384 lines
47 KiB
XML
384 lines
47 KiB
XML
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<mods:title id="7FC3B329C0CCA0926AA584827A3D719E">Description of three new species of the genus Chromaphyosemion Radda, 1971 (Cyprinodontiformes: Nothobranchiidae) from the coastal plains of Cameroon with a preliminary review of the Chromaphyosemion splendopleure complex.</mods:title>
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<mods:roleTerm id="0C80C540DC131DCB34CA7CD4DA08271C">Author</mods:roleTerm>
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<mods:namePart id="B44835489F54684E03E4603751CEFF8F">Rainer Sonnenberg</mods:namePart>
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<mods:typeOfResource id="E630BE310009F8D35F2F13F0D0448734">text</mods:typeOfResource>
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<mods:title id="95DF67C4547217FF7F9384399A4A2C5E">Zootaxa</mods:title>
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<mods:date id="481B5FF2E57B9FF3A7E76668D49DE9F3">2007</mods:date>
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<mods:detail id="B614D574F6249BE4F14B172A9AD765BF" type="volume">
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<mods:number id="2D605BCBD0F019039BCFD04BAD28A0A7">1591</mods:number>
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<mods:start id="8C4D2758E243E4D76CDEEF8B5A917DDA">1</mods:start>
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<mods:classification id="4136CA53207F2F3F05454F84E681EC36">journal article</mods:classification>
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<mods:identifier id="7A42060FBD8F72B92650C038BE6ADF67" type="ZooBank-PUB">z01591p001</mods:identifier>
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<treatment id="D0A1B27ACEC5634C34425A5BE3702A98" ID-DOI="http://doi.org/10.5281/zenodo.6248383" ID-GBIF-Taxon="125136720" ID-Zenodo-Dep="6248383" LSID="urn:lsid:plazi:treatment:D0A1B27ACEC5634C34425A5BE3702A98" httpUri="http://treatment.plazi.org/id/D0A1B27ACEC5634C34425A5BE3702A98" lastPageNumber="11" pageNumber="7">
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<subSubSection id="409AEA44AFC652A314F9C9EC4CC09BB6" type="nomenclature">
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<paragraph id="EB7E7474C78E7A0A2295B57481B6431C" pageNumber="7">
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<taxonomicName id="111330138568852CDC69B538C3231494" ID-CoL="5YGJQ" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" species="koungueense" status="sp. nov.">Chromaphyosemion koungueense</taxonomicName>
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<taxonomicNameLabel id="3DACC520F6FF65DB3F6717DDA532FEBB" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="83732F9336BE236D8D55C66DC0456901" type="description">
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<paragraph id="E2CD61A3667A809391A07212A4B26295" pageNumber="7">(Figs. 3-6)</paragraph>
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</subSubSection>
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<subSubSection id="CECE3232F379FC9BF0B4E07D6E4EA743" type="reference_group">
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<paragraph id="FAC70C7820C5F5334EF66C0C94CEDF4F" pageNumber="7">
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<taxonomicName id="DDDD15D914D8ED925C480C9DD600BF38" LSID-ZBK="urn:lsid:zoobank.org:act:78C80644-7E92-46C8-8753-3D4BA9CB628B" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" lsidName-ZBK="Aphyosemion bivittatum hollyi Myers 1933:184" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" species="bivittatum">Aphyosemion bivittatum</taxonomicName>
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(non
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<bibRefCitation id="D1E532F7861799A05ED2F4F7C87416E6" author="Loennberg, E." pagination="179 - 195" refId="ref17678" refString="Loennberg, E. (1895) Notes on fishes collected in the Cameroons by Mr. Y. Sjoestedt. Oefversigt af Kongliga VetenskapsAkademiens Foerhandlingar, 3, 179 - 195." type="journal article" year="1895">
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<normalizedToken id="99C2BEF8E79D401B11277E1256CBEC79" originalValue="Lönnberg">Loennberg</normalizedToken>
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, 1895
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</bibRefCitation>
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),
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<bibRefCitation id="27A71683DEA748191502F0335587F31B" author="Scheel, J. J." refId="ref18680" refString="Scheel, J. J. (1968) Rivulins of the Old World. TFH Publications, New Jersey, 480 pp." type="book" year="1968">Scheel 1968: 117-127</bibRefCitation>
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(in part, distribution, morphology, taxonomy, synonymy), 1974: 47-98 (in part, BIV-DI strain, BIV-NN strain, color pattern, crossing experiments, distribution, karyotypes).
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</paragraph>
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<paragraph id="3175F4F46DC0BF7BAA5859935815CC3C" pageNumber="7">
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<taxonomicName id="3D2067E4BE550C2D3030367B1C6DA0FC" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" species="bitaeniatum" subGenus="Chromaphyosemion">Aphyosemion (Chromaphyosemion) bitaeniatum</taxonomicName>
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(non
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<bibRefCitation id="1EE6EE50A068CFB9DBA45CE20ED3EC55" author="Ahl, E." pagination="135 - 145" refId="ref16192" refString="Ahl, E. (1924) Neue afrikanische Zahnkarpfen aus dem zoologischen Museum Berlin. Zoologischer Anzeiger, 61, 135 - 145." type="journal article" year="1924">Ahl, 1924</bibRefCitation>
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),
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<bibRefCitation id="092882ABD4CC11DD246897B8A1063CE7" author="Scheel, J. J." refId="ref18754" refString="Scheel, J. J. (1990) Atlas of the Killifishes of the Old World. TFH Publication, Neptune City, New Jersey, 448 pp." type="book" year="1990">Scheel 1990: 168-173</bibRefCitation>
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(in part, distribution, karyotypes, taxonomy).
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</paragraph>
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<paragraph id="88049B7EAFD6A37B966827FDDB959C1D" pageNumber="7">
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<taxonomicName id="457AB6F3C96DECF7C18B2C7DE0D6C87E" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" species="splendopleure" subGenus="Chromaphyosemion">Aphyosemion (Chromaphyosemion) splendopleure</taxonomicName>
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(non
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<bibRefCitation id="4B6C9D7AE4420106800EC0E109FC0B79" author="Bruening, C." pagination="356" refId="ref16519" refString="Bruening, C. (1929 a) Der gruene Leuchtaugenfisch und der afrikanische Glanzkaerpfling. Wochenschrift fuer Aquarien- und Terrarienkunde, 26, 356." type="journal article" year="1929">
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<normalizedToken id="CA0A4909659BCB59FEF5639A3132F2F8" originalValue="Brüning">Bruening</normalizedToken>
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, 1929a
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</bibRefCitation>
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),
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<bibRefCitation id="963BB7BCC9A55B27B6920B02C0593830" author="Agnese, J. - F. & Zentz, F. & Legros, O. & Sellos, D." pagination="332 - 346" refId="ref16129" refString="Agnese, J. - F., Zentz, F., Legros, O. & Sellos, D. (2006) Phylogenetic relationships and phylogeography of the killifish species of the subgenus Chromaphyosemion (Radda, 1971) in West Africa, inferred from mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 40, 332 - 346." type="journal article" year="2006">
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<normalizedToken id="F5957557032D75E2BC681FF4565FF5D4" originalValue="Agnèse">Agnese</normalizedToken>
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et al. 2006
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</bibRefCitation>
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(phylogeny, collection localities, taxonomy),
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<bibRefCitation id="1821DCDE4A6084ACFF84EC9E3953E5C1" author="Amiet, J. L." refId="ref16267" refString="Amiet, J. L. (1987) Le Genre Aphyosemion Myers (Pisces, Teleostei, Cyprinodontiformes). Faune du Cameroun, Volume 2. Sciences Nat, Compiegne, France, 262 pp." type="book" year="1987">Amiet 1987: 49</bibRefCitation>
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, 176, plate 36, figs. 13 and 14 (in part, collection localities, color pattern),
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<bibRefCitation id="E98D31506C422F5129B8BEF162B9CFF8" author="Huber, J. H." pagination="15 - 23" refId="ref17026" refString="Huber, J. H. (1998 a) Aphyosemion alpha n. sp. - Eine neue Art der Untergattung Chromaphyosemion mit einem ausgepraegtem Faerbungsmuster und einer besonders suedlichen Verbreitung. Das Aquarium, 350, 15 - 23." type="journal article" year="1998">Huber 1998a: 20-21</bibRefCitation>
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(in part, taxonomy, distribution), 2000: 471 (in part, collection localities),
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<bibRefCitation id="F393FF28CE260F0667DFDEAB97814CB9" author="Legros, O. & Zentz, F." pagination="1 - 28" refId="ref17585" refString="Legros, O. & Zentz, F. (2007) Aphyosemion malumbresi n. sp., un nouveau Chromaphyosemion (Teleostei: Aplocheilidae) originaire de Guinee Equatoriale continentale. AKFB Killi-Contact, no. 6, decembre 2006, 1 - 28." type="journal article" year="2007">Legros & Zentz 2007: 20-21</bibRefCitation>
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(phylogeny),
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<bibRefCitation id="9D2B08B33CF41E74E5C0D4164D7CC7A0" author="Radda, A. C. & Puerzl, E." refId="ref18353" refString="Radda, A. C. & Puerzl, E. (1982) Killifische aus aller Welt. Feldfuehrer der Cyprinodontiformes der Laender der Bucht von Biafra (SO-Nigeria, West-Kamerun). Band 2. Verlag Otto Hofmann, Wien, 72 pp." type="book" year="1982">
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Radda &
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<normalizedToken id="BB923FC901879F288C3029982BA20C30" originalValue="Pürzl">Puerzl</normalizedToken>
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1982: 24-25
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</bibRefCitation>
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(in part, distribution),
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<bibRefCitation id="9CC52648DA95DBE64017618B3AF5A394" author="Radda, A. C. & Puerzl, E." refId="ref18399" refString="Radda, A. C. & Puerzl, E. (1983) Killifische aus aller Welt. Feldfuehrer der Cyprinodontiformes der Laender der Kuesten- laender Zentralafrikas I Ostkamerun. Band 3. Verlag Otto Hofmann, Wien, 80 pp." type="book" year="1983">
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Radda &
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<normalizedToken id="7660EB4845412E59C2E5C62FC9711263" originalValue="Pürzl">Puerzl</normalizedToken>
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1983: 20-21
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</bibRefCitation>
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(in part, distribution) 1987: 70 (in part, distribution),
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<bibRefCitation id="6667E1C670E476A17762984AFB55B118" author="Seegers, L." pagination="155 - 163" refId="ref18837" refString="Seegers, L. (1981) Chromaphyosemion - kleine Farbwunder aus Westafrika. Die Aquarien- und Terrarienzeitschrift, 34, 155 - 163." type="journal article" year="1981">Seegers 1981: 159</bibRefCitation>
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, 161 (in part, distribution), 1986: 52-54 (in part, distribution), 1997: 122 (color photo, male from Douala),
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<bibRefCitation id="D0A5D77E16BE1BFDFC143D134A1FD32E" author="Wildekamp, R. H." refId="ref19762" refString="Wildekamp, R. H. (1993) A World of Killies. Atlas of the Oviparous Cyprinodontiform Fishes of the World. Volume 1. The American Killifish Association, Mishawaka, Indiana, 311 pp." type="book" year="1993">Wildekamp 1993: 277-279</bibRefCitation>
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(in part, distribution).
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</paragraph>
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<paragraph id="F2ADA0A0821E0FF43BB45BD8ECF8D84C" pageNumber="7">
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<taxonomicName id="5FEED291274EE2F31049C3B87860EA94" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" species="splendopleure" subGenus="Chromaphyosemion">Aphyosemion (Chromaphyosemion) splendopleure</taxonomicName>
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phenotype Dizangue (non
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<bibRefCitation id="B521218021A972C100F5D717C3CEFFF8" author="Bruening, C." pagination="356" refId="ref16519" refString="Bruening, C. (1929 a) Der gruene Leuchtaugenfisch und der afrikanische Glanzkaerpfling. Wochenschrift fuer Aquarien- und Terrarienkunde, 26, 356." type="journal article" year="1929">
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<normalizedToken id="92ED14222B0E876A5142A13FE8FE77EB" originalValue="Brüning">Bruening</normalizedToken>
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, 1929a
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</bibRefCitation>
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),
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<bibRefCitation id="836BF7867AC0DA8456B1BD1CBC72858B" author="Eberl, W." refId="ref16675" refString="Eberl, W. (1996) Die Untergattung Chromaphyosemion. Deutsche Killifisch Gemeinschaft Journal, Supplementheft 4, 88 pp." type="book" year="1996">Eberl 1996: 50</bibRefCitation>
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(color pattern, collection localities, color photo p.1),
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<bibRefCitation id="F772FAFBD1E13439B1DB55F024125E92" author="Eigelshofen, W." pagination="8 - 13" refId="ref16697" refString="Eigelshofen, W. (1998) Chromaphyosemion. Beeindruckend in Form und Farbe. Aquaristik Aktuell, 9 - 10, 8 - 13." type="journal article" year="1998">Eigelshofen 1998: 10-12</bibRefCitation>
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(distribution, color photo).
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</paragraph>
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<paragraph id="476D56173292D39C0C3F531FE0443735" pageNumber="7">
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<taxonomicName id="4DDCC0A7D34AF4812C7E52FA1C9173A8" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" species="splendopleure">Chromaphyosemion splendopleure</taxonomicName>
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phenotype 'Dizangue ' (non
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<bibRefCitation id="66D55BCDD640D00C62D98A86A12F6DB9" author="Bruening, C." pagination="356" refId="ref16519" refString="Bruening, C. (1929 a) Der gruene Leuchtaugenfisch und der afrikanische Glanzkaerpfling. Wochenschrift fuer Aquarien- und Terrarienkunde, 26, 356." type="journal article" year="1929">
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<normalizedToken id="D2229E64EA793EC1031118C17ED3A3E6" originalValue="Brüning">Bruening</normalizedToken>
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, 1929a
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</bibRefCitation>
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),
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<bibRefCitation id="62C220168FD89C383593194901F4A9F7" author="Legros, O. & Zentz, F. & Agnese, J. F." pagination="3 - 30" refId="ref17632" refString="Legros, O., Zentz, F. & Agnese, J. F. (2005) Description de deux nouveaux Chromaphyosemion (Teleostei: Aplocheilidae) du sud de la Plaine littorale camerounaise. AKFB Killi-Contact, 33, 3 - 30." type="journal article" year="2005">Legros et al. 2005:16-17</bibRefCitation>
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(color pattern, distribution, phylogeny).
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</paragraph>
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<paragraph id="287A0D4F481F752683A4D6CBF9FA3105" pageNumber="7">
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<taxonomicName id="CD656B302A3AC97A82DDA398893E5288" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" species="splendopleure">Chromaphyosemion sp. aff. splendopleure</taxonomicName>
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'Dizangue',
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<bibRefCitation id="BE495DC03170D78C64C9139AE4E9A137" author="Sonnenberg, R." pagination="79 - 94" refId="ref19032" refString="Sonnenberg, R. (2000) The Distribution of Chromaphyosemion Radda, 1971 (Teleostei: Cyprinodontiformes) on the Coastal Plains of West and Central Africa. In Rheinwald, G. (Ed.) Isolated Vertebrate Communities in the Tropics. Proceedings of the 4 th International Symposium, Bonn. Bonner zoologische Monographien 46, pp. 79 - 94." type="journal article" year="2000">Sonnenberg 2000: 81</bibRefCitation>
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, 85 (color pattern, distribution, color photo Nr. 3).
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</paragraph>
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</subSubSection>
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<subSubSection id="85B3B20E45541CE58346E6538A442457" type="materials_examined">
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<paragraph id="CB9739EE576F44B2DDD51AFF18065B9F" pageNumber="7">
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<materialsCitation id="EDDB1F259EA80139566046124C1963CF" ID-GBIF-Occurrence="1416639495" collectionCode="ZFMK" collectorName="U. Kaempf, R. Sonnenberg, A. Traenkner" country="Cameroon" latitude="3.81025" location="at the road from Edea to Dizangue and Ndonga, in the Koungue, a small river in the forest near the village Koungue Ndonga, collection locality CMM 3" longitude="9.91775" specimenCode="ZFMK 39949" typeStatus="holotype">
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<typeStatus id="F1D53AFD16B00CD6A03753AD683A244F" type="holotype">Holotype</typeStatus>
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.
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<specimenCode id="1D5792DFC6B038B62CA62452FB902BC0">
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<collectionCode id="5A12FFD415B849E41B6939B8049EF902">ZFMK</collectionCode>
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39949
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</specimenCode>
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, male (24.3 mm SL),
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<collectingCountry id="F08A4E18DD85CD644F5A383A40CD454E">Cameroon</collectingCountry>
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,
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<location id="8EA23C00FCE27F3B1B5C846C68622AF5" LSID="urn:lsid:plazi:treatment:D0A1B27ACEC5634C34425A5BE3702A98:8EA23C00FCE27F3B1B5C846C68622AF5" country="Cameroon" latitude="3.81025" longitude="9.91775" name="at the road from Edea to Dizangue and Ndonga, in the Koungue, a small river in the forest near the village Koungue Ndonga, collection locality CMM 3">
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at the road from Edea to Dizangue and Ndonga, in the
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<normalizedToken id="F5DA94B6823F58935302435C439E3FAE" originalValue="Koungué">Koungue</normalizedToken>
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, a small river in the forest near the village
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<normalizedToken id="C61D7952E9B34F525175997F47DACBC0" originalValue="Koungué">Koungue</normalizedToken>
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Ndonga
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</location>
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(
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<geoCoordinate id="CC2AA9CB265084CAC8D19A4EA20707CF" direction="north" orientation="latitude" value="3.81025">03°48'36.9'' N</geoCoordinate>
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,
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<geoCoordinate id="A876922221905F47465C1D7DE83268E8" direction="east" orientation="longitude" value="9.91775">09°55'03.9'' E</geoCoordinate>
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),
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<collectorName id="43331F7E7D60DE012721A0EFCE6C458F">
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U.
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<normalizedToken id="4B28F30153E26EFA7F41592014AE1EBF" originalValue="Kämpf">Kaempf</normalizedToken>
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, R. Sonnenberg, and A.
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<normalizedToken id="445C49DEC737556E52BEB499CF709DFC" originalValue="Tränkner">Traenkner</normalizedToken>
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</collectorName>
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leg., collection locality CMM 3.
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</materialsCitation>
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</paragraph>
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<paragraph id="732D599A64039E7EC7ED4A9824EE6FD2" pageNumber="7">
|
|
<materialsCitation id="C2F61BA6F9E4C5C27A19AAA0F8E3C0BC" ID-GBIF-Occurrence="1416639500" collectionCode="ZFMK" collectorName="U. Kaempf, R. Sonnenberg, A. Traenkner" country="Cameroon" latitude="3.81025" location="at the road from Edea to Dizangue and Ndonga, in the Koungue, a small river in the forest near the village Koungue Ndonga, collection locality CMM 3" longitude="9.91775" specimenCode="ZFMK 39950 - 39955" typeStatus="paratype">
|
|
<typeStatus id="09501E53E76521334C53232B9F1FE975" type="paratype">Paratypes</typeStatus>
|
|
.
|
|
<specimenCode id="B0E245D69952B9B9365D1DAAEA55676A">
|
|
<collectionCode id="D76028B9D39C177527F3B2A2568F99DF">ZFMK</collectionCode>
|
|
39950-39955
|
|
</specimenCode>
|
|
(2 males, 3 females, 1 juvenile),
|
|
<typeStatus id="FDBF35CB2E8C207784B042B32DB9AF55" type="paratopotype">paratopotypes</typeStatus>
|
|
,
|
|
<location id="EF47700712F3FAF2E35BE20C28ADA660" LSID="urn:lsid:plazi:treatment:D0A1B27ACEC5634C34425A5BE3702A98:EF47700712F3FAF2E35BE20C28ADA660" country="Cameroon" latitude="3.81025" longitude="9.91775" name="at the road from Edea to Dizangue and Ndonga, in the Koungue, a small river in the forest near the village Koungue Ndonga, collection locality CMM 3">collected with the holotype</location>
|
|
.
|
|
</materialsCitation>
|
|
</paragraph>
|
|
<paragraph id="87B24B6BF65C6C677F72B18E8A5B3348" pageNumber="7">
|
|
<materialsCitation id="7CBD91D0F3C298B48DE3C97420523809" ID-GBIF-Occurrence="1416639514" collectionCode="ZFMK" collectorName="H. Kullmann and M. Voelker" country="Cameroon" latitude="3.7135556" location="a small creek crossing the road Mbambou to Mouanko, some kilometers in oil palm plantation in direction Mouanko, collection locality KV 03/14" longitude="9.857834" specimenCode="ZFMK 39956 - 39957" typeStatus="paratype">
|
|
<specimenCode id="3A0DFF3C38F4B4698528E3A7DB8CA366">
|
|
<collectionCode id="9606E34F15A0124D467A3E903A70BF83">ZFMK</collectionCode>
|
|
39956-39957
|
|
</specimenCode>
|
|
(2 males),
|
|
<collectingCountry id="FBEA4197C95473D8317758D2E4B146CF">Cameroon</collectingCountry>
|
|
,
|
|
<location id="D276AADF6695B11DFA53A9C4C241FC21" LSID="urn:lsid:plazi:treatment:D0A1B27ACEC5634C34425A5BE3702A98:D276AADF6695B11DFA53A9C4C241FC21" country="Cameroon" latitude="3.7135556" longitude="9.857834" name="a small creek crossing the road Mbambou to Mouanko, some kilometers in oil palm plantation in direction Mouanko, collection locality KV 03/14">a small creek crossing the road Mbambou to Mouanko, some kilometers in oil palm plantation in direction Mouanko</location>
|
|
(
|
|
<geoCoordinate id="8DAE5882E891E0D1FB3B555ADE336431" direction="north" orientation="latitude" value="3.7135556">03°42'48.8'' N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="EA94F62CE26270A40EBF178BB61903D1" direction="east" orientation="longitude" value="9.857834">09°51'28.2'' E</geoCoordinate>
|
|
),
|
|
<collectorName id="886CFF4DEDCF4D05022D3BB19A25E451">
|
|
H. Kullmann and M.
|
|
<normalizedToken id="9551E82BFD4A57CDD42BEDC3CA0719A4" originalValue="Völker">Voelker</normalizedToken>
|
|
</collectorName>
|
|
leg., collection locality KV 03/14.
|
|
</materialsCitation>
|
|
</paragraph>
|
|
<paragraph id="3C71F7B324538C20B44BE72D78EFC6E9" pageNumber="7">
|
|
<materialsCitation id="6FEA592C5B8DD1C796A420E38F000772" ID-GBIF-Occurrence="1416639497" collectionCode="ZFMK" collectorName="U. Kaempf, R. Sonnenberg and A. Traenkner" country="Cameroon" latitude="3.541361" location="near the road from Edea to Kribi, collection locality CMM 8" longitude="10.086694" specimenCode="ZFMK 39958 - 39959" typeStatus="paratype">
|
|
<specimenCode id="08CB203E277A15AB2E7FEC01B90FBD1E">
|
|
<collectionCode id="6D0D9945E444805C23368044A1ADBB6F">ZFMK</collectionCode>
|
|
39958-39959
|
|
</specimenCode>
|
|
(2 males),
|
|
<collectingCountry id="749549DD8BDE0986A7D568B0180E8369">Cameroon</collectingCountry>
|
|
,
|
|
<location id="9FCA2241F57E339609DF7A0619E91301" LSID="urn:lsid:plazi:treatment:D0A1B27ACEC5634C34425A5BE3702A98:9FCA2241F57E339609DF7A0619E91301" country="Cameroon" latitude="3.541361" longitude="10.086694" name="near the road from Edea to Kribi, collection locality CMM 8">near the road from Edea to Kribi</location>
|
|
(
|
|
<geoCoordinate id="BBBEACB9FAC2660012540CE0EA0CAE0E" direction="north" orientation="latitude" value="3.541361">03°32'28.9'' N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="88D49892CFE65190E19AAD0534A3D54B" direction="east" orientation="longitude" value="10.086694">10°05'12.1'' E</geoCoordinate>
|
|
),
|
|
<collectorName id="F0E27CC5698F87785798F59780C7C8B1">
|
|
U.
|
|
<normalizedToken id="F64D62616C5415844B1933EF32269E59" originalValue="Kämpf">Kaempf</normalizedToken>
|
|
, R. Sonnenberg and A.
|
|
<normalizedToken id="2FD409DA80B0A51EC837CC98B50E5FDB" originalValue="Tränkner">Traenkner</normalizedToken>
|
|
</collectorName>
|
|
leg., collection locality CMM 8.
|
|
</materialsCitation>
|
|
</paragraph>
|
|
<paragraph id="20A18DFAF5D450DED1857F06435E3B61" pageNumber="7">
|
|
<materialsCitation id="9FE76DC95B4F14E33E7AE2A205C4EC8B" ID-GBIF-Occurrence="1416639510" collectionCode="ZFMK" collectorName="H. Kullmann, B. Misof and R. Sonnenberg" country="Cameroon" latitude="3.7216945" location="a small creek crossing the road from Edea to Kribi, near the village Koukoue, collection locality C 01/77" longitude="10.107472" specimenCode="ZFMK 39960 - 39964" typeStatus="paratype">
|
|
<specimenCode id="6D505A93737486F3AD44AFA2D332F429">
|
|
<collectionCode id="68B9EC5C06178951F8000D238AA6B8CA">ZFMK</collectionCode>
|
|
39960-39964
|
|
</specimenCode>
|
|
(3 males, 2 females),
|
|
<collectingCountry id="D85F765B6A22951BDCB965CF2FE0182C">Cameroon</collectingCountry>
|
|
,
|
|
<location id="C0FBA0365A75709EE751D3AE18E9677D" LSID="urn:lsid:plazi:treatment:D0A1B27ACEC5634C34425A5BE3702A98:C0FBA0365A75709EE751D3AE18E9677D" country="Cameroon" latitude="3.7216945" longitude="10.107472" name="a small creek crossing the road from Edea to Kribi, near the village Koukoue, collection locality C 01/77">
|
|
a small creek crossing the road from Edea to Kribi, near the village
|
|
<normalizedToken id="8B0172FA6E0A13B86C913511DC0CBF3C" originalValue="Koukoué">Koukoue</normalizedToken>
|
|
</location>
|
|
(
|
|
<geoCoordinate id="99CECAF7F5A4C912897B6E3B9BF1DDF4" direction="north" orientation="latitude" value="3.7216945">03°43'18.1'' N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="0FA164320654D340AEF97C346D86F19D" direction="east" orientation="longitude" value="10.107472">10°06'26.9'' E</geoCoordinate>
|
|
),
|
|
<collectorName id="88E5CA0CAA4A070538D0E4A31E69337E">H. Kullmann, B. Misof and R. Sonnenberg</collectorName>
|
|
leg., collection locality C 01/77.
|
|
</materialsCitation>
|
|
</paragraph>
|
|
<paragraph id="D2135DCDE1A913586CA463D62FFB4665" pageNumber="7">
|
|
<materialsCitation id="6DCB591461C716ACC032D8BC8DF6E217" ID-GBIF-Occurrence="1416639496" collectionCode="ZFMK" collectorName="H. Kullmann, B. Misof and R. Sonnenberg" country="Cameroon" latitude="3.608111" location="a small creek near the road Edea to Kribi, collection locality C 01/78" longitude="10.107278" specimenCode="ZFMK 39965 - 39969" typeStatus="paratype">
|
|
<specimenCode id="A823182AA55854EEECBD38A51F11A3E2">
|
|
<collectionCode id="AE267C790E834879844154D8AB9E02F5">ZFMK</collectionCode>
|
|
39965-39969
|
|
</specimenCode>
|
|
(4 males, 1 female),
|
|
<collectingCountry id="FB5B32A0DC6B0E6C8409BF7D8727182F">Cameroon</collectingCountry>
|
|
,
|
|
<location id="99C63B0E9BB926526F4A3857C2462381" LSID="urn:lsid:plazi:treatment:D0A1B27ACEC5634C34425A5BE3702A98:99C63B0E9BB926526F4A3857C2462381" country="Cameroon" latitude="3.608111" longitude="10.107278" name="a small creek near the road Edea to Kribi, collection locality C 01/78">a small creek near the road Edea to Kribi</location>
|
|
(
|
|
<geoCoordinate id="5F67A56CD38BB18B0E2F4277A90D9BC8" direction="north" orientation="latitude" value="3.608111">03°36'29.2'' N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="1673E03DA234B1C1E55852B49A31EDC5" direction="east" orientation="longitude" value="10.107278">10°06'26.2'' E</geoCoordinate>
|
|
),
|
|
<collectorName id="1BF84474E4DDCF610D6A2026CADD1DED">H. Kullmann, B. Misof and R. Sonnenberg</collectorName>
|
|
leg., collection locality C 01/78.
|
|
</materialsCitation>
|
|
</paragraph>
|
|
<paragraph id="12FC5BAC8790F03FB25A38C56D87CBBA" pageNumber="7">
|
|
<materialsCitation id="A9940801D6DA23883DF99D3FAA902F1F" ID-GBIF-Occurrence="1416639498" collectionCode="CENAREST" collectorName="H. Kullmann, B. Misof and R. Sonnenberg" country="Cameroon" latitude="3.7216945" location="a small creek crossing the road from Edea to Kribi, near the village Koukoue, collection locality C 01/77" longitude="10.107472" typeStatus="paratype">
|
|
<collectionCode id="E9AC0F76CFBEDD0B60986A2ABC6C0CEA">CENAREST</collectionCode>
|
|
, Batoke,
|
|
<collectingCountry id="3FCCE79C2BABD59408CED4A365A7483F">Cameroon</collectingCountry>
|
|
(1 male, 1 female),
|
|
<location id="8BC0AC9E768B085E9A3A8FC7D2EA1409" LSID="urn:lsid:plazi:treatment:D0A1B27ACEC5634C34425A5BE3702A98:8BC0AC9E768B085E9A3A8FC7D2EA1409" country="Cameroon" latitude="3.7216945" longitude="10.107472" name="a small creek crossing the road from Edea to Kribi, near the village Koukoue, collection locality C 01/77">same collection data as ZFMK 39960-39964, collection locality C 01/77</location>
|
|
.
|
|
</materialsCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="2E2769D71AFAA561C9D4595B3FDF472F" type="diagnosis">
|
|
<paragraph id="EDA6B2917BA5BC29B4774A939F3BF9F6" lastPageNumber="8" pageNumber="7">
|
|
Diagnosis.
|
|
<taxonomicName id="B5983442E54ED9FEEB4DD50A4EDC4E95" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" species="koungueense">Chromaphyosemion koungueense</taxonomicName>
|
|
is distinguished from its closest relatives,
|
|
<taxonomicName id="6BB2C7DD3B7BEC37E6E7FAD725E531F5" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" species="omega" status="sp. nov.">C. omega</taxonomicName>
|
|
<taxonomicNameLabel id="82D2BB6E26A6777CB9E3541A800C0A62" rank="species">sp. nov.</taxonomicNameLabel>
|
|
and
|
|
<taxonomicName id="57A575E1FFACCFA68AFEE7377317B184" LSID-ZBK="urn:lsid:zoobank.org:act:58293C95-476D-42FC-B52D-62A8938702E3" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" lsidName-ZBK="Chromaphyosemion punctulatum Legros, Zentz & Agnèse 2005" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" species="punctulatum">C. punctulatum</taxonomicName>
|
|
, by the bluish-green color on side of males and copper-brown metallic scales on dorsal part of sides; with bluish-green to yellow-greenish unpaired fins, on dorsal and caudal fins with red dots or streaks in basal and streaks in distal parts, almost no red patterns on pelvic and anal fins except for small red stripes on fin rays in anal fin in some specimens versus bluish-orange on sides with metallic greenish to copper scales on dorsal sides in combination with mainly orange fins with greenish center in dorsal and caudal and nearly complete orange anal and pelvic fins in
|
|
<taxonomicName id="4B6DD8FFC69C3D10FBBAD1CFA6301642" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" species="omega" status="sp. nov.">C. omega</taxonomicName>
|
|
<taxonomicNameLabel id="49CB5698A1D35454C89F5766E8126BA6" rank="species">sp. nov.</taxonomicNameLabel>
|
|
or greenish sides with bluish-green metallic scales on dorsal part of sides, red dots on greenish to yellowish-green anal fin in combination with mainly red dots in dorsal and caudal fin in
|
|
<taxonomicName id="A6A9EDE6A274058930042AC2E10209A9" LSID-ZBK="urn:lsid:zoobank.org:act:58293C95-476D-42FC-B52D-62A8938702E3" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" lsidName-ZBK="Chromaphyosemion punctulatum Legros, Zentz & Agnèse 2005" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" species="punctulatum">C. punctulatum</taxonomicName>
|
|
. Lateral dark stripes in males more prominent than in
|
|
<taxonomicName id="81338628CB2927EAEC7806F87DA364FC" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" species="omega" status="sp. nov.">C. omega</taxonomicName>
|
|
<taxonomicNameLabel id="18E55B68964086F8CFB7ED30EF98C2C0" rank="species">sp. nov.</taxonomicNameLabel>
|
|
and
|
|
<taxonomicName id="C1AA7278781AB6094B389C2160FD8D32" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" sensu="lato" species="splendopleure" status="s.l.">C. splendopleure</taxonomicName>
|
|
<taxonomicNameLabel id="764D09F9DF14887EA2819ABACBA73E12" sensu="lato">s.l.</taxonomicNameLabel>
|
|
It is distinguished from
|
|
<taxonomicName id="641749291AF12F56734C163CB7B66E4D" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" species="loennbergii">C. loennbergii</taxonomicName>
|
|
by the greenish body color and greenish color of pelvic and anal fins, if yellow to yellow-orange then only in lower half versus metallic copper color on scales on dorsal part of sides and bluish on sides in males, with blue in lower half and orange in upper half of anal fin. It is distinguished from
|
|
<taxonomicName id="B6B137AF50E7837149CE4B89058F9563" LSID-ZBK="urn:lsid:zoobank.org:act:494F6568-0395-45FD-B6C4-99E11F027244" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" lsidName-ZBK="Chromaphyosemion melanogaster Legros, Zentz & Agnèse 2005" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" species="melanogaster">C. melanogaster</taxonomicName>
|
|
by the more bluish-green color on side of males together with the orange anterior ventral color of body and orange in caudal and anal fins versus green color on sides and not orange on anterior ventral body, unpaired fins green. The species can be distinguished from
|
|
<taxonomicName id="9D4C383BF9779E355D743C40822FCC9F" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="7" phylum="Chordata" rank="species" sensu="lato" species="splendopleure" status="s.l.">C. splendopleure</taxonomicName>
|
|
<taxonomicNameLabel id="56C59844800F5A740F824426A6004D38" sensu="lato">s.l.</taxonomicNameLabel>
|
|
by the sides being bluish-green with greenish color of anal fin and marginal blue band versus reddish, orange or yellow with bluish or pinkish on sides in adult males, orange or yellowgreenish anal fin with greenish or orange marginal band. It is distinguished by all remaining
|
|
<taxonomicName id="5B7D09C75735662C0A9F290E2BE24389" genus="Chromphyosemion" pageNumber="7" rank="genus">Chromphyosemion</taxonomicName>
|
|
species by the unique combination of color patterns.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="871EE31F92DA3B63EAD5A6146C383375" type="description">
|
|
<paragraph id="E7EC2B1168B21AB93FC0A5945D71C2E5" pageNumber="8">
|
|
Description. See Figures 3-6 for general appearance and Table 2 for morphometric data of the type series. Shows all characters apomorphic for
|
|
<taxonomicName id="B12FEAD83D526A9966C5CF598DC8CCEA" LSID-ZBK="urn:lsid:zoobank.org:act:10DF0F9D-4060-4E5F-A156-DFE88550256D" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" lsidName-ZBK="Chromaphyosemion Aphyosemion Radda 1971:118" order="Cyprinodontiformes" pageNumber="8" phylum="Chordata" rank="genus">Chromaphyosemion</taxonomicName>
|
|
, strong sexual dimorphism, adult males more colorful and usually larger than females and having larger fins with extended fin rays. Slender, laterally slightly rounded and elongate small nothobranchiid species, females anteriorly more rounded than males. Dorsal profile straight on head, from nape to caudal peduncle slightly convex, greatest body height anterior to base of dorsal and anal fin. Ventral profile slightly convex from head to end of anal fin base, caudal peduncle slightly concave on ventral. Snout slightly rounded, mouth directed upwards, lower jaw longer than upper, posterior end of mouth at same level as center of eye. Dorsal and anal fins in males pointed, dorsal fin with elongated fin streamer, anal fin trapezoid, only slightly pointed. Caudal fin in males with middle lobe; elongated fin streamers with same color as in dorsal fin. Fins in females generally smaller and rounded. Unpaired fins usually covered basally with an opaque mucus film. Dorsal and anal fins posterior to mid-body First ray of dorsal fin in front or above 1st or 2nd anal fin ray. Dorsal fin with 11-13 rays, anal fin with 13-14 rays. Pectoral fins extending over origin of pelvic fins, pelvic fins not or just reaching anal fin.
|
|
</paragraph>
|
|
<paragraph id="C281EFC016A8B635A233B9CB5D847168" pageNumber="9">Scales cycloid, body and head entirely scaled, except ventral surface of head. Frontal squamation of G- type, scales on mid-longitudinal series 24-26 + 2-3 on caudal fin base. Transverse row of scales above pelvic fins 8-9, circum-peduncular scale row 12-13. Open frontal neuromast system with two separate grooves, preopercular neuromast system with 6 pores.</paragraph>
|
|
<paragraph id="DCA2D0EF3661DC24C501D9C154EE9360" pageNumber="9">Coloration. Live specimens. Males (Figs. 3-5). Back brownish, scale border darker, some with red dots, upper rows of scales on sides from approximately area of greatest body height to end of caudal peduncle reflecting metallic copper to greenish. On sides two parallel black stripes, the upper one starting on lower lip through center of eye straight to dorsal part of caudal peduncle, the lower stripe starting on throat, running directly under eye to ventral part of caudal peduncle, both ending at caudal fin base. Black stripes can completely disappear depending on mood and social status of individual or only traces are seen from upper one on anterior half of body, lower one from anterior base of anal fin to base of caudal fin on ventral caudal peduncle; in fully colored males a dark marking of upper stripe left directly behind opercle. Sides pale grey to beige, many scales with dark border or red dot. Anterior ventral part in males often orange, on lower dark stripe scales with metallic bluish to greenish sheen. In fully colored males, throat and belly at anterior part of body orange, above and on posterior sides metallic bluish-green or greenish, ventrally posterior lower black stripe often visible.</paragraph>
|
|
<paragraph id="F7604E75529A75858E203DFB0F0AAA1D" lastPageNumber="11" pageNumber="10">Dorsal fin in center bluish-green or greenish, on anterior part sometimes with copper or orange area, posterior green. On basal part red dots between fin rays, in most specimens fusing to streaks in upper half of fin, fin streamer yellow to orange. Anal fin greenish, on basal part blue in several specimens, submarginal band red, marginal band blue, in central part of anal fin sometimes with a yellowish-green or yellowish-orange area, often restricted between fin rays. Usually without red dots or streaks in anal fin, in some specimens with basal row of red dots. Caudal fin upper and lower margin blue, in lower part with submarginal red band, in upper part blue margin interrupted by red streaks. Fin in center green or bluish-green, in most specimens one not sharply separated yellowish-green to orange band above submarginal red band. Between fin rays red streaks in posterior central part, in anterior part quite variable, varying from red dots to streaks, red pattern ranging between fine to rather broad markings. Orange fin streamers in adult males, in young fish sometimes whitish to yellow. Pelvic fins similarly colored as anal fin, greenish to yellowish-green with red submarginal and blue marginal band. Pectoral fins hyaline with yellow to greenish hue, more intensive on outer fin border. Females (Fig. 6). Back brown, dorsally first three rows of scales with metallic copper or greenish color, similar but less pronounced as in males; scales with dark border, some with red dot, on sides two parallel black stripes, between stripes pale greyish to bluish, ventrally greyish to whitish. Some specimens with small red dots on scales, more or less regularly in rows. Orange area on opercle between lateral stripes. All fins hyaline, coloration similar to fins of males, metallic bluish-greenish. Dark or red dots in dorsal, dots or streaks in caudal fin. Anal fin greenish with dark submarginal region and marginal blue band. Pelvic fins hyaline greenish with bluish margin. Pectoral fins hyaline with little dark pigmentation.</paragraph>
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<paragraph id="2596BE7FA9EF7A8BC0676B5C962F8A67" pageNumber="11">Preserved in ethanol. Males. Back and sides brown, both lateral stripes dark brown, belly light brown to whitish. Scales on back with dark brown border forming reticulated pattern, on back and sides sometimes traces of red on scales. Border of opercle ventral to lower stripe dark, below lower lip whitish to beige. Unpaired and pelvic fins hyaline greyish with dark pigmentation between fin rays, dark margins on unpaired and pelvic fins. Pectoral fins hyaline.</paragraph>
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<paragraph id="B4427DC96FDB6980440CEECD4F710C96" pageNumber="11">Females. Similar to males and juveniles. Body brownish, scales on back with dark border, belly beige to whitish. Fins hyaline greyish, little dark pigmentation and no dark margin on anal fin.</paragraph>
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</subSubSection>
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<subSubSection id="71A1983F4E64753FDEB379DFD1DCB4AA" type="etymology">
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<paragraph id="49751ED4FA2C801B4F179F164684C150" pageNumber="11">
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Etymology. This species is named after the river and the nearby village
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<normalizedToken id="A348DF8D15C1E841D93982F12E6C46D7" originalValue="Koungué">Koungue</normalizedToken>
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, the collection locality of the holotype. With this name, I also want to acknowledge the friendly welcome and invaluable help of the
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<normalizedToken id="2FD474BBE1536A8E903416A0AB3F3B24" originalValue="Koungué">Koungue</normalizedToken>
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villagers.
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</paragraph>
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</subSubSection>
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<subSubSection id="1279E07573B7A0CFEA98B173A7DB978B" type="distribution">
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<paragraph id="93C1293679B9E0836E631DF57DB1F5AC" pageNumber="11">
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Distribution and habitat.
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<taxonomicName id="CA07ACBFB8C97F0981806FC9F3FD4C38" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" species="koungueense">C. koungueense</taxonomicName>
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is endemic in Cameroon and known from the area north of Kribi up to Douala at the Wouri (see Map 1). South of the Sanaga it is replaced in the east by
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<taxonomicName id="E51E1E1DD1FCC03FABEC3728B5CEB0B0" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" species="loennbergii">C. loennbergii</taxonomicName>
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, north of the Sanaga by
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<taxonomicName id="0A2263F5025DF4A1D860E7279BDF4315" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" species="omega" status="sp. nov.">C. omega</taxonomicName>
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<taxonomicNameLabel id="EC48CBF1DD928BCB609783A0674EA88E" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="8FD348713FDC2080F227EBD9CA1BBA02" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" species="riggenbachi">C. riggenbachi</taxonomicName>
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|
. In the northern part of its distribution area it is replaced by
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<taxonomicName id="E411AD66F45EC31C98EA56DFDA864BA4" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" sensu="lato" species="splendopleure" status="s.l.">C. splendopleure</taxonomicName>
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<taxonomicNameLabel id="4FC211B0DA0FACD4A2E49F44CC9BF312" sensu="lato">s.l.</taxonomicNameLabel>
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|
probably at the Wouri, in the south it is currently not clear if it is in direct contact with
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|
<taxonomicName id="21742164CCC907767BFED68E4E363275" LSID-ZBK="urn:lsid:zoobank.org:act:494F6568-0395-45FD-B6C4-99E11F027244" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" lsidName-ZBK="Chromaphyosemion melanogaster Legros, Zentz & Agnèse 2005" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" species="melanogaster">C. melanogaster</taxonomicName>
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or
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<taxonomicName id="18BA37EC274B9B411A5DF6531A40ECBB" LSID-ZBK="urn:lsid:zoobank.org:act:58293C95-476D-42FC-B52D-62A8938702E3" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" lsidName-ZBK="Chromaphyosemion punctulatum Legros, Zentz & Agnèse 2005" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" species="punctulatum">C. punctulatum</taxonomicName>
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|
. Around Kribi the latter two species are known together with
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|
<taxonomicName id="E02B69E11FDE69DB13FECAA154F89267" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" species="loennbergii">C. loennbergii</taxonomicName>
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but the distribution of
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|
<taxonomicName id="8BECE13AE9E6928CCC9906187699F272" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" species="koungueense">C. koungueense</taxonomicName>
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for this area is unknown in detail.
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</paragraph>
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|
</subSubSection>
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|
<subSubSection id="DE64CCE60F22B64174EEE96355C60235" type="biology_ecology">
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<paragraph id="1DB4B1DA90F761B947B122C67CD603A6" pageNumber="11">
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|
The type locality is a small river in the forest, about 3 m wide and 0.4 m deep. It has a muddy bottom on sand, covered with decomposing leaves. The water temperature was 26°C, air temperature 28°C, pH 4.9 and conductivity ~10 µS/cm. Syntopic fish species were
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<taxonomicName id="7B496E682907991D06FAF265B4FE0795" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" species="infrafasciatus">Epiplatys infrafasciatus</taxonomicName>
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|
, 'Aphyosemion' ahli,
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|
<taxonomicName id="0342815CA27FA96B5570D0C59B4E5D79" LSID-ZBK="urn:lsid:zoobank.org:act:3A07CDFA-2136-40A4-8CC8-16EC4E5036D6" class="Actinopterygii" family="Poeciliidae" genus="Kribia" kingdom="Animalia" lsidName-ZBK="Kribia nana elongata Blache & Miton 1961:399" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" species="nana">Kribia nana</taxonomicName>
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, juvenile
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|
<taxonomicName id="0E011382106CC0C8C326A565F0091AAB" class="Actinopterygii" family="Cichlidae" genus="Benitochromis" kingdom="Animalia" order="Perciformes" pageNumber="11" phylum="Chordata" rank="species" species="finleyi">Benitochromis cf. finleyi</taxonomicName>
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and
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|
<taxonomicName id="1566AAC05A1DB260E9769573821208F8" LSID-ZBK="urn:lsid:zoobank.org:act:6C67C5A6-A29C-4FF3-8B0B-C22F586A4980" class="Actinopterygii" family="Poeciliidae" genus="Barboides" kingdom="Animalia" lsidName-ZBK="Barboides gracilis Brüning 1929:759" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" species="gracilis">Barboides gracilis</taxonomicName>
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.
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</paragraph>
|
|
</subSubSection>
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|
<subSubSection id="3B58708911F976BB6BC1FD09C6091529" type="discussion">
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|
<paragraph id="4DFEB1A870281A0E09C85E7EF4A82B3F" lastPageNumber="12" pageNumber="11">
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|
Remarks. This species is comparatively variable in color patterns compared to
|
|
<taxonomicName id="19DA8CF61A81BE074D26B3933AB84852" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" species="omega" status="sp. nov.">C. omega</taxonomicName>
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|
<taxonomicNameLabel id="B5F7FC44F3BAE510DF70AB5D06D11073" rank="species">sp. nov.</taxonomicNameLabel>
|
|
and
|
|
<taxonomicName id="77292C7B6F014BFDD5DB69DA91F7CD72" LSID-ZBK="urn:lsid:zoobank.org:act:58293C95-476D-42FC-B52D-62A8938702E3" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" lsidName-ZBK="Chromaphyosemion punctulatum Legros, Zentz & Agnèse 2005" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" species="punctulatum">C. punctulatum</taxonomicName>
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|
as closely related species. Despite this the isolated populations of Bioko and Bimbia (Figs. 7-8) are tentatively excluded here as there are probably significant differences in color patterns, but not as clear as in comparison with
|
|
<taxonomicName id="776359EC24D7D7367F809E3C0005FBA2" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" species="omega" status="sp. nov.">C. omega</taxonomicName>
|
|
<taxonomicNameLabel id="DE23A5D9A64C5854832D04EDB4EFC990" rank="species">sp. nov.</taxonomicNameLabel>
|
|
Additionally they are completely allopatric and separated from the distribution area of
|
|
<taxonomicName id="56EE9A2CCB88312C59CEAA40BBD22FA8" class="Actinopterygii" family="Nothobranchiidae" genus="Chromaphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="11" phylum="Chordata" rank="species" species="koungueense">C. koungueense</taxonomicName>
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|
by the Atlantic Ocean or the Wouri estuary respectively. Instead of the premature inclusion in the latter species, the taxonomic status of these populations should be reconsidered after detailed studies including, e.g., mate-choice experiments.
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|
</paragraph>
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|
</subSubSection>
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</treatment>
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</document> |