147 lines
11 KiB
XML
147 lines
11 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.669.11773" ID-GBIF-Dataset="e2f4b6e8-d909-4231-867a-08808dfc66ca" ID-Pensoft-Pub="1313-2970-669-113" ID-ZBK="13D2ECC5A53C4A19A00C406E940BADD2" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-669-113" ModsDocOrigin="ZooKeys 669" ModsDocTitle="Revision of the Neotropical water scavenger beetle genus Tobochares Short & García, 2007 (Coleoptera, Hydrophilidae, Acidocerinae)" checkinTime="1492859332780" checkinUser="pensoft" docAuthor="Kohlenberg, Alex T. & Short, Andrew Edward Z." docDate="2017" docId="D1E75077473CA150D0D32105719BD027" docLanguage="en" docName="ZooKeys 669: 113-146" docOrigin="ZooKeys 669" docSource="http://dx.doi.org/10.3897/zookeys.669.11773" docTitle="Tobochares canthus Kohlenberg & Short, 2017, sp. n." docType="treatment" docUuid="A2946B0F-ED87-4890-848E-7D2DF2CF41EA" docUuidSource="ZooBank" docVersion="5" lastPageNumber="119" masterDocId="FFB58A38FFA2FFE95A3EFF9EA741B933" masterDocTitle="Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae)" masterLastPageNumber="146" masterPageNumber="113" pageNumber="117" updateTime="1643557144543" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the Neotropical water scavenger beetle genus Tobochares Short & Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kohlenberg, Alex T.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Short, Andrew Edward Z.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>669</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>113</mods:start>
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<mods:end>146</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.669.11773</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.669.11773</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-669-113</mods:identifier>
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<mods:identifier type="ZBK">13D2ECC5A53C4A19A00C406E940BADD2</mods:identifier>
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<mods:identifier type="ZooBank">13D2ECC5A53C4A19A00C406E940BADD2</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="128696892" LSID="urn:lsid:zoobank.org:act:A2946B0F-ED87-4890-848E-7D2DF2CF41EA" httpUri="http://treatment.plazi.org/id/D1E75077473CA150D0D32105719BD027" lastPageId="6" lastPageNumber="119" pageId="4" pageNumber="117">
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<subSubSection pageId="4" pageNumber="117" type="nomenclature">
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<paragraph pageId="4" pageNumber="117">
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<taxonomicName LSID="http://zoobank.org/A2946B0F-ED87-4890-848E-7D2DF2CF41EA" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares canthus" order="Coleoptera" pageId="4" pageNumber="117" phylum="Arthropoda" rank="species" species="canthus">Tobochares canthus</taxonomicName>
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<taxonomicNameLabel pageId="4" pageNumber="117">sp. n.</taxonomicNameLabel>
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Figs 3D, 6
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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, 7D, 9A, 10A, 12D, 13B, 14H, 15, 18
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="118" type="type material">
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<paragraph pageId="5" pageNumber="118">
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<pageBreakToken pageId="5" pageNumber="118" start="start">Type</pageBreakToken>
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material.
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</paragraph>
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<paragraph pageId="5" pageNumber="118">
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Holotype (male): "VENEZULEA: Amazonas State/
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="5.386783">5°23.207'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-67.615364">67°36.922'W</geoCoordinate>
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, 125m/ Tobogan de la Selva; 8.viii.2008/ leg. A. Short, M.
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<normalizedToken originalValue="García">Garcia</normalizedToken>
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, L. Joly/ AS-08-080b; old
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<normalizedToken originalValue="“tobogancito”">"tobogancito"</normalizedToken>
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/ on seepage area w/ detritus", "[barcode]/ SEMC0877726/ KUNHM-ENT" (MIZA). Paratypes (28): VENEZUELA: Amazonas: same data as type (28 exs., SEMC, MALUZ, MIZA; includes 1 male mounted on SEM stub).
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</paragraph>
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<caption pageId="5" pageNumber="118">
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<paragraph pageId="5" pageNumber="118">
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Figure 6. Front and lateral view of heads of
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="5" pageNumber="118" phylum="Arthropoda" rank="genus">Tobochares</taxonomicName>
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spp.
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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<taxonomicName genus="T." lsidName="T. pallidus" pageId="5" pageNumber="118" rank="species" species="pallidus">T. pallidus</taxonomicName>
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sp. n.
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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<taxonomicName genus="T." lsidName="T. canthus" pageId="5" pageNumber="118" rank="species" species="canthus">T. canthus</taxonomicName>
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sp. n.
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<normalizedToken originalValue="E–F">E-F</normalizedToken>
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<taxonomicName genus="T." lsidName="T. emarginatus" pageId="5" pageNumber="118" rank="species" species="emarginatus">T. emarginatus</taxonomicName>
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sp. n.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="118" type="differential diagnosis">
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<paragraph pageId="5" pageNumber="118">Differential diagnosis.</paragraph>
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<paragraph pageId="5" pageNumber="118">
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The lack of impressed striae on the elytra (Fig. 3D), strongly emarginated eye (Figs 6C-D), lack of spicules on the surface of the abdominal ventrites, and extremely broad glabrous patch on the metaventrite (Fig. 10A) all serve to easily separate this species from other
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="5" pageNumber="118" phylum="Arthropoda" rank="genus">Tobochares</taxonomicName>
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with the exception of
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<taxonomicName genus="T." lsidName="T. emarginatus" pageId="5" pageNumber="118" rank="species" species="emarginatus">T. emarginatus</taxonomicName>
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. It may be separated from
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<taxonomicName genus="T." lsidName="T. emarginatus" pageId="5" pageNumber="118" rank="species" species="emarginatus">T. emarginatus</taxonomicName>
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by the paler dorsal coloration and the shape of the aedeagus (Fig. 14H).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="119" pageId="5" pageNumber="118" type="description">
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<paragraph pageId="5" pageNumber="118">Description.</paragraph>
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<paragraph lastPageId="6" lastPageNumber="119" pageId="5" pageNumber="118">
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Size and form: Body length 1.7-2.0 mm. Body elongate oval, moderately dorsoventrally compressed. Color and punctation. Dorsum of head dark brown, frons darker (nearly black) laterally and around eyes, anterolateral margins
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<pageBreakToken pageId="6" pageNumber="119" start="start">of</pageBreakToken>
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clypeus with paler preocular patches (Fig. 6
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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); maxillary palps distinctly pale (Fig. 7D). Pronotum dark brown with the lateral margins paler; elytra dark brown, slightly paler at lateral margins and posteriorly. Meso- and metathoracic ventrites and abdominal ventrites very dark brown, with prosternum slightly paler; legs, labial palps, and antennae distinctly paler. Ground punctation on head, pronotum and elytra moderately fine. Head. Eyes measuring ~90µm anteroposteriorly, continuous with outline of head, emarginate at lateral margin, narrowing to about a fourth of the width (Fig. 6
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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). Thorax. Elytra with punctures loosely organized into rows. Metafemora mostly glabrous on ventral face (Fig. 12D). Elevation of mesoventrite forming a low transverse carina, not quite elevated to the same plane as the ventral surface of the mesocoxae (Fig. 9A). Metaventrite with distinct median ovoid glabrous area that is more than half of the total metaventrite length, and nearly as wide as it is long (Fig. 10A). Abdomen. Abdominal ventrites uniformly and densely pubescent (Fig. 13B). Aedeagus (Fig. 14H) with parameres narrow, slightly less than half as wide as the median lobe; generally straight and parallel sided along entire length; apex of parameres not expanded, slightly tapered to a blunt tip on the medial corner; apex of median lobe distinctly surpassing the apex of the parameres; gonopore situated just below the apex of the median lobe.
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</paragraph>
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<caption pageId="6" pageNumber="119">
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<paragraph pageId="6" pageNumber="119">
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Figure 7. Ventral view of heads of
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<taxonomicName class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="genus">Tobochares</taxonomicName>
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spp. A
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<taxonomicName genus="T." lsidName="T. sipaliwini" pageId="6" pageNumber="119" rank="species" species="sipaliwini">T. sipaliwini</taxonomicName>
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B
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<taxonomicName genus="T." lsidName="T. kasikasima" pageId="6" pageNumber="119" rank="species" species="kasikasima">T. kasikasima</taxonomicName>
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C
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<taxonomicName genus="T." lsidName="T. pallidus" pageId="6" pageNumber="119" rank="species" species="pallidus">T. pallidus</taxonomicName>
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sp. n. D
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<taxonomicName genus="T." lsidName="T. canthus" pageId="6" pageNumber="119" rank="species" species="canthus">T. canthus</taxonomicName>
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sp. n.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="119" type="etymology">
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<paragraph pageId="6" pageNumber="119">Etymology.</paragraph>
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<paragraph pageId="6" pageNumber="119">
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Named after the condition of the eyes, in which a lateral
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<taxonomicName lsidName="canthus" pageId="6" pageNumber="119" rank="species" species="canthus">canthus</taxonomicName>
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of the frons partially divides them into lower and upper surfaces.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="119" type="distribution">
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<paragraph pageId="6" pageNumber="119">Distribution.</paragraph>
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<paragraph pageId="6" pageNumber="119">Known only from the type locality in Venezuela (Fig. 15).</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="119" type="biology_ecology">
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<paragraph pageId="6" pageNumber="119">Biology.</paragraph>
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<paragraph pageId="6" pageNumber="119">
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This species was collected on a rock seepage along the margin of the Rio Coromoto (Fig. 18
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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). The seepage drains water from the surrounding area and is not fed directly by the creek itself. The rock surface had scattered leaves and detritus, as well as algae in some patches.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |