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<document id="DA319C69301725FF760E90C0AB055D9D" ID-DOI="10.11646/zootaxa.5339.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="8297316" ID-ZooBank="6C4BD5F2-951D-4B34-8AC0-158C609CC07B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1693306588916" checkinUser="plazi" docAuthor="Bilton, David T. &amp; Bird, Matthew S." docDate="2023" docId="D20987A0FFCE1C4FD293FDF7FBCD0595" docLanguage="en" docName="zootaxa.5339.2.6.pdf" docOrigin="Zootaxa 5339 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5339.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eupotemus tuberculatus Bilton &amp; Bird 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="197" masterDocId="2E30FFD8FFCF1C4DD204FFD4FF980444" masterDocTitle="Eupotemus tuberculatus sp. nov., the first hooded shore beetle confirmed from southern Africa (Coleoptera, Epimetopidae)" masterLastPageNumber="200" masterPageNumber="196" pageNumber="197" updateTime="1693306979188" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="A7C91D7DDE690E27E6EA59F6B4E3F8B3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7B6D856FCD5B772C76FFAD12D6F07197">
<mods:title id="D5B973FDCF9AF95D946A4617294130B3">Eupotemus tuberculatus sp. nov., the first hooded shore beetle confirmed from southern Africa (Coleoptera, Epimetopidae)</mods:title>
</mods:titleInfo>
<mods:name id="25564FE6B45EF058956115F5B4A4F6D4" type="personal">
<mods:role id="6338464A9724BC4D075D208CAECDF882">
<mods:roleTerm id="F1A222ACC86DEC0F2C60AA0CFA73B6AB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6A7AE0AABB4588756893A4E072FB97F1">Bilton, David T.</mods:namePart>
<mods:nameIdentifier id="C67F7D5C9F083881C1FB0BB4702DA8C2" type="ORCID">0000-0001-9163-1008</mods:nameIdentifier>
<mods:affiliation id="195693A0091B9D1F4A3EEC7C2EC2C39F">School of Biological and Marine Sciences, University of Plymouth, Drake Circus, Plymouth PL 4 8 AA, UK. &amp; Department of Zoology, University of Johannesburg, PO Box 524, Auckland Park, Johannesburg 2006, South Africa. https: // orcid. org / 0000 - 0001 - 9163 - 1008</mods:affiliation>
</mods:name>
<mods:name id="F882BBBD1682928688C7EA8DD2F66648" type="personal">
<mods:role id="85AE7C2BB98CFC81C1BFE72A8A4AA3AA">
<mods:roleTerm id="FEE8D66A7C06623144A95115EA33F5C2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A9B92021DAA9E50CC3935504095F2DD9">Bird, Matthew S.</mods:namePart>
<mods:nameIdentifier id="E82BE61582A70E221233CA18F9C264CA" type="ORCID">0000-0001-9163-1008</mods:nameIdentifier>
<mods:affiliation id="49310B9326A283F8925D3EA339A89834">Department of Zoology, University of Johannesburg, PO Box 524, Auckland Park, Johannesburg 2006, South Africa. https: // orcid. org / 0000 - 0001 - 9163 - 1008</mods:affiliation>
</mods:name>
<mods:typeOfResource id="7007365C546AFA8001A5FAEA6A6E94C8">text</mods:typeOfResource>
<mods:relatedItem id="C06BDC4DB13695F6522128AA1BA1E919" type="host">
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<mods:title id="FE47686B14A7EB2810987FAAFF5BE9B8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E288CBC3F8B3ACC37D3F956541A97930">
<mods:date id="2A11ECFA0732033AD3ACF94F4DFA0011">2023</mods:date>
<mods:detail id="946EEE37EBD2ED868660A256CA72B31D" type="pubDate">
<mods:number id="B63D71B2821336BF7F1AE7ECF902B55B">2023-08-29</mods:number>
</mods:detail>
<mods:detail id="99316EA5D382E45E3ADE63F4D59A8721" type="volume">
<mods:number id="E10ACECAE30022BAA237BD50885FEDE6">5339</mods:number>
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<mods:number id="E40B01D2B1314B587ED239FB0030176A">2</mods:number>
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<mods:start id="5390BEAEEEEA2AE8DA49059C175FD2D4">196</mods:start>
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<mods:url id="60E894B1110C5719DE50F14430EE89CA">http://dx.doi.org/10.11646/zootaxa.5339.2.6</mods:url>
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<mods:classification id="62DEE7577213FDD3EA2F48CF023345E0">journal article</mods:classification>
<mods:identifier id="2C51FD53F036EAE06BEDCCF019C62697" type="DOI">10.11646/zootaxa.5339.2.6</mods:identifier>
<mods:identifier id="BE39F043A45409C6366E36E54B731BB0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A1F9255341EE4F515B7E6FAF04AC2898" type="Zenodo-Dep">8297316</mods:identifier>
<mods:identifier id="9B4ED3B6AA6CE12BEA3BC34CD1CC437C" type="ZooBank">6C4BD5F2-951D-4B34-8AC0-158C609CC07B</mods:identifier>
</mods:mods>
<treatment id="D20987A0FFCE1C4FD293FDF7FBCD0595" LSID="urn:lsid:plazi:treatment:D20987A0FFCE1C4FD293FDF7FBCD0595" httpUri="http://treatment.plazi.org/id/D20987A0FFCE1C4FD293FDF7FBCD0595" lastPageId="2" lastPageNumber="197" pageId="1" pageNumber="197">
<subSubSection id="12BA653DFFCE1C4CD293FDF7FDBC0679" box="[151,548,547,574]" pageId="1" pageNumber="197" type="nomenclature">
<paragraph id="5A1F36B6FFCE1C4CD293FDF7FDBC0679" blockId="1.[151,548,547,609]" box="[151,548,547,574]" pageId="1" pageNumber="197">
<heading id="015781DAFFCE1C4CD293FDF7FDBC0679" bold="true" box="[151,548,547,574]" fontSize="11" level="1" pageId="1" pageNumber="197" reason="1">
<emphasis id="68D4EAA4FFCE1C4CD293FDF7FDBC0679" bold="true" box="[151,548,547,574]" pageId="1" pageNumber="197">
<taxonomicName id="9DA04D35FFCE1C4CD293FDF7FE27067A" authority="Bilton &amp; Bird, 2023" authorityName="Bilton &amp; Bird" authorityYear="2023" box="[151,447,547,574]" class="Insecta" family="Epimetopidae" genus="Eupotemus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="tuberculatus" status="sp. nov.">
<emphasis id="68D4EAA4FFCE1C4CD293FDF7FE27067A" bold="true" box="[151,447,547,574]" italics="true" pageId="1" pageNumber="197">Eupotemus tuberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73E757DFFFCE1C4CD3C2FDF7FDBC0679" box="[454,548,547,573]" pageId="1" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="12BA653DFFCE1C4CD293FD93FE8A0625" box="[151,274,583,609]" pageId="1" pageNumber="197" type="description">
<paragraph id="5A1F36B6FFCE1C4CD293FD93FE8A0625" blockId="1.[151,548,547,609]" box="[151,274,583,609]" pageId="1" pageNumber="197">
(
<figureCitation id="C29B2A33FFCE1C4CD29BFD93FE910625" box="[159,265,583,609]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[152,255,1939,1963]" captionStartId-1="3.[152,255,1874,1898]" captionTargetBox-0="[188,1398,499,1900]" captionTargetBox-1="[287,1300,181,1850]" captionTargetId-0="figure-165@2.[175,1411,486,1913]" captionTargetId-1="figure-28@3.[287,1300,181,1850]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Eupotemus tuberculatus sp. nov., holotype.A) dorsal habitus; B) lateral habitus; C) fronto-dorsal view; D) aedeagus dorsal; E) aedeadus lateral; F) aedeagus ventral. Scale bars A–C) = 0.5 mm; D–F) = 100 μm." captionText-1="FIGURE 2. Eupotemus carinaticollis group, confirmed distribution and habitat. A) Distribution of E. carinaticollis group species, red symbol = E. taianus Fikáček et al., 2021, green symbols = E. carinaticollis (Basilewsky, 1956), yellow symbol = E. uluguru Fikáček et al., 2021, blue star = E. tuberculatus sp. nov.; B) Type locality of E. tuberculatus sp. nov. South Africa, Northern Province, Magaliesberg, Mountain Sanctuary Park, margins of mountain stream, 1430 m – red arrow indicates area where specimens were found." figureDoi-0="http://doi.org/10.5281/zenodo.8297318" figureDoi-1="http://doi.org/10.5281/zenodo.8297322" httpUri-0="https://zenodo.org/record/8297318/files/figure.png" httpUri-1="https://zenodo.org/record/8297322/files/figure.png" pageId="1" pageNumber="197">Figs. 1–2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="12BA653DFFCE1C4CD293FD5BFC56077D" pageId="1" pageNumber="197" type="materials_examined">
<paragraph id="5A1F36B6FFCE1C4CD293FD5BFD7E068A" blockId="1.[151,1437,655,2014]" pageId="1" pageNumber="197">
<emphasis id="68D4EAA4FFCE1C4CD293FD5BFEA206ED" bold="true" box="[151,314,655,681]" pageId="1" pageNumber="197">Type locality.</emphasis>
<materialsCitation id="EAC83CEBFFCE1C4CD340FD5BFD7A068A" country="South Africa" elevation="1430" location="Mountain Sanctuary Park" municipality="Magaliesberg" pageId="1" pageNumber="197" specimenCount="1" stateProvince="North West">
<emphasis id="68D4EAA4FFCE1C4CD340FD5BFE7806ED" bold="true" box="[324,480,655,681]" pageId="1" pageNumber="197">
<collectingCountry id="22B77626FFCE1C4CD340FD5BFE7806ED" box="[324,480,655,681]" name="South Africa" pageId="1" pageNumber="197">South Africa</collectingCountry>
</emphasis>
,
<collectingRegion id="9864F854FFCE1C4CD3F6FD5BFD7E06ED" box="[498,742,655,682]" country="South Africa" name="North West" pageId="1" pageNumber="197">North West Province</collectingRegion>
,
<collectingMunicipality id="BA7BACCCFFCE1C4CD0F2FD5BFC0906ED" box="[758,913,655,681]" pageId="1" pageNumber="197">Magaliesberg</collectingMunicipality>
,
<location id="5F7F606DFFCE1C4CD1A4FD5BFB5106ED" LSID="urn:lsid:plazi:treatment:D20987A0FFCE1C4FD293FDF7FBCD0595:5F7F606DFFCE1C4CD1A4FD5BFB5106ED" box="[928,1225,655,681]" country="South Africa" municipality="Magaliesberg" name="Mountain Sanctuary Park" pageId="1" pageNumber="197" stateProvince="North West">Mountain Sanctuary Park</location>
, margins of small mountain stream, -25.837929°, 27.468740°,
<quantity id="9D589B53FFCE1C4CD08EFD67FD7A068A" box="[650,738,691,718]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="1" pageNumber="197" unit="m" value="1430.0">
<elevation id="D18DD185FFCE1C4CD08EFD67FD7A068A" box="[650,738,691,718]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="1" pageNumber="197" unit="m" value="1430.0">1430 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="5A1F36B6FFCE1C4CD2C3FD03FA9D0751" blockId="1.[151,1437,655,2014]" pageId="1" pageNumber="197">
<emphasis id="68D4EAA4FFCE1C4CD2C3FD03FEE306B5" bold="true" box="[199,379,727,753]" pageId="1" pageNumber="197">Type material.</emphasis>
<materialsCitation id="EAC83CEBFFCE1C4CD383FD03FA9D0751" collectingDate="2023-02-16" collectionCode="T, AMG" collectorName="Bilton" country="South Africa" elevation="1430" location="Magaliesberg" municipality="Mountain Sanctuary Park" pageId="1" pageNumber="197" specimenCount="2" typeStatus="holotype">
<emphasis id="68D4EAA4FFCE1C4CD383FD03FE6C06B5" bold="true" box="[391,500,727,753]" pageId="1" pageNumber="197">
<typeStatus id="851B8814FFCE1C4CD383FD03FE6C06B5" box="[391,500,727,753]" pageId="1" pageNumber="197">Holotype</typeStatus>
</emphasis>
(male): “
<date id="2E1E1076FFCE1C4CD06DFD03FD4606B5" box="[617,734,727,753]" pageId="1" pageNumber="197" value="2023-02-16">
<collectingDate id="3E5AE99EFFCE1C4CD06DFD03FD4606B5" box="[617,734,727,753]" pageId="1" pageNumber="197" value="2023-02-16">16/ii/2023</collectingDate>
</date>
<collectingCountry id="22B77626FFCE1C4CD0EFFD03FC1806B5" box="[747,896,727,753]" name="South Africa" pageId="1" pageNumber="197">South Africa</collectingCountry>
NWP//
<location id="5F7F606DFFCE1C4CD1E2FD03FBE606B5" LSID="urn:lsid:plazi:treatment:D20987A0FFCE1C4FD293FDF7FBCD0595:5F7F606DFFCE1C4CD1E2FD03FBE606B5" box="[998,1150,727,753]" country="South Africa" municipality="Mountain Sanctuary Park" name="Magaliesberg" pageId="1" pageNumber="197">Magaliesberg</location>
– stream in//
<collectingMunicipality id="BA7BACCCFFCE1C4CD72BFD03FED90751" pageId="1" pageNumber="197">Mountain Sanctuary Park</collectingMunicipality>
// -25.837929°, 27.468740°,
<quantity id="9D589B53FFCE1C4CD086FD2FFD4E0751" box="[642,726,763,789]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="1" pageNumber="197" unit="m" value="1430.0">
<elevation id="D18DD185FFCE1C4CD086FD2FFD4E0751" box="[642,726,763,789]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="1" pageNumber="197" unit="m" value="1430.0">1430 m</elevation>
</quantity>
// D
<collectionCode id="3CB1AE73FFCE1C4CD10CFD28FC810752" box="[776,793,764,790]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="197">T</collectionCode>
<collectorName id="F7555360FFCE1C4CD11BFD2FFCFD0751" box="[799,869,763,789]" pageId="1" pageNumber="197">Bilton</collectorName>
leg.” and red
<typeStatus id="851B8814FFCE1C4CD607FD2FFBFD0751" box="[1027,1125,763,789]" pageId="1" pageNumber="197">holotype</typeStatus>
label (
<collectionCode id="3CB1AE73FFCE1C4CD6B6FD2FFB620751" box="[1202,1274,763,789]" country="South Africa" httpUri="http://grbio.org/cool/b9qz-7eka" name="Albany Museum" pageId="1" pageNumber="197">AMG</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="5A1F36B6FFCE1C4CD2C3FCCBFC56077D" blockId="1.[151,1437,655,2014]" box="[199,974,799,826]" pageId="1" pageNumber="197">
<materialsCitation id="EAC83CEBFFCE1C4CD2C3FCCBFC52077D" box="[199,970,799,826]" collectingDate="2023-02-16" collectionCode="CBP" collectorName="Bilton" country="South Africa" elevation="1" location="Magaliesberg" municipality="Mountain Sanctuary Park" pageId="1" pageNumber="197" specimenCount="2" typeStatus="paratype">
<emphasis id="68D4EAA4FFCE1C4CD2C3FCCBFEAC077D" bold="true" box="[199,308,799,825]" pageId="1" pageNumber="197">
<typeStatus id="851B8814FFCE1C4CD2C3FCCBFEAC077D" box="[199,308,799,825]" pageId="1" pageNumber="197">Paratype</typeStatus>
</emphasis>
:
<quantity id="9D589B53FFCE1C4CD347FCCBFEFE077D" box="[323,358,799,825]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="197" unit="m" value="1.0">
<elevation id="D18DD185FFCE1C4CD347FCCBFEFE077D" box="[323,358,799,825]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="197" unit="m" value="1.0">1m</elevation>
</quantity>
#, same data as holotype and red
<typeStatus id="851B8814FFCE1C4CD0DEFCF4FCA2077E" box="[730,826,800,826]" pageId="1" pageNumber="197">paratype</typeStatus>
label (
<collectionCode id="3CB1AE73FFCE1C4CD183FCCBFC59077D" box="[903,961,799,825]" pageId="1" pageNumber="197">CBP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="12BA653DFFCE1C4CD2C3FC97FB35013D" pageId="1" pageNumber="197" type="description">
<paragraph id="5A1F36B6FFCE1C4CD2C3FC97FEAC07C6" blockId="1.[151,1437,655,2014]" pageId="1" pageNumber="197">
<emphasis id="68D4EAA4FFCE1C4CD2C3FC97FEC10719" bold="true" box="[199,345,835,861]" pageId="1" pageNumber="197">Description.</emphasis>
Size:
<typeStatus id="851B8814FFCE1C4CD3A4FC97FD950719" box="[416,525,835,861]" pageId="1" pageNumber="197">Holotype</typeStatus>
: BL
<quantity id="9D589B53FFCE1C4CD046FC97FD32071A" box="[578,682,835,862]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="197" unit="mm" value="2.9">2.90 mm</quantity>
; EL
<quantity id="9D589B53FFCE1C4CD0DBFC97FCDE071A" box="[735,838,835,862]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="197" unit="mm" value="1.9">1.90 mm</quantity>
; EW
<quantity id="9D589B53FFCE1C4CD181FC97FC75071A" box="[901,1005,835,862]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="197" unit="mm" value="1.6">1.60 mm</quantity>
.
<typeStatus id="851B8814FFCE1C4CD1FCFC90FBC6071A" box="[1016,1118,836,862]" pageId="1" pageNumber="197">Paratype</typeStatus>
: BL
<quantity id="9D589B53FFCE1C4CD690FC97FB63071A" box="[1172,1275,835,862]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="1" pageNumber="197" unit="mm" value="2.75">2.75 mm</quantity>
; EL
<quantity id="9D589B53FFCE1C4CD734FC97FA00071A" box="[1328,1432,835,862]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="197" unit="mm" value="1.8">1.80 mm</quantity>
; EW
<quantity id="9D589B53FFCE1C4CD2CDFCB3FEA807C6" box="[201,304,871,898]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="197" unit="mm" value="1.5">1.50 mm</quantity>
.
</paragraph>
<paragraph id="5A1F36B6FFCE1C4CD2C3FC58FE610149" blockId="1.[151,1437,655,2014]" pageId="1" pageNumber="197">
Dorsum (
<figureCitation id="C29B2A33FFCE1C4CD336FC5FFEEC07E1" box="[306,372,907,933]" captionStart="FIGURE 1" captionStartId="2.[152,255,1939,1963]" captionTargetBox="[188,1398,499,1900]" captionTargetId="figure-165@2.[175,1411,486,1913]" captionTargetPageId="2" captionText="FIGURE 1. Eupotemus tuberculatus sp. nov., holotype.A) dorsal habitus; B) lateral habitus; C) fronto-dorsal view; D) aedeagus dorsal; E) aedeadus lateral; F) aedeagus ventral. Scale bars A–C) = 0.5 mm; D–F) = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8297318" httpUri="https://zenodo.org/record/8297318/files/figure.png" pageId="1" pageNumber="197">Fig. 1</figureCitation>
) predominantly dark chestnut brown to black; palpi and antennae straw yellow, last antennal segment darker; femora reddish, tarsi orange-brown. Front of head and pronotal and elytral margins paler. Venter dark reddish brown; labium paler.
<typeStatus id="851B8814FFCE1C4CD010FC00FDEE07AA" box="[532,630,980,1006]" pageId="1" pageNumber="197">Paratype</typeStatus>
is somewhat paler due to tenerality. Habitus and sculpture as in
<figureCitation id="C29B2A33FFCE1C4CD73EFC07FF350055" captionStart="FIGURE 1" captionStartId="2.[152,255,1939,1963]" captionTargetBox="[188,1398,499,1900]" captionTargetId="figure-165@2.[175,1411,486,1913]" captionTargetPageId="2" captionText="FIGURE 1. Eupotemus tuberculatus sp. nov., holotype.A) dorsal habitus; B) lateral habitus; C) fronto-dorsal view; D) aedeagus dorsal; E) aedeadus lateral; F) aedeagus ventral. Scale bars A–C) = 0.5 mm; D–F) = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8297318" httpUri="https://zenodo.org/record/8297318/files/figure.png" pageId="1" pageNumber="197">Fig. 1A– C</figureCitation>
. Lateral ridge of the pronotum narrowly interrupted in middle. Ridge on elytral interval 3 complete throughout. Ridge on interval 5 interrupted anteriorly and posteriorly: short ridge present around shoulder, followed by interruption over approx. 1/6 of elytral length; ridge present over following 1/2 of elytral length, lower posteriorly; interrupted again after posterior declivity, with distinct, isolated tubercle approx. halfway to elytral apex. Ridge on elytral interval 7 complete to posterior declivity, then obsolete, with isolated tubercle(s) posteriorly, final one being continuous with tubercles on interval 9. Elytral interval 9 with two short ridges, one positioned appro x. 1/3, the other approx. 2/3 along elytral length; apically with tubercles continuous with those on interval 7. Elytral punctures connected by low elevated line.
</paragraph>
<paragraph id="5A1F36B6FFCE1C4CD2C3FAC3FB35013D" blockId="1.[151,1437,655,2014]" pageId="1" pageNumber="197">
Aedeagus (
<figureCitation id="C29B2A33FFCE1C4CD340FAC3FE230175" box="[324,443,1303,1329]" captionStart="FIGURE 1" captionStartId="2.[152,255,1939,1963]" captionTargetBox="[188,1398,499,1900]" captionTargetId="figure-165@2.[175,1411,486,1913]" captionTargetPageId="2" captionText="FIGURE 1. Eupotemus tuberculatus sp. nov., holotype.A) dorsal habitus; B) lateral habitus; C) fronto-dorsal view; D) aedeagus dorsal; E) aedeadus lateral; F) aedeagus ventral. Scale bars A–C) = 0.5 mm; D–F) = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.8297318" httpUri="https://zenodo.org/record/8297318/files/figure.png" pageId="1" pageNumber="197">Fig. 1D–F</figureCitation>
):
<quantity id="9D589B53FFCE1C4CD3CAFAC3FDE80176" box="[462,624,1303,1330]" metricMagnitude="-4" metricUnit="m" metricValue="8.85" metricValueMax="9.2" metricValueMin="8.5" pageId="1" pageNumber="197" unit="mm" value="0.885" valueMax="0.92" valueMin="0.85">0.85–0.92 mm</quantity>
long. Parameres ca. 1.9× longer than phallobase, constricted subapically, widened at apex. Median lobe 3.5× longer than wide, apical part narrowing in apical fourth, sides of median lobe narrowing to apex in a straight line. Phallobase basally with narrow symmetrical manubrium.
</paragraph>
</subSubSection>
<subSubSection id="12BA653DFFCE1C4CD2C3FA57FC430341" pageId="1" pageNumber="197" type="diagnosis">
<paragraph id="5A1F36B6FFCE1C4CD2C3FA57FC430341" blockId="1.[151,1437,655,2014]" pageId="1" pageNumber="197">
<emphasis id="68D4EAA4FFCE1C4CD2C3FA57FE5701D9" bold="true" box="[199,463,1411,1437]" pageId="1" pageNumber="197">Differential diagnosis.</emphasis>
The new species clearly belongs to the
<taxonomicName id="9DA04D35FFCE1C4CD197FA50FB3401D9" box="[915,1196,1411,1437]" class="Insecta" family="Epimetopidae" genus="Eupotemus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="carinaticollis">
<emphasis id="68D4EAA4FFCE1C4CD197FA50FB3401D9" box="[915,1196,1411,1437]" italics="true" pageId="1" pageNumber="197">Eupotemus carinaticollis</emphasis>
</taxonomicName>
group
<emphasis id="68D4EAA4FFCE1C4CD6FAFA50FAA401D9" box="[1278,1340,1412,1437]" italics="true" pageId="1" pageNumber="197">sensu</emphasis>
<bibRefCitation id="3E314B47FFCE1C4CD740FA57FEBB0185" author="Fikacek, M. &amp; Matsumoto, K. &amp; Perkins, P. D. &amp; Prokin, A. &amp; Sazhnez, A. &amp; Litovkin, S. &amp; Jach, M. A." pageId="1" pageNumber="197" pagination="1 - 34" refId="ref2086" refString="Fikacek, M., Matsumoto, K., Perkins, P. D., Prokin, A., Sazhnez, A., Litovkin, S. &amp; Jach, M. A. (2021) The family Epimetopidae (Coleoptera: Hydrophiloidea): review of current knowledge, genus-level phylogeny, and taxonomic revision of Eupotemus. Acta Entomologica Musei Nationalis Pragae, 61, 1 - 34. https: // doi. org / 10.37520 / aemnp. 2021.001" type="journal article" year="2021">
Fikáček
<emphasis id="68D4EAA4FFCE1C4CD293FA7CFF570185" box="[151,207,1447,1473]" italics="true" pageId="1" pageNumber="197">et al.</emphasis>
(2021)
</bibRefCitation>
, based on the lateral ridge of the pronotum being interrupted in the middle and similarities in aedeagal anatomy. Based on the
<typeStatus id="851B8814FFCE1C4CD39AFA1FFD9E01A1" box="[414,518,1483,1509]" pageId="1" pageNumber="197">Holotype</typeStatus>
and
<typeStatus id="851B8814FFCE1C4CD03AFA18FD3B01A2" box="[574,675,1484,1510]" pageId="1" pageNumber="197">Paratype</typeStatus>
,
<taxonomicName id="9DA04D35FFCE1C4CD0AAFA18FCC101A1" authority="Bilton &amp; Bird, 2023" authorityName="Bilton &amp; Bird" authorityYear="2023" box="[686,857,1483,1509]" class="Insecta" family="Epimetopidae" genus="Eupotemus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="tuberculatus" status="sp. nov.">
<emphasis id="68D4EAA4FFCE1C4CD0AAFA18FCC101A1" box="[686,857,1483,1509]" italics="true" pageId="1" pageNumber="197">E. tuberculatus</emphasis>
</taxonomicName>
<emphasis id="68D4EAA4FFCE1C4CD165FA1FFC2301A1" bold="true" box="[865,955,1483,1509]" pageId="1" pageNumber="197">
<taxonomicNameLabel id="73E757DFFFCE1C4CD165FA1FFC2301A1" box="[865,955,1483,1509]" pageId="1" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
would be readily distinguished from other member of the group by the strong posterior elytral tubercles and the pattern of development of elytral ridges. Posterior tubercles are present in both known specimens, but slightly less strongly developed in the
<typeStatus id="851B8814FFCE1C4CD700F9C0FAFC026A" box="[1284,1380,1556,1582]" pageId="1" pageNumber="197">paratype</typeStatus>
than the
<typeStatus id="851B8814FFCE1C4CD2C6F9E3FEB00215" box="[194,296,1591,1617]" pageId="1" pageNumber="197">holotype</typeStatus>
. Given that
<bibRefCitation id="3E314B47FFCE1C4CD3B1F9E3FD3D0215" author="Fikacek, M. &amp; Matsumoto, K. &amp; Perkins, P. D. &amp; Prokin, A. &amp; Sazhnez, A. &amp; Litovkin, S. &amp; Jach, M. A." box="[437,677,1591,1617]" pageId="1" pageNumber="197" pagination="1 - 34" refId="ref2086" refString="Fikacek, M., Matsumoto, K., Perkins, P. D., Prokin, A., Sazhnez, A., Litovkin, S. &amp; Jach, M. A. (2021) The family Epimetopidae (Coleoptera: Hydrophiloidea): review of current knowledge, genus-level phylogeny, and taxonomic revision of Eupotemus. Acta Entomologica Musei Nationalis Pragae, 61, 1 - 34. https: // doi. org / 10.37520 / aemnp. 2021.001" type="journal article" year="2021">
Fikáček
<emphasis id="68D4EAA4FFCE1C4CD013F9ECFDC90215" box="[535,593,1591,1617]" italics="true" pageId="1" pageNumber="197">et al.</emphasis>
(2021)
</bibRefCitation>
note that elytral sculpture can be variable in long series, the most reliable means of identification is the form of the aedeagus. This has the same basic design as seen in other members of the
<taxonomicName id="9DA04D35FFCE1C4CD2E1F9ABFEE202DD" box="[229,378,1663,1689]" class="Insecta" family="Epimetopidae" genus="Eupotemus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="carinaticollis">
<emphasis id="68D4EAA4FFCE1C4CD2E1F9ABFEE202DD" box="[229,378,1663,1689]" italics="true" pageId="1" pageNumber="197">carinaticollis</emphasis>
</taxonomicName>
group, and shares paramere apices which are flattened in dorsal and ventral views with
<taxonomicName id="9DA04D35FFCE1C4CD780F954FD9202FA" authority="(Basilewsky, 1956)" baseAuthorityName="Basilewsky" baseAuthorityYear="1956" class="Insecta" family="Epimetopidae" genus="Eupotemus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="carinaticollis">
<emphasis id="68D4EAA4FFCE1C4CD780F954FEB402F9" italics="true" pageId="1" pageNumber="197">E. carinaticollis</emphasis>
(
<bibRefCitation id="3E314B47FFCE1C4CD33EF977FD9902FA" author="Basilewsky, P." box="[314,513,1699,1726]" pageId="1" pageNumber="197" pagination="172 - 174" refId="ref2036" refString="Basilewsky, P. (1956) Contributions a lʼetude de la faune entomologique du Ruanda-Urundi (Mission P. Basilewsky 1953). LXXXII. Coleoptera Georyssidae. Annales du Musee Royal du Congo Belge, Serie in 8, Sciences Zoologiques, 51, 172 - 174." type="journal article" year="1956">Basilewsky, 1956</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="9DA04D35FFCE1C4CD03BF970FC0C02F9" authority="Fikacek et al., 2021" authorityName="Fikacek" authorityYear="2021" box="[575,916,1699,1726]" class="Insecta" family="Epimetopidae" genus="Eupotemus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="taianus">
<emphasis id="68D4EAA4FFCE1C4CD03BF970FD3702F9" box="[575,687,1700,1725]" italics="true" pageId="1" pageNumber="197">E. taianus</emphasis>
<bibRefCitation id="3E314B47FFCE1C4CD0B2F977FC0C02F9" author="Fikacek, M. &amp; Matsumoto, K. &amp; Perkins, P. D. &amp; Prokin, A. &amp; Sazhnez, A. &amp; Litovkin, S. &amp; Jach, M. A." box="[694,916,1699,1726]" pageId="1" pageNumber="197" pagination="1 - 34" refId="ref2086" refString="Fikacek, M., Matsumoto, K., Perkins, P. D., Prokin, A., Sazhnez, A., Litovkin, S. &amp; Jach, M. A. (2021) The family Epimetopidae (Coleoptera: Hydrophiloidea): review of current knowledge, genus-level phylogeny, and taxonomic revision of Eupotemus. Acta Entomologica Musei Nationalis Pragae, 61, 1 - 34. https: // doi. org / 10.37520 / aemnp. 2021.001" type="journal article" year="2021">
Fikáček
<emphasis id="68D4EAA4FFCE1C4CD110F970FCD402F9" box="[788,844,1699,1725]" italics="true" pageId="1" pageNumber="197">et al.</emphasis>
, 2021
</bibRefCitation>
</taxonomicName>
. Compared to these species
<taxonomicName id="9DA04D35FFCE1C4CD6CFF970FAED02F9" authority="Bilton &amp; Bird, 2023" authorityName="Bilton &amp; Bird" authorityYear="2023" box="[1227,1397,1699,1725]" class="Insecta" family="Epimetopidae" genus="Eupotemus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="tuberculatus" status="sp. nov.">
<emphasis id="68D4EAA4FFCE1C4CD6CFF970FAED02F9" box="[1227,1397,1699,1725]" italics="true" pageId="1" pageNumber="197">E. tuberculatus</emphasis>
</taxonomicName>
<emphasis id="68D4EAA4FFCE1C4CD77FF977FF5002A5" bold="true" pageId="1" pageNumber="197">
<taxonomicNameLabel id="73E757DFFFCE1C4CD77FF977FF5002A5" pageId="1" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a less pronounced lateral constriction in the parameres, and a somewhat broader, more rounded apex to the median lobe. For comparative features of aedeagi in the group see
<tableCitation id="1722030DFFCE1C4CD185F93FFC4D0341" box="[897,981,1771,1797]" captionStart="TABLE 1" captionStartId="4.[152,244,152,176]" captionText="TABLE 1. Aedeagal features of the Eupotemus carinaticollis group (after Fikáček et al. (2021))." pageId="1" pageNumber="197">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="12BA653DFFCE1C4CD2C3F8DBFB22036D" box="[199,1210,1807,1834]" pageId="1" pageNumber="197" type="etymology">
<paragraph id="5A1F36B6FFCE1C4CD2C3F8DBFB22036D" blockId="1.[151,1437,655,2014]" box="[199,1210,1807,1834]" pageId="1" pageNumber="197">
<emphasis id="68D4EAA4FFCE1C4CD2C3F8DBFED6036D" bold="true" box="[199,334,1807,1833]" pageId="1" pageNumber="197">Etymology.</emphasis>
Named in reference to the tuberculate elytral dorsum, particularly posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="12BA653DFFCE1C4FD2C3F8E7FBCD0595" lastPageId="2" lastPageNumber="198" pageId="1" pageNumber="197" type="distribution">
<paragraph id="5A1F36B6FFCE1C4FD2C3F8E7FBCD0595" blockId="1.[151,1437,655,2014]" lastBlockId="2.[151,1437,151,465]" lastPageId="2" lastPageNumber="198" pageId="1" pageNumber="197">
<emphasis id="68D4EAA4FFCE1C4CD2C3F8E7FE600309" bold="true" box="[199,504,1843,1869]" pageId="1" pageNumber="197">Distribution and ecology.</emphasis>
This is the first epimetopid confirmed from southern Africa. Geographically, the closest known species are
<taxonomicName id="9DA04D35FFCE1C4CD3BCF88CFDF30335" baseAuthorityName="Basilewsky" baseAuthorityYear="1956" box="[440,619,1879,1905]" class="Insecta" family="Epimetopidae" genus="Eupotemus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="carinaticollis">
<emphasis id="68D4EAA4FFCE1C4CD3BCF88CFDF30335" box="[440,619,1879,1905]" italics="true" pageId="1" pageNumber="197">E. carinaticollis</emphasis>
</taxonomicName>
from
<collectingCountry id="22B77626FFCE1C4CD0A8F883FB9E0335" box="[684,1030,1879,1905]" name="Democratic Republic of the Congo" pageId="1" pageNumber="197">Democratic Republic of Congo</collectingCountry>
and
<collectingCountry id="22B77626FFCE1C4CD63DF883FB0D0335" box="[1081,1173,1879,1905]" name="Burundi" pageId="1" pageNumber="197">Burundi</collectingCountry>
and
<taxonomicName id="9DA04D35FFCE1C4CD6CCF88CFE8F03D1" authority="Fikacek et al., 2021" authorityName="Fikacek" authorityYear="2021" class="Insecta" family="Epimetopidae" genus="Eupotemus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="uluguru">
<emphasis id="68D4EAA4FFCE1C4CD6CCF88CFAA70335" box="[1224,1343,1879,1905]" italics="true" pageId="1" pageNumber="197">E. uluguru</emphasis>
<bibRefCitation id="3E314B47FFCE1C4CD740F883FE8F03D1" author="Fikacek, M. &amp; Matsumoto, K. &amp; Perkins, P. D. &amp; Prokin, A. &amp; Sazhnez, A. &amp; Litovkin, S. &amp; Jach, M. A." pageId="1" pageNumber="197" pagination="1 - 34" refId="ref2086" refString="Fikacek, M., Matsumoto, K., Perkins, P. D., Prokin, A., Sazhnez, A., Litovkin, S. &amp; Jach, M. A. (2021) The family Epimetopidae (Coleoptera: Hydrophiloidea): review of current knowledge, genus-level phylogeny, and taxonomic revision of Eupotemus. Acta Entomologica Musei Nationalis Pragae, 61, 1 - 34. https: // doi. org / 10.37520 / aemnp. 2021.001" type="journal article" year="2021">
Fikáček
<emphasis id="68D4EAA4FFCE1C4CD293F8A8FF4903D1" box="[151,209,1915,1941]" italics="true" pageId="1" pageNumber="197">et al.</emphasis>
, 2021
</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="22B77626FFCE1C4CD35AF8AFFE5B03D1" box="[350,451,1915,1941]" name="Tanzania" pageId="1" pageNumber="197">Tanzania</collectingCountry>
(
<figureCitation id="C29B2A33FFCE1C4CD3D7F8AFFDB603D1" box="[467,558,1915,1941]" captionStart="FIGURE 2" captionStartId="3.[152,255,1874,1898]" captionTargetBox="[287,1300,181,1850]" captionTargetId="figure-28@3.[287,1300,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Eupotemus carinaticollis group, confirmed distribution and habitat. A) Distribution of E. carinaticollis group species, red symbol = E. taianus Fikáček et al., 2021, green symbols = E. carinaticollis (Basilewsky, 1956), yellow symbol = E. uluguru Fikáček et al., 2021, blue star = E. tuberculatus sp. nov.; B) Type locality of E. tuberculatus sp. nov. South Africa, Northern Province, Magaliesberg, Mountain Sanctuary Park, margins of mountain stream, 1430 m – red arrow indicates area where specimens were found." figureDoi="http://doi.org/10.5281/zenodo.8297322" httpUri="https://zenodo.org/record/8297322/files/figure.png" pageId="1" pageNumber="197">Fig. 2A</figureCitation>
).
<bibRefCitation id="3E314B47FFCE1C4CD047F8AFFD6D03D1" author="Hebauer, F." box="[579,757,1915,1941]" pageId="1" pageNumber="197" pagination="7 - 16" refId="ref2174" refString="Hebauer, F. (1999) Neue und wenig bekannte Hydrophiloidea aus dem sudlichen Afrika (Coleoptera, Hydrophiloidea). Acta Coleopterologica, 15, 7 - 16." type="journal article" year="1999">Hebauer (1999)</bibRefCitation>
notes a single specimen from
<collectingCountry id="22B77626FFCE1C4CD654F8AFFB5003D1" box="[1104,1224,1915,1941]" name="Zimbabwe" pageId="1" pageNumber="197">Zimbabwe</collectingCountry>
he attributed to
<taxonomicName id="9DA04D35FFCE1C4CD780F8A8FEB403FD" baseAuthorityName="Basilewsky" baseAuthorityYear="1956" class="Insecta" family="Epimetopidae" genus="Eupotemus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="carinaticollis">
<emphasis id="68D4EAA4FFCE1C4CD780F8A8FEB403FD" italics="true" pageId="1" pageNumber="197">E. carinaticollis</emphasis>
</taxonomicName>
in the Hungarian Natural History Museum, Budapest. Assuming this record is correct, the specimen may belong to
<taxonomicName id="9DA04D35FFCE1C4CD340F810FE770399" authority="Bilton &amp; Bird, 2023" authorityName="Bilton &amp; Bird" authorityYear="2023" box="[324,495,1987,2013]" class="Insecta" family="Epimetopidae" genus="Eupotemus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="197" phylum="Arthropoda" rank="species" species="tuberculatus" status="sp. nov.">
<emphasis id="68D4EAA4FFCE1C4CD340F810FE770399" box="[324,495,1987,2013]" italics="true" pageId="1" pageNumber="197">E. tuberculatus</emphasis>
</taxonomicName>
<emphasis id="68D4EAA4FFCE1C4CD3FCF817FDCB0399" bold="true" box="[504,595,1987,2013]" pageId="1" pageNumber="197">
<taxonomicNameLabel id="73E757DFFFCE1C4CD3FCF817FDCB0399" box="[504,595,1987,2013]" pageId="1" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, or another, as yet undescribed, species.
<bibRefCitation id="3E314B47FFCE1C4CD62CF817FA8F0399" author="Fikacek, M. &amp; Matsumoto, K. &amp; Perkins, P. D. &amp; Prokin, A. &amp; Sazhnez, A. &amp; Litovkin, S. &amp; Jach, M. A." box="[1064,1303,1987,2013]" pageId="1" pageNumber="197" pagination="1 - 34" refId="ref2086" refString="Fikacek, M., Matsumoto, K., Perkins, P. D., Prokin, A., Sazhnez, A., Litovkin, S. &amp; Jach, M. A. (2021) The family Epimetopidae (Coleoptera: Hydrophiloidea): review of current knowledge, genus-level phylogeny, and taxonomic revision of Eupotemus. Acta Entomologica Musei Nationalis Pragae, 61, 1 - 34. https: // doi. org / 10.37520 / aemnp. 2021.001" type="journal article" year="2021">
Fikáček
<emphasis id="68D4EAA4FFCE1C4CD68EF810FB5C0399" box="[1162,1220,1987,2013]" italics="true" pageId="1" pageNumber="197">et al.</emphasis>
(2021)
</bibRefCitation>
did not see this specimen, despite examining material from Budapest, and enquiries about this specimen by DTB have proved fruitless.
<emphasis id="68D4EAA4FFCD1C4FD2FBFF6FFE300491" box="[255,424,187,213]" italics="true" pageId="2" pageNumber="198">
E,
<taxonomicName id="9DA04D35FFCD1C4FD319FF6FFE300491" authority="Bilton &amp; Bird, 2023" authorityName="Bilton &amp; Bird" authorityYear="2023" box="[285,424,187,213]" class="Insecta" family="Epimetopidae" genus="Eupotemus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="tuberculatus" status="sp. nov.">tuberculatus</taxonomicName>
</emphasis>
<emphasis id="68D4EAA4FFCD1C4FD3AAFF6FFD9F0491" bold="true" box="[430,519,187,213]" pageId="2" pageNumber="198">
<taxonomicNameLabel id="73E757DFFFCD1C4FD3AAFF6FFD9F0491" box="[430,519,187,213]" pageId="2" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known currently from a single stream within a private nature reserve (Mountain Sanctuary Park) in the Magaliesberg, north of Johannesburg,
<collectingCountry id="22B77626FFCD1C4FD131FF0BFC5D04BD" box="[821,965,223,249]" name="South Africa" pageId="2" pageNumber="198">South Africa</collectingCountry>
.The Magaliesberg Formation is an ancient (ca. 2.1 Ga) mountain range (
<bibRefCitation id="3E314B47FFCD1C4FD3E6FED7FD2F0559" author="Parizot, M. &amp; Eriksson, P. G. &amp; Aifa, T. &amp; Sarkar, S. &amp; Banerjee, S. &amp; Catuneanu, O. &amp; Altermann, W. &amp; Bumby, A. J. &amp; Bordy, E. M. &amp; van Rooy, J. L." box="[482,695,258,285]" pageId="2" pageNumber="198" pagination="274 - 296" refId="ref2456" refString="Parizot, M., Eriksson, P. G., Aifa, T., Sarkar, S., Banerjee, S., Catuneanu, O., Altermann, W., Bumby, A. J., Bordy, E. M. &amp; van Rooy, J. L. (2005) Suspected microbial mat-related crack-like sedimentary structures in the Palaeoproterozoic Magaliesberg Formation sandstones, South Africa. Precambrian Research, 138, 274 - 296. https: // doi. org / 10.1016 / j. precamres. 2005.05.007" type="journal article" year="2005">
Parizot
<emphasis id="68D4EAA4FFCD1C4FD03FFED7FDEC0559" box="[571,628,259,285]" italics="true" pageId="2" pageNumber="198">et al.</emphasis>
2005
</bibRefCitation>
), extending for approximately
<quantity id="9D589B53FFCD1C4FD616FED6FBFF0559" box="[1042,1127,258,285]" metricMagnitude="5" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="198" unit="km" value="150.0">150 km</quantity>
from Pretoria in the east to Rustenburg in the west. Due to the patchy nature of the geology of the region, headwater streams can have either quartzitic (sandstone) or dolomitic origin, with the collection site being a typical small perennial dolomitic stream with approximately neutral pH. Both specimens were washed from marginal sand and gravels in a small lateral ebayment with partial shade (
<figureCitation id="C29B2A33FFCD1C4FD3EBFE47FDD305E9" box="[495,587,403,429]" captionStart="FIGURE 2" captionStartId="3.[152,255,1874,1898]" captionTargetBox="[287,1300,181,1850]" captionTargetId="figure-28@3.[287,1300,181,1850]" captionTargetPageId="3" captionText="FIGURE 2. Eupotemus carinaticollis group, confirmed distribution and habitat. A) Distribution of E. carinaticollis group species, red symbol = E. taianus Fikáček et al., 2021, green symbols = E. carinaticollis (Basilewsky, 1956), yellow symbol = E. uluguru Fikáček et al., 2021, blue star = E. tuberculatus sp. nov.; B) Type locality of E. tuberculatus sp. nov. South Africa, Northern Province, Magaliesberg, Mountain Sanctuary Park, margins of mountain stream, 1430 m – red arrow indicates area where specimens were found." figureDoi="http://doi.org/10.5281/zenodo.8297322" httpUri="https://zenodo.org/record/8297322/files/figure.png" pageId="2" pageNumber="198">Fig. 2B</figureCitation>
).
<taxonomicName id="9DA04D35FFCD1C4FD067FE47FC8805E9" authority="Bilton &amp; Bird, 2023" authorityName="Bilton &amp; Bird" authorityYear="2023" box="[611,784,403,429]" class="Insecta" family="Epimetopidae" genus="Eupotemus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="tuberculatus" status="sp. nov.">
<emphasis id="68D4EAA4FFCD1C4FD067FE47FC8805E9" box="[611,784,403,429]" italics="true" pageId="2" pageNumber="198">E. tuberculatus</emphasis>
</taxonomicName>
<emphasis id="68D4EAA4FFCD1C4FD11FFE47FCE005E9" bold="true" box="[795,888,403,429]" pageId="2" pageNumber="198">
<taxonomicNameLabel id="73E757DFFFCD1C4FD11FFE47FCE005E9" box="[795,888,403,429]" pageId="2" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was taken together with
<taxonomicName id="9DA04D35FFCD1C4FD6A6FE47FEA90595" authority="Kuwert, 1890" authorityName="Kuwert" authorityYear="1890" class="Insecta" family="Hydrophilidae" genus="Laccobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="praecipuus">
<emphasis id="68D4EAA4FFCD1C4FD6A6FE47FA0405E8" box="[1186,1436,403,429]" italics="true" pageId="2" pageNumber="198">Laccobius praecipuus</emphasis>
Kuwert, 1890
</taxonomicName>
(
<taxonomicName id="9DA04D35FFCD1C4FD344FE63FE7D0595" authorityName="Latreille" authorityYear="1802" box="[320,485,439,465]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
) and
<taxonomicName id="9DA04D35FFCD1C4FD027FE63FC3D0595" authority="Perkins, 2014" authorityName="Perkins" authorityYear="2014" box="[547,933,439,465]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="species" species="biseptosa">
<emphasis id="68D4EAA4FFCD1C4FD027FE63FC9C0595" box="[547,772,439,465]" italics="true" pageId="2" pageNumber="198">Hydraena biseptosa</emphasis>
Perkins, 2014
</taxonomicName>
(
<taxonomicName id="9DA04D35FFCD1C4FD1B7FE63FBD10595" authorityName="Mulsant" authorityYear="1844" box="[947,1097,439,465]" class="Insecta" family="Hydraenidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="198" phylum="Arthropoda" rank="family">Hydraenidae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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