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135 lines
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<document ID-CLB-Dataset="256667" ID-DOI="http://dx.doi.org/10.3897/phytokeys.225.99356" ID-GBIF-Dataset="9edaba47-012e-4c83-a606-8a760dc24de6" ID-PMC="PMC10209509" ID-Pensoft-Pub="1314-2003-225-115" ID-Pensoft-UUID="AB75E2E9609C5A7583F34B48EA61B976" ID-PubMed="37252326" ModsDocID="1314-2003-225-115" checkinTime="1682624511249" checkinUser="pensoft" docAuthor="Khamar, Hamid, Oualidi, Jalal El, Touhami, Amina Ouazzani & Civeyrel, Laure" docDate="2023" docId="D119486BD6BD512495A3C54682B8E33E" docLanguage="en" docName="PhytoKeys 225: 115-152" docOrigin="PhytoKeys 225" docPubDate="2023-04-27" docSource="http://dx.doi.org/10.3897/phytokeys.225.99356" docTitle="Verbascum tibesticum (Quézel) Hub. - Mor., in Bauhinia 5 (1): 15. 1973." docType="treatment" docVersion="5" id="AB75E2E9609C5A7583F34B48EA61B976" lastPageNumber="115" masterDocId="AB75E2E9609C5A7583F34B48EA61B976" masterDocTitle="Nomenclature and typification in Verbascum (Scrophulariaceae) from North Africa" masterLastPageNumber="152" masterPageNumber="115" pageNumber="115" updateTime="1732936468263" updateUser="ExternalLinkService">
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<mods:title id="EBAE7F1DA6B3DA30E4DDC690D01F6B0B">Nomenclature and typification in Verbascum (Scrophulariaceae) from North Africa</mods:title>
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<mods:namePart id="97664BFEC2BC837537591679205BF241">Khamar, Hamid</mods:namePart>
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<mods:affiliation id="9C64A638C768431CA7C534B0F88C1512">Department of Botany and Plant Ecology, Scientific Institute, University Mohammed V in Rabat, B. P. 703, Rabat 10106, Morocco</mods:affiliation>
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<mods:nameIdentifier id="E42B0099BFA4EEEEB611B157BC7985F7" type="email">khamarhamid90@gmail.com</mods:nameIdentifier>
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<mods:namePart id="8283D2EF006159E5FB42511AA4B0B0BC">Oualidi, Jalal El</mods:namePart>
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<mods:affiliation id="100499FC21986CBB1231D04F4314B318">Department of Botany and Plant Ecology, Scientific Institute, University Mohammed V in Rabat, B. P. 703, Rabat 10106, Morocco</mods:affiliation>
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<mods:namePart id="D29FFFAAEE0DF5A94409C1D809D4F6BA">Touhami, Amina Ouazzani</mods:namePart>
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<mods:affiliation id="CDECCB3CAAED5780ECD2AA8690F2C696">Plant, Animal Productions and Agro-industry Laboratory, Department of Biology, Faculty of Sciences, Ibn Tofail University, B. P. 133 14000, Kenitra, Morocco</mods:affiliation>
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<mods:namePart id="0E462151ED702AF592E6D28213A1ED83">Civeyrel, Laure</mods:namePart>
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<mods:affiliation id="F67AF426F183E8B21584E6F00503B225">Laboratoire Evolution & Diversite Biologique (EDB UMR 5174), Universite de Toulouse, CNRS, IRD, UPS. 118 route de Narbonne, Bat 4 R 1, 31062 Toulouse cedex 9, France</mods:affiliation>
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<mods:title id="22449C1B5F41256EE38463851C179277">PhytoKeys</mods:title>
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<mods:date id="8058F31C9E097213111C1FEAB61BC288">2023</mods:date>
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<mods:number id="DCF8E2CD27E7F24FB60740B0F349931C">2023-04-27</mods:number>
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<mods:number id="C6B84203CFF84555663366A0FE7D5BA2">225</mods:number>
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<mods:classification id="E9814F35C57ACF39B03EF4F5844B43B4">journal article</mods:classification>
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<mods:identifier id="3C662B0BC2905FEB68345E6B7AB46CB3" type="DOI">http://dx.doi.org/10.3897/phytokeys.225.99356</mods:identifier>
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<treatment id="D119486BD6BD512495A3C54682B8E33E" ID-GBIF-Taxon="209579275" LSID="urn:lsid:plazi:treatment:D119486BD6BD512495A3C54682B8E33E" httpUri="http://treatment.plazi.org/id/D119486BD6BD512495A3C54682B8E33E" lastPageNumber="115" pageId="0" pageNumber="115" scope_family="Scrophulariaceae">
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<subSubSection id="FE4AE107ADED9F3DAC14D12EC1D2EA24" pageId="0" pageNumber="115" type="nomenclature">
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<paragraph id="1FEACC270E29442458EA80866F99364D" pageId="0" pageNumber="115">
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<taxonomicName id="DF4A8128023099B70386E7D593351810" LSID="D119486B-D6BD-5124-95A3-C54682B8E33E" authority="(Quezel) Hub. - Mor., in Bauhinia 5 (1): 15. 1973." authorityName="(Quézel) Hub. - Mor., in Bauhinia 5 (1): 15. 1973." class="Dicotyledoneae" family="Scrophulariaceae" genus="Verbascum" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Verbascum tibesticum" order="Lamiales" pageId="0" pageNumber="115" rank="species" species="tibesticum">
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Verbascum tibesticum (
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<normalizedToken id="7EDBAE5802706CC2ADD65094492A969D" originalValue="Quézel">Quezel</normalizedToken>
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) Hub.-Mor., in Bauhinia 5(1): 15. 1973.
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="A137D6434D71CB0990F2B9789AF52491" pageId="0" pageNumber="115" type="reference_group">
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<paragraph id="ADAAFAE87ECDBE704D6D54EDD6E376C1" pageId="0" pageNumber="115">
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<taxonomicName id="41DF6E5D4A93BAA09F5C3A5AC053B0AF" class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia tibestica" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="tibestica">Celsia tibestica</taxonomicName>
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≡
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<taxonomicName id="126247F7B3E570180E232D8BB1790FA0" class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia tibestica" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="tibestica">Celsia tibestica</taxonomicName>
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<normalizedToken id="12EC3A62977D7C72D15FD6D35ADBF117" originalValue="Quézel">Quezel</normalizedToken>
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, in Bull. Soc. Hist. Nat. Afrique N. 48: 95. 1957. Type: [Tchad]. Emi Koussi, Koudou. 2000 m, September-Novembre 1956,
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<emphasis id="C33208A260FCCEC6BCFF47DFED1FF249" italics="true" pageId="0" pageNumber="115">
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P.
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<normalizedToken id="6955949390ACB88AA7CC3476FF2A0438" originalValue="Quézel">Quezel</normalizedToken>
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, s n.
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</emphasis>
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(Lectotype, designated here: AIX [AIX000033]!; isolectotype: AIX [AIX000034]!). [image of lectotype available at https://plants.jstor.org/stable/viewer/10.5555/al.ap.specimen.aix000033]
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</paragraph>
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</subSubSection>
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<subSubSection id="9E6128F85C3621D99E5BAA2F402FB858" pageId="0" pageNumber="115" type="notes">
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<paragraph id="A436178A686292392E992AFCE5362AD9" pageId="0" pageNumber="115">Notes.</paragraph>
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<paragraph id="8B8EA00DC0615A6828A7945773AC0849" pageId="0" pageNumber="115">
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The type material of this species was collected during
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<normalizedToken id="AB27CB2F84D89DB2EBF1804C89BA5C39" originalValue="Quézel’s">Quezel's</normalizedToken>
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expedition to Borkou and to Tibesti in September-November 1956. In the protologue,
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<bibRefCitation id="E3AD8A39DF6E1B1E8465C27932FF1F09" author="Quezel, P" journalOrPublisher="Institut de Recherches Sahariennes" pageId="0" pageNumber="115" pagination="99 - 357" refId="B95" refString="Quezel, P, 1957. Mission Botanique au Tibesti. Contribution a l'etude de la flore et de la vegetation du Borkou et du Tibesti. Memoires. Institut de Recherches Sahariennes 4: 99 - 357" title="Mission Botanique au Tibesti. Contribution a l'etude de la flore et de la vegetation du Borkou et du Tibesti. Memoires." volume="4" year="1957">
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<normalizedToken id="B7975CCCCD285D9DF08D862730CA5347" originalValue="Quézel">Quezel</normalizedToken>
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(1957)
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</bibRefCitation>
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did not designate a holotype nor did he mention the name of the herbarium where the original material was housed. We have traced three specimens at the herbarium of
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<normalizedToken id="020060CFD4FCDC8137DA6C2338BC260A" originalValue="Muséum">Museum</normalizedToken>
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<normalizedToken id="6C3272C1F7ACD861B3CD72ADD7FD30F3" originalValue="d’Histoire">d'Histoire</normalizedToken>
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Naturelle
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<normalizedToken id="42558988C83919C1A65A2FF01E6E467F" originalValue="d’Aix-en-Provence">d'Aix-en-Provence</normalizedToken>
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(AIX), France. The sheet AIX000033 contains a well conserved plant with mature fruit and rosette and an original label handwritten by P.
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<normalizedToken id="23DA2CF54F3518C8E36744BA9FD20E15" originalValue="Quèzel">Quezel</normalizedToken>
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that reads "Mission Botanique de
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<normalizedToken id="9CCD18AB829D785316F243816B34F3DB" originalValue="l’Institut">l'Institut</normalizedToken>
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de Recherches Sahariennes de
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<normalizedToken id="420DAF8015CBE479C5384950277CF119" originalValue="l’Université">l'Universite</normalizedToken>
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<normalizedToken id="1F5B7A33E10C6E80BABE372153B30A1F" originalValue="d’Alger">d'Alger</normalizedToken>
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au Borkou et au Tibesti /
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<taxonomicName id="545E0D2C7384180DE467A34CA957FBCD" class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia tibestica" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="tibestica">
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<emphasis id="AC98CE8DB3B5AF097FAAF42127274775" italics="true" pageId="0" pageNumber="115">Celsia tibestica</emphasis>
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</taxonomicName>
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nov; sp. / Emi Koussi, Koudou. 2000 m / Sept-November 1956, Dr. P.
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<normalizedToken id="8D747DD6B1D727D5EA863DAF5CC6B9E0" originalValue="Quézel”">Quezel"</normalizedToken>
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. The sheet AIX000034 bears only a rosette and an original label handwritten by P.
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<normalizedToken id="588DBE91C32888B42F2C1FA72950A475" originalValue="Quèzel">Quezel</normalizedToken>
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reading "Mission Botanique de
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<normalizedToken id="77FCDB6FE60D6A97B973EBA1BC49EAE7" originalValue="l’Institut">l'Institut</normalizedToken>
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de Recherches Sahariennes de
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<normalizedToken id="25864648D4CA128BEA15D1C4AF6FF3F1" originalValue="l’Université">l'Universite</normalizedToken>
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<normalizedToken id="A1C2DA6AB107DBE366C23B4679F47370" originalValue="d’Alger">d'Alger</normalizedToken>
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au Borkou et au Tibesti /
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<taxonomicName id="9B7524C291718AD02BDE3D972BBAF8B3" class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Celsia tibestica" order="Lamiales" pageId="0" pageNumber="115" phylum="Tracheophyta" rank="species" species="tibestica">
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<emphasis id="F90633F3E4C5A5C5BB4B0475D4B82A5D" italics="true" pageId="0" pageNumber="115">Celsia tibestica</emphasis>
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</taxonomicName>
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Qz. / Koudou / Sept-November 1956, Dr. P.
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<normalizedToken id="39E31D940AC5235545D57AE4A1884DA3" originalValue="Quézel”">Quezel"</normalizedToken>
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. Both specimens are in accordance with the protologue, and based on Art. 9.6 of the ICN (
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<bibRefCitation id="7A7D229FECC8C52656070740DD175C1D" DOI="https://doi.org/10.12705/Code.2018" author="Turland, NJ" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="115" refId="B108" refString="Turland, NJ, Wiersema, JH, Barrie, FR, Greuter, W, Hawksworth, DL, Herendeen, PS, Knapp, S, Kusber, WH, Li, DZ, Marhold, K, May, TW, McNeill, J, Monro, AM, Prado, J, Price, MJ, Smith, GF [Eds], 2018. International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten. https://doi.org/10.12705/Code.2018" title="International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten." url="https://doi.org/10.12705/Code.2018" year="2018">Turland et al. 2018</bibRefCitation>
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) both must be treated as syntypes. The sheet bearing the barcode AIX000033 is designated here as the lectotype, because it is in better condition than the other specimen and shows all relevant characters mentioned in the protologue.
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</paragraph>
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</subSubSection>
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</treatment>
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