httpUri="https://zenodo.org/record/5563611/files/figure.png" pageId="27" pageNumber="198">Fig. 21C</figureCitation>
). Large syncoxa with outer spinules as shown; with three endites; proximal endite bilobed, each lobe with one seta; middle and distal endites elongate, the latter slightly longer, with three spinulose setae each. Basis drawn out into strong claw, with strong spinulose spine and two slender setae, one of which arising from elongate setophore. Endopod one-segmented, with six slender setae (one arising basally, two medially, and three apically).
</paragraph>
<caption id="0ECE77B47F3DFFE2C099F8DCFE1454E9" ID-DOI="http://doi.org/10.5281/zenodo.5563609" ID-Zenodo-Dep="5563609" httpUri="https://zenodo.org/record/5563609/files/figure.png" pageId="28" pageNumber="199" startId="28.[151,250,1875,1899]" targetBox="[161,1425,193,1838]" targetPageId="28">
<paragraph id="5A0E273C7F3DFFE2C099F8DCFE1454E9" blockId="28.[151,1437,1875,1971]" pageId="28" pageNumber="199">
<emphasis id="68C5FB2E7F3DFFE2C099F8DCFEF45431" bold="true" box="[151,291,1875,1899]" pageId="28" pageNumber="199">FIGURE 20.</emphasis>
<taxonomicName id="9DB15CBF7F3DFFE2C122F8DCFE315431" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[300,486,1875,1899]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="28" pageNumber="199" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">
<emphasis id="68C5FB2E7F3DFFE2C122F8DCFE315431" box="[300,486,1875,1899]" italics="true" pageId="28" pageNumber="199">Delavalia reducta</emphasis>
</taxonomicName>
<emphasis id="68C5FB2E7F3DFFE2C1E1F8DCFD945431" bold="true" box="[495,579,1875,1899]" pageId="28" pageNumber="199">
<taxonomicNameLabel id="73F646557F3DFFE2C1E1F8DCFD945431" box="[495,579,1875,1899]" pageId="28" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A, rostrum and antennule (microphotograph available in http://metadata. icmyl.unam.mx/handle/20.500.12201/10625); B, antenna (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10618).
</paragraph>
</caption>
<caption id="0ECE77B47F3CFFE3C099F89AFDFE542F" ID-DOI="http://doi.org/10.5281/zenodo.5563611" ID-Zenodo-Dep="5563611" httpUri="https://zenodo.org/record/5563611/files/figure.png" pageId="29" pageNumber="200" startId="29.[151,250,1813,1837]" targetBox="[159,1431,191,1782]" targetPageId="29">
<paragraph id="5A0E273C7F3CFFE3C099F89AFDFE542F" blockId="29.[151,1437,1813,1909]" pageId="29" pageNumber="200">
<emphasis id="68C5FB2E7F3CFFE3C099F89AFEF35477" bold="true" box="[151,292,1813,1837]" pageId="29" pageNumber="200">FIGURE 21.</emphasis>
<taxonomicName id="9DB15CBF7F3CFFE3C120F89AFE3E5477" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[302,489,1813,1837]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="29" pageNumber="200" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">
<emphasis id="68C5FB2E7F3CFFE3C120F89AFE3E5477" box="[302,489,1813,1837]" italics="true" pageId="29" pageNumber="200">Delavalia reducta</emphasis>
</taxonomicName>
<emphasis id="68C5FB2E7F3CFFE3C1FDF89AFD9F5477" bold="true" box="[499,584,1813,1837]" pageId="29" pageNumber="200">
<taxonomicNameLabel id="73F646557F3CFFE3C1FDF89AFD9F5477" box="[499,584,1813,1837]" pageId="29" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A, mandible (microphotograph available in http://metadata.icmyl.unam. mx/handle/20.500.12201/10622); B, maxillule; C, maxilla; D, maxilliped (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10616).
</paragraph>
</caption>
<paragraph id="5A0E273C7F3FFFE0C0C9FF18FCCF521B" blockId="30.[151,1437,151,1365]" pageId="30" pageNumber="201">
Maxilliped (
<figureCitation id="C28A3BB97F3FFFE0C15FFF18FE6F53EB" box="[337,440,151,177]" captionStart="FIGURE 21" captionStartId="29.[151,250,1813,1837]" captionTargetBox="[159,1431,191,1782]" captionTargetId="figure-15@29.[151,1436,181,1789]" captionTargetPageId="29" captionText="FIGURE 21. Delavalia reducta sp. nov., female: A, mandible (microphotograph available in http://metadata.icmyl.unam. mx/handle/20.500.12201/10622); B, maxillule; C, maxilla; D, maxilliped (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10616)." figureDoi="http://doi.org/10.5281/zenodo.5563611" httpUri="https://zenodo.org/record/5563611/files/figure.png" pageId="30" pageNumber="201">Fig. 21D</figureCitation>
) subchelate. Syncoxa rectangular; about two times as long as wide; with medial spinules as shown; with one bare and two spinulose strong elements, of which bare seta and one spinulose element at the same level, the other arising distally from long pedestal. Basis shorter than syncoxa, oval, with some outer spinules, with one anterior and one posterior inner spinular row, with two slender distal setae subequal in length. Endopod one-segmented, with one claw-like element and one seta.
</paragraph>
<paragraph id="5A0E273C7F3FFFE0C0C9FEC4FCE95197" blockId="30.[151,1437,151,1365]" pageId="30" pageNumber="201">
P1 (
<figureCitation id="C28A3BB97F3FFFE0C0F6FEC4FEB3523F" box="[248,356,331,357]" captionStart="FIGURE 22" captionStartId="31.[151,250,1933,1957]" captionTargetBox="[157,1427,532,1903]" captionTargetId="figure-205@31.[151,1436,525,1910]" captionTargetPageId="31" captionText="FIGURE 22. Delavalia reducta sp. nov., female: A, P1, anterior (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10620); B, P2, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10621)." figureDoi="http://doi.org/10.5281/zenodo.5563613" httpUri="https://zenodo.org/record/5563613/files/figure.png" pageId="30" pageNumber="201">Fig. 22A</figureCitation>
). Coxa massive, 1.6 times as wide as long; with two outer, and one medial subdistal row of spinules. Basis with outer and inner pinnate spines, with spinules at base of endopod and at base of inner spine, and with few inner setules. Exopod three-segmented, longer than endopod; no pores detected on exopodal segments; EXP1 longest, EXP3 shortest; all segments without outer nor inner acute distal processes; EXP1 and EXP2 with longitudinal row of outer spinules, EXP1 without, EXP2 with inner seta; EXP3 with spinules at the base of setae/ spines, with two outer spines and two apical elements, of which outermost apical element spine-like, innermost apical element pinnate and rat-tail like in distal half. Endopod two-segmented, reaching middle of EXP2, segments without inner nor outer acute distal processes; no pores detected on endopodal segments; ENP1 barely reaching tip of EXP1, 1.5 times as long as wide, visibly longer than ENP2, with outer and distal spinules, without armature; ENP2 small, rectangular, about 1.4 times as long as wide, and 0.7 times as long as ENP1, with few outer spinules, with one inner and one apical seta, both strong and plumose.
</paragraph>
<paragraph id="5A0E273C7F3FFFE0C0C9FD58FD6B566B" blockId="30.[151,1437,151,1365]" pageId="30" pageNumber="201">
P2P4 (
<figureCitation id="C28A3BB97F3FFFE0C12FFD58FE4551AB" box="[289,402,727,753]" captionStart="FIGURE 22" captionStartId="31.[151,250,1933,1957]" captionTargetBox="[157,1427,532,1903]" captionTargetId="figure-205@31.[151,1436,525,1910]" captionTargetPageId="31" captionText="FIGURE 22. Delavalia reducta sp. nov., female: A, P1, anterior (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10620); B, P2, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10621)." figureDoi="http://doi.org/10.5281/zenodo.5563613" httpUri="https://zenodo.org/record/5563613/files/figure.png" pageId="30" pageNumber="201">Figs. 22B</figureCitation>
,
<figureCitation id="C28A3BB97F3FFFE0C190FD58FE2451AB" box="[414,499,727,753]" captionStart="FIGURE 23" captionStartId="32.[151,250,1873,1897]" captionTargetBox="[162,1426,191,1839]" captionTargetId="figure-20@32.[151,1436,181,1849]" captionTargetPageId="32" captionText="FIGURE 23. Delavalia reducta sp. nov., female: A, P3, anterior (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10617); B, P4, anterior (microphotograph available in http://metadata.icmyl.unam. mx/handle/20.500.12201/10623); C, P5, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10624)." figureDoi="http://doi.org/10.5281/zenodo.5563615" httpUri="https://zenodo.org/record/5563615/files/figure.png" pageId="30" pageNumber="201">23AB</figureCitation>
). Intercoxal sclerite (not shown) not transversely elongate, trapezoidal, with strong pointed process on distal outer corners, without surface ornamentation. Coxa with outer spinules proximally and subdistally. Basis with strong acute process between rami and at inner distal corner, the latter larger, of P2 largest, of P4 smallest; seemingly without spinular ornamentation. Exopod three-segmented, of P3 slightly shorter than, of P4 visibly longer than ENP, relative length of P2 EXP uncertain due to bad condition of endopod; pores detected on last exopodal segments only; EXP1 and EXP2 with outer distal process (of P4 less developed), with longitudinal row of outer spinules, of P2 and P3 with, of P4 seemingly without inner distal frill, with inner seta; EXP3 with distal processes as shown, with few spinules proximally, with three outer spines and two apical setae, of P2 with two, of P3 and P4 with three inner setae. Endopod three-segmented (P2 badly damaged and second and third endopodal segments lost during dissection), of P2 most probably longer than, of P3 slightly longer than EXP, of P4 reaching proximal third of EXP3; pores present on the last endopodal segments of P3 and P4 (P2 ENP23 unknown); P3P4 ENP1 shortest, ENP3 longest; ENP1 and ENP2 (of P2 unknown) with outer acute and inner small distal process, outer process of ENP2 (at least of P3 and P4) visibly longer, outer process of P4 ENP2 bifurcate, most probably aberrant, distal processes of ENP3 as shown; ENP1 with inner element, of P2 a slender seta, of P3 and P4 a stiff element with inner margin pinnate, of P4 very long; armature of P2 ENP2 unknown (most probably two setae), P3 and P4 ENP2 with one inner seta; P2 ENP3 unknown, P3P4 ENP3 with one apical outer spine, two apical elements, and three (P3), or two (P4) inner setae.
</paragraph>
<paragraph id="5A0E273C7F3FFFE0C0C9FAB4FD67560F" blockId="30.[151,1437,151,1365]" box="[199,688,1339,1365]" pageId="30" pageNumber="201">Setal formula of swimming legs as follows:</paragraph>
<paragraph id="5A0E273C7F3FFFE0C205FA06FA9556AB" pageId="30" pageNumber="201">
<table id="28B1D59C7F3F0001C0A8FA06FA9556AB" box="[166,1346,1417,1521]" colsContinueFrom="25.[166,1340,985,1089]" colsContinueIn="42.[166,1347,481,585]" gridcols="5" gridrows="3" pageId="30" pageNumber="201">
<tr id="E481257E7F3F0001C0A8FA06FA9556FB" box="[166,1346,1417,1441]" gridrow="0" pageId="30" pageNumber="201" rowspan-0="1">
<th id="A7504C027F3F0001C1FDFA06FDE856FB" box="[499,575,1417,1441]" gridcol="1" gridrow="0" pageId="30" pageNumber="201">P1</th>
<th id="A7504C027F3F0001C2FAFA06FCE856FB" box="[756,831,1417,1441]" gridcol="2" gridrow="0" pageId="30" pageNumber="201">P2</th>
<th id="A7504C027F3F0001C3FBFA06FB9656FB" box="[1013,1089,1417,1441]" gridcol="3" gridrow="0" pageId="30" pageNumber="201">P3</th>
<th id="A7504C027F3F0001C4F8FA06FA9556FB" box="[1270,1346,1417,1441]" gridcol="4" gridrow="0" pageId="30" pageNumber="201">P4</th>
</tr>
<tr id="E481257E7F3F0001C0A8FA3EFA955693" box="[166,1346,1457,1481]" gridrow="1" pageId="30" pageNumber="201">
<th id="A7504C027F3F0001C0A8FA3EFF015693" box="[166,214,1457,1481]" gridcol="0" gridrow="1" pageId="30" pageNumber="201">EXP</th>
<td id="A7504C027F3F0001C1FDFA3EFDE85693" box="[499,575,1457,1481]" gridcol="1" gridrow="1" pageId="30" pageNumber="201">0,1,022</td>
<td id="A7504C027F3F0001C2FAFA3EFCE85693" box="[756,831,1457,1481]" gridcol="2" gridrow="1" pageId="30" pageNumber="201">1,1,223</td>
<td id="A7504C027F3F0001C3FBFA3EFB965693" box="[1013,1089,1457,1481]" gridcol="3" gridrow="1" pageId="30" pageNumber="201">1,1,323</td>
<td id="A7504C027F3F0001C4F8FA3EFA955693" box="[1270,1346,1457,1481]" gridcol="4" gridrow="1" pageId="30" pageNumber="201">1,1,323</td>
</tr>
<tr id="E481257E7F3F0001C0A8FA56FA9556AB" box="[166,1346,1497,1521]" gridrow="2" pageId="30" pageNumber="201">
<th id="A7504C027F3F0001C0A8FA56FF0156AB" box="[166,214,1497,1521]" gridcol="0" gridrow="2" pageId="30" pageNumber="201">ENP</th>
<td id="A7504C027F3F0001C1FDFA56FDE856AB" box="[499,575,1497,1521]" gridcol="1" gridrow="2" pageId="30" pageNumber="201">0,110</td>
<td id="A7504C027F3F0001C2FAFA56FCE856AB" box="[756,831,1497,1521]" gridcol="2" gridrow="2" pageId="30" pageNumber="201">1,?,?</td>
<td id="A7504C027F3F0001C3FBFA56FB9656AB" box="[1013,1089,1497,1521]" gridcol="3" gridrow="2" pageId="30" pageNumber="201">1,1,321</td>
<td id="A7504C027F3F0001C4F8FA56FA9556AB" box="[1270,1346,1497,1521]" gridcol="4" gridrow="2" pageId="30" pageNumber="201">1,1,221</td>
</tr>
</table>
</paragraph>
<paragraph id="5A0E273C7F3FFFE0C0C9F9AEFC2555D9" blockId="30.[151,1437,1569,2028]" pageId="30" pageNumber="201">
P5 (
<figureCitation id="C28A3BB97F3FFFE0C0FBF9AEFE895561" box="[245,350,1569,1595]" captionStart="FIGURE 23" captionStartId="32.[151,250,1873,1897]" captionTargetBox="[162,1426,191,1839]" captionTargetId="figure-20@32.[151,1436,181,1849]" captionTargetPageId="32" captionText="FIGURE 23. Delavalia reducta sp. nov., female: A, P3, anterior (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10617); B, P4, anterior (microphotograph available in http://metadata.icmyl.unam. mx/handle/20.500.12201/10623); C, P5, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10624)." figureDoi="http://doi.org/10.5281/zenodo.5563615" httpUri="https://zenodo.org/record/5563615/files/figure.png" pageId="30" pageNumber="201">Fig. 23C</figureCitation>
). Baseoendopod pentagonal, transversely elongate; endopodal lobe poorly-developed, with four setae, of which outermost shortest and set closely to adjacent seta, all setae naked, except for innermost seta pinnate. Exopod oval, with some outer spinules proximally, with five setae as shown.
</paragraph>
<paragraph id="5A0E273C7F3FFFE0C0C9F902FD455591" blockId="30.[151,1437,1569,2028]" pageId="30" pageNumber="201">
P6 (
<figureCitation id="C28A3BB97F3FFFE0C0F9F902FE8A55FD" box="[247,349,1677,1703]" captionStart="FIGURE 19" captionStartId="26.[151,250,1911,1935]" captionTargetBox="[163,1428,319,1879]" captionTargetId="figure-56@26.[151,1436,309,1887]" captionTargetPageId="26" captionText="FIGURE 19. Delavalia reducta sp. nov., female:A, habitus, dorsal; B, anal somite and caudal rami, dorsal; C, urosome, ventral (P5-bearing somite omitted)." figureDoi="http://doi.org/10.5281/zenodo.5563607" httpUri="https://zenodo.org/record/5563607/files/figure.png" pageId="30" pageNumber="201">Fig. 19C</figureCitation>
) represented by a minute flap covering ventrolateral genital aperture, fused to somite, without surface ornamentation, with one slender seta.
</paragraph>
<paragraph id="5A0E273C7F3FFFE0C0C9F95AFE5F55B5" blockId="30.[151,1437,1569,2028]" box="[199,392,1749,1775]" pageId="30" pageNumber="201">
<emphasis id="68C5FB2E7F3FFFE0C0C9F95AFEDC55B5" bold="true" box="[199,267,1749,1775]" pageId="30" pageNumber="201">Male.</emphasis>
Unknown.
</paragraph>
<paragraph id="5A0E273C7F3FFFE0C0C9F977FB1A5449" blockId="30.[151,1437,1569,2028]" box="[199,1229,1784,1811]" pageId="30" pageNumber="201">
<emphasis id="68C5FB2E7F3FFFE0C0C9F977FE9A5448" bold="true" box="[199,333,1784,1810]" pageId="30" pageNumber="201">Variability.</emphasis>
No variability was detected in the single female found in the sediment samples.
</paragraph>
</subSubSection>
<subSubSection id="12AB74B77F3FFFDCC0C9F892FDF3521A" lastPageId="34" lastPageNumber="205" pageId="30" pageNumber="201" type="discussion">
<paragraph id="5A0E273C7F3FFFE1C0C9F892FAF052AF" blockId="30.[151,1437,1569,2028]" lastBlockId="31.[151,1437,150,501]" lastPageId="31" lastPageNumber="202" pageId="30" pageNumber="201">
<emphasis id="68C5FB2E7F3FFFE0C0C9F892FEEC546D" bold="true" box="[199,315,1821,1847]" pageId="30" pageNumber="201">Remarks.</emphasis>
The genus
<taxonomicName id="9DB15CBF7F3FFFE0C1CCF892FDE7546D" authorityName="Brady" authorityYear="1869" box="[450,560,1821,1847]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="30" pageNumber="201" phylum="Arthropoda" rank="genus">
<emphasis id="68C5FB2E7F3FFFE0C1CCF892FDE7546D" box="[450,560,1821,1847]" italics="true" pageId="30" pageNumber="201">Delavalia</emphasis>
</taxonomicName>
is the most diverse genus within
<taxonomicName id="9DB15CBF7F3FFFE0C3BFF892FBEC546D" authorityName="Brady" authorityYear="1880" box="[945,1083,1821,1847]" class="Hexanauplia" family="Miraciidae" kingdom="Animalia" order="Harpacticoida" pageId="30" pageNumber="230" phylum="Arthropoda" rank="subFamily" subFamily="Stenheliinae">Stenheliinae</taxonomicName>
both in number of species and morphologically. Due to the latter, the monophyly of the genus has been questioned repeatedly (
<emphasis id="68C5FB2E7F3FFFE0C4FBF8CDFACA5401" box="[1269,1309,1858,1883]" italics="true" pageId="30" pageNumber="201">e.g.</emphasis>
<bibRefCitation id="3E205ACD7F3FFFE0C526F8CEFEFC5425" author="Karanovic, T. &amp; Kim, K." pageId="30" pageNumber="201" pagination="1 - 96" refId="ref30851" refString="Karanovic, T. &amp; Kim, K. (2014) New insights into polyphyly of the harpacticoid genus Delavalia (Crustacea, Copepoda) through morphological and molecular study of an unprecedented diversity of sympatric species in a small South Korean bay. Zootaxa, 3783 (1), 1 - 96. https: // doi. org / 10.11646 / zootaxa. 3783.1.1" type="journal article" year="2014">Karanovic &amp; Kim 2014</bibRefCitation>
;
<bibRefCitation id="3E205ACD7F3FFFE0C139F8EAFDD45425" author="Mu, F. &amp; Huys, R." box="[311,515,1893,1919]" pageId="30" pageNumber="201" pagination="179 - 206" refId="ref31326" refString="Mu, F. &amp; Huys, R. (2002) New species of Stenhelia (Copepoda, Harpacticoida, Diosaccidae) from the Bohai Sea (China) with notes on subgeneric division and phylogenetic relationships. Cahiers de Biologie marine, 43, 179 - 206. https: // doi. org / 10.21411 / CBM. A. C 482 EC 6 A" type="journal article" year="2002">Mu &amp; Huys 2002</bibRefCitation>
;
<bibRefCitation id="3E205ACD7F3FFFE0C200F8EAFD4B5425" author="Willen, E." box="[526,668,1893,1919]" pageId="30" pageNumber="201" refId="ref32556" refString="Willen, E. (2000) Phylogeny of the Thalestridimorpha Lang, 1944 (Crustacea, Copepoda). Cuvillier Verlag, Gottingen, 233 pp." type="book" year="2000">Willen 2000</bibRefCitation>
,
<bibRefCitation id="3E205ACD7F3FFFE0C2A6F8EAFD345425" author="Willen, E." box="[680,739,1893,1919]" pageId="30" pageNumber="201" pagination="27 - 42" refId="ref32584" refString="Willen, E. (2002) Notes on the systematic position of the Stenheliinae (Copepoda, Harpacticoida) within the Thalestridimorpha and description of two new species from Motupore Island, Papua New Guinea. Cahiers de Biologie marine, 43, 27 - 42. https: // doi. org / 10.21411 / CBM. A. FCF 07 EC 3" type="journal article" year="2002">2002</bibRefCitation>
,
<bibRefCitation id="3E205ACD7F3FFFE0C2E1F8EAFCFB5425" author="Willen, E." box="[751,812,1893,1919]" pageId="30" pageNumber="201" pagination="1691 - 1711" refId="ref32649" refString="Willen, E. (2003) A new species of Stenhelia (Copepoda, Harpacticoida) from a hydrothermal, active, submarine volcano in the New Ireland Fore-Arc system (Papua New Guinea) with notes on deep sea colonization within the Stenheliinae. Journal of natural History, 37, 1691 - 1711. https: // doi. org / 10.1080 / 00222930110114437" type="journal article" year="2003">2003</bibRefCitation>
). Several studies have focused their efforts towards the monophyly of the genus and several authors have contributed importantly (see
<bibRefCitation id="3E205ACD7F3FFFE0C401F806FA8954F9" author="Gomez, S. &amp; Cruz-Barraza, J. A." box="[1039,1374,1929,1955]" pageId="30" pageNumber="201" pagination="117 - 150" refId="ref30471" refString="Gomez, S. &amp; Cruz-Barraza, J. A. (2021) On some deep-sea Stenheliinae from the Gulf of California and the west coast of the Baja California Peninsula (Mexico): Wellstenhelia euterpoides sp. nov., and Wellstenvalia wellsi gen. et sp. nov. (Copepoda: Harpacticoida: Miraciidae). Zootaxa, 5051 (1), 117 - 150. https: // doi. org / 10.11646 / zootaxa. 5051.1.10" type="journal article" year="2021">Gómez &amp; Cruz-Barraza 2021</bibRefCitation>
). The disparity in maxilliped structure (
<bibRefCitation id="3E205ACD7F3FFFE0C20BF822FC98549D" author="Gomez, S. &amp; Cruz-Barraza, J. A." box="[517,847,1965,1991]" pageId="30" pageNumber="201" pagination="117 - 150" refId="ref30471" refString="Gomez, S. &amp; Cruz-Barraza, J. A. (2021) On some deep-sea Stenheliinae from the Gulf of California and the west coast of the Baja California Peninsula (Mexico): Wellstenhelia euterpoides sp. nov., and Wellstenvalia wellsi gen. et sp. nov. (Copepoda: Harpacticoida: Miraciidae). Zootaxa, 5051 (1), 117 - 150. https: // doi. org / 10.11646 / zootaxa. 5051.1.10" type="journal article" year="2021">Gómez &amp; Cruz-Barraza 2021</bibRefCitation>
), the different positions of caudal setae I and II (either subdistally or more proximally) (
<bibRefCitation id="3E205ACD7F3FFFE0C207F85EFC8E54B1" author="Gomez, S. &amp; Cruz-Barraza, J. A." box="[521,857,2001,2027]" pageId="30" pageNumber="201" pagination="117 - 150" refId="ref30471" refString="Gomez, S. &amp; Cruz-Barraza, J. A. (2021) On some deep-sea Stenheliinae from the Gulf of California and the west coast of the Baja California Peninsula (Mexico): Wellstenhelia euterpoides sp. nov., and Wellstenvalia wellsi gen. et sp. nov. (Copepoda: Harpacticoida: Miraciidae). Zootaxa, 5051 (1), 117 - 150. https: // doi. org / 10.11646 / zootaxa. 5051.1.10" type="journal article" year="2021">Gómez &amp; Cruz-Barraza 2021</bibRefCitation>
), the disparity in structure and armature of the male P5 (
<bibRefCitation id="3E205ACD7F3EFFE1C0CAFF18FE4653EB" author="Mu, F. &amp; Huys, R." box="[196,401,151,177]" pageId="31" pageNumber="202" pagination="179 - 206" refId="ref31326" refString="Mu, F. &amp; Huys, R. (2002) New species of Stenhelia (Copepoda, Harpacticoida, Diosaccidae) from the Bohai Sea (China) with notes on subgeneric division and phylogenetic relationships. Cahiers de Biologie marine, 43, 179 - 206. https: // doi. org / 10.21411 / CBM. A. C 482 EC 6 A" type="journal article" year="2002">Mu &amp; Huys 2002</bibRefCitation>
;
<bibRefCitation id="3E205ACD7F3EFFE1C192FF18FDFC53EB" author="Willen, E." box="[412,555,151,177]" pageId="31" pageNumber="202" pagination="1691 - 1711" refId="ref32649" refString="Willen, E. (2003) A new species of Stenhelia (Copepoda, Harpacticoida) from a hydrothermal, active, submarine volcano in the New Ireland Fore-Arc system (Papua New Guinea) with notes on deep sea colonization within the Stenheliinae. Journal of natural History, 37, 1691 - 1711. https: // doi. org / 10.1080 / 00222930110114437" type="journal article" year="2003">Willen 2003</bibRefCitation>
), the different shapes of the anal operculum (
<bibRefCitation id="3E205ACD7F3EFFE1C427FF18FB2153EB" author="Mu, F. &amp; Huys, R." box="[1065,1270,151,177]" pageId="31" pageNumber="202" pagination="179 - 206" refId="ref31326" refString="Mu, F. &amp; Huys, R. (2002) New species of Stenhelia (Copepoda, Harpacticoida, Diosaccidae) from the Bohai Sea (China) with notes on subgeneric division and phylogenetic relationships. Cahiers de Biologie marine, 43, 179 - 206. https: // doi. org / 10.21411 / CBM. A. C 482 EC 6 A" type="journal article" year="2002">Mu &amp; Huys 2002</bibRefCitation>
;
<bibRefCitation id="3E205ACD7F3EFFE1C50FFF18FA4753EB" author="Willen, E." box="[1281,1424,151,177]" pageId="31" pageNumber="202" pagination="1691 - 1711" refId="ref32649" refString="Willen, E. (2003) A new species of Stenhelia (Copepoda, Harpacticoida) from a hydrothermal, active, submarine volcano in the New Ireland Fore-Arc system (Papua New Guinea) with notes on deep sea colonization within the Stenheliinae. Journal of natural History, 37, 1691 - 1711. https: // doi. org / 10.1080 / 00222930110114437" type="journal article" year="2003">Willen 2003</bibRefCitation>
), the different shapes of caudal seta I (either a spine or a small seta) (
<bibRefCitation id="3E205ACD7F3EFFE1C3ABFF34FB2B538F" author="Gomez, S. &amp; Cruz-Barraza, J. A." box="[933,1276,187,213]" pageId="31" pageNumber="202" pagination="117 - 150" refId="ref30471" refString="Gomez, S. &amp; Cruz-Barraza, J. A. (2021) On some deep-sea Stenheliinae from the Gulf of California and the west coast of the Baja California Peninsula (Mexico): Wellstenhelia euterpoides sp. nov., and Wellstenvalia wellsi gen. et sp. nov. (Copepoda: Harpacticoida: Miraciidae). Zootaxa, 5051 (1), 117 - 150. https: // doi. org / 10.11646 / zootaxa. 5051.1.10" type="journal article" year="2021">Gómez &amp; Cruz-Barraza 2021</bibRefCitation>
), the different shapes of the 2-segmented endopod of P1, the disparity in swimming leg pattern, and morphology of the dimorphic endopod of the male P2 (
<bibRefCitation id="3E205ACD7F3EFFE1C1BDFE8CFDAA5247" author="Mu, F. &amp; Huys, R." box="[435,637,259,285]" pageId="31" pageNumber="202" pagination="179 - 206" refId="ref31326" refString="Mu, F. &amp; Huys, R. (2002) New species of Stenhelia (Copepoda, Harpacticoida, Diosaccidae) from the Bohai Sea (China) with notes on subgeneric division and phylogenetic relationships. Cahiers de Biologie marine, 43, 179 - 206. https: // doi. org / 10.21411 / CBM. A. C 482 EC 6 A" type="journal article" year="2002">Mu &amp; Huys 2002</bibRefCitation>
) in different species of
<taxonomicName id="9DB15CBF7F3EFFE1C386FE8CFC215247" authorityName="Brady" authorityYear="1869" box="[904,1014,259,285]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="31" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="68C5FB2E7F3EFFE1C386FE8CFC215247" box="[904,1014,259,285]" italics="true" pageId="31" pageNumber="202">Delavalia</emphasis>
</taxonomicName>
support the di- or polyphyletic status of the genus (
<bibRefCitation id="3E205ACD7F3EFFE1C123FEA8FE20521B" author="Mu, F. &amp; Huys, R." box="[301,503,295,321]" pageId="31" pageNumber="202" pagination="179 - 206" refId="ref31326" refString="Mu, F. &amp; Huys, R. (2002) New species of Stenhelia (Copepoda, Harpacticoida, Diosaccidae) from the Bohai Sea (China) with notes on subgeneric division and phylogenetic relationships. Cahiers de Biologie marine, 43, 179 - 206. https: // doi. org / 10.21411 / CBM. A. C 482 EC 6 A" type="journal article" year="2002">Mu &amp; Huys 2002</bibRefCitation>
). Some authors have subdivided the genus
<taxonomicName id="9DB15CBF7F3EFFE1C3DBFEA8FB94521B" authorityName="Brady" authorityYear="1869" box="[981,1091,295,321]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="31" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="68C5FB2E7F3EFFE1C3DBFEA8FB94521B" box="[981,1091,295,321]" italics="true" pageId="31" pageNumber="202">Delavalia</emphasis>
</taxonomicName>
with diagnostic purposes only.
<bibRefCitation id="3E205ACD7F3EFFE1C099FEC4FEE4523F" author="Willen, E." box="[151,307,331,357]" pageId="31" pageNumber="202" pagination="1691 - 1711" refId="ref32649" refString="Willen, E. (2003) A new species of Stenhelia (Copepoda, Harpacticoida) from a hydrothermal, active, submarine volcano in the New Ireland Fore-Arc system (Papua New Guinea) with notes on deep sea colonization within the Stenheliinae. Journal of natural History, 37, 1691 - 1711. https: // doi. org / 10.1080 / 00222930110114437" type="journal article" year="2003">Willen (2003)</bibRefCitation>
subdivided the genus
<taxonomicName id="9DB15CBF7F3EFFE1C227FEC4FD40523F" authorityName="Brady" authorityYear="1869" box="[553,663,331,357]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="31" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="68C5FB2E7F3EFFE1C227FEC4FD40523F" box="[553,663,331,357]" italics="true" pageId="31" pageNumber="202">Delavalia</emphasis>
</taxonomicName>
into a number of groups and subgroups based on the shape of the anal operculum, on the combination of a specialized setation pattern on the female P5 and presence/reduction/absence of the distal inner setae on P2P4 EXP3, shape of the male and female P5, and reduction of the setation of swimming legs. In their key to the genera of
<taxonomicName id="9DB15CBF7F3EFFE1C203FE38FD4D528B" authorityName="Brady" authorityYear="1880" box="[525,666,439,465]" class="Hexanauplia" family="Miraciidae" kingdom="Animalia" order="Harpacticoida" pageId="31" pageNumber="230" phylum="Arthropoda" rank="subFamily" subFamily="Stenheliinae">Stenheliinae</taxonomicName>
,
<bibRefCitation id="3E205ACD7F3EFFE1C2ABFE38FCAC528B" author="Huys, R. &amp; Mu, F. H." box="[677,891,439,465]" pageId="31" pageNumber="202" pagination="51 - 68" refId="ref30681" refString="Huys, R. &amp; Mu, F. H. (2008) Description of a new species of Onychostenhelia Ito (Copepoda, Harpacticoida, Miraciidae) from the Bohai Sea, China. Zootaxa, 1706 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 1706.1.2" type="journal article" year="2008">Huys &amp; Mu (2008)</bibRefCitation>
subdivided the genus
<taxonomicName id="9DB15CBF7F3EFFE1C47AFE38FB35528B" authorityName="Brady" authorityYear="1869" box="[1140,1250,439,465]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="31" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="68C5FB2E7F3EFFE1C47AFE38FB35528B" box="[1140,1250,439,465]" italics="true" pageId="31" pageNumber="202">Delavalia</emphasis>
</taxonomicName>
into four groups based on the segmentation pattern of the antennary exopod and number of outer spines on P2P4 EXP3.
</paragraph>
<caption id="0ECE77B47F3EFFE1C099F802FE1454B7" ID-DOI="http://doi.org/10.5281/zenodo.5563613" ID-Zenodo-Dep="5563613" httpUri="https://zenodo.org/record/5563613/files/figure.png" pageId="31" pageNumber="202" startId="31.[151,250,1933,1957]" targetBox="[157,1427,532,1903]" targetPageId="31">
<paragraph id="5A0E273C7F3EFFE1C099F802FE1454B7" blockId="31.[151,1437,1933,2029]" pageId="31" pageNumber="202">
<emphasis id="68C5FB2E7F3EFFE1C099F802FEF054FF" bold="true" box="[151,295,1933,1957]" pageId="31" pageNumber="202">FIGURE 22.</emphasis>
<taxonomicName id="9DB15CBF7F3EFFE1C13AF802FE2654FF" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[308,497,1933,1957]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="31" pageNumber="202" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">
<emphasis id="68C5FB2E7F3EFFE1C13AF802FE2654FF" box="[308,497,1933,1957]" italics="true" pageId="31" pageNumber="202">Delavalia reducta</emphasis>
</taxonomicName>
<emphasis id="68C5FB2E7F3EFFE1C1F0F802FD8254FF" bold="true" box="[510,597,1933,1957]" pageId="31" pageNumber="202">
<taxonomicNameLabel id="73F646557F3EFFE1C1F0F802FD8254FF" box="[510,597,1933,1957]" pageId="31" pageNumber="202" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A, P1, anterior (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10620); B, P2, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10621).
</paragraph>
</caption>
<caption id="0ECE77B47F01FFDEC099F8DEFE14548F" ID-DOI="http://doi.org/10.5281/zenodo.5563615" ID-Zenodo-Dep="5563615" httpUri="https://zenodo.org/record/5563615/files/figure.png" pageId="32" pageNumber="203" startId="32.[151,250,1873,1897]" targetBox="[162,1426,191,1839]" targetPageId="32">
<paragraph id="5A0E273C7F01FFDEC099F8DEFE14548F" blockId="32.[151,1437,1873,2005]" pageId="32" pageNumber="203">
<emphasis id="68C5FB2E7F01FFDEC099F8DEFEF05433" bold="true" box="[151,295,1873,1897]" pageId="32" pageNumber="203">FIGURE 23.</emphasis>
<taxonomicName id="9DB15CBF7F01FFDEC13AF8DEFE265433" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[308,497,1873,1897]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="32" pageNumber="203" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">
<emphasis id="68C5FB2E7F01FFDEC13AF8DEFE265433" box="[308,497,1873,1897]" italics="true" pageId="32" pageNumber="203">Delavalia reducta</emphasis>
</taxonomicName>
<emphasis id="68C5FB2E7F01FFDEC1F0F8DEFD825433" bold="true" box="[510,597,1873,1897]" pageId="32" pageNumber="203">
<taxonomicNameLabel id="73F646557F01FFDEC1F0F8DEFD825433" box="[510,597,1873,1897]" pageId="32" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A, P3, anterior (microphotograph available in http://metadata.icmyl. unam.mx/handle/20.500.12201/10617); B, P4, anterior (microphotograph available in http://metadata.icmyl.unam. mx/handle/20.500.12201/10623); C, P5, anterior (microphotograph available in http://metadata.icmyl.unam.mx/ handle/20.500.12201/10624).
</paragraph>
</caption>
<paragraph id="5A0E273C7F00FFDFC0C9FF18FE4951B3" blockId="33.[151,1437,151,2022]" pageId="33" pageNumber="204">
The species described above were attributed to
<bibRefCitation id="3E205ACD7F00FFDFC2EFFF18FC4553EB" author="Willen, E." box="[737,914,151,177]" pageId="33" pageNumber="204" pagination="1691 - 1711" refId="ref32649" refString="Willen, E. (2003) A new species of Stenhelia (Copepoda, Harpacticoida) from a hydrothermal, active, submarine volcano in the New Ireland Fore-Arc system (Papua New Guinea) with notes on deep sea colonization within the Stenheliinae. Journal of natural History, 37, 1691 - 1711. https: // doi. org / 10.1080 / 00222930110114437" type="journal article" year="2003">Willens (2003)</bibRefCitation>
<taxonomicName id="9DB15CBF7F00FFDFC394FF18FBF953EB" baseAuthorityName="Boeck" baseAuthorityYear="1873" box="[922,1070,151,177]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longicaudata">
<emphasis id="68C5FB2E7F00FFDFC394FF18FBF953EB" box="[922,1070,151,177]" italics="true" pageId="33" pageNumber="204">longicaudata</emphasis>
</taxonomicName>
-group based on the apomorphic endopod of P
<quantity id="9D498AD97F00FFDFC13EFF34FE80538F" box="[304,343,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="33" pageNumber="204" unit="in" value="1.0">1 in</quantity>
which ENP1 is longer than ENP2, the latter being short and with one multiplumose and flagellate apical seta. To the
<taxonomicName id="9DB15CBF7F00FFDFC165FF51FE2853A2" baseAuthorityName="Boeck" baseAuthorityYear="1873" box="[363,511,222,248]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longicaudata">
<emphasis id="68C5FB2E7F00FFDFC165FF51FE2853A2" box="[363,511,222,248]" italics="true" pageId="33" pageNumber="204">longicaudata</emphasis>
</taxonomicName>
-group belong
<emphasis id="68C5FB2E7F00FFDFC2A8FF50FD1953A2" box="[678,718,223,248]" italics="true" pageId="33" pageNumber="204">De.</emphasis>
<taxonomicName id="9DB15CBF7F00FFDFC2DBFF51FC8353A2" authority=", De." authorityName="De." box="[725,852,222,249]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="noodti">
<emphasis id="68C5FB2E7F00FFDFC2DBFF51FCCA53A2" box="[725,797,222,248]" italics="true" pageId="33" pageNumber="204">noodti</emphasis>
,
<emphasis id="68C5FB2E7F00FFDFC322FF50FC8353A2" box="[812,852,223,248]" italics="true" pageId="33" pageNumber="204">De.</emphasis>
</taxonomicName>
<taxonomicName id="9DB15CBF7F00FFDFC355FF51FC2E53A2" authority=", De." authorityName="De." box="[859,1017,222,249]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="islandica">
<emphasis id="68C5FB2E7F00FFDFC355FF51FC1553A2" box="[859,962,222,248]" italics="true" pageId="33" pageNumber="204">islandica</emphasis>
,
<emphasis id="68C5FB2E7F00FFDFC3DFFF50FC2E53A2" box="[977,1017,223,248]" italics="true" pageId="33" pageNumber="204">De.</emphasis>
</taxonomicName>
<taxonomicName id="9DB15CBF7F00FFDFC40EFF51FBBE53A2" authority=", De." authorityName="De." box="[1024,1129,222,249]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="lima">
<emphasis id="68C5FB2E7F00FFDFC40EFF51FBE553A2" box="[1024,1074,222,248]" italics="true" pageId="33" pageNumber="204">lima</emphasis>
,
<emphasis id="68C5FB2E7F00FFDFC44FFF50FBBE53A2" box="[1089,1129,223,248]" italics="true" pageId="33" pageNumber="204">De.</emphasis>
</taxonomicName>
<taxonomicName id="9DB15CBF7F00FFDFC47FFF51FAC753A2" authority=", De." authorityName="De." box="[1137,1296,222,249]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="diegensis">
<emphasis id="68C5FB2E7F00FFDFC47FFF51FB0E53A2" box="[1137,1241,222,248]" italics="true" pageId="33" pageNumber="204">diegensis</emphasis>
,
<emphasis id="68C5FB2E7F00FFDFC4E6FF50FAC753A2" box="[1256,1296,223,248]" italics="true" pageId="33" pageNumber="204">De.</emphasis>
</taxonomicName>
<taxonomicName id="9DB15CBF7F00FFDFC519FF51FF685241" authority=", De." authorityName="De." class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longipilosa">
<emphasis id="68C5FB2E7F00FFDFC519FF51FA4253A2" box="[1303,1429,222,248]" italics="true" pageId="33" pageNumber="204">longipilosa</emphasis>
,
<emphasis id="68C5FB2E7F00FFDFC099FE8DFF685241" box="[151,191,258,283]" italics="true" pageId="33" pageNumber="204">De.</emphasis>
</taxonomicName>
<taxonomicName id="9DB15CBF7F00FFDFC0C8FE8DFE465241" authority=", De." authorityName="De." box="[198,401,258,284]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longicaudata">
<emphasis id="68C5FB2E7F00FFDFC0C8FE8DFE8D5246" box="[198,346,258,284]" italics="true" pageId="33" pageNumber="204">longicaudata</emphasis>
,
<emphasis id="68C5FB2E7F00FFDFC167FE8DFE465241" box="[361,401,258,283]" italics="true" pageId="33" pageNumber="204">De.</emphasis>
</taxonomicName>
<taxonomicName id="9DB15CBF7F00FFDFC196FE8DFD965241" authority=", De." authorityName="De." box="[408,577,258,284]" class="Hexanauplia" family="Miraciidae" genus="Stenhelia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="coineauae">
<emphasis id="68C5FB2E7F00FFDFC196FE8DFDDC5241" box="[408,523,258,283]" italics="true" pageId="33" pageNumber="204">coineauae</emphasis>
,
<emphasis id="68C5FB2E7F00FFDFC217FE8DFD965241" box="[537,577,258,283]" italics="true" pageId="33" pageNumber="204">De.</emphasis>
</taxonomicName>
<taxonomicName id="9DB15CBF7F00FFDFC246FE8DFD2F5241" authority=", De." authorityName="De." box="[584,760,258,284]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="68C5FB2E7F00FFDFC246FE8DFD165246" box="[584,705,258,284]" italics="true" pageId="33" pageNumber="204">intermedia</emphasis>
,
<emphasis id="68C5FB2E7F00FFDFC2DEFE8DFD2F5241" box="[720,760,258,283]" italics="true" pageId="33" pageNumber="204">De.</emphasis>
</taxonomicName>
<taxonomicName id="9DB15CBF7F00FFDFC2F1FE8DFC755246" baseAuthorityName="Wells" baseAuthorityYear="1965" box="[767,930,258,284]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="mastigochaeta">
<emphasis id="68C5FB2E7F00FFDFC2F1FE8DFC755246" box="[767,930,258,284]" italics="true" pageId="33" pageNumber="204">mastigochaeta</emphasis>
</taxonomicName>
and
<emphasis id="68C5FB2E7F00FFDFC3D4FE8DFBD55241" box="[986,1026,258,283]" italics="true" pageId="33" pageNumber="204">De.</emphasis>
<taxonomicName id="9DB15CBF7F00FFDFC407FE8DFAE55246" authority="(Willen 2003)" baseAuthorityName="Willen" baseAuthorityYear="2003" box="[1033,1330,258,284]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="nuwukensis">
<emphasis id="68C5FB2E7F00FFDFC407FE8DFB5D5246" box="[1033,1162,258,284]" italics="true" pageId="33" pageNumber="204">nuwukensis</emphasis>
(
<bibRefCitation id="3E205ACD7F00FFDFC497FE8DFAFC5246" author="Willen, E." box="[1177,1323,258,284]" pageId="33" pageNumber="204" pagination="1691 - 1711" refId="ref32649" refString="Willen, E. (2003) A new species of Stenhelia (Copepoda, Harpacticoida) from a hydrothermal, active, submarine volcano in the New Ireland Fore-Arc system (Papua New Guinea) with notes on deep sea colonization within the Stenheliinae. Journal of natural History, 37, 1691 - 1711. https: // doi. org / 10.1080 / 00222930110114437" type="journal article" year="2003">Willen 2003</bibRefCitation>
)
</taxonomicName>
. Some of these are marine shallow water species (
<emphasis id="68C5FB2E7F00FFDFC256FEA9FD785265" box="[600,687,294,319]" italics="true" pageId="33" pageNumber="204">e.g. De.</emphasis>
<taxonomicName id="9DB15CBF7F00FFDFC2BBFEAAFCBE5265" authority=", De." authorityName="De." box="[693,873,293,320]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longipilosa">
<emphasis id="68C5FB2E7F00FFDFC2BBFEAAFCE45265" box="[693,819,293,319]" italics="true" pageId="33" pageNumber="204">longipilosa</emphasis>
,
<emphasis id="68C5FB2E7F00FFDFC34FFEA9FCBE5265" box="[833,873,294,319]" italics="true" pageId="33" pageNumber="204">De.</emphasis>
</taxonomicName>
<taxonomicName id="9DB15CBF7F00FFDFC361FEAAFBEF5265" authority=", De." authorityName="De." box="[879,1080,293,320]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longicaudata">
<emphasis id="68C5FB2E7F00FFDFC361FEAAFBD45265" box="[879,1027,293,319]" italics="true" pageId="33" pageNumber="204">longicaudata</emphasis>
,
<emphasis id="68C5FB2E7F00FFDFC41EFEA9FBEF5265" box="[1040,1080,294,319]" italics="true" pageId="33" pageNumber="204">De.</emphasis>
</taxonomicName>
<taxonomicName id="9DB15CBF7F00FFDFC431FEA9FB305265" authority=", De." authorityName="De." box="[1087,1255,294,320]" class="Hexanauplia" family="Miraciidae" genus="Stenhelia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="coineauae">
<emphasis id="68C5FB2E7F00FFDFC431FEA9FB655265" box="[1087,1202,294,319]" italics="true" pageId="33" pageNumber="204">coineauae</emphasis>
,
<emphasis id="68C5FB2E7F00FFDFC4B1FEA9FB305265" box="[1215,1255,294,319]" italics="true" pageId="33" pageNumber="204">De.</emphasis>
</taxonomicName>
<taxonomicName id="9DB15CBF7F00FFDFC4E0FEAAFA4A5265" authority=", De." authorityName="De." box="[1262,1437,293,320]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="68C5FB2E7F00FFDFC4E0FEAAFAB05265" box="[1262,1383,293,319]" italics="true" pageId="33" pageNumber="204">intermedia</emphasis>
,
<emphasis id="68C5FB2E7F00FFDFC57BFEA9FA4A5265" box="[1397,1437,294,319]" italics="true" pageId="33" pageNumber="204">De.</emphasis>
</taxonomicName>
<taxonomicName id="9DB15CBF7F00FFDFC099FEC7FEED5238" baseAuthorityName="Wells" baseAuthorityYear="1965" box="[151,314,328,354]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="mastigochaeta">
<emphasis id="68C5FB2E7F00FFDFC099FEC7FEED5238" box="[151,314,328,354]" italics="true" pageId="33" pageNumber="204">mastigochaeta</emphasis>
</taxonomicName>
and
<emphasis id="68C5FB2E7F00FFDFC17BFEC6FDF15238" box="[373,550,328,354]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC1ABFEC7FDF15238" baseAuthorityName="Willen" baseAuthorityYear="2003" box="[421,550,328,354]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="nuwukensis">nuwukensis</taxonomicName>
</emphasis>
), but some have been described from the deep sea.
<taxonomicName id="9DB15CBF7F00FFDFC472FEC7FAEC5238" box="[1148,1339,328,354]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="noodti">
<emphasis id="68C5FB2E7F00FFDFC472FEC7FAEC5238" box="[1148,1339,328,354]" italics="true" pageId="33" pageNumber="204">Delavalia noodti</emphasis>
</taxonomicName>
and
<emphasis id="68C5FB2E7F00FFDFC57BFEC6FF2952DC" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC099FEE3FF2952DC" box="[151,254,364,390]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="islandica">islandica</taxonomicName>
</emphasis>
were described from the Iceland-Faroe Ridge at
<quantity id="9D498AD97F00FFDFC320FEE3FCA152DD" box="[814,886,364,391]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="33" pageNumber="204" unit="m" value="500.0">500 m</quantity>
depth (
<bibRefCitation id="3E205ACD7F00FFDFC3C2FEE3FBA952DC" author="Schriever, G." box="[972,1150,364,390]" pageId="33" pageNumber="204" pagination="27 - 34" refId="ref31709" refString="Schriever, G. (1982) Neue Harpacticoidea (Crustacea, Copepoda) aus dem Nordatlantik. II. Vier neue Arten der Familien Diosaccidae und Ameiridae. &quot; Meteor &quot; Forschungsergebnisse, Reihe D, 35, 27 - 34." type="journal article" year="1982">Schriever 1982</bibRefCitation>
),
<emphasis id="68C5FB2E7F00FFDFC49DFEE2FB2252DC" box="[1171,1269,364,390]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC4CDFEE3FB2252DC" box="[1219,1269,364,390]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="lima">lima</taxonomicName>
</emphasis>
was described from the
<collectingCountry id="22A667AC7F00FFDFC0F3FE1FFEE752F0" box="[253,304,400,426]" name="Peru" pageId="33" pageNumber="204">Peru</collectingCountry>
Trench at
<quantity id="9D498AD97F00FFDFC1A8FE1FFE3B52F0" box="[422,492,400,426]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="33" pageNumber="204" unit="m" value="920.0">920 m</quantity>
depth (
<bibRefCitation id="3E205ACD7F00FFDFC231FE00FCB652F0" author="Becker, K. - H. &amp; Schriever, G." box="[575,865,399,426]" pageId="33" pageNumber="204" pagination="38 - 62" refId="ref29056" refString="Becker, K. - H. &amp; Schriever, G. (1979) Eidonomie und Taxonomie abyssaler Harpacticoidea (Crustacea, Copepoda) Teil III. 13 neue Tiefsee-Copepoda Harpacticoidea der Familien Canuellidae, Cerviniidae, Tisbidae, Thalestridae, Diosaccidae und Ameiridae. &quot; Meteor &quot; Forschungsergebnisse, Reihe D, 31, 38 - 62." type="journal article" year="1979">Becker &amp; Schriever 1979</bibRefCitation>
),
<emphasis id="68C5FB2E7F00FFDFC37AFE1FFBDD52F3" box="[884,1034,399,425]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC3ACFE00FBDD52F3" box="[930,1034,399,425]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="diegensis">diegensis</taxonomicName>
</emphasis>
was found in the San Diego Trough at
<quantity id="9D498AD97F00FFDFC0BCFE3CFED95297" box="[178,270,435,461]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="33" pageNumber="204" unit="m" value="1200.0">1,200 m</quantity>
depth (
<bibRefCitation id="3E205ACD7F00FFDFC16FFE3CFD835297" author="Thistle, D. &amp; Coull, B. C." box="[353,596,435,461]" pageId="33" pageNumber="204" pagination="63 - 72" refId="ref32287" refString="Thistle, D. &amp; Coull, B. C. (1979) A revised key to Stenhelia (Stenhelia) (Copepoda: Harpacticoida) including two new species from the Pacific. Zoological Journal of the Linnean Society, 66, 63 - 72. https: // doi. org / 10.1111 / j. 1096 - 3642.1979. tb 01901. x" type="journal article" year="1979">Thistle &amp; Coull 1979</bibRefCitation>
), and an undescribed species (
<taxonomicName id="9DB15CBF7F00FFDFC3A8FE3CFBB45297" box="[934,1123,435,461]" class="Hexanauplia" family="Miraciidae" genus="Stenhelia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="undefined-6">
<emphasis id="68C5FB2E7F00FFDFC3A8FE3CFBD95297" box="[934,1038,435,461]" italics="true" pageId="33" pageNumber="204">Stenhelia</emphasis>
spec.
<quantity id="9D498AD97F00FFDFC45BFE3CFBB45297" box="[1109,1123,435,461]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="33" pageNumber="204" unit="in" value="6.0">6</quantity>
</taxonomicName>
in
<bibRefCitation id="3E205ACD7F00FFDFC48BFE3CFAF95297" author="Willen, E." box="[1157,1326,435,461]" pageId="33" pageNumber="204" pagination="1691 - 1711" refId="ref32649" refString="Willen, E. (2003) A new species of Stenhelia (Copepoda, Harpacticoida) from a hydrothermal, active, submarine volcano in the New Ireland Fore-Arc system (Papua New Guinea) with notes on deep sea colonization within the Stenheliinae. Journal of natural History, 37, 1691 - 1711. https: // doi. org / 10.1080 / 00222930110114437" type="journal article" year="2003">Willen (2003))</bibRefCitation>
, probably related to
<emphasis id="68C5FB2E7F00FFDFC10AFE58FEB452AA" box="[260,355,470,496]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC13FFE59FEB452AA" box="[305,355,470,496]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="lima">lima</taxonomicName>
</emphasis>
, was found in the
<collectingCountry id="22A667AC7F00FFDFC227FE59FDAC52AA" box="[553,635,470,496]" name="Angola" pageId="33" pageNumber="204">Angola</collectingCountry>
Basin at
<quantity id="9D498AD97F00FFDFC2EEFE59FCED52AB" box="[736,826,470,497]" metricMagnitude="3" metricUnit="m" metricValue="5.389" pageId="33" pageNumber="204" unit="m" value="5389.0">5,389 m</quantity>
depth (
<bibRefCitation id="3E205ACD7F00FFDFC385FE59FBCD52AB" author="Willen, E." box="[907,1050,470,497]" pageId="33" pageNumber="204" pagination="1691 - 1711" refId="ref32649" refString="Willen, E. (2003) A new species of Stenhelia (Copepoda, Harpacticoida) from a hydrothermal, active, submarine volcano in the New Ireland Fore-Arc system (Papua New Guinea) with notes on deep sea colonization within the Stenheliinae. Journal of natural History, 37, 1691 - 1711. https: // doi. org / 10.1080 / 00222930110114437" type="journal article" year="2003">Willen 2003</bibRefCitation>
).Also, in their appendix A,
<bibRefCitation id="3E205ACD7F00FFDFC544FE58FEF4514E" author="George, K. &amp; Veit-Kohler, G. &amp; Martinez Arbizu, P. &amp; Seifried, S. &amp; Rose, A. &amp; Willen, E. &amp; Brohldick, K. &amp; Corgosinho, P. H. &amp; Drewes, J. &amp; Menzel, L. &amp; Moura, G. &amp; Schminke, H. K." pageId="33" pageNumber="204" pagination="57 - 73" refId="ref29916" refString="George, K. H, Veit-Kohler, G., Martinez Arbizu, P., Seifried, S., Rose, A., Willen, E., Brohldick, K., Corgosinho, P. H., Drewes, J., Menzel, L., Moura, G. &amp; Schminke, H. K. (2014). Community structure and species diversity of Harpacticoida (Crustacea: Copepoda) at two sites in the deep sea of the Angola Basin (Southeast Atlantic). Organisms, Diversity &amp; Evolution, 14, 57 - 73. https: // doi. org / 10.1007 / s 13127 - 013 - 0154 - 2" type="journal article" year="2014">
George
<emphasis id="68C5FB2E7F00FFDFC099FE74FF1E514E" box="[151,201,506,532]" italics="true" pageId="33" pageNumber="204">et al</emphasis>
. (2014)
</bibRefCitation>
reported four unidentified species of
<taxonomicName id="9DB15CBF7F00FFDFC2C1FE75FCEA514E" authorityName="Brady" authorityYear="1869" box="[719,829,506,532]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="68C5FB2E7F00FFDFC2C1FE75FCEA514E" box="[719,829,506,532]" italics="true" pageId="33" pageNumber="204">Delavalia</emphasis>
</taxonomicName>
found in deep-sea sediment samples (
<quantity id="9D498AD97F00FFDFC4E0FE75FA9A514E" box="[1262,1357,506,533]" metricMagnitude="3" metricUnit="m" metricValue="5.389" pageId="33" pageNumber="204" unit="m" value="5389.0">5,389 m</quantity>
depth) from the
<collectingCountry id="22A667AC7F00FFDFC0F4FD92FE9B516D" box="[250,332,541,567]" name="Angola" pageId="33" pageNumber="204">Angola</collectingCountry>
Basin; it is conceivable that one of
<bibRefCitation id="3E205ACD7F00FFDFC2D9FD92FC1E516D" author="George, K. &amp; Veit-Kohler, G. &amp; Martinez Arbizu, P. &amp; Seifried, S. &amp; Rose, A. &amp; Willen, E. &amp; Brohldick, K. &amp; Corgosinho, P. H. &amp; Drewes, J. &amp; Menzel, L. &amp; Moura, G. &amp; Schminke, H. K." box="[727,969,541,568]" pageId="33" pageNumber="204" pagination="57 - 73" refId="ref29916" refString="George, K. H, Veit-Kohler, G., Martinez Arbizu, P., Seifried, S., Rose, A., Willen, E., Brohldick, K., Corgosinho, P. H., Drewes, J., Menzel, L., Moura, G. &amp; Schminke, H. K. (2014). Community structure and species diversity of Harpacticoida (Crustacea: Copepoda) at two sites in the deep sea of the Angola Basin (Southeast Atlantic). Organisms, Diversity &amp; Evolution, 14, 57 - 73. https: // doi. org / 10.1007 / s 13127 - 013 - 0154 - 2" type="journal article" year="2014">
Georges
<emphasis id="68C5FB2E7F00FFDFC34FFD91FCA5516D" box="[833,882,541,567]" italics="true" pageId="33" pageNumber="204">et al</emphasis>
. (2014)
</bibRefCitation>
species of
<taxonomicName id="9DB15CBF7F00FFDFC44DFD92FB66516D" authorityName="Brady" authorityYear="1869" box="[1091,1201,541,567]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="68C5FB2E7F00FFDFC44DFD92FB66516D" box="[1091,1201,541,567]" italics="true" pageId="33" pageNumber="204">Delavalia</emphasis>
</taxonomicName>
could be conspecific to
<bibRefCitation id="3E205ACD7F00FFDFC0BAFDCEFEB35101" author="Willen, E." box="[180,356,577,603]" pageId="33" pageNumber="204" pagination="1691 - 1711" refId="ref32649" refString="Willen, E. (2003) A new species of Stenhelia (Copepoda, Harpacticoida) from a hydrothermal, active, submarine volcano in the New Ireland Fore-Arc system (Papua New Guinea) with notes on deep sea colonization within the Stenheliinae. Journal of natural History, 37, 1691 - 1711. https: // doi. org / 10.1080 / 00222930110114437" type="journal article" year="2003">Willens (2003)</bibRefCitation>
<taxonomicName id="9DB15CBF7F00FFDFC162FDCEFE035101" authorityName="Boeck" authorityYear="1865" box="[364,468,577,603]" class="Hexanauplia" family="Miraciidae" genus="Stenhelia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="68C5FB2E7F00FFDFC162FDCEFE035101" box="[364,468,577,603]" italics="true" pageId="33" pageNumber="204">Stenhelia</emphasis>
</taxonomicName>
“spec 6”. The depth distribution of this and some other lineages (see Gómez &amp; Cruz- Barraza 2021; Gómez 2021) support the view that deep-sea harpacticoids may have originated from shallow-water ancestral stocks as suggested earlier by
<bibRefCitation id="3E205ACD7F00FFDFC255FD07FCEE51F8" author="Mu, F. &amp; Huys, R." box="[603,825,648,674]" pageId="33" pageNumber="204" pagination="179 - 206" refId="ref31326" refString="Mu, F. &amp; Huys, R. (2002) New species of Stenhelia (Copepoda, Harpacticoida, Diosaccidae) from the Bohai Sea (China) with notes on subgeneric division and phylogenetic relationships. Cahiers de Biologie marine, 43, 179 - 206. https: // doi. org / 10.21411 / CBM. A. C 482 EC 6 A" type="journal article" year="2002">Mu &amp; Huys (2002)</bibRefCitation>
. The shallow-water species
<emphasis id="68C5FB2E7F00FFDFC476FD06FAEB51F8" box="[1144,1340,648,674]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC4A6FD07FAEB51F8" box="[1192,1340,648,674]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longicaudata">longicaudata</taxonomicName>
</emphasis>
and
<emphasis id="68C5FB2E7F00FFDFC57BFD06FECF519F" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC099FD24FECF519F" baseAuthorityName="Willen" baseAuthorityYear="2003" box="[151,280,683,709]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="nuwukensis">nuwukensis</taxonomicName>
</emphasis>
are assumed here to be the most primitive species of the
<taxonomicName id="9DB15CBF7F00FFDFC391FD24FBE4519F" box="[927,1075,683,709]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longicaudata">
<emphasis id="68C5FB2E7F00FFDFC391FD24FBE4519F" box="[927,1075,683,709]" italics="true" pageId="33" pageNumber="204">longicaudata</emphasis>
</taxonomicName>
-group based on the retention of four setae on P1 ENP2.
</paragraph>
<paragraph id="5A0E273C7F00FFDFC0C9FD7DFEC95735" blockId="33.[151,1437,151,2022]" pageId="33" pageNumber="204">
The deep-sea
<emphasis id="68C5FB2E7F00FFDFC169FD7CFE3A5056" box="[359,493,754,780]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC196FD7DFE3A5056" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[408,493,754,780]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">reducta</taxonomicName>
</emphasis>
<emphasis id="68C5FB2E7F00FFDFC1F8FD7DFD855056" bold="true" box="[502,594,754,780]" pageId="33" pageNumber="204">
<taxonomicNameLabel id="73F646557F00FFDFC1F8FD7DFD855056" box="[502,594,754,780]" pageId="33" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is unique within the genus by the autapomorphic P1 ENP2 with reduced armature from three to two well-developed and densely setulose setae. Amongst the species of the
<taxonomicName id="9DB15CBF7F00FFDFC4F1FC99FA44506A" box="[1279,1427,790,816]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longicaudata">
<emphasis id="68C5FB2E7F00FFDFC4F1FC99FA44506A" box="[1279,1427,790,816]" italics="true" pageId="33" pageNumber="204">longicaudata</emphasis>
</taxonomicName>
- group, the loss of the inner seta of P1 ENP1 is shared with the deep-sea
<emphasis id="68C5FB2E7F00FFDFC3DBFCB5FB9A5009" box="[981,1101,825,851]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC40BFCB6FB9A5009" box="[1029,1101,825,851]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="noodti">noodti</taxonomicName>
</emphasis>
only, but a close relationship between these two species seems unlikely given the remarkable differences in the general structure and setation of the antennary exopod (one-segmented and with two setae in
<emphasis id="68C5FB2E7F00FFDFC343FC0EFC1250C0" box="[845,965,896,922]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC373FC0FFC1250C0" box="[893,965,896,922]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="noodti">noodti</taxonomicName>
</emphasis>
, and typical for the genus in
<emphasis id="68C5FB2E7F00FFDFC519FC0EFA4A50C0" box="[1303,1437,896,922]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC546FC0FFA4A50C0" authorityName="Gómez" authorityYear="2021" box="[1352,1437,896,922]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="reducta">reducta</taxonomicName>
</emphasis>
(three-segmented with one, one, and four setae on the first, second and third segments, respectively), armature formula of the swimming legs (P1 ENP2 with three setae in
<emphasis id="68C5FB2E7F00FFDFC332FC47FC6450BB" box="[828,947,967,993]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC365FC48FC6450BB" box="[875,947,967,993]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="noodti">noodti</taxonomicName>
</emphasis>
, but two setae only in
<emphasis id="68C5FB2E7F00FFDFC4BFFC47FAE250BB" box="[1201,1333,967,993]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC4EEFC48FAE250BB" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[1248,1333,967,993]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">reducta</taxonomicName>
</emphasis>
<emphasis id="68C5FB2E7F00FFDFC532FC48FA4250BB" bold="true" box="[1340,1429,967,993]" pageId="33" pageNumber="204">
<taxonomicNameLabel id="73F646557F00FFDFC532FC48FA4250BB" box="[1340,1429,967,993]" pageId="33" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, loss of the inner seta on P2 ENP
<quantity id="9D498AD97F00FFDFC203FC64FDE3575E" box="[525,564,1002,1029]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="33" pageNumber="204" unit="in" value="1.0">1 in</quantity>
<emphasis id="68C5FB2E7F00FFDFC232FC64FC99575E" box="[572,846,1002,1029]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC265FC65FD64575E" box="[619,691,1002,1028]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="noodti">noodti</taxonomicName>
—De.
<taxonomicName id="9DB15CBF7F00FFDFC308FC65FC99575E" box="[774,846,1002,1028]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="noodti">noodti</taxonomicName>
</emphasis>
also lacks the inner seta on P2 ENP2 and possesses two inner setae on P2 ENP3, but these segments are unknown for
<emphasis id="68C5FB2E7F00FFDFC39EFB80FBC15772" box="[912,1046,1038,1064]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC3CFFB81FBC15772" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[961,1046,1038,1064]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">reducta</taxonomicName>
</emphasis>
<emphasis id="68C5FB2E7F00FFDFC410FB81FBAE5772" bold="true" box="[1054,1145,1038,1064]" pageId="33" pageNumber="204">
<taxonomicNameLabel id="73F646557F00FFDFC410FB81FBAE5772" box="[1054,1145,1038,1064]" pageId="33" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
—, presence of two inner setae on P3 and P4 EXP
<quantity id="9D498AD97F00FFDFC1A4FBBEFE055711" box="[426,466,1073,1099]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="33" pageNumber="204" unit="in" value="3.0">3 in</quantity>
<emphasis id="68C5FB2E7F00FFDFC1D7FBBDFD875711" box="[473,592,1073,1099]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC206FBBEFD875711" box="[520,592,1073,1099]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="noodti">noodti</taxonomicName>
</emphasis>
, but three in
<emphasis id="68C5FB2E7F00FFDFC2EBFBBDFCBE5711" box="[741,873,1073,1099]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC31AFBBEFCBE5711" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[788,873,1073,1099]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="reducta" status="sp. nov.">reducta</taxonomicName>
</emphasis>
<emphasis id="68C5FB2E7F00FFDFC361FBBEFC1E5711" bold="true" box="[879,969,1073,1099]" pageId="33" pageNumber="204">
<taxonomicNameLabel id="73F646557F00FFDFC361FBBEFC1E5711" box="[879,969,1073,1099]" pageId="33" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and loss of the inner seta on P3 ENP
<quantity id="9D498AD97F00FFDFC578FBBEFA4B5711" box="[1398,1436,1073,1099]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="33" pageNumber="204" unit="in" value="1.0">1 in</quantity>
<emphasis id="68C5FB2E7F00FFDFC099FBD9FED95735" box="[151,270,1109,1135]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC0C8FBDAFED95735" box="[198,270,1109,1135]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="noodti">noodti</taxonomicName>
</emphasis>
).
</paragraph>
<paragraph id="5A0E273C7F00FFDFC0C9FBF7FB5C567A" blockId="33.[151,1437,151,2022]" pageId="33" pageNumber="204">
The deep-sea
<taxonomicName id="9DB15CBF7F00FFDFC16AFBF7FDFA57C8" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[356,557,1144,1170]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="asetosa" status="sp. nov.">
<emphasis id="68C5FB2E7F00FFDFC16AFBF7FDFA57C8" box="[356,557,1144,1170]" italics="true" pageId="33" pageNumber="204">Delavalia asetosa</emphasis>
</taxonomicName>
<emphasis id="68C5FB2E7F00FFDFC23BFBF7FD5857C8" bold="true" box="[565,655,1144,1170]" pageId="33" pageNumber="204">
<taxonomicNameLabel id="73F646557F00FFDFC23BFBF7FD5857C8" box="[565,655,1144,1170]" pageId="33" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and the shallow-water
<emphasis id="68C5FB2E7F00FFDFC398FBF6FBBF57C8" box="[918,1128,1144,1170]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC3CBFBF7FBBF57C8" baseAuthorityName="Wells" baseAuthorityYear="1965" box="[965,1128,1144,1170]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="mastigochaeta">mastigochaeta</taxonomicName>
</emphasis>
are the only species within the
<taxonomicName id="9DB15CBF7F00FFDFC0CCFB13FE8157EC" box="[194,342,1180,1206]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longicaudata">
<emphasis id="68C5FB2E7F00FFDFC0CCFB13FE8157EC" box="[194,342,1180,1206]" italics="true" pageId="33" pageNumber="204">longicaudata</emphasis>
</taxonomicName>
-group without inner armature on P1 EXP2. Amongst the species of the
<taxonomicName id="9DB15CBF7F00FFDFC482FB13FAF757EC" box="[1164,1312,1180,1206]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longicaudata">
<emphasis id="68C5FB2E7F00FFDFC482FB13FAF757EC" box="[1164,1312,1180,1206]" italics="true" pageId="33" pageNumber="204">longicaudata</emphasis>
</taxonomicName>
-group, the deep-sea species
<taxonomicName id="9DB15CBF7F00FFDFC158FB30FDC85783" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[342,543,1215,1241]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="asetosa" status="sp. nov.">
<emphasis id="68C5FB2E7F00FFDFC158FB30FDC85783" box="[342,543,1215,1241]" italics="true" pageId="33" pageNumber="204">Delavalia asetosa</emphasis>
</taxonomicName>
<emphasis id="68C5FB2E7F00FFDFC22BFB30FDAA5783" bold="true" box="[549,637,1215,1241]" pageId="33" pageNumber="204">
<taxonomicNameLabel id="73F646557F00FFDFC22BFB30FDAA5783" box="[549,637,1215,1241]" pageId="33" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="68C5FB2E7F00FFDFC284FB4FFCD75783" box="[650,768,1215,1241]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC2B6FB30FCD75783" box="[696,768,1215,1241]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="noodti">noodti</taxonomicName>
</emphasis>
,
<emphasis id="68C5FB2E7F00FFDFC302FB4FFC425783" box="[780,917,1215,1241]" italics="true" pageId="33" pageNumber="204">
D,
<taxonomicName id="9DB15CBF7F00FFDFC320FB30FC425783" box="[814,917,1215,1241]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="islandica">islandica</taxonomicName>
</emphasis>
, and the shallow water species
<emphasis id="68C5FB2E7F00FFDFC4FFFB4FFA4B5783" box="[1265,1436,1215,1241]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC510FB30FA4B5783" box="[1310,1436,1215,1241]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longipilosa">longipilosa</taxonomicName>
</emphasis>
share the reduction of the inner armature of P3 EXP3 from three to two setae. However,
<emphasis id="68C5FB2E7F00FFDFC461FB6CFB2557A6" box="[1135,1266,1251,1276]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC490FB6CFB2557A6" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[1182,1266,1251,1276]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="asetosa" status="sp. nov.">asetosa</taxonomicName>
</emphasis>
<emphasis id="68C5FB2E7F00FFDFC4F6FB6CFA8657A7" bold="true" box="[1272,1361,1251,1277]" pageId="33" pageNumber="204">
<taxonomicNameLabel id="73F646557F00FFDFC4F6FB6CFA8657A7" box="[1272,1361,1251,1277]" pageId="33" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
seems to be unique within the
<taxonomicName id="9DB15CBF7F00FFDFC1AEFA89FDE3567A" box="[416,564,1286,1312]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longicaudata">
<emphasis id="68C5FB2E7F00FFDFC1AEFA89FDE3567A" box="[416,564,1286,1312]" italics="true" pageId="33" pageNumber="204">longicaudata</emphasis>
</taxonomicName>
-group by the loss of inner armature on P3P4 EXP1.
</paragraph>
<paragraph id="5A0E273C7F00FFDFC0C9FAA5FB6B5505" blockId="33.[151,1437,151,2022]" pageId="33" pageNumber="204">
Amongst the species of the
<taxonomicName id="9DB15CBF7F00FFDFC20BFAA5FD4E561E" box="[517,665,1322,1348]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longicaudata">
<emphasis id="68C5FB2E7F00FFDFC20BFAA5FD4E561E" box="[517,665,1322,1348]" italics="true" pageId="33" pageNumber="204">longicaudata</emphasis>
</taxonomicName>
-group, only four species, the shallow-water
<emphasis id="68C5FB2E7F00FFDFC498FAA5FAED5619" box="[1174,1338,1322,1347]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC4C9FAA5FAED5619" box="[1223,1338,1322,1347]" class="Hexanauplia" family="Miraciidae" genus="Stenhelia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="coineauae">coineauae</taxonomicName>
</emphasis>
, and the deep-sea
<emphasis id="68C5FB2E7F00FFDFC10EFAC1FE10563D" box="[256,455,1357,1383]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC121FAC2FE10563D" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[303,455,1357,1383]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="californiensis" status="sp. nov.">californiensis</taxonomicName>
</emphasis>
<emphasis id="68C5FB2E7F00FFDFC1C1FAC2FDFE563D" bold="true" box="[463,553,1357,1383]" pageId="33" pageNumber="204">
<taxonomicNameLabel id="73F646557F00FFDFC1C1FAC2FDFE563D" box="[463,553,1357,1383]" pageId="33" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="68C5FB2E7F00FFDFC239FAC1FD1A563D" box="[567,717,1357,1383]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC269FAC2FD1A563D" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[615,717,1357,1383]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="profunda" status="sp. nov.">profunda</taxonomicName>
</emphasis>
<emphasis id="68C5FB2E7F00FFDFC2DDFAC2FCFA563D" bold="true" box="[723,813,1357,1383]" pageId="33" pageNumber="204">
<taxonomicNameLabel id="73F646557F00FFDFC2DDFAC2FCFA563D" box="[723,813,1357,1383]" pageId="33" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<emphasis id="68C5FB2E7F00FFDFC365FAC1FC1A563D" box="[875,973,1357,1383]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC395FAC2FC1A563D" box="[923,973,1357,1383]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="lima">lima</taxonomicName>
</emphasis>
share exactly the same armature formula of P1P4 (P1 EXP/ENP: 0,1,022/1,111; P2 EXP/ENP: 1,1,223/1,2,121; P3 EXP/ENP: 1,1,323/1,1,321; P4 EXP/ ENP: 1,1,323/1,1,221). All the other species display some reduction in the armature complement of one or more swimming legs. Within this core of species,
<emphasis id="68C5FB2E7F00FFDFC29FFA37FC8E568B" box="[657,857,1463,1489]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC2CFFA38FC8E568B" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[705,857,1463,1489]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="californiensis" status="sp. nov.">californiensis</taxonomicName>
</emphasis>
<emphasis id="68C5FB2E7F00FFDFC36CFA37FC6A5688" bold="true" box="[866,957,1464,1490]" pageId="33" pageNumber="204">
<taxonomicNameLabel id="73F646557F00FFDFC36CFA37FC6A5688" box="[866,957,1464,1490]" pageId="33" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="68C5FB2E7F00FFDFC3F8FA37FB4E568B" box="[1014,1177,1464,1489]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC428FA37FB4E568B" box="[1062,1177,1464,1489]" class="Hexanauplia" family="Miraciidae" genus="Stenhelia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="coineauae">coineauae</taxonomicName>
</emphasis>
display a combination of six and five setae on the exopod and endopodal lobe, respectively, of the female P5.
<taxonomicName id="9DB15CBF7F00FFDFC46CFA54FAEA56AF" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[1122,1341,1499,1525]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="profunda" status="sp. nov.">
<emphasis id="68C5FB2E7F00FFDFC46CFA54FAEA56AF" box="[1122,1341,1499,1525]" italics="true" pageId="33" pageNumber="204">Delavalia profunda</emphasis>
</taxonomicName>
<emphasis id="68C5FB2E7F00FFDFC54DFA54FA4B56AF" bold="true" box="[1347,1436,1499,1525]" pageId="33" pageNumber="204">
<taxonomicNameLabel id="73F646557F00FFDFC54DFA54FA4B56AF" box="[1347,1436,1499,1525]" pageId="33" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
retained the plesiomorphic armature complement of six setae on the exopod of the female P5, but the armature of the endopodal lobe is reduced from five to four setae, while the armature complement of the female P5 EXP and baseoendopod is reduced from six to five and from five to four setae, respectively, in
<emphasis id="68C5FB2E7F00FFDFC45AF9C9FB6B5505" box="[1108,1212,1605,1631]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC48DF9CAFB6F5505" box="[1155,1208,1605,1631]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="lima">lima</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="5A0E273C7F00FFDCC0C9F9E6FDF3521A" blockId="33.[151,1437,151,2022]" lastBlockId="34.[151,1437,151,320]" lastPageId="34" lastPageNumber="205" pageId="33" pageNumber="204">
The polyphyletic status of
<taxonomicName id="9DB15CBF7F00FFDFC1FCF9E6FDB755D9" authorityName="Brady" authorityYear="1869" box="[498,608,1641,1667]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="68C5FB2E7F00FFDFC1FCF9E6FDB755D9" box="[498,608,1641,1667]" italics="true" pageId="33" pageNumber="204">Delavalia</emphasis>
</taxonomicName>
is evident from the disparity in the structure and armature complement of a number of appendages both in the female and male (see above). The monophyly of the
<taxonomicName id="9DB15CBF7F00FFDFC474F903FAD955FC" box="[1146,1294,1676,1702]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longicaudata">
<emphasis id="68C5FB2E7F00FFDFC474F903FAD955FC" box="[1146,1294,1676,1702]" italics="true" pageId="33" pageNumber="204">longicaudata</emphasis>
</taxonomicName>
-group is not clear yet.
<bibRefCitation id="3E205ACD7F00FFDFC10DF93FFE775590" author="Willen, E." box="[259,416,1712,1738]" pageId="33" pageNumber="204" pagination="1691 - 1711" refId="ref32649" refString="Willen, E. (2003) A new species of Stenhelia (Copepoda, Harpacticoida) from a hydrothermal, active, submarine volcano in the New Ireland Fore-Arc system (Papua New Guinea) with notes on deep sea colonization within the Stenheliinae. Journal of natural History, 37, 1691 - 1711. https: // doi. org / 10.1080 / 00222930110114437" type="journal article" year="2003">Willen (2003)</bibRefCitation>
commented on the apomorphic status of the P1 ENP, and noted that in the groundpattern of this taxon the male P5 EXP is discrete, the armature complement of the female P5 is complete and that the setae on the third exopodal segment of swimming legs are not reduced. Although the shape and structure of the maxilliped seem to be constant within this group, other appendages need some attention. For example, the mandible of
<emphasis id="68C5FB2E7F00FFDFC57BF894FEDD540D" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC099F8B1FEDD540D" box="[151,266,1854,1879]" class="Hexanauplia" family="Miraciidae" genus="Stenhelia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="coineauae">coineauae</taxonomicName>
</emphasis>
seems to possess a two-segmented exopod and four setae on the basis, and the exopod and endopod of the maxillule appear as discrete in
<bibRefCitation id="3E205ACD7F00FFDFC229F8EEFD685421" author="Soyer, J." box="[551,703,1889,1915]" pageId="33" pageNumber="204" pagination="263 - 280" refId="ref32164" refString="Soyer, J. (1971) Contribution a l'etude des Copepodes Harpacticoides de Mediterranee occidentale. 5. Stenhelia (Delavalia) coineauae n. sp., Stenhelia (D.) bocqueti n. sp. et Typhlamphiascus bouligandi n. sp. (Diosaccidae, Sars). Vie et Milieu, (A), 22, 263 - 280." type="journal article" year="1971">Soyer (1971)</bibRefCitation>
, the maxillulary rami of
<emphasis id="68C5FB2E7F00FFDFC3D3F8EDFB975421" box="[989,1088,1889,1915]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC400F8EEFB975421" box="[1038,1088,1889,1915]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="lima">lima</taxonomicName>
</emphasis>
and
<emphasis id="68C5FB2E7F00FFDFC474F8EDFAC45421" box="[1146,1299,1889,1915]" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC4A5F8EEFAC45421" box="[1195,1299,1889,1915]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="diegensis">diegensis</taxonomicName>
</emphasis>
appear also as discrete in
<bibRefCitation id="3E205ACD7F00FFDFC13AF80BFDB054C5" author="Becker, K. - H. &amp; Schriever, G." box="[308,615,1924,1951]" pageId="33" pageNumber="204" pagination="38 - 62" refId="ref29056" refString="Becker, K. - H. &amp; Schriever, G. (1979) Eidonomie und Taxonomie abyssaler Harpacticoidea (Crustacea, Copepoda) Teil III. 13 neue Tiefsee-Copepoda Harpacticoidea der Familien Canuellidae, Cerviniidae, Tisbidae, Thalestridae, Diosaccidae und Ameiridae. &quot; Meteor &quot; Forschungsergebnisse, Reihe D, 31, 38 - 62." type="journal article" year="1979">Becker &amp; Schriever (1979)</bibRefCitation>
and
<bibRefCitation id="3E205ACD7F00FFDFC291F80BFC7154C5" author="Thistle, D. &amp; Coull, B. C." box="[671,934,1924,1951]" pageId="33" pageNumber="204" pagination="63 - 72" refId="ref32287" refString="Thistle, D. &amp; Coull, B. C. (1979) A revised key to Stenhelia (Stenhelia) (Copepoda: Harpacticoida) including two new species from the Pacific. Zoological Journal of the Linnean Society, 66, 63 - 72. https: // doi. org / 10.1111 / j. 1096 - 3642.1979. tb 01901. x" type="journal article" year="1979">Thistle &amp; Coull (1979)</bibRefCitation>
, respectively, and the female antenna of
<emphasis id="68C5FB2E7F00FFDFC57BF80AFF085498" italics="true" pageId="33" pageNumber="204">
De.
<taxonomicName id="9DB15CBF7F00FFDFC099F827FF085498" box="[151,223,1960,1986]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="noodti">noodti</taxonomicName>
</emphasis>
seems to bear a basis with a one-segmented exopod with two setae and a two-segmented endopod with two inner setae on the first segment (
<bibRefCitation id="3E205ACD7F00FFDFC20FF844FD0D54BC" author="Schriever, G." box="[513,730,1995,2022]" pageId="33" pageNumber="204" pagination="27 - 34" refId="ref31709" refString="Schriever, G. (1982) Neue Harpacticoidea (Crustacea, Copepoda) aus dem Nordatlantik. II. Vier neue Arten der Familien Diosaccidae und Ameiridae. &quot; Meteor &quot; Forschungsergebnisse, Reihe D, 35, 27 - 34." type="journal article" year="1982">Schriever 1982: 28</bibRefCitation>
,
<figureCitation id="C28A3BB97F00FFDFC2EBF844FCF554BC" box="[741,802,1995,2022]" captionStart="FIGURE 1" captionStartId="2.[151,250,1540,1564]" captionTargetBox="[163,1425,237,1513]" captionTargetId="figure-48@2.[163,1428,237,1519]" captionTargetPageId="2" captionText="FIGURE 1. Sampling locations visited during oceanographic cruises Talud IV (circles), Talud X (stars), Talud XV (squares) and Talud XVIB (triangles). Full figures represent positive collection of Delavalia profunda sp. nov. (Stn 24 —Talud XV), Delavalia californiensis sp. nov. (Stn 20 —Talud XV), Delavalia asetosa sp. nov. (Stn 19 —Talud IV), Delavalia reducta sp. nov. (Stn 25 —Talud XVIB), Archaeohuysia huysi gen. et sp. nov. (Stn 21 —Talud XVIB), Diarthropodella prima gen. et sp. nov. (Stn 15 —Talud X—, and Stn 19 —Talud IV), and Diarthropodella secunda sp. nov. (Stn 18 —Talud X). Insert shows location of stations 15, 18 and 19 visited during Talud X cruise." figureDoi="http://doi.org/10.5281/zenodo.5563571" httpUri="https://zenodo.org/record/5563571/files/figure.png" pageId="33" pageNumber="204">Fig. 1</figureCitation>
(A2)). The female of all the species of the
<taxonomicName id="9DB15CBF7F00FFDFC4F1F844FA4454BF" box="[1279,1427,1995,2021]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="204" phylum="Arthropoda" rank="species" species="longicaudata">
<emphasis id="68C5FB2E7F00FFDFC4F1F844FA4454BF" box="[1279,1427,1995,2021]" italics="true" pageId="33" pageNumber="204">longicaudata</emphasis>
</taxonomicName>
- group have been described, but the male is known only for the shallow-water
<emphasis id="68C5FB2E7F03FFDCC409FF18FBF853EA" box="[1031,1071,151,176]" italics="true" pageId="34" pageNumber="205">De.</emphasis>
<taxonomicName id="9DB15CBF7F03FFDCC438FF18FB3C53EA" authority=", De." authorityName="De." box="[1078,1259,151,177]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="205" phylum="Arthropoda" rank="species" species="longipilosa">
<emphasis id="68C5FB2E7F03FFDCC438FF18FB6353EB" box="[1078,1204,151,177]" italics="true" pageId="34" pageNumber="205">longipilosa</emphasis>
,
<emphasis id="68C5FB2E7F03FFDCC4CDFF18FB3C53EA" box="[1219,1259,151,176]" italics="true" pageId="34" pageNumber="205">De.</emphasis>
</taxonomicName>
<taxonomicName id="9DB15CBF7F03FFDCC4FCFF18FAB253EA" box="[1266,1381,151,176]" class="Hexanauplia" family="Miraciidae" genus="Stenhelia" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="205" phylum="Arthropoda" rank="species" species="coineauae">
<emphasis id="68C5FB2E7F03FFDCC4FCFF18FAB253EA" box="[1266,1381,151,176]" italics="true" pageId="34" pageNumber="205">coineauae</emphasis>
</taxonomicName>
, and
<emphasis id="68C5FB2E7F03FFDCC099FF34FEBC538E" box="[151,363,186,212]" italics="true" pageId="34" pageNumber="205">
De.
<taxonomicName id="9DB15CBF7F03FFDCC0C6FF35FEBC538E" baseAuthorityName="Wells" baseAuthorityYear="1965" box="[200,363,186,212]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="205" phylum="Arthropoda" rank="species" species="mastigochaeta">mastigochaeta</taxonomicName>
</emphasis>
, and for the deep-sea
<emphasis id="68C5FB2E7F03FFDCC262FF34FD33538E" box="[620,740,186,212]" italics="true" pageId="34" pageNumber="205">
De.
<taxonomicName id="9DB15CBF7F03FFDCC292FF35FD33538E" box="[668,740,186,212]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="205" phylum="Arthropoda" rank="species" species="noodti">noodti</taxonomicName>
</emphasis>
and
<emphasis id="68C5FB2E7F03FFDCC310FF34FC30538E" box="[798,999,186,212]" italics="true" pageId="34" pageNumber="205">
De.
<taxonomicName id="9DB15CBF7F03FFDCC341FF35FC30538E" authority="Gómez, 2021" authorityName="Gómez" authorityYear="2021" box="[847,999,186,212]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="205" phylum="Arthropoda" rank="species" species="californiensis" status="sp. nov.">californiensis</taxonomicName>
</emphasis>
<emphasis id="68C5FB2E7F03FFDCC3FEFF35FB9C538E" bold="true" box="[1008,1099,186,212]" pageId="34" pageNumber="205">
<taxonomicNameLabel id="73F646557F03FFDCC3FEFF35FB9C538E" box="[1008,1099,186,212]" pageId="34" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Although the disparity in the structure and shape of the male P2 ENP in the species of
<taxonomicName id="9DB15CBF7F03FFDCC327FF51FC4053A2" authorityName="Brady" authorityYear="1869" box="[809,919,222,248]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="205" phylum="Arthropoda" rank="genus">
<emphasis id="68C5FB2E7F03FFDCC327FF51FC4053A2" box="[809,919,222,248]" italics="true" pageId="34" pageNumber="205">Delavalia</emphasis>
</taxonomicName>
is indicative of the polyphyletic status of the genus (
<bibRefCitation id="3E205ACD7F03FFDCC0E4FE8DFE6C5246" author="Mu, F. &amp; Huys, R." box="[234,443,258,284]" pageId="34" pageNumber="205" pagination="179 - 206" refId="ref31326" refString="Mu, F. &amp; Huys, R. (2002) New species of Stenhelia (Copepoda, Harpacticoida, Diosaccidae) from the Bohai Sea (China) with notes on subgeneric division and phylogenetic relationships. Cahiers de Biologie marine, 43, 179 - 206. https: // doi. org / 10.21411 / CBM. A. C 482 EC 6 A" type="journal article" year="2002">Mu &amp; Huys 2002</bibRefCitation>
), the male P2 ENP seems to be similar in those species of the
<taxonomicName id="9DB15CBF7F03FFDCC49AFE8DFAFF5246" box="[1172,1320,258,284]" class="Hexanauplia" family="Miraciidae" genus="Delavalia" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="205" phylum="Arthropoda" rank="species" species="longicaudata">
<emphasis id="68C5FB2E7F03FFDCC49AFE8DFAFF5246" box="[1172,1320,258,284]" italics="true" pageId="34" pageNumber="205">longicaudata</emphasis>
</taxonomicName>
-group for which the male has been described.
</paragraph>
</subSubSection>
</treatment>
</document>