559 lines
56 KiB
XML
559 lines
56 KiB
XML
<document id="6CCC640A8255095B5A9B871C590A7A62" ID-DOI="10.13133/2284-4880/765" ID-ISSN="2284-4880" ID-Zenodo-Dep="12762154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721238703073" checkinUser="felipe" docAuthor="Gardini, Giulio" docDate="2022" docId="D16187C89C7F421E531BFEE5885D7DF3" docLanguage="en" docName="FragEntomol.54.1.119-132.pdf" docOrigin="Fragmenta entomologica (Frag. Entomol., Roma) 54 (1)" docSource="http://dx.doi.org/10.13133/2284-4880/765" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Acanthocreagris cyrnea Gardini, 2022, sp. n." docType="treatment" docVersion="2" lastPageNumber="121" masterDocId="2D58FFB09C7E421D502BFFD7895F7C5F" masterDocTitle="Two new species of Acanthocreagris from Corsica and mainland France, and notes on some congeneric species (Pseudoscorpiones: Neobisiidae)" masterLastPageNumber="132" masterPageNumber="119" pageNumber="120" updateTime="1721239365388" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:titleInfo id="322A3483E95CCAD4A2942DBF7E27113E">
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<mods:title id="07037541E653B9AFAD3BA891C532B194">Two new species of Acanthocreagris from Corsica and mainland France, and notes on some congeneric species (Pseudoscorpiones: Neobisiidae)</mods:title>
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<mods:name id="82B956806E6C1BA77B61B52A0BFB804D" type="personal">
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<mods:namePart id="A4B4186D1744F958C42C172CCC1FD607">Gardini, Giulio</mods:namePart>
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<mods:title id="6A07058CED5A18E9DC8E682346D1AA22">Fragmenta entomologica</mods:title>
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</mods:titleInfo>
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<mods:part id="9A46A9EDD068B9435A1BF6E80961ED45">
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<mods:date id="10C80331679E17B59CFAB937CBDC0030">2022</mods:date>
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<mods:detail id="D9DFCB88627AD93FCDA9C86792FDC130" type="series">
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<mods:title id="DE187BC08DBDA15CA9A6E6ED508FAA2A">Frag. Entomol., Roma</mods:title>
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<mods:number id="FF2C25D0FC4D20197E1C49547B1F92EB">2022-05-15</mods:number>
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<mods:number id="34C7253928A5384780A708DE1AE48C09">54</mods:number>
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<mods:start id="E1DD48B5A64DAEE0BBEEFC3E37A9F4F3">119</mods:start>
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<mods:end id="E05551A1C79D5B0DBFA0893EB7C86FA1">132</mods:end>
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<mods:url id="454A794E91F38102A26A1DD6DDCED77C">http://dx.doi.org/10.13133/2284-4880/765</mods:url>
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<mods:classification id="65283FF7B4E05E09E749CC64541D4C18">journal article</mods:classification>
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<mods:identifier id="E1BD0EBA6A6F4EFDC836EC59619151B8" type="DOI">10.13133/2284-4880/765</mods:identifier>
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<mods:identifier id="4045530B407169C9C7BEDC0EE0306EBA" type="ISSN">2284-4880</mods:identifier>
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</mods:mods>
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<treatment id="D16187C89C7F421E531BFEE5885D7DF3" LSID="urn:lsid:plazi:treatment:D16187C89C7F421E531BFEE5885D7DF3" httpUri="http://treatment.plazi.org/id/D16187C89C7F421E531BFEE5885D7DF3" lastPageId="3" lastPageNumber="121" pageId="1" pageNumber="120">
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<subSubSection id="11D265559C7F421C531BFEE58DC47D13" box="[816,1179,306,333]" pageId="1" pageNumber="120" type="nomenclature">
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<paragraph id="597736DE9C7F421C531BFEE58DC47D13" blockId="1.[816,1179,306,333]" box="[816,1179,306,333]" pageId="1" pageNumber="120">
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<heading id="023F81B29C7F421C531BFEE58DC47D13" bold="true" box="[816,1179,306,333]" fontSize="11" level="1" pageId="1" pageNumber="120" reason="1">
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<emphasis id="6BBCEACC9C7F421C531BFEE58DC47D13" bold="true" box="[816,1179,306,333]" pageId="1" pageNumber="120">
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<taxonomicName id="9EC84D5D9C7F421C531BFEE58D0E7D12" authorityName="Gardini" authorityYear="2022" box="[816,1105,306,333]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="120" phylum="Arthropoda" rank="species" species="cyrnea" status="sp. nov.">
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<emphasis id="6BBCEACC9C7F421C531BFEE58D0E7D12" bold="true" box="[816,1105,306,333]" italics="true" pageId="1" pageNumber="120">Acanthocreagris cyrnea</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="708F57B79C7F421C5472FEE58DC47D13" box="[1113,1179,306,332]" pageId="1" pageNumber="120" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="11D265559C7F421C531BFEA38C577E13" pageId="1" pageNumber="120" type="materials_examined">
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<paragraph id="597736DE9C7F421C531BFEA38C577E13" blockId="1.[816,1436,372,588]" pageId="1" pageNumber="120">
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<emphasis id="6BBCEACC9C7F421C531BFEA38A8A7DD3" bold="true" box="[816,981,372,396]" pageId="1" pageNumber="120">Type material.</emphasis>
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<materialsCitation id="E9A03C839C7F421C53F1FEA38A927D93" collectingDate="2019-07-07" country="France" latitude="42.717224" location="Grotta di Punta Vecchiaia" longLatPrecision="19" longitude="9.324445" pageId="1" pageNumber="120" specimenCount="1" specimenCount-female="1" stateProvince="Corse" typeStatus="holotype">
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<emphasis id="6BBCEACC9C7F421C53F1FEA38D167DD3" bold="true" box="[986,1097,372,396]" pageId="1" pageNumber="120">
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<collectingCountry id="21DF764E9C7F421C53F1FEA38D167DD3" box="[986,1097,372,396]" name="France" pageId="1" pageNumber="120">FRANCE</collectingCountry>
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</emphasis>
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:
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<emphasis id="6BBCEACC9C7F421C547EFEA28DF67DD3" box="[1109,1193,373,396]" italics="true" pageId="1" pageNumber="120">
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<collectingRegion id="9B0CF83C9C7F421C547EFEA28DF67DD3" box="[1109,1193,373,396]" country="France" name="Corse" pageId="1" pageNumber="120">Corsica</collectingRegion>
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</emphasis>
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:
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<typeStatus id="8673887C9C7F421C549DFEA38C467DD3" box="[1206,1305,372,396]" pageId="1" pageNumber="120" type="holotype">Holotype</typeStatus>
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<specimenCount id="4FCEFD579C7F421C5534FEA28C6C7DD3" box="[1311,1331,373,396]" count="1" pageId="1" pageNumber="120" type="female">♀</specimenCount>
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, Patrimo- nio,
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<location id="5C1760059C7F421C5374FE438D307DF3" LSID="urn:lsid:plazi:treatment:D16187C89C7F421E531BFEE5885D7DF3:5C1760059C7F421C5374FE438D307DF3" box="[863,1135,404,428]" country="France" latitude="42.717224" longLatPrecision="19" longitude="9.324445" name="Grotta di Punta Vecchiaia" pageId="1" pageNumber="120" stateProvince="Corse">Grotta di Punta Vecchiaia</location>
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(
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<geoCoordinate id="3CFC50199C7F421C5456FE438C5F7DF3" box="[1149,1280,404,428]" degrees="42" direction="north" minutes="43" orientation="latitude" pageId="1" pageNumber="120" precision="15" seconds="02" value="42.717224">42°43’02”N</geoCoordinate>
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<geoCoordinate id="3CFC50199C7F421C552CFE438C247DF3" box="[1287,1403,404,428]" degrees="9" direction="east" minutes="19" orientation="longitude" pageId="1" pageNumber="120" precision="15" seconds="28" value="9.324445">9°19’28”E</geoCoordinate>
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), 0 m,
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<date id="2D76101E9C7F421C537EFE628A927D93" box="[853,973,436,460]" pageId="1" pageNumber="120" value="2019-07-07">
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<collectingDate id="3D32E9F69C7F421C537EFE628A927D93" box="[853,973,436,460]" pageId="1" pageNumber="120" value="2019-07-07">7 Jul 2019</collectingDate>
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</date>
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||
</materialsCitation>
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,
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<materialsCitation id="E9A03C839C7F421C53F0FE638AC87DB3" collectionCode="MNHN" country="France" location="Ligue Insulaire Speleologique" pageId="1" pageNumber="120" specimenCount="1" stateProvince="Corse" typeStatus="holotype">
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<location id="5C1760059C7F421C53F0FE638C7C7D93" LSID="urn:lsid:plazi:treatment:D16187C89C7F421E531BFEE5885D7DF3:5C1760059C7F421C53F0FE638C7C7D93" box="[987,1315,436,460]" country="France" name="Ligue Insulaire Speleologique" pageId="1" pageNumber="120" stateProvince="Corse">Ligue Insulaire Spéléologique</location>
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<collectingRegion id="9B0CF83C9C7F421C5506FE638C347D93" box="[1325,1387,436,460]" country="France" name="Corse" pageId="1" pageNumber="120">Corse</collectingRegion>
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leg. (
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<collectionCode id="3FD9AE1B9C7F421C5313FE038ACF7DB3" box="[824,912,468,492]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="120">MNHN</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="E9A03C839C7F421C5389FE028C5A7E13" collectingDate="2019-04-27" collectorName="J. Lips & Coll. G. Gardini & Genoa" country="France" county="Oletta" elevation="120" latitude="42.658432" location="Grotte Cast" longLatPrecision="1" longitude="9.298281" municipality="Massif de Castiglione" pageId="1" pageNumber="120" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
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<typeStatus id="8673887C9C7F421C5389FE028D517DB2" box="[930,1038,469,493]" pageId="1" pageNumber="120" type="paratype">Paratypes</typeStatus>
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:
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||
<specimenCount id="4FCEFD579C7F421C5432FE038D1D7DB3" box="[1049,1090,468,492]" count="1" pageId="1" pageNumber="120" type="male">1 ♂</specimenCount>
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(with both galeae and left pedipalp broken)
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||
<specimenCount id="4FCEFD579C7F421C53E1FE238AB47E53" box="[970,1003,500,524]" count="1" pageId="1" pageNumber="120" type="female">1♀</specimenCount>
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,
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<collectingCounty id="B0164E529C7F421C53D5FE238D1D7E53" box="[1022,1090,500,524]" pageId="1" pageNumber="120">Oletta</collectingCounty>
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,
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||
<collectingMunicipality id="B913ACA49C7F421C5479FE238C177E53" box="[1106,1352,500,524]" pageId="1" pageNumber="120">Massif de Castiglione</collectingMunicipality>
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||
,
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||
<location id="5C1760059C7F421C5573FE238A007E73" LSID="urn:lsid:plazi:treatment:D16187C89C7F421E531BFEE5885D7DF3:5C1760059C7F421C5573FE238A007E73" country="France" county="Oletta" latitude="42.658432" longLatPrecision="1" longitude="9.298281" municipality="Massif de Castiglione" name="Grotte Cast" pageId="1" pageNumber="120">Grotte Cast</location>
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||
1 (
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||
<geoCoordinate id="3CFC50199C7F421C53A5FDC38D6C7E73" box="[910,1075,532,556]" degrees="42" direction="north" minutes="39" orientation="latitude" pageId="1" pageNumber="120" precision="1" seconds="30.35" value="42.658432">42°39’30.35”N</geoCoordinate>
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<geoCoordinate id="3CFC50199C7F421C546BFDC38D887E73" box="[1088,1239,532,556]" degrees="9" direction="east" minutes="17" orientation="longitude" pageId="1" pageNumber="120" precision="1" seconds="53.81" value="9.298281">9°17’53.81”E</geoCoordinate>
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),
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||
<quantity id="9E309B3B9C7F421C54DBFDC38C637E73" box="[1264,1340,532,556]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="120" unit="m" value="120.0">
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<elevation id="D2E5D1ED9C7F421C54DBFDC38C637E73" box="[1264,1340,532,556]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="120" unit="m" value="120.0">120 m</elevation>
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</quantity>
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,
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||
<date id="2D76101E9C7F421C5566FDC38A377E13" pageId="1" pageNumber="120" value="2019-04-27">
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||
<collectingDate id="3D32E9F69C7F421C5566FDC38A377E13" pageId="1" pageNumber="120" value="2019-04-27">27 Apr 2019</collectingDate>
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||
</date>
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||
,
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||
<collectorName id="F43D53089C7F421C5358FDE28AE57E13" box="[883,954,564,588]" pageId="1" pageNumber="120">J. Lips</collectorName>
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||
leg. (
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<collectorName id="F43D53089C7F421C53DCFDE38DF87E13" box="[1015,1191,564,588]" pageId="1" pageNumber="120">Coll. G. Gardini</collectorName>
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,
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<collectorName id="F43D53089C7F421C5499FDE38DA37E13" box="[1202,1276,564,588]" pageId="1" pageNumber="120">Genoa</collectorName>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="11D265559C7F421F531BFDA38BCD7ED3" lastPageId="2" lastPageNumber="121" pageId="1" pageNumber="120" type="diagnosis">
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||
<paragraph id="597736DE9C7F421F531BFDA38BCD7ED3" blockId="1.[816,1437,628,717]" lastBlockId="2.[151,772,180,652]" lastPageId="2" lastPageNumber="121" pageId="1" pageNumber="120">
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||
<emphasis id="6BBCEACC9C7F421C531BFDA38AC17ED3" bold="true" box="[816,926,628,652]" pageId="1" pageNumber="120">Diagnosis</emphasis>
|
||
(
|
||
<specimenCount id="4FCEFD579C7F421C539BFDA38A987ED3" box="[944,967,628,652]" count="1" pageId="1" pageNumber="120" type="male">♂</specimenCount>
|
||
<specimenCount id="4FCEFD579C7F421C53ECFDA38A887ED3" box="[967,983,628,652]" count="1" pageId="1" pageNumber="120" type="female">♀</specimenCount>
|
||
).
|
||
<materialsCitation id="E9A03C839C7F421F53C4FDA38BD07ED3" country="France" lastPageId="2" lastPageNumber="121" location="French" municipality="Italian" pageId="1" pageNumber="120" specimenCount="27" specimenCount-female="14" specimenCount-male="13">
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||
A subterranean troglomorphic
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||
<taxonomicName id="9EC84D5D9C7F421C5517FDA38AD77EF3" authorityName="Mahnert" authorityYear="1974" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="120" phylum="Arthropoda" rank="genus">
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||
<emphasis id="6BBCEACC9C7F421C5517FDA38AD77EF3" italics="true" pageId="1" pageNumber="120">Acanthocreagris</emphasis>
|
||
</taxonomicName>
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||
from
|
||
<collectingCountry id="21DF764E9C7F421C53E5FD438D407EF3" box="[974,1055,660,684]" name="France" pageId="1" pageNumber="120">Corsica</collectingCountry>
|
||
that differs from other
|
||
<location id="5C1760059C7F421C5508FD438C327EF3" LSID="urn:lsid:plazi:treatment:D16187C89C7F421E531BFEE5885D7DF3:5C1760059C7F421C5508FD438C327EF3" box="[1315,1389,660,684]" country="France" municipality="Italian" name="French" pageId="1" pageNumber="120">French</location>
|
||
and
|
||
<collectingMunicipality id="B913ACA49C7F421C5318FD638A287E93" box="[819,887,692,716]" pageId="1" pageNumber="120">Italian</collectingMunicipality>
|
||
species of the genus in the following combination of characters: no eyes or eye-spots; galea (
|
||
<specimenCount id="4FCEFD579C7C421F525FFF638BDA7C93" box="[628,645,180,204]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
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||
) long, apically with three branches, reaching the apex of the cheliceral movable finger; trochanter with 4 spiniform setae with simple and acuminate tips; pedipalpal femur gradually clavate, length 1.20 (
|
||
<specimenCount id="4FCEFD579C7C421F5181FEE388E07D13" box="[426,447,308,332]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 1.23 (
|
||
<specimenCount id="4FCEFD579C7C421F5205FEE38B607D13" box="[558,575,308,332]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) mm, 5.33 (
|
||
<specimenCount id="4FCEFD579C7C421F52E3FEE38B827D13" box="[712,733,308,332]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 5.12 (
|
||
<specimenCount id="4FCEFD579C7C421F50F2FE8389B57D33" box="[217,234,340,364]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) times as long as broad; patella 1.02 (
|
||
<specimenCount id="4FCEFD579C7C421F52BBFE838BFA7D33" box="[656,677,340,364]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 1.09 (
|
||
<specimenCount id="4FCEFD579C7C421F508AFEA389ED7DD3" box="[161,178,372,396]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) mm, 3.84 (
|
||
<specimenCount id="4FCEFD579C7C421F5114FEA3880B7DD3" box="[319,340,372,396]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 3.89 (
|
||
<specimenCount id="4FCEFD579C7C421F51ECFEA388877DD3" box="[455,472,372,396]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) ×, ratio between club and pedicel 1.73 (
|
||
<specimenCount id="4FCEFD579C7C421F5100FE43881F7DF3" box="[299,320,404,428]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 1.66 (
|
||
<specimenCount id="4FCEFD579C7C421F519BFE43889D7DF3" box="[432,450,404,428]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
); chela with pedicel 1.95 (
|
||
<specimenCount id="4FCEFD579C7C421F52CDFE438BA47DF3" box="[742,763,404,428]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 2.0 (
|
||
<specimenCount id="4FCEFD579C7C421F50C6FE6389A17D93" box="[237,254,436,460]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) mm, 4.81 (
|
||
<specimenCount id="4FCEFD579C7C421F51A5FE6388FC7D93" box="[398,419,436,460]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 4.44 (
|
||
<specimenCount id="4FCEFD579C7C421F5233FE638B767D93" box="[536,553,436,460]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) ×; movable chelal finger 1.18 (
|
||
<specimenCount id="4FCEFD579C7C421F510BFE03886A7DB3" box="[288,309,468,492]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 1.20 (
|
||
<specimenCount id="4FCEFD579C7C421F5181FE0388E47DB3" box="[426,443,468,492]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) mm; ratio between movable finger and hand with pedicel 1.34 (
|
||
<specimenCount id="4FCEFD579C7C421F5236FE238B6D7E53" box="[541,562,500,524]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 1.25 (
|
||
<specimenCount id="4FCEFD579C7C421F528CFE238BE67E53" box="[679,697,500,524]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
), ratio between femur and movable finger 1.01 (
|
||
<specimenCount id="4FCEFD579C7C421F5270FDC38B2F7E73" box="[603,624,532,556]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 1.02 (
|
||
<specimenCount id="4FCEFD579C7C421F52CAFDC38BAC7E73" box="[737,755,532,556]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
), ratio between pedipalpal femur and carapace 1.36 (
|
||
<specimenCount id="4FCEFD579C7C421F52ECFDE38B837E13" box="[711,732,564,588]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 1.38 (
|
||
<specimenCount id="4FCEFD579C7C421F50F3FD8389B57E33" box="[216,234,596,620]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
); fixed and movable chelal fingers with 99 (
|
||
<specimenCount id="4FCEFD579C7C421F52E3FD838B827E33" box="[712,733,596,620]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 96 (
|
||
<specimenCount id="4FCEFD579C7C421F50E8FDA3898B7ED3" box="[195,212,628,652]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) and 90 (
|
||
<specimenCount id="4FCEFD579C7C421F5117FDA3880E7ED3" box="[316,337,628,652]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 87 (
|
||
<specimenCount id="4FCEFD579C7C421F5183FDA388E67ED3" box="[424,441,628,652]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) teeth, respectively
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="0DB766569C7F421C50A3F8958D8A7BD4" ID-DOI="http://doi.org/10.5281/zenodo.12762156" ID-Zenodo-Dep="12762156" httpUri="https://zenodo.org/record/12762156/files/figure.png" pageId="1" pageNumber="120" startId="1.[136,174,1858,1877]" targetBox="[138,1427,746,1846]" targetPageId="1" targetType="figure">
|
||
<paragraph id="597736DE9C7F421C50A3F8958D8A7BD4" blockId="1.[136,1437,1858,1931]" pageId="1" pageNumber="120">
|
||
<emphasis id="6BBCEACC9C7F421C50A3F89589BB7B09" bold="true" box="[136,228,1858,1878]" pageId="1" pageNumber="120">Figs 1–8 –</emphasis>
|
||
<taxonomicName id="9EC84D5D9C7F421C50C2F89588E97B0A" authorityName="Gardini" authorityYear="2022" box="[233,438,1858,1877]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="1" pageNumber="120" phylum="Arthropoda" rank="species" species="cyrnea" status="sp. nov.">
|
||
<emphasis id="6BBCEACC9C7F421C50C2F89588E97B0A" box="[233,438,1858,1877]" italics="true" pageId="1" pageNumber="120">Acanthocreagris cyrnea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="708F57B79C7F421C5190F89488B47B09" box="[443,491,1859,1878]" pageId="1" pageNumber="120" rank="species">sp. n.</taxonomicNameLabel>
|
||
, ♀ holotype:
|
||
<emphasis id="6BBCEACC9C7F421C5274F8958B357B0A" bold="true" box="[607,618,1858,1877]" pageId="1" pageNumber="120">1</emphasis>
|
||
, epistome of carapace;
|
||
<emphasis id="6BBCEACC9C7F421C531FF8958A607B0A" bold="true" box="[820,831,1858,1877]" pageId="1" pageNumber="120">2</emphasis>
|
||
, right chelicera, dorsal view;
|
||
<emphasis id="6BBCEACC9C7F421C5416F8958D177B0A" bold="true" box="[1085,1096,1858,1877]" pageId="1" pageNumber="120">3</emphasis>
|
||
, galea of rigt movable cheliceral finger, dorsal (left) and antiaxial (right) view;
|
||
<emphasis id="6BBCEACC9C7F421C51F4F88A88B77B2F" bold="true" box="[479,488,1885,1904]" pageId="1" pageNumber="120">4</emphasis>
|
||
, anterolateral process of right coxa I;
|
||
<emphasis id="6BBCEACC9C7F421C5312F88A8A1D7B2F" bold="true" box="[825,834,1885,1904]" pageId="1" pageNumber="120">5</emphasis>
|
||
, trochanter, femur and patella of right pedipalp, dorsal view;
|
||
<emphasis id="6BBCEACC9C7F421C5574F88A8C377B2F" bold="true" box="[1375,1384,1885,1904]" pageId="1" pageNumber="120">6</emphasis>
|
||
, right pedipalpal chela, dorsal view;
|
||
<emphasis id="6BBCEACC9C7F421C51A7F8AF88C87BD4" bold="true" box="[396,407,1912,1931]" pageId="1" pageNumber="120">7</emphasis>
|
||
, same, antiaxial view;
|
||
<emphasis id="6BBCEACC9C7F421C5270F8AF8B397BD4" bold="true" box="[603,614,1912,1931]" pageId="1" pageNumber="120">8</emphasis>
|
||
, right leg IV, antiaxial view. Scale lines: 0.2 mm (2, 4–8); 0.1 mm (1, 3).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="11D265559C7C421F50BCFD638A5A7F73" pageId="2" pageNumber="121" type="etymology">
|
||
<paragraph id="597736DE9C7C421F50BCFD638A5A7F73" blockId="2.[151,773,692,812]" pageId="2" pageNumber="121">
|
||
<emphasis id="6BBCEACC9C7C421F50BCFD63884C7E93" bold="true" box="[151,275,692,716]" pageId="2" pageNumber="121">Etymology.</emphasis>
|
||
It takes its name from Cyrnus or Cyrnos (adj.: Cyrneus, -a, -um), one of the sons of Heracles, tutelary deity of
|
||
<collectingCountry id="21DF764E9C7C421F509FFD2388597F53" box="[180,262,756,780]" name="France" pageId="2" pageNumber="121">Corsica</collectingCountry>
|
||
, who chose the island as his residence and from whom the island took its name (Herodotus, Historiae 1.167).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="11D265559C7C421F50BCFC838C7B7993" pageId="2" pageNumber="121" type="description">
|
||
<paragraph id="597736DE9C7C421F50BCFC838D567873" blockId="2.[151,772,852,1932]" lastBlockId="2.[831,1452,180,1484]" pageId="2" pageNumber="121">
|
||
<emphasis id="6BBCEACC9C7C421F50BCFC8388D17F33" bold="true" box="[151,398,852,876]" pageId="2" pageNumber="121">Description of adults</emphasis>
|
||
(
|
||
<specimenCount id="4FCEFD579C7C421F5189FC8388E67F33" box="[418,441,852,876]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
<specimenCount id="4FCEFD579C7C421F5192FC8388967F33" box="[441,457,852,876]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
). Carapace, chelicera, pedipalps and palpal coxae red-brown. Carapace 1.62 (
|
||
<specimenCount id="4FCEFD579C7C421F52CDFCA38BA47FD3" box="[742,763,884,908]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 1.56 (
|
||
<specimenCount id="4FCEFD579C7C421F5128FC43884B7FF3" box="[259,276,916,940]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) times as long as broad, without eyes or eye-spots, anterior margin with slightly prominent and broadly rounded epistome (Fig. 1); 22 macrosetae, anterior and posterior rows with 4 and 6 macrosetae, respectively. Chaetotaxy of tergites I–X 6–7:7–8:7:8–9:8– 9:9:9:10–12:10–12:9. Chaetotaxy of sternites II–XI (
|
||
<specimenCount id="4FCEFD579C7C421F52CDFBE38BA47813" box="[742,763,1076,1100]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) 16:13:9:12:13:13:14:12:10:9, genital atrium with 3+3 setae; median genital sac reaching sternite V; chaetotaxy of sternites II–XI (
|
||
<specimenCount id="4FCEFD579C7C421F516EFB43880978F3" box="[325,342,1172,1196]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) 11:12:6:14:15:17:16:15:11:8; sternites III and IV (
|
||
<specimenCount id="4FCEFD579C7C421F5132FB63886F7893" box="[281,304,1204,1228]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
<specimenCount id="4FCEFD579C7C421F511BFB63881F7893" box="[304,320,1204,1228]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) each with 3 or 4 microsetae in front on each stigma, sternites VI–VIII with 2 discal setae; anal cone with 2+2 setae. Chelicera (Fig. 2) 1.95 (
|
||
<specimenCount id="4FCEFD579C7C421F52B1FB238BF07953" box="[666,687,1268,1292]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 2.0 (
|
||
<specimenCount id="4FCEFD579C7C421F5089FAC389EC7973" box="[162,179,1300,1324]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) times as long as broad, palm with 6 setae; fixed finger with worn out teeth (
|
||
<specimenCount id="4FCEFD579C7C421F51A8FAE388C77913" box="[387,408,1332,1356]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 13 subequal teeth (
|
||
<specimenCount id="4FCEFD579C7C421F52B4FAE38BEE7913" box="[671,689,1332,1356]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
), movable finger with worn out teeth (
|
||
<specimenCount id="4FCEFD579C7C421F5223FA838B427933" box="[520,541,1364,1388]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 10 (
|
||
<specimenCount id="4FCEFD579C7C421F52AAFA838BCD7933" box="[641,658,1364,1388]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) teeth, of which the 3-4 distal larger;
|
||
<emphasis id="6BBCEACC9C7C421F51EEFAA2888279D3" box="[453,477,1397,1420]" italics="true" pageId="2" pageNumber="121">gs</emphasis>
|
||
ratio 0.74 (
|
||
<specimenCount id="4FCEFD579C7C421F5277FAA38B2E79D3" box="[604,625,1396,1420]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 0.70 (
|
||
<specimenCount id="4FCEFD579C7C421F52CAFAA38BAC79D3" box="[737,755,1396,1420]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
), galea long, apically with three branches (
|
||
<specimenCount id="4FCEFD579C7C421F5248FA438B2979F3" box="[611,630,1428,1452]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
; both galeae broken in
|
||
<specimenCount id="4FCEFD579C7C421F512FFA6388467993" box="[260,281,1460,1484]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) (Fig. 3); rallum with 8 blades, the four distal ones laterally pinnate, the four proximal ones (apparently) smooth, of these the first shorter; serrulae interior and exterior with about 20 and 25 blades, respectively. Manducatory process with 3 setae. Coxal setae: pedipalp 7–8, I 5–6, II 6, III 5–6, IV 7; anterolateral process of coxa I sharp, apically pointed (Fig. 4). Pedipalp (Figs 5–7): trochanter with granular surface, 3.33 (
|
||
<specimenCount id="4FCEFD579C7C421F5209F9438B687AF3" box="[546,567,1684,1708]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 3.40 (
|
||
<specimenCount id="4FCEFD579C7C421F528DF9438BE87AF3" box="[678,695,1684,1708]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) times as long as broad, without tubercles on paraxial face, with 4 spiniform setae with simple and acuminate tips; femur 5.33 (
|
||
<specimenCount id="4FCEFD579C7C421F50FCF92389B37B53" box="[215,236,1780,1804]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 5.12 (
|
||
<specimenCount id="4FCEFD579C7C421F517DF92388387B53" box="[342,359,1780,1804]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) times as long as broad, gradually clavate, with granular surface as in fig. 5; patella 3.84 (
|
||
<specimenCount id="4FCEFD579C7C421F52E2F8C38B817B73" box="[713,734,1812,1836]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 3.89 (
|
||
<specimenCount id="4FCEFD579C7C421F50FCF8E389B77B13" box="[215,232,1844,1868]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) times as long as broad, club elongate and slightly granular on paraxial face, pedicel granular, ratio between club and pedicel 1.73 (
|
||
<specimenCount id="4FCEFD579C7C421F51BCF8A388F37BD3" box="[407,428,1908,1932]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 1.66 (
|
||
<specimenCount id="4FCEFD579C7C421F520BF8A38B6D7BD3" box="[544,562,1908,1932]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
), ratio X/Y = 0.22 (
|
||
<specimenCount id="4FCEFD579C7C421F5363FF638A027C93" box="[840,861,180,204]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 0.20 (
|
||
<specimenCount id="4FCEFD579C7C421F53E4FF638ABE7C93" box="[975,993,180,204]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
); chela with pedicel 4.81 (
|
||
<specimenCount id="4FCEFD579C7C421F5522FF638C417C93" box="[1289,1310,180,204]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 4.44 (
|
||
<specimenCount id="4FCEFD579C7C421F55BBFF638CFE7C93" box="[1424,1441,180,204]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) times as long as broad; hand of chela with pedicel 2.17 (
|
||
<specimenCount id="4FCEFD579C7C421F5363FF238A027D53" box="[840,861,244,268]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 2.13 (
|
||
<specimenCount id="4FCEFD579C7C421F53E7FF238A827D53" box="[972,989,244,268]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) times as long as broad, elongated oval in dorsal view with a slightly convex paraxial profile, with granular surface as in figs 6–7; fixed chelal finger with 99 (
|
||
<specimenCount id="4FCEFD579C7C421F5363FE838A027D33" box="[840,861,340,364]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 96 (
|
||
<specimenCount id="4FCEFD579C7C421F539DFE838A987D33" box="[950,967,340,364]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) contiguous teeth with dental canals, venom duct short, nodus ramosus subterminal, subterminal sensillum as in fig. 7; movable chelal finger with 90 (
|
||
<specimenCount id="4FCEFD579C7C421F557AFE438C397DF3" box="[1361,1382,404,428]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 87 (
|
||
<specimenCount id="4FCEFD579C7C421F5361FE638A047D93" box="[842,859,436,460]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) contiguous teeth with dental canals (Fig. 7) reaching back near
|
||
<emphasis id="6BBCEACC9C7C421F539FFE038A9E7DB3" box="[948,961,468,492]" italics="true" pageId="2" pageNumber="121">b</emphasis>
|
||
, the distal 14 teeth pointed; movable chelal finger with four sensilla, two of which subterminal (Fig. 7), the others at the level of the tenth (
|
||
<specimenCount id="4FCEFD579C7C421F54FFFDC38DB67E73" box="[1236,1257,532,556]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or the eighth and eleventh tooth (
|
||
<specimenCount id="4FCEFD579C7C421F53CCFDE38AA77E13" box="[999,1016,564,588]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) from the trichobothrium
|
||
<emphasis id="6BBCEACC9C7C421F5524FDE38C787E13" box="[1295,1319,564,588]" italics="true" pageId="2" pageNumber="121">sb</emphasis>
|
||
; trichobothria as in figs 6–7, relative position of trichobothria along chelal axis (
|
||
<specimenCount id="4FCEFD579C7C421F53E8FDA38A897ED3" box="[963,982,628,652]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
):
|
||
<emphasis id="6BBCEACC9C7C421F53C7FDA28AA07ED3" box="[1004,1023,629,652]" italics="true" pageId="2" pageNumber="121">et</emphasis>
|
||
0.11/
|
||
<emphasis id="6BBCEACC9C7C421F5417FDA28D147ED3" box="[1084,1099,629,652]" italics="true" pageId="2" pageNumber="121">it</emphasis>
|
||
0.20/
|
||
<emphasis id="6BBCEACC9C7C421F54A3FDA28DF97ED3" box="[1160,1190,629,652]" italics="true" pageId="2" pageNumber="121">est</emphasis>
|
||
0.25/
|
||
<emphasis id="6BBCEACC9C7C421F54C8FDA28DA37ED3" box="[1251,1276,629,652]" italics="true" pageId="2" pageNumber="121">ist</emphasis>
|
||
0.34/
|
||
<emphasis id="6BBCEACC9C7C421F5511FDA38C067ED3" box="[1338,1369,628,652]" italics="true" pageId="2" pageNumber="121">isb</emphasis>
|
||
0.63/
|
||
<emphasis id="6BBCEACC9C7C421F55BDFDA38CF47ED3" box="[1430,1451,628,652]" italics="true" pageId="2" pageNumber="121">ib</emphasis>
|
||
0.80/
|
||
<emphasis id="6BBCEACC9C7C421F535DFD438AC57EF3" box="[886,922,660,684]" italics="true" pageId="2" pageNumber="121">esb</emphasis>
|
||
0.86/
|
||
<emphasis id="6BBCEACC9C7C421F53F1FD438AAC7EF3" box="[986,1011,660,684]" italics="true" pageId="2" pageNumber="121">eb</emphasis>
|
||
0.93/
|
||
<emphasis id="6BBCEACC9C7C421F5418FD428D657EF3" box="[1075,1082,661,684]" italics="true" pageId="2" pageNumber="121">t</emphasis>
|
||
0.23/
|
||
<emphasis id="6BBCEACC9C7C421F5451FD428DD37EF3" box="[1146,1164,661,684]" italics="true" pageId="2" pageNumber="121">st</emphasis>
|
||
0.36/
|
||
<emphasis id="6BBCEACC9C7C421F54E7FD438DBB7EF3" box="[1228,1252,660,684]" italics="true" pageId="2" pageNumber="121">sb</emphasis>
|
||
0.65/
|
||
<emphasis id="6BBCEACC9C7C421F550FFD438C6E7EF3" box="[1316,1329,660,684]" italics="true" pageId="2" pageNumber="121">b</emphasis>
|
||
0.86; ratio between movable finger and hand of chela with pedicel 1.34 (
|
||
<specimenCount id="4FCEFD579C7C421F5354FD038ACB7EB3" box="[895,916,724,748]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 1.25 (
|
||
<specimenCount id="4FCEFD579C7C421F5429FD038D4B7EB3" box="[1026,1044,724,748]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
); ratio between pedipalpal femur and movable finger 1.01 (
|
||
<specimenCount id="4FCEFD579C7C421F5402FD238D617F53" box="[1065,1086,756,780]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) 1.02 (
|
||
<specimenCount id="4FCEFD579C7C421F54A4FD238DFE7F53" box="[1167,1185,756,780]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
); ratio between pedipalpal femur and carapace 1.36 (
|
||
<specimenCount id="4FCEFD579C7C421F54A0FCC38DFF7F73" box="[1163,1184,788,812]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) 1.38 (
|
||
<specimenCount id="4FCEFD579C7C421F54DEFCC38C587F73" box="[1269,1287,788,812]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
). Leg IV (Fig. 8): trochanter 2.25 (
|
||
<specimenCount id="4FCEFD579C7C421F5435FCE38D6A7F13" box="[1054,1077,820,844]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
<specimenCount id="4FCEFD579C7C421F541EFCE38D1A7F13" box="[1077,1093,820,844]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) times as long as deep, femur + patella 3.86 (
|
||
<specimenCount id="4FCEFD579C7C421F53FAFC838AB97F33" box="[977,998,852,876]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 4.34 (
|
||
<specimenCount id="4FCEFD579C7C421F5472FC838D357F33" box="[1113,1130,852,876]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) times as long as deep, tibia 6.30 (
|
||
<specimenCount id="4FCEFD579C7C421F53AAFCA38AC97FD3" box="[897,918,884,908]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 7.30 (
|
||
<specimenCount id="4FCEFD579C7C421F5421FCA38D447FD3" box="[1034,1051,884,908]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) times, basitarsus 3.38 (
|
||
<specimenCount id="4FCEFD579C7C421F551BFCA38C187FD3" box="[1328,1351,884,908]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
<specimenCount id="4FCEFD579C7C421F556CFCA38C087FD3" box="[1351,1367,884,908]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) times, telotarsus 7.84 (
|
||
<specimenCount id="4FCEFD579C7C421F53DDFC438D547FF3" box="[1014,1035,916,940]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 6.71 (
|
||
<specimenCount id="4FCEFD579C7C421F54ADFC438DC87FF3" box="[1158,1175,916,940]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) times as long as deep, tactile seta in the proximal half (TS = 0.23–0.29), ratio between basitarsus and telotarsus 0.60–0.65, subterminal seta furcate or trifid, slightly dentate, claws with a very small dorsal tooth.
|
||
</paragraph>
|
||
<paragraph id="597736DE9C7C421F534EFBE28C7B7993" blockId="2.[831,1452,180,1484]" pageId="2" pageNumber="121">
|
||
<emphasis id="6BBCEACC9C7C421F534EFBE28AA27813" box="[869,1021,1077,1100]" italics="true" pageId="2" pageNumber="121">Measurements</emphasis>
|
||
(mm). Body length 2.6 (
|
||
<specimenCount id="4FCEFD579C7C421F553BFBE38C7B7813" box="[1296,1316,1076,1100]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 3.0 (
|
||
<specimenCount id="4FCEFD579C7C421F55A1FBE38CC37813" box="[1418,1436,1076,1100]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
). Carapace 0.88 × 0.54 anteriorly (
|
||
<specimenCount id="4FCEFD579C7C421F54BCFB838DF47833" box="[1175,1195,1108,1132]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 0.89 × 0.57 anteriorly (
|
||
<specimenCount id="4FCEFD579C7C421F534FFBA38A2978D3" box="[868,886,1140,1164]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
). Chelicera 0.48 × 0.245 (
|
||
<specimenCount id="4FCEFD579C7C421F54ACFBA38DC478D3" box="[1159,1179,1140,1164]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 0.52 × 0.26 (
|
||
<specimenCount id="4FCEFD579C7C421F5566FBA38C0178D3" box="[1357,1374,1140,1164]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
), movable finger length 0.32 (
|
||
<specimenCount id="4FCEFD579C7C421F541CFB438D1478F3" box="[1079,1099,1172,1196]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 0.35 (
|
||
<specimenCount id="4FCEFD579C7C421F5498FB438D9A78F3" box="[1203,1221,1172,1196]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
). Pedipalp: trochanter 0.65 × 0.195 (
|
||
<specimenCount id="4FCEFD579C7C421F53FBFB638ABB7893" box="[976,996,1204,1228]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 0.68 × 0.20 (
|
||
<specimenCount id="4FCEFD579C7C421F54B8FB638DFA7893" box="[1171,1189,1204,1228]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
); femur 1.20 × 0.225 (
|
||
<specimenCount id="4FCEFD579C7C421F55A5FB638CFC7893" box="[1422,1443,1204,1228]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 1.23 × 0.24 (
|
||
<specimenCount id="4FCEFD579C7C421F53CCFB038AA678B3" box="[999,1017,1236,1260]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
); patella 1.02 × 0.265 (
|
||
<specimenCount id="4FCEFD579C7C421F54DAFB038C5A78B3" box="[1265,1285,1236,1260]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 1.09 × 0.28 (
|
||
<specimenCount id="4FCEFD579C7C421F5361FB238A037953" box="[842,860,1268,1292]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
); chela with pedicel 1.95 × 0.405 (
|
||
<specimenCount id="4FCEFD579C7C421F54FAFB238DBA7953" box="[1233,1253,1268,1292]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 2.0 × 0.45 (
|
||
<specimenCount id="4FCEFD579C7C421F55BBFB238CFE7953" box="[1424,1441,1268,1292]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
) (depth 0.385
|
||
<specimenCount id="4FCEFD579C7C421F53ECFAC28A827973" box="[967,989,1301,1324]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
, 0.415
|
||
<specimenCount id="4FCEFD579C7C421F540CFAC38D657973" box="[1063,1082,1300,1324]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
); hand with pedicel length 0.88 (
|
||
<specimenCount id="4FCEFD579C7C421F55A5FAC38CFC7973" box="[1422,1443,1300,1324]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 0.96 (
|
||
<specimenCount id="4FCEFD579C7C421F53B3FAE38AF57913" box="[920,938,1332,1356]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
); movable finger length 1.18 (
|
||
<specimenCount id="4FCEFD579C7C421F54C9FAE38DA97913" box="[1250,1270,1332,1356]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 1.20 (
|
||
<specimenCount id="4FCEFD579C7C421F5577FAE38C317913" box="[1372,1390,1332,1356]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
). Leg IV: trochanter 0.36 × 0.16 (
|
||
<specimenCount id="4FCEFD579C7C421F544EFA838D267933" box="[1125,1145,1364,1388]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 0.35 × 0.155 (
|
||
<specimenCount id="4FCEFD579C7C421F5568FA838C0A7933" box="[1347,1365,1364,1388]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
); femur + patella 0.87 × 0.225 (
|
||
<specimenCount id="4FCEFD579C7C421F541EFAA38D1679D3" box="[1077,1097,1396,1420]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 0.89 × 0.205 (
|
||
<specimenCount id="4FCEFD579C7C421F5521FAA38C4379D3" box="[1290,1308,1396,1420]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
); tibia 0.82 × 0.13 (
|
||
<specimenCount id="4FCEFD579C7C421F5350FA438AD079F3" box="[891,911,1428,1452]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 0.84 × 0.115 (
|
||
<specimenCount id="4FCEFD579C7C421F5466FA438D0079F3" box="[1101,1119,1428,1452]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
); basitarsus 0.305 × 0.09 (
|
||
<specimenCount id="4FCEFD579C7C421F555EFA438CD279F3" box="[1397,1421,1428,1452]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
<specimenCount id="4FCEFD579C7C421F55A7FA438CC379F3" box="[1420,1436,1428,1452]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
); telotarsus 0.51 × 0.065 (
|
||
<specimenCount id="4FCEFD579C7C421F5417FA638D0F7993" box="[1084,1104,1460,1484]" count="1" pageId="2" pageNumber="121" type="male">♂</specimenCount>
|
||
) or 0.47 × 0.07 (
|
||
<specimenCount id="4FCEFD579C7C421F5528FA638C4A7993" box="[1283,1301,1460,1484]" count="1" pageId="2" pageNumber="121" type="female">♀</specimenCount>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="11D265559C7C421E5314FA23885D7DF3" lastPageId="3" lastPageNumber="122" pageId="2" pageNumber="121" type="discussion">
|
||
<paragraph id="597736DE9C7C421E5314FA23885D7DF3" blockId="2.[831,1453,1524,1932]" lastBlockId="3.[136,757,180,428]" lastPageId="3" lastPageNumber="122" pageId="2" pageNumber="121">
|
||
<emphasis id="6BBCEACC9C7C421F5314FA238AF27A53" bold="true" box="[831,941,1524,1548]" pageId="2" pageNumber="121">Remarks.</emphasis>
|
||
Two species of the genus
|
||
<taxonomicName id="9EC84D5D9C7C421F54F8FA238CDD7A53" authorityName="Mahnert" authorityYear="1974" box="[1235,1410,1524,1548]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="121" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6BBCEACC9C7C421F54F8FA238CDD7A53" box="[1235,1410,1524,1548]" italics="true" pageId="2" pageNumber="121">Acanthocreagris</emphasis>
|
||
</taxonomicName>
|
||
are currently known from
|
||
<collectingCountry id="21DF764E9C7C421F5400F9C38D217A73" box="[1067,1150,1556,1580]" name="France" pageId="2" pageNumber="121">Corsica</collectingCountry>
|
||
,
|
||
<taxonomicName id="9EC84D5D9C7C421F54ACF9C38C207A73" authority="Mahnert, 1974" authorityName="Mahnert" authorityYear="1974" box="[1159,1407,1556,1580]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="corsa">
|
||
<emphasis id="6BBCEACC9C7C421F54ACF9C38D807A73" box="[1159,1247,1556,1580]" italics="true" pageId="2" pageNumber="121">A. corsa</emphasis>
|
||
<bibRefCitation id="3D594B2F9C7C421F54CFF9C38C207A73" author="Mahnert V." box="[1252,1407,1556,1580]" pageId="2" pageNumber="121" pagination="845 - 885" refId="ref12847" refString="Mahnert V. 1974. Acanthocreagris nov. gen. mit Bemerkungen zur Gattung Microcreagris (Pseudoscorpiones, Neobisiidae). (Uber griechische Pseudoskorpione IV). Revue suisse de Zoologie, 81: 845 - 885." type="book chapter" year="1974">Mahnert, 1974</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9EC84D5D9C7C421F5314F9E38D0C7A13" authority="Mahnert, 1978" authorityName="Mahnert" authorityYear="1978" box="[831,1107,1588,1612]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="aelleni">
|
||
<emphasis id="6BBCEACC9C7C421F5314F9E38AF87A13" box="[831,935,1588,1612]" italics="true" pageId="2" pageNumber="121">A. aelleni</emphasis>
|
||
<bibRefCitation id="3D594B2F9C7C421F539BF9E38D0C7A13" author="Mahnert V." box="[944,1107,1588,1612]" pageId="2" pageNumber="121" pagination="381 - 384" refId="ref12906" refString="Mahnert V. 1978. Pseudoskorpione (Arachnida) aus der Hohle Sisco (Korsika). Revue suisse de Zoologie, 85: 381 - 384." type="book chapter" year="1978">Mahnert, 1978</bibRefCitation>
|
||
</taxonomicName>
|
||
: the first eyed and epigean (see above), the second, described on a male from the Cave of Sisco, anophthalmic with moderate adaptive characters to the subterranean environment. These species have the cheliceral galea apically branched, with two or four branches (
|
||
<taxonomicName id="9EC84D5D9C7C421F5363F9038AF97AB3" authorityName="Mahnert" authorityYear="1974" box="[840,934,1748,1772]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="corsa">
|
||
<emphasis id="6BBCEACC9C7C421F5363F9038AF97AB3" box="[840,934,1748,1772]" italics="true" pageId="2" pageNumber="121">A. corsa</emphasis>
|
||
</taxonomicName>
|
||
), or with two branches (
|
||
<taxonomicName id="9EC84D5D9C7C421F54EEF9038C707AB3" authorityName="Mahnert" authorityYear="1978" box="[1221,1327,1748,1772]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="aelleni">
|
||
<emphasis id="6BBCEACC9C7C421F54EEF9038C707AB3" box="[1221,1327,1748,1772]" italics="true" pageId="2" pageNumber="121">A. aelleni</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="9EC84D5D9C7C421F5560F9038AB77B53" authorityName="Gardini" authorityYear="2022" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="cyrnea" status="sp. nov.">
|
||
<emphasis id="6BBCEACC9C7C421F5560F9038AB77B53" italics="true" pageId="2" pageNumber="121">Acanthocreagris cyrnea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="708F57B79C7C421F53DAF9228D727B53" box="[1009,1069,1781,1804]" pageId="2" pageNumber="121" rank="species">sp. n.</taxonomicNameLabel>
|
||
is to be related to
|
||
<taxonomicName id="9EC84D5D9C7C421F552BF9238C377B53" authorityName="Mahnert" authorityYear="1978" box="[1280,1384,1780,1804]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="aelleni">
|
||
<emphasis id="6BBCEACC9C7C421F552BF9238C377B53" box="[1280,1384,1780,1804]" italics="true" pageId="2" pageNumber="121">A. aelleni</emphasis>
|
||
</taxonomicName>
|
||
, from which it differs chiefly in the higher degree of troglomorphic modifications (comparisons between males): length of pedipalpal femur
|
||
<quantity id="9E309B3B9C7C421F53D1F8838D077B33" box="[1018,1112,1876,1900]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="121" unit="mm" value="1.2">1.20 mm</quantity>
|
||
(5.33 times as long as broad) in
|
||
<taxonomicName id="9EC84D5D9C7C421F5314F8A38AFA7BD3" authorityName="Gardini" authorityYear="2022" box="[831,933,1908,1932]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="cyrnea">
|
||
<emphasis id="6BBCEACC9C7C421F5314F8A38AFA7BD3" box="[831,933,1908,1932]" italics="true" pageId="2" pageNumber="121">A. cyrnea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="9E309B3B9C7C421F5399F8A38D507BD3" box="[946,1039,1908,1932]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="2" pageNumber="121" unit="mm" value="0.79">0.79 mm</quantity>
|
||
(4.45 ×) in
|
||
<taxonomicName id="9EC84D5D9C7C421F54A6F8A38DAD7BD3" authorityName="Mahnert" authorityYear="1978" box="[1165,1266,1908,1932]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="121" phylum="Arthropoda" rank="species" species="aelleni">
|
||
<emphasis id="6BBCEACC9C7C421F54A6F8A38DAD7BD3" box="[1165,1266,1908,1932]" italics="true" pageId="2" pageNumber="121">A. aelleni</emphasis>
|
||
</taxonomicName>
|
||
; length of patella
|
||
<quantity id="9E309B3B9C7D421E50A3FF6389B67C93" box="[136,233,180,204]" metricMagnitude="-3" metricUnit="m" metricValue="1.02" pageId="3" pageNumber="122" unit="mm" value="1.02">1.02 mm</quantity>
|
||
(3.84 ×) in
|
||
<taxonomicName id="9EC84D5D9C7D421E515DFF6388807C93" authorityName="Gardini" authorityYear="2022" box="[374,479,180,204]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="cyrnea">
|
||
<emphasis id="6BBCEACC9C7D421E515DFF6388807C93" box="[374,479,180,204]" italics="true" pageId="3" pageNumber="122">A. cyrnea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="9E309B3B9C7D421E51C4FF638B0E7C93" box="[495,593,180,204]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="3" pageNumber="122" unit="mm" value="0.68">0.68 mm</quantity>
|
||
(3.04 ×) in
|
||
<taxonomicName id="9EC84D5D9C7D421E52F6FF63898F7CB3" authorityName="Mahnert" authorityYear="1978" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="aelleni">
|
||
<emphasis id="6BBCEACC9C7D421E52F6FF63898F7CB3" italics="true" pageId="3" pageNumber="122">A. aelleni</emphasis>
|
||
</taxonomicName>
|
||
; ratio between club and pedicel
|
||
<quantity id="9E309B3B9C7D421E5219FF038B217CB3" box="[562,638,212,236]" metricMagnitude="-2" metricUnit="m" metricValue="4.3942" pageId="3" pageNumber="122" unit="in" value="1.73">1.73 in</quantity>
|
||
<taxonomicName id="9EC84D5D9C7D421E52A3FF038BB27CB3" authorityName="Gardini" authorityYear="2022" box="[648,749,212,236]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="cyrnea">
|
||
<emphasis id="6BBCEACC9C7D421E52A3FF038BB27CB3" box="[648,749,212,236]" italics="true" pageId="3" pageNumber="122">A. cyrnea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="9E309B3B9C7D421E50A3FF23898A7D53" box="[136,213,244,268]" metricMagnitude="-2" metricUnit="m" metricValue="5.308599999999999" pageId="3" pageNumber="122" unit="in" value="2.09">2.09 in</quantity>
|
||
<emphasis id="6BBCEACC9C7D421E50F5FF2388107D53" box="[222,335,244,268]" italics="true" pageId="3" pageNumber="122">
|
||
<taxonomicName id="9EC84D5D9C7D421E50F5FF2388177D53" authorityName="Mahnert" authorityYear="1978" box="[222,328,244,268]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="aelleni">A. aelleni</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
length of chela with pedicel
|
||
<quantity id="9E309B3B9C7D421E52B8FF238BAB7D53" box="[659,756,244,268]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="3" pageNumber="122" unit="mm" value="1.95">1.95 mm</quantity>
|
||
(4.81 ×) in
|
||
<taxonomicName id="9EC84D5D9C7D421E512BFEC388397D73" authorityName="Gardini" authorityYear="2022" box="[256,358,276,300]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="cyrnea">
|
||
<emphasis id="6BBCEACC9C7D421E512BFEC388397D73" box="[256,358,276,300]" italics="true" pageId="3" pageNumber="122">A. cyrnea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="9E309B3B9C7D421E5158FEC3888E7D73" box="[371,465,276,300]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="3" pageNumber="122" unit="mm" value="1.42">1.42 mm</quantity>
|
||
(4.32 ×) in
|
||
<taxonomicName id="9EC84D5D9C7D421E527BFEC38BEA7D73" authorityName="Mahnert" authorityYear="1978" box="[592,693,276,300]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="aelleni">
|
||
<emphasis id="6BBCEACC9C7D421E527BFEC38BEA7D73" box="[592,693,276,300]" italics="true" pageId="3" pageNumber="122">A. aelleni</emphasis>
|
||
</taxonomicName>
|
||
; ratio between pedipalpal femur and carapace
|
||
<quantity id="9E309B3B9C7D421E521DFEE38BDF7D13" box="[566,640,308,332]" metricMagnitude="-2" metricUnit="m" metricValue="3.4544" pageId="3" pageNumber="122" unit="in" value="1.36">1.36 in</quantity>
|
||
<taxonomicName id="9EC84D5D9C7D421E52A3FEE38BB27D13" authorityName="Gardini" authorityYear="2022" box="[648,749,308,332]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="cyrnea">
|
||
<emphasis id="6BBCEACC9C7D421E52A3FEE38BB27D13" box="[648,749,308,332]" italics="true" pageId="3" pageNumber="122">A. cyrnea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="9E309B3B9C7D421E50A3FE83898B7D33" box="[136,212,340,364]" metricMagnitude="-2" metricUnit="m" metricValue="2.9972" pageId="3" pageNumber="122" unit="in" value="1.18">1.18 in</quantity>
|
||
<taxonomicName id="9EC84D5D9C7D421E50F6FE83881C7D33" authorityName="Mahnert" authorityYear="1978" box="[221,323,340,364]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="aelleni">
|
||
<emphasis id="6BBCEACC9C7D421E50F6FE83881C7D33" box="[221,323,340,364]" italics="true" pageId="3" pageNumber="122">A. aelleni</emphasis>
|
||
</taxonomicName>
|
||
. The relative position of trichobothrium
|
||
<emphasis id="6BBCEACC9C7D421E50A1FEA389F67DD3" box="[138,169,372,396]" italics="true" pageId="3" pageNumber="122">isb</emphasis>
|
||
along chelal axis is
|
||
<quantity id="9E309B3B9C7D421E5154FEA388967DD3" box="[383,457,372,396]" metricMagnitude="-2" metricUnit="m" metricValue="1.6002" pageId="3" pageNumber="122" unit="in" value="0.63">0.63 in</quantity>
|
||
<taxonomicName id="9EC84D5D9C7D421E51E4FEA38B6A7DD3" authorityName="Gardini" authorityYear="2022" box="[463,565,372,396]" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="cyrnea">
|
||
<emphasis id="6BBCEACC9C7D421E51E4FEA38B6A7DD3" box="[463,565,372,396]" italics="true" pageId="3" pageNumber="122">A. cyrnea</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<specimenCount id="4FCEFD579C7D421E526DFEA38B077DD3" box="[582,600,372,396]" count="1" pageId="3" pageNumber="122" type="female">♀</specimenCount>
|
||
), ca
|
||
<quantity id="9E309B3B9C7D421E52A6FEA38B887DD3" box="[653,727,372,396]" metricMagnitude="-2" metricUnit="m" metricValue="1.8288" pageId="3" pageNumber="122" unit="in" value="0.72">0.72 in</quantity>
|
||
<taxonomicName id="9EC84D5D9C7D421E52F6FEA3898F7DF3" authorityName="Mahnert" authorityYear="1978" class="Arachnida" family="Neobisiidae" genus="Acanthocreagris" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="122" phylum="Arthropoda" rank="species" species="aelleni">
|
||
<emphasis id="6BBCEACC9C7D421E52F6FEA3898F7DF3" italics="true" pageId="3" pageNumber="122">A. aelleni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<specimenCount id="4FCEFD579C7D421E50F4FE4389AA7DF3" box="[223,245,404,428]" count="1" pageId="3" pageNumber="122" type="male">♂</specimenCount>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |