175 lines
13 KiB
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175 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.672.10212" ID-GBIF-Dataset="62682ac8-f1bf-4e19-a4e3-0c24a5764441" ID-PMC="PMC5527339" ID-Pensoft-Pub="1313-2970-672-1" ID-PubMed="28769665" ID-ZBK="48810F7697B940B29A75DB0F98C5311F" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-672-1" ModsDocOrigin="ZooKeys 672" ModsDocTitle="New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae)" checkinTime="1493896456945" checkinUser="pensoft" docAuthor="Chanabun, Ratmanee, Inkavilay, Khamla & Panha, Somsak" docDate="2017" docId="D19F4D5966A6F9EF119FBC1730D3B83E" docLanguage="en" docName="ZooKeys 672: 1-34" docOrigin="ZooKeys 672" docSource="http://dx.doi.org/10.3897/zookeys.672.10212" docTitle="Glyphidrilus sekongensis Chanabun & Panha, sp. n." docType="treatment" docUuid="F9F57BA6-C6DC-46EB-8D14-9427E9ADD3E7" docUuidSource="ZooBank" docVersion="5" lastPageNumber="12" masterDocId="C91DFFAFFFFFFFF8F80DFFDC7D30FF88" masterDocTitle="New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae)" masterLastPageNumber="34" masterPageNumber="1" pageNumber="12" updateTime="1668164443210" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species of semi-aquatic freshwater earthworm genus Glyphidrilus Horst, 1889 from Thailand and Laos (Oligochaeta, Almidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chanabun, Ratmanee</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Inkavilay, Khamla</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>672</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>34</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.672.10212</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.672.10212</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-672-1</mods:identifier>
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<mods:identifier type="ZBK">48810F7697B940B29A75DB0F98C5311F</mods:identifier>
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<mods:identifier type="ZooBank">48810F7697B940B29A75DB0F98C5311F</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="128835290" LSID="urn:lsid:zoobank.org:act:F9F57BA6-C6DC-46EB-8D14-9427E9ADD3E7" httpUri="http://treatment.plazi.org/id/D19F4D5966A6F9EF119FBC1730D3B83E" lastPageNumber="12" pageId="11" pageNumber="12">
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<subSubSection pageId="11" pageNumber="12" type="nomenclature">
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<paragraph pageId="11" pageNumber="12">
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<taxonomicName LSID="http://zoobank.org/F9F57BA6-C6DC-46EB-8D14-9427E9ADD3E7" authority="Chanabun & Panha" class="Clitellata" family="Almidae" genus="Glyphidrilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphidrilus sekongensis" order="Opisthopora" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="sekongensis">Glyphidrilus sekongensis Chanabun & Panha</taxonomicName>
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<taxonomicNameLabel pageId="11" pageNumber="12">sp. n.</taxonomicNameLabel>
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Figs 1, 9, Table 1
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="type material">
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<paragraph pageId="11" pageNumber="12">Type material.</paragraph>
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<paragraph pageId="11" pageNumber="12">
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Holotype: One adult (CUMZ 3422) in a stream at Ban Kiangkong, Lamarm, Sekong, Laos (
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.5585">15°33'30.6"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="106.32205">106°19'19.4"E</geoCoordinate>
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), 472 meters elevation on 17 October 2013. Paratypes: 2 adults and 11 juveniles (CUMZ 3423) all same collection data as holotype.
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</paragraph>
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<caption pageId="11" pageNumber="12">
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<paragraph pageId="11" pageNumber="12">
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Figure 9. Morphology of holotype (CUMZ 3422) of
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<taxonomicName class="Clitellata" family="Almidae" genus="Glyphidrilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphidrilus sekongensis" order="Haplotaxida" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="sekongensis">Glyphidrilus sekongensis</taxonomicName>
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sp. n. A external ventral view, annular clitellum in
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<normalizedToken originalValue="XVI–XXXVII">XVI-XXXVII</normalizedToken>
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B internal dorsal view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="diagnosis">
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<paragraph pageId="11" pageNumber="12">Diagnosis.</paragraph>
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<paragraph pageId="11" pageNumber="12">
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<taxonomicName class="Clitellata" family="Almidae" genus="Glyphidrilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphidrilus sekongensis" order="Haplotaxida" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="sekongensis">Glyphidrilus sekongensis</taxonomicName>
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sp. n. is a small sized earthworm with the clitellar wings on the lateral side of the body in
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<normalizedToken originalValue="XXV–XXXI">XXV-XXXI</normalizedToken>
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. Clitellum in XVI,
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<normalizedToken originalValue="XVII–XXXVI">XVII-XXXVI</normalizedToken>
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, XXXVII. Female pores, male pores and spermathecal pores not visible. Genital markings: paired or asymmetrical on bc in XVI,
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<normalizedToken originalValue="XVII–XXIV">XVII-XXIV</normalizedToken>
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and XXXII. Four pairs of seminal vesicles in
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<normalizedToken originalValue="IX–XII">IX-XII</normalizedToken>
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. Intestinal origin in XVI. Ovaries in
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<normalizedToken originalValue="XIII–XIV">XIII-XIV</normalizedToken>
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. Spermathecae between 12/13-15/16.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="description">
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<paragraph pageId="11" pageNumber="12">Description of holotype.</paragraph>
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<paragraph pageId="11" pageNumber="12">
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Dimensions: body length 134 mm, diameter 2.4 mm in segment VIII, 2.3 mm before the clitellar wing in segment XXIV, 2.4 mm after wing in segment XXXII in clitellar region; body cylindrical in anterior part, quadrangular in transverse section behind clitellum. 329 segments. Body color pale brown with variations from red to pink on adjacent tissues of wing portions in different individuals of newly collected specimens. The posterior end on the dorsal surface is considerably broader than the ventral. Clitellar wing on ventro-lateral part of clitellum in
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<normalizedToken originalValue="XXV–XXXI">XXV-XXXI</normalizedToken>
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, 4.0 mm in height, and 0.5 mm in width on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular in
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<normalizedToken originalValue="XVI–XXXVII">XVI-XXXVII</normalizedToken>
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. Four pairs of setae per segment from II, setal formula aa:ab:bc:cd:dd = 1.0:0.3:1.0:0.5:1.3 in segment VIII and 1.8:0.8:1.3:0.6:1.5 in postclitellar segments. Female pores, male pores, and spermathecal pores not visible. Genital markings: laterally paired or asymmetrical on bc in
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<normalizedToken originalValue="XVI–XXIV">XVI-XXIV</normalizedToken>
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, XXXII.
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</paragraph>
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<paragraph pageId="11" pageNumber="12">
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Septa 5/6-8/9 thicker than between other segments, 9/10-14/15 thick and 15/16 to the last others segment, 9/10-14/15 thick and 15/16 to the last segment thin. Gizzard small, globular in VIII. Intestine enlarged from XVI. Hearts, four pairs in
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<normalizedToken originalValue="VIII–XI">VIII-XI</normalizedToken>
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, first in VIII and last in XI. A pair of holonephridia in each segment from segment XIII onwards. Seminal vesicles, four pairs in IX, X, XI, XII. Ovaries, two pairs in XIII and XIV. Testis free in X and XI. Prostate and accessory glands absent. Spermathecae sessile and diverticula absent, small elongated oval or globular between 12/13-15/16, about 0.1-0.3 mm in diameter, two to seven on each side per segment.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="variation">
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<paragraph pageId="11" pageNumber="12">Variations.</paragraph>
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<paragraph pageId="11" pageNumber="12">
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Body lengths of adult (n = 3) and juvenile paratypes (n = 11). Adults ranged from 90-134 mm (109.6
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<normalizedToken originalValue="±">+/-</normalizedToken>
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12.2), with 237-337 segments. Juveniles ranged from 86+-130 mm (104.2
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<normalizedToken originalValue="±">+/-</normalizedToken>
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14.0), with 209+-329 segments. Wings in
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<normalizedToken originalValue="XXV–XXXI">XXV-XXXI</normalizedToken>
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, clitellum in XVI,
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<normalizedToken originalValue="XVII–XXXVI">XVII-XXXVI</normalizedToken>
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, XXXVII. Genital markings: laterally paired or asymmetrical on bc in XVI,
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<normalizedToken originalValue="XVII–XXIV">XVII-XXIV</normalizedToken>
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, XXXII.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="distribution">
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<paragraph pageId="11" pageNumber="12">Distribution.</paragraph>
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<paragraph pageId="11" pageNumber="12">The new species was found in a stream at Ban Kiangkong, Lamarm, Sekong, Laos. Soils are slightly sandy mixed with black organic matter.</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="etymology">
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<paragraph pageId="11" pageNumber="12">Etymology.</paragraph>
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<paragraph pageId="11" pageNumber="12">The species was named after Sekong, the type locality in Laos.</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="remarks">
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<paragraph pageId="11" pageNumber="12">Remarks.</paragraph>
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<paragraph pageId="11" pageNumber="12">
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(see Table 1).
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<taxonomicName class="Clitellata" family="Almidae" genus="Glyphidrilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphidrilus sekongensis" order="Haplotaxida" pageId="11" pageNumber="12" phylum="Annelida" rank="species" species="sekongensis">Glyphidrilus sekongensis</taxonomicName>
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sp. n. differs from other species by the locations of external and internal characteristics such as wings, clitellum, the arrangement of genital markings, and spermathecae. The new species differs from
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<taxonomicName lsidName="G. vangviengensis" pageId="11" pageNumber="12" rank="species" species="vangviengensis">G. vangviengensis</taxonomicName>
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Chanabun & Panha, 2011 from Song River, Vientiane, Laos by the latter having a shorter clitellum in XIX,
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<normalizedToken originalValue="XX–XXXV">XX-XXXV</normalizedToken>
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, XXXVI, XXXVII, widely paired genital markings in bc XVIII, XIX, XX,
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<normalizedToken originalValue="XXI–XXIV">XXI-XXIV</normalizedToken>
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, XXXIII, XXXIV, paired on aa in XII, XIII, XIV, XV, and lacks spermathecae. Differences from
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<taxonomicName lsidName="G. yunnanensis" pageId="11" pageNumber="12" rank="species" species="yunnanensis">G. yunnanensis</taxonomicName>
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Chen & Xu, 1977 reported from China are that
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<taxonomicName lsidName="G. yunnanensis" pageId="11" pageNumber="12" rank="species" species="yunnanensis">G. yunnanensis</taxonomicName>
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has longer wings in
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<normalizedToken originalValue="XXII–XXXII">XXII-XXXII</normalizedToken>
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, clitellum in
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<normalizedToken originalValue="XVIII–XXXVIII">XVIII-XXXVIII</normalizedToken>
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, and lacks spermathecae. Differences from
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<taxonomicName lsidName="G. mekongensis" pageId="11" pageNumber="12" rank="species" species="mekongensis">G. mekongensis</taxonomicName>
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Panha & Chanabun, 2012 are that
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<taxonomicName lsidName="G. mekongensis" pageId="11" pageNumber="12" rank="species" species="mekongensis">G. mekongensis</taxonomicName>
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has longer wings in
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<normalizedToken originalValue="XXIV–½">XXIV-1/2</normalizedToken>
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XXXIII, XXXIII, XXXIV,
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<normalizedToken originalValue="½">1/2</normalizedToken>
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XXXV, and lacks spermathecae. It differs from
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<taxonomicName lsidName="G. huailuangensis" pageId="11" pageNumber="12" rank="species" species="huailuangensis">G. huailuangensis</taxonomicName>
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Chanabun & Panha, 2013 recorded from Najahlauy National Park, Ubon Ratchathani, northeast Thailand by the latter having a larger body size, clitellum in XII, XIII,
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<normalizedToken originalValue="XVI–XXXII">XVI-XXXII</normalizedToken>
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, XXXIII, and lacks spermathecae. It differs from
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<taxonomicName lsidName="G. champasakensis" pageId="11" pageNumber="12" rank="species" species="champasakensis">G. champasakensis</taxonomicName>
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sp. n. from Mekong River at Ban Khonkhen, Champasak, Laos by the latter having longer wings in XXIII,
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<normalizedToken originalValue="XXIV–½">XXIV-1/2</normalizedToken>
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XXXII, XXXII, XXXIII, longer clitellum in XIX,
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<normalizedToken originalValue="XX–XLIX">XX-XLIX</normalizedToken>
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, L, LI, LII, and spermathecae between 14/15-19/20.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |