treatments-xml/data/D1/18/87/D11887F6C454FFB7C6D0832480C9F9A3.xml
2024-06-21 12:52:41 +02:00

282 lines
34 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="39B1066280FA018BF57213DA88AFCCF8" ID-CLB-Dataset="46798" ID-DOI="10.11646/zootaxa.4078.1.22" ID-GBIF-Dataset="f71df488-ad28-4308-aed3-27540ae6ab4e" ID-ISSN="1175-5326" ID-Zenodo-Dep="255899" ID-ZooBank="C45311A2-E7EA-448F-9F38-79E145E53674" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460361558807" checkinUser="plazi" docAuthor="Grossi, Paschoal C., Santos, Mariana D. &amp; Almeida, Lúcia M." docDate="2016" docId="D11887F6C454FFB7C6D0832480C9F9A3" docLanguage="en" docName="zootaxa.4078.1.22.pdf" docOrigin="Zootaxa 4078 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyclocephala everardoi Grossi, Santos &amp; Almeida, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="250" masterDocId="2D21FF8EC450FFB2C64786128148FF9B" masterDocTitle="Two new species of Cyclocephala (Coleoptera: Scarabaeoidea: Melolonthidae) from Minas Gerais State, Brazil" masterLastPageNumber="251" masterPageNumber="245" pageNumber="249" updateTime="1698294628747" updateUser="ExternalLinkService">
<mods:mods id="974214C98768A38451C289410FA79468" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2A704476D77D55BC7EEF6579F2A73808">
<mods:title id="73EE91F1803E3103CC33F1A8A4636A42">Two new species of Cyclocephala (Coleoptera: Scarabaeoidea: Melolonthidae) from Minas Gerais State, Brazil</mods:title>
</mods:titleInfo>
<mods:name id="B3464502BB48D0FB78C7B25E6AAACE07" type="personal">
<mods:role id="09712E594CF9CBAD6DC0160894E61F7A">
<mods:roleTerm id="9E0318121C319388450B8EF2ACC94B1B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5F58235C02AE63832188E62986BA2B35">Grossi, Paschoal C.</mods:namePart>
</mods:name>
<mods:name id="E74905F14A1F673EA17511067EC51996" type="personal">
<mods:role id="E7501A0AE559EB3D61B497443E2381CC">
<mods:roleTerm id="4039FC09385A03AB1D93A3E31C97266D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="92A201082D04C23A74192F39902EDA31">Santos, Mariana D.</mods:namePart>
</mods:name>
<mods:name id="A70329FE65BEC60DC1154CED253603D2" type="personal">
<mods:role id="850557329A5AAAC845618CCE27676202">
<mods:roleTerm id="7341681A8EF59E2D0F1E89B67C8D2AF5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="61CCA4009FB72DDAAFD235EB304BD13A">Almeida, Lúcia M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="73233FEC8F993125D3B930AAD7A36DEC">text</mods:typeOfResource>
<mods:relatedItem id="705616B423A48D173AA774E1426F0E83" type="host">
<mods:titleInfo id="F67CB03E174B73076856FD2DE7FAEAB4">
<mods:title id="14E6854621CCA45A9EF88DAD1B56D537">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E493053F17C638675C64B0465B261063">
<mods:date id="5AC34416AF645B899F3F144BFFFD0DF7">2016</mods:date>
<mods:detail id="39D45DF910DA80CBAEF6EED0238B8776" type="volume">
<mods:number id="8CAB8F64280389D8FE66ACFCDA1FCAF3">4078</mods:number>
</mods:detail>
<mods:detail id="8A51B80E9F667AFC06854627BF5E0E43" type="issue">
<mods:number id="18A27F8C87F46B70A374BFA1662118CD">1</mods:number>
</mods:detail>
<mods:extent id="66169AAEF43F844093C0006372D19A94" unit="page">
<mods:start id="3C12C0FF38FA0A7836C5E71106C51522">245</mods:start>
<mods:end id="BC50B9DC180EA3748C72B10AAA755164">251</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3539890ED6853C7DC5DE2570137EFBBB">journal article</mods:classification>
<mods:identifier id="9A4D2AEF212141096B7BA356FAD125BA" type="CLB-Dataset">46798</mods:identifier>
<mods:identifier id="F6F38069AE5F154A3ACA2696B6851B5D" type="DOI">10.11646/zootaxa.4078.1.22</mods:identifier>
<mods:identifier id="E34B0360C0321CEB62DF92F0722B2FB2" type="GBIF-Dataset">f71df488-ad28-4308-aed3-27540ae6ab4e</mods:identifier>
<mods:identifier id="9BD59C6C02CAF018B41A196DAF9AE340" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DE1C7AD1C91DC343ACE8583831890CA3" type="Zenodo-Dep">255899</mods:identifier>
<mods:identifier id="159B20305314702A0A6BE95B907F7A28" type="ZooBank">C45311A2-E7EA-448F-9F38-79E145E53674</mods:identifier>
</mods:mods>
<treatment id="D11887F6C454FFB7C6D0832480C9F9A3" ID-DOI="http://doi.org/10.5281/zenodo.5659770" ID-GBIF-Taxon="119409861" ID-Zenodo-Dep="5659770" LSID="urn:lsid:plazi:treatment:D11887F6C454FFB7C6D0832480C9F9A3" httpUri="http://treatment.plazi.org/id/D11887F6C454FFB7C6D0832480C9F9A3" lastPageId="5" lastPageNumber="250" pageId="4" pageNumber="249">
<subSubSection id="11AB656BC454FFB6C6D08324803BFAEA" pageId="4" pageNumber="249" type="nomenclature">
<paragraph id="590E36E0C454FFB6C6D08324835FFACB" blockId="4.[151,535,1334,1394]" box="[151,535,1334,1360]" pageId="4" pageNumber="249">
<heading id="0246818CC454FFB6C6D08324835FFACB" bold="true" box="[151,535,1334,1360]" fontSize="11" level="1" pageId="4" pageNumber="249" reason="1">
<emphasis id="6BC5EAF2C454FFB6C6D08324835FFACB" bold="true" box="[151,535,1334,1360]" pageId="4" pageNumber="249">
<taxonomicName id="9EB14D63C454FFB6C6D0832480FBFACB" box="[151,435,1334,1360]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="everardoi" status="sp. nov.">
<emphasis id="6BC5EAF2C454FFB6C6D0832480FBFACB" bold="true" box="[151,435,1334,1360]" italics="true" pageId="4" pageNumber="249">Cyclocephala everardoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C454FFB6C7FD8324835FFACB" box="[442,535,1334,1360]" pageId="4" pageNumber="249" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="590E36E0C454FFB6C6D0834B803BFAEA" blockId="4.[151,535,1334,1394]" box="[151,371,1369,1394]" pageId="4" pageNumber="249">
(
<figureCitation id="C18A2A65C454FFB6C6D9834B804DFAEA" box="[158,261,1369,1394]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="4" pageNumber="249">Figures 1</figureCitation>
G, 2AE)
</paragraph>
</subSubSection>
<subSubSection id="11AB656BC454FFB7C6D083B6818FFE13" lastPageId="5" lastPageNumber="250" pageId="4" pageNumber="249" type="materials_examined">
<paragraph id="590E36E0C454FFB6C6D083B682E2FA7B" blockId="4.[151,1436,1440,2008]" pageId="4" pageNumber="249">
<emphasis id="6BC5EAF2C454FFB6C6D083B68030FA26" bold="true" box="[151,376,1444,1469]" pageId="4" pageNumber="249">Material examined</emphasis>
.
<typeStatus id="860A8842C454FFB6C7C183B680A6FA26" box="[390,494,1444,1469]" pageId="4" pageNumber="249" type="holotype">Holotype</typeStatus>
male labeled: “
<collectingCountry id="21A67670C454FFB6C4DC83B683AAFA26" box="[667,738,1444,1469]" name="Brazil" pageId="4" pageNumber="249">Brasil</collectingCountry>
, MG,
<quantity id="9E499B05C454FFB6C56283B68215FA26" box="[805,861,1444,1469]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="249" unit="km" value="8.0">8 km</quantity>
ao N de/Cristália,
<date id="2D0F1020C454FFB6C26A83B685EAFA26" box="[1069,1186,1444,1469]" pageId="4" pageNumber="249" value="2010-02-16">16.ii.2010</date>
, luz/
<geoCoordinate id="3C855027C454FFB6C29F83B68465FA27" box="[1240,1325,1440,1469]" direction="south" orientation="latitude" pageId="4" pageNumber="249" precision="925" value="-16.65">16o39'S</geoCoordinate>
,
<geoCoordinate id="3C855027C454FFB6C37F83B684D0FA26" box="[1336,1432,1440,1469]" direction="west" orientation="longitude" pageId="4" pageNumber="249" precision="925" value="-42.883335">42o53'W</geoCoordinate>
,
<quantity id="9E499B05C454FFB6C6D083D58191FA7B" box="[151,217,1479,1504]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="4" pageNumber="249" unit="m" value="720.0">720m</quantity>
,/Grossi, Parizotto &amp; Melo/ao amanhecer (56: a.m.)” (
<collectionCode id="3FA0AE25C454FFB6C50683DA82D7FA7B" box="[833,927,1480,1504]" pageId="4" pageNumber="249">CERPE</collectionCode>
).
</paragraph>
<paragraph id="590E36E0C454FFB7C68083FE818FFE13" blockId="4.[151,1436,1440,2008]" lastBlockId="5.[151,1436,151,464]" lastPageId="5" lastPageNumber="250" pageId="4" pageNumber="249">
<emphasis id="6BC5EAF2C454FFB6C68083FE8032F99E" bold="true" box="[199,378,1516,1541]" pageId="4" pageNumber="249">
<typeStatus id="860A8842C454FFB6C68083FE807CF99E" box="[199,308,1516,1541]" pageId="4" pageNumber="249" type="holotype">Holotype</typeStatus>
male
</emphasis>
.
<emphasis id="6BC5EAF2C454FFB6C7C983FE836BF99E" box="[398,547,1516,1541]" italics="true" pageId="4" pageNumber="249">Total length:</emphasis>
<quantity id="9E499B05C454FFB6C46883FE83D4F99F" box="[559,668,1516,1540]" metricMagnitude="-3" metricUnit="m" metricValue="9.01" pageId="4" pageNumber="249" unit="mm" value="9.01">9.01 mm</quantity>
.
<emphasis id="6BC5EAF2C454FFB6C4EB83FE82EDF99E" box="[684,933,1516,1541]" italics="true" pageId="4" pageNumber="249">Total pronotal width:</emphasis>
<quantity id="9E499B05C454FFB6C5F483FE8557F99F" box="[947,1055,1516,1541]" metricMagnitude="-3" metricUnit="m" metricValue="3.59" pageId="4" pageNumber="249" unit="mm" value="3.59">3.59 mm</quantity>
.
<emphasis id="6BC5EAF2C454FFB6C27783FE853AF99E" box="[1072,1138,1516,1541]" italics="true" pageId="4" pageNumber="249">Body:</emphasis>
Dorsal surface glabrous, ventrally densely setose, setae distinctly elongate. Color brownish red, with lighter areas on clypeal apex, pronotum, scutellum (
<figureCitation id="C18A2A65C454FFB6C7DD802680ACF9D7" box="[410,484,1588,1613]" captionStart="FIGURE 2" captionStartId="3.[151,250,1966,1988]" captionTargetBox="[215,1381,209,1933]" captionTargetId="figure@3.[187,1400,193,1945]" captionTargetPageId="3" captionText="FIGURE 2. Cyclocephala everardoi sp. nov. A. holotype male, dorsal habitus; B. head pronotum, and protibiae in detail; C. antennal lamellae; D. anterior protarsus; E. parameres in caudal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/255901/files/figure.png" pageId="4" pageNumber="249">Fig. 2</figureCitation>
A), legs, and venter.
<emphasis id="6BC5EAF2C454FFB6C4A380268262F9D6" box="[740,810,1588,1613]" italics="true" pageId="4" pageNumber="249">Head:</emphasis>
Shape transverse with abrupt narrowing to clypeus, sparsely punctate; punctures larger on clypeus and on sides; clypeus narrowly parabolic, ⅓ of head width, with elongated punctures, anteriorly concave, and with Cshaped punctures, apex reflexed and bordered (
<figureCitation id="C18A2A65C454FFB6C375806E8433F90F" box="[1330,1403,1660,1685]" captionStart="FIGURE 2" captionStartId="3.[151,250,1966,1988]" captionTargetBox="[215,1381,209,1933]" captionTargetId="figure@3.[187,1400,193,1945]" captionTargetPageId="3" captionText="FIGURE 2. Cyclocephala everardoi sp. nov. A. holotype male, dorsal habitus; B. head pronotum, and protibiae in detail; C. antennal lamellae; D. anterior protarsus; E. parameres in caudal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/255901/files/figure.png" pageId="4" pageNumber="249">Fig. 2</figureCitation>
B). Frontoclypeal suture obsolete at middle, with 2 lateral, transverse impressions. Antennae with lamellae extremely developed, almost
<date id="2D0F1020C454FFB6C72C80D680CFF947" box="[363,391,1732,1756]" pageId="4" pageNumber="249">3x</date>
longer than antennomeres 37 together, and strongly and densely punctate with a spongiose aspect, and with scattered setae (
<figureCitation id="C18A2A65C454FFB6C44F80F58303F89B" box="[520,587,1767,1792]" captionStart="FIGURE 2" captionStartId="3.[151,250,1966,1988]" captionTargetBox="[215,1381,209,1933]" captionTargetId="figure@3.[187,1400,193,1945]" captionTargetPageId="3" captionText="FIGURE 2. Cyclocephala everardoi sp. nov. A. holotype male, dorsal habitus; B. head pronotum, and protibiae in detail; C. antennal lamellae; D. anterior protarsus; E. parameres in caudal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/255901/files/figure.png" pageId="4" pageNumber="249">Fig. 2</figureCitation>
C); first lamella externally with longitudinal row of dense setae; eyes large, wider than clypeal width (
<figureCitation id="C18A2A65C454FFB6C7FE811E80B3F8BF" box="[441,507,1804,1829]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="4" pageNumber="249">Figs 1</figureCitation>
G2B).
<emphasis id="6BC5EAF2C454FFB6C41E811E83ADF8BE" box="[601,741,1804,1829]" italics="true" pageId="4" pageNumber="249">Mouthparts:</emphasis>
Mandibles slender, apices upturned and externally curved and with elongated setae, basal half laterally crenulated; last maxillar palpomere slightly longer than the preceding 2 together.
<emphasis id="6BC5EAF2C454FFB6C74181478037F8F7" box="[262,383,1877,1900]" italics="true" pageId="4" pageNumber="249">Pronotum:</emphasis>
Surface weakly and sparsely punctate, punctures stronger posteriorly; posterior margin slightly projected backward, rounded, with no bead; sides with long setae, weakly crenulated. Scutellum punctate; punctures small to large, some elongate.
<emphasis id="6BC5EAF2C454FFB6C418818E83E6F82E" box="[607,686,1948,1973]" italics="true" pageId="4" pageNumber="249">Elytra:</emphasis>
Surface distinctly and irregularly punctate; striae not well defined, slightly wrinkled near suture.
<emphasis id="6BC5EAF2C454FFB6C7A481D3836AF843" box="[483,546,1985,2008]" italics="true" pageId="4" pageNumber="249">Legs:</emphasis>
Protibiae with 3 distinct teeth, teeth decreasing in size from apex to base; apical tooth twice as long as middle tooth, middle tooth more than
<date id="2D0F1020C455FFB7C516868A8225FF2B" box="[849,877,152,176]" pageId="5" pageNumber="250">3x</date>
longer than basal tooth (
<figureCitation id="C18A2A65C455FFB7C2CA8685859EFF2B" box="[1165,1238,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1966,1988]" captionTargetBox="[215,1381,209,1933]" captionTargetId="figure@3.[187,1400,193,1945]" captionTargetPageId="3" captionText="FIGURE 2. Cyclocephala everardoi sp. nov. A. holotype male, dorsal habitus; B. head pronotum, and protibiae in detail; C. antennal lamellae; D. anterior protarsus; E. parameres in caudal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/255901/files/figure.png" pageId="5" pageNumber="250">Figs 2</figureCitation>
AB); inner claw widely bifid, forming a right angle (
<figureCitation id="C18A2A65C455FFB7C47586AE833FFF4F" box="[562,631,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1966,1988]" captionTargetBox="[215,1381,209,1933]" captionTargetId="figure@3.[187,1400,193,1945]" captionTargetPageId="3" captionText="FIGURE 2. Cyclocephala everardoi sp. nov. A. holotype male, dorsal habitus; B. head pronotum, and protibiae in detail; C. antennal lamellae; D. anterior protarsus; E. parameres in caudal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/255901/files/figure.png" pageId="5" pageNumber="250">Fig. 2</figureCitation>
D).
<emphasis id="6BC5EAF2C455FFB7C4E486AF83B8FF4F" box="[675,752,189,212]" italics="true" pageId="5" pageNumber="250">Venter:</emphasis>
Anterior prosternal margin sinuous and crenulated by setae insertion, distinctly trilobed at middle, with lateral lobes more prominent than middle lobe, conspicuous (
<figureCitation id="C18A2A65C455FFB7C37286CD843DFF63" box="[1333,1397,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="5" pageNumber="250">Fig. 1</figureCitation>
G). Prosternal process high, exceeding coxal height.
<emphasis id="6BC5EAF2C455FFB7C4FF87168262FE86" box="[696,810,260,285]" italics="true" pageId="5" pageNumber="250">Genitalia:</emphasis>
Parameres symmetrical with elongated parameres, with 2 lateral projections, one basal and one at middle; basal larger, middle one projecting backwards; apices diverging internally, externally with small serrations; subapex with a slightly diverging, rounded tooth at external angle (
<figureCitation id="C18A2A65C455FFB7C328875E81EEFE13" captionStart="FIGURE 2" captionStartId="3.[151,250,1966,1988]" captionTargetBox="[215,1381,209,1933]" captionTargetId="figure@3.[187,1400,193,1945]" captionTargetPageId="3" captionText="FIGURE 2. Cyclocephala everardoi sp. nov. A. holotype male, dorsal habitus; B. head pronotum, and protibiae in detail; C. antennal lamellae; D. anterior protarsus; E. parameres in caudal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/255901/files/figure.png" pageId="5" pageNumber="250">Fig. 2</figureCitation>
E).
</paragraph>
</subSubSection>
<subSubSection id="11AB656BC455FFB7C680878680C9F9A3" pageId="5" pageNumber="250" type="etymology">
<paragraph id="590E36E0C455FFB7C6808786857AFE4B" blockId="5.[151,1436,151,464]" pageId="5" pageNumber="250">
<emphasis id="6BC5EAF2C455FFB7C6808786800EFE36" bold="true" box="[199,326,404,429]" pageId="5" pageNumber="250">Etymology</emphasis>
. This species is named in honor of the dedicated entomologist, Everardo José Grossi, father of the first author, a doctor with a passion for beetles and an expert in rhinoceros beetles.
</paragraph>
<caption id="0DCE6668C455FFB7C6D084128583FD8C" ID-Table-UUID="0DCE6668C455FFB7C6D084128583FD8C" box="[151,1227,511,535]" httpUri="http://table.plazi.org/id/0DCE6668C455FFB7C6D084128583FD8C" pageId="5" pageNumber="250" targetBox="[151,1408,562,1199]" targetIsTable="true" targetPageId="5">
<paragraph id="590E36E0C455FFB7C6D084128583FD8C" blockId="5.[151,1227,511,535]" box="[151,1227,511,535]" pageId="5" pageNumber="250">
TABLE 1. Main characters to differentiate the two new species of
<taxonomicName id="9EB14D63C455FFB7C5C487ED855FFD8C" box="[899,1047,511,535]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="genus">
<emphasis id="6BC5EAF2C455FFB7C5C487ED855FFD8C" bold="true" box="[899,1047,511,535]" italics="true" pageId="5" pageNumber="250">Cyclocephala</emphasis>
</taxonomicName>
described here.
</paragraph>
</caption>
<paragraph id="590E36E0C455FFB7C6D884208053FB34" pageId="5" pageNumber="250">
<table id="2BB1C440C455004DC6D0842084C8FB34" box="[151,1408,562,1199]" gridcols="3" gridrows="9" pageId="5" pageNumber="250">
<tr id="E78134A2C455004DC6D0842084C8FDD3" box="[151,1408,562,584]" gridrow="0" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D08420802DFDD3" box="[151,357,562,584]" gridcol="0" gridrow="0" pageId="5" pageNumber="250">
<emphasis id="6BC5EAF2C455FFB7C6D884208040FDD3" bold="true" box="[159,264,562,584]" pageId="5" pageNumber="250">Character</emphasis>
</th>
<th id="A4505DDEC455004DC7B1842082E5FDD3" box="[502,941,562,584]" gridcol="1" gridrow="0" pageId="5" pageNumber="250">
<emphasis id="6BC5EAF2C455FFB7C7B18420838EFDD3" bold="true" box="[502,710,562,584]" pageId="5" pageNumber="250">
<taxonomicName id="9EB14D63C455FFB7C7B18420833CFDD3" box="[502,628,562,584]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C7B18420833CFDD3" bold="true" box="[502,628,562,584]" italics="true" pageId="5" pageNumber="250">C. machadoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C43D8420838EFDD3" box="[634,710,562,584]" pageId="5" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="A4505DDEC455004DC581842084C8FDD3" box="[966,1408,562,584]" gridcol="2" gridrow="0" pageId="5" pageNumber="250">
<emphasis id="6BC5EAF2C455FFB7C581842085D9FDD3" bold="true" box="[966,1169,562,584]" pageId="5" pageNumber="250">
<taxonomicName id="9EB14D63C455FFB7C58184208577FDD3" box="[966,1087,562,584]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="everardoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C58184208577FDD3" bold="true" box="[966,1087,562,584]" italics="true" pageId="5" pageNumber="250">C. everardoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C202842085D9FDD3" box="[1093,1169,562,584]" pageId="5" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="E78134A2C455004DC6D0844884C8FDF4" box="[151,1408,602,623]" gridrow="1" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D08448802DFDF4" box="[151,357,602,623]" gridcol="0" gridrow="1" pageId="5" pageNumber="250">Color</th>
<td id="A4505DDEC455004DC7B1844882E5FDF4" box="[502,941,602,623]" gridcol="1" gridrow="1" pageId="5" pageNumber="250">Testaceous with black head</td>
<td id="A4505DDEC455004DC581844884C8FDF4" box="[966,1408,602,623]" gridcol="2" gridrow="1" pageId="5" pageNumber="250">Brownish red</td>
</tr>
<tr id="E78134A2C455004DC6D0849084C8FD2F" box="[151,1408,642,692]" gridrow="2" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D08490802DFD2F" box="[151,357,642,692]" gridcol="0" gridrow="2" pageId="5" pageNumber="250">Clypeus</th>
<td id="A4505DDEC455004DC7B1849082E5FD2F" box="[502,941,642,692]" gridcol="1" gridrow="2" pageId="5" pageNumber="250">Almost impunctate, strongly reflexed.</td>
<td id="A4505DDEC455004DC581849084C8FD2F" box="[966,1408,642,692]" gridcol="2" gridrow="2" pageId="5" pageNumber="250">With elongated and Cshaped punctures, slightly reflexed</td>
</tr>
<tr id="E78134A2C455004DC6D084D584C8FD61" box="[151,1408,711,762]" gridrow="3" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D084D5802DFD61" box="[151,357,711,762]" gridcol="0" gridrow="3" pageId="5" pageNumber="250">Frontoclypeal suture</th>
<td id="A4505DDEC455004DC7B184D582E5FD61" box="[502,941,711,762]" gridcol="1" gridrow="3" pageId="5" pageNumber="250">Obsolete, lacking impression</td>
<td id="A4505DDEC455004DC58184D584C8FD61" box="[966,1408,711,762]" gridcol="2" gridrow="3" pageId="5" pageNumber="250">Obsolete at middle and with two transverse impressions</td>
</tr>
<tr id="E78134A2C455004DC6D0851F84C8FCA4" box="[151,1408,781,831]" gridrow="4" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D0851F802DFCA4" box="[151,357,781,831]" gridcol="0" gridrow="4" pageId="5" pageNumber="250">Pronotum</th>
<td id="A4505DDEC455004DC7B1851F82E5FCA4" box="[502,941,781,831]" gridcol="1" gridrow="4" pageId="5" pageNumber="250">Surface moderately and uniformly punctate, smooth at center line</td>
<td id="A4505DDEC455004DC581851F84C8FCA4" box="[966,1408,781,831]" gridcol="2" gridrow="4" pageId="5" pageNumber="250">Surface weakly and sparsely punctate, punctures posteriorly stronger</td>
</tr>
<tr id="E78134A2C455004DC6D0854084C8FC1F" box="[151,1408,850,900]" gridrow="5" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D08540802DFC1F" box="[151,357,850,900]" gridcol="0" gridrow="5" pageId="5" pageNumber="250">Elytra</th>
<td id="A4505DDEC455004DC7B1854082E5FC1F" box="[502,941,850,900]" gridcol="1" gridrow="5" pageId="5" pageNumber="250">Striae distinct, marked by lines of punctures; interstriae almost impunctate</td>
<td id="A4505DDEC455004DC581854084C8FC1F" box="[966,1408,850,900]" gridcol="2" gridrow="5" pageId="5" pageNumber="250">Striae indistinct and surface nearly totally, uniformly punctate</td>
</tr>
<tr id="E78134A2C455004DC6D0858584C8FC7C" box="[151,1408,919,999]" gridrow="6" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D08585802DFC7C" box="[151,357,919,999]" gridcol="0" gridrow="6" pageId="5" pageNumber="250">Protibiae</th>
<td id="A4505DDEC455004DC7B1858582E5FC7C" box="[502,941,919,999]" gridcol="1" gridrow="6" pageId="5" pageNumber="250">Basal tooth weak, and more than 3x smaller than middle tooth</td>
<td id="A4505DDEC455004DC581858584C8FC7C" box="[966,1408,919,999]" gridcol="2" gridrow="6" pageId="5" pageNumber="250">Basal tooth more developed, and distinctly projected, three times smaller than middle tooth</td>
</tr>
<tr id="E78134A2C455004DC6D085E884C8FBB7" box="[151,1408,1018,1068]" gridrow="7" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D085E8802DFBB7" box="[151,357,1018,1068]" gridcol="0" gridrow="7" pageId="5" pageNumber="250">Aedeagi</th>
<td id="A4505DDEC455004DC7B185E882E5FBB7" box="[502,941,1018,1068]" gridcol="1" gridrow="7" pageId="5" pageNumber="250">Parameres with apices parallel; middle tooth small, projecting laterally</td>
<td id="A4505DDEC455004DC58185E884C8FBB7" box="[966,1408,1018,1068]" gridcol="2" gridrow="7" pageId="5" pageNumber="250">Parameres with apices diverging; middle tooth more developed, projecting backwards</td>
</tr>
<tr id="E78134A2C455004DC6D0828784C8FB34" box="[151,1408,1173,1199]" gridrow="8" pageId="5" pageNumber="250" rowspan-1="1" rowspan-2="1">
<th id="A4505DDEC455004DC6D08287802DFB34" box="[151,357,1173,1199]" gridcol="0" gridrow="8" pageId="5" pageNumber="250">Discussion</th>
</tr>
</table>
</paragraph>
<paragraph id="590E36E0C455FFB7C6D082CE80C9F9A3" blockId="5.[151,1436,1244,1592]" pageId="5" pageNumber="250">
The specimens were collected with UV light.
<taxonomicName id="9EB14D63C455FFB7C4F682CE8289FB6E" box="[689,961,1244,1269]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="everardoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C4F682CE8289FB6E" box="[689,961,1244,1269]" italics="true" pageId="5" pageNumber="250">Cyclocephala everardoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C58B82CE8561FB6E" box="[972,1065,1244,1269]" pageId="5" pageNumber="250" rank="species">
<emphasis id="6BC5EAF2C455FFB7C58B82CE8561FB6E" bold="true" box="[972,1065,1244,1269]" pageId="5" pageNumber="250">sp. nov.</emphasis>
</taxonomicNameLabel>
was taken at dawn in
<date id="2D0F1020C455FFB7C37082CE819AFA83" pageId="5" pageNumber="250" value="2010-02">February 2010</date>
, between 5:00 and 6:00 A.M., in an area of sandy and rupestrian Cerrado fields (
<figureCitation id="C18A2A65C455FFB7C2C982ED859BFA83" box="[1166,1235,1279,1304]" captionStart="FIGURE 3" captionStartId="4.[151,250,1212,1234]" captionTargetBox="[177,1408,243,1176]" captionTargetId="figure@4.[171,1416,193,1190]" captionTargetPageId="4" captionText="FIGURE 3. General aspect of the region of Cristália municipality, Minas Gerais State, Brazil, where the holotype of C. everardoi sp. nov. was collected." httpUri="https://zenodo.org/record/255902/files/figure.png" pageId="5" pageNumber="250">Fig. 3</figureCitation>
A).
<taxonomicName id="9EB14D63C455FFB7C34582ED804DFAA6" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C34582ED804DFAA6" italics="true" pageId="5" pageNumber="250">Cyclocephala machadoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C74A8336802FFAA6" box="[269,359,1316,1341]" pageId="5" pageNumber="250" rank="species">
<emphasis id="6BC5EAF2C455FFB7C74A8336802FFAA6" bold="true" box="[269,359,1316,1341]" pageId="5" pageNumber="250">sp. nov.</emphasis>
</taxonomicNameLabel>
lacks other information. Both
<typeStatus id="860A8842C455FFB7C48483368278FAA6" box="[707,816,1316,1341]" pageId="5" pageNumber="250" type="holotype">holotypes</typeStatus>
are pinned; in
<taxonomicName id="9EB14D63C455FFB7C59C83378520FAA6" box="[987,1128,1316,1341]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="everardoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C59C83378520FAA6" box="[987,1128,1316,1341]" italics="true" pageId="5" pageNumber="250">C. everardoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C23783368582FAA6" box="[1136,1226,1316,1341]" pageId="5" pageNumber="250" rank="species">
<emphasis id="6BC5EAF2C455FFB7C23783368582FAA6" bold="true" box="[1136,1226,1316,1341]" pageId="5" pageNumber="250">sp. nov.</emphasis>
</taxonomicNameLabel>
part of the middle tooth of left protibia is missing, while in
<taxonomicName id="9EB14D63C455FFB7C42F835A83BFFAFB" box="[616,759,1351,1376]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C42F835A83BFFAFB" box="[616,759,1351,1376]" italics="true" pageId="5" pageNumber="250">C. machadoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C54783558212FAFB" box="[768,858,1351,1376]" pageId="5" pageNumber="250" rank="species">
<emphasis id="6BC5EAF2C455FFB7C54783558212FAFB" bold="true" box="[768,858,1351,1376]" pageId="5" pageNumber="250">sp. nov.</emphasis>
</taxonomicNameLabel>
part of the right first and second antennal lamellae are missing. They were initially confused with
<taxonomicName id="9EB14D63C455FFB7C4E0837E85C9FA1F" authority="Erichson 1847" authorityName="Erichson" authorityYear="1847" box="[679,1153,1388,1413]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="macrophylla">
<emphasis id="6BC5EAF2C455FFB7C4E0837E829DFA1E" box="[679,981,1388,1413]" italics="true" pageId="5" pageNumber="250">Cyclocephala macrophylla</emphasis>
Erichson 1847
</taxonomicName>
.
<taxonomicName id="9EB14D63C455FFB7C2CB837E84D3FA1E" box="[1164,1435,1388,1413]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C2CB837E84D3FA1E" box="[1164,1435,1388,1413]" italics="true" pageId="5" pageNumber="250">Cyclocephala machadoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C6D0839D81B9FA33" box="[151,241,1423,1448]" pageId="5" pageNumber="250" rank="species">
<emphasis id="6BC5EAF2C455FFB7C6D0839D81B9FA33" bold="true" box="[151,241,1423,1448]" pageId="5" pageNumber="250">sp. nov.</emphasis>
</taxonomicNameLabel>
is similar to
<taxonomicName id="9EB14D63C455FFB7C7C383828359FA33" box="[388,529,1423,1448]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="everardoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C7C383828359FA33" box="[388,529,1423,1448]" italics="true" pageId="5" pageNumber="250">C. everardoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C45F839D833AFA33" box="[536,626,1423,1448]" pageId="5" pageNumber="250" rank="species">
<emphasis id="6BC5EAF2C455FFB7C45F839D833AFA33" bold="true" box="[536,626,1423,1448]" pageId="5" pageNumber="250">sp. nov.</emphasis>
</taxonomicNameLabel>
and was also confused with this. In comparing them, we concluded that they were similar but distinct species and they are diagnosed in the
<tableCitation id="1433035BC455FFB7C5DD83A682A7FA57" box="[922,1007,1460,1485]" captionStart="TABLE 1" captionStartId="5.[151,239,512,535]" captionTargetBox="[151,1408,562,1199]" captionTargetPageId="5" captionText="TABLE 1. Main characters to differentiate the two new species of Cyclocephala described here." httpUri="http://table.plazi.org/id/0DCE6668C455FFB7C6D084128583FD8C" pageId="5" pageNumber="250" tableUuid="0DCE6668C455FFB7C6D084128583FD8C">Table 1</tableCitation>
. We also believe that they compose a new species group within
<emphasis id="6BC5EAF2C455FFB7C78383C5832DFA6B" box="[452,613,1495,1520]" italics="true" pageId="5" pageNumber="250">
<taxonomicName id="9EB14D63C455FFB7C78383C58328FA6B" box="[452,608,1495,1520]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="genus">Cyclocephala</taxonomicName>
,
</emphasis>
being diagnosed by the short body length, extremely elongate antennal lamellae, eyes very developed, clypeus elongate and truncate, somewhat quadrate and inner claw of anterior tarsi with a wide incision.
</paragraph>
</subSubSection>
</treatment>
</document>