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<document id="4CF3D44D88612B42DC2EAF7540601EDB" ID-DOI="10.5252/geodiversitas2023v45a9" ID-ISSN="1638-9395" ID-Zenodo-Dep="8037926" ID-ZooBank="urn:lsid:zoobank.org:pub:1C430978-5EE6-49AE-AF7C-23C710161CB7" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1686745894455" checkinUser="felipe" docAuthor="Bronnert, Constance &amp; Métais, Grégoire" docDate="2023" docId="D066B24B521AB67EFF24F98DD8A3FE65" docLanguage="en" docName="Geodiversitas.45.9.277-326.pdf" docOrigin="Geodiversitas 45 (9)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Hallensia Franzen &amp; Haubold 1986" docType="treatment" docVersion="4" lastPageNumber="303" masterDocId="2C5FCA335206B662FFF8FF96DB4AFF94" masterDocTitle="Early Eocene hippomorph perissodactyls (Mammalia) from the Paris Basin" masterLastPageNumber="326" masterPageNumber="277" pageNumber="303" updateTime="1686841665420" updateUser="jonas" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="C2938BECF68248D8A9A8254B5A34F2DD">Early Eocene hippomorph perissodactyls (Mammalia) from the Paris Basin</mods:title>
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<mods:namePart id="C8021042BC21809E87E8C6A9C3FC475B">Bronnert, Constance</mods:namePart>
<mods:affiliation id="2C3EF95869C72A7C21AFECCA5783DDB8">Muséum national d’Histoire naturelle, Centre de Recherche en Paléontologie – Paris (CR 2 P, UMR 7207), CNRS / MNHN / Sorbonne Université, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:namePart id="4CB288AAC904B819357F97AABD1B4E9C">Métais, Grégoire</mods:namePart>
<mods:affiliation id="C6B9284B2748F06C6EC29CB766AE992E">Muséum national d’Histoire naturelle, Centre de Recherche en Paléontologie – Paris (CR 2 P, UMR 7207), CNRS / MNHN / Sorbonne Université, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:date id="DE383B9BAE11DCCBD695A750D1DC5492">2023</mods:date>
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Genre
<taxonomicName id="9FCF78DE521AB67EFEDDF98DD9FAF9A1" authority="Franzen &amp; Haubold, 1986" authorityName="Franzen &amp; Haubold" authorityYear="1986" box="[293,688,1563,1589]" class="Mammalia" family="Lophialetidae" genus="Hallensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="28" pageNumber="303" phylum="Chordata" rank="genus">
<emphasis id="6ABBDF4F521AB67EFEDDF98DDACDF9A1" box="[293,391,1563,1589]" italics="true" pageId="28" pageNumber="303">Hallensia</emphasis>
<bibRefCitation id="3C5E7EAC521AB67EFE76F98DD9FAF9A1" author="Franzen &amp; Haubold" box="[398,688,1563,1589]" firstAuthor="Franzen" pageId="28" pageNumber="303" pagination="159 - 170" refId="ref33472" refString="FRANZEN J. L. &amp; HAUBOLD H. 1986. - The middle Eocene of European mammalian stratigraphy. Definition of the Geiseltalian. Modern Geology 10 (2 - 3): 159 - 170." type="journal article" year="1986">Franzen &amp; Haubold, 1986</bibRefCitation>
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<taxonomicName id="9FCF78DE521AB67EFF7CF9C2D958F9F8" authority="Franzen &amp; Haubold, 1986: 37" authorityName="Franzen &amp; Haubold" authorityPageNumber="37" authorityYear="1986" box="[132,530,1620,1644]" class="Mammalia" family="Lophialetidae" genus="Hallensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="28" pageNumber="303" phylum="Chordata" rank="genus">
<emphasis id="6ABBDF4F521AB67EFF7CF9C2DB97F9F8" box="[132,221,1620,1644]" italics="true" pageId="28" pageNumber="303">Hallensia</emphasis>
<bibRefCitation id="3C5E7EAC521AB67EFF1BF9C2D958F9F8" author="Franzen &amp; Haubold" box="[227,530,1620,1644]" firstAuthor="Franzen" page="37" pageId="28" pageNumber="303" pagination="159 - 170" refId="ref33472" refString="FRANZEN J. L. &amp; HAUBOLD H. 1986. - The middle Eocene of European mammalian stratigraphy. Definition of the Geiseltalian. Modern Geology 10 (2 - 3): 159 - 170." type="journal article" year="1986">Franzen &amp; Haubold, 1986: 37</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<typeStatus id="8774BDFF521AB67EFF7CF912DBFFF90D" box="[132,181,1668,1691]" pageId="28" pageNumber="303">TYPE</typeStatus>
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. —
<taxonomicName id="9FCF78DE521AB67EFED7F912D9A3F908" authority="Franzen &amp; Haubold, 1986" authorityName="Franzen &amp; Haubold" authorityYear="1986" box="[303,745,1668,1692]" class="Mammalia" family="Lophialetidae" genus="Hallensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="28" pageNumber="303" phylum="Chordata" rank="species" species="matthesi">
<emphasis id="6ABBDF4F521AB67EFED7F912DA97F908" box="[303,477,1668,1692]" italics="true" pageId="28" pageNumber="303">Hallensia matthesi</emphasis>
<bibRefCitation id="3C5E7EAC521AB67EFE1BF912D9A3F908" author="Franzen &amp; Haubold" box="[483,745,1668,1692]" firstAuthor="Franzen" pageId="28" pageNumber="303" pagination="159 - 170" refId="ref33472" refString="FRANZEN J. L. &amp; HAUBOLD H. 1986. - The middle Eocene of European mammalian stratigraphy. Definition of the Geiseltalian. Modern Geology 10 (2 - 3): 159 - 170." type="journal article" year="1986">Franzen &amp; Haubold, 1986</bibRefCitation>
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by original designation.
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. —
<taxonomicName id="9FCF78DE521AB67EFEA1F958D9EDF972" authority=", Franzen, 1990" authorityName=", Franzen" authorityYear="1990" box="[345,679,1742,1766]" class="Mammalia" family="Lophialetidae" genus="Hallensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="28" pageNumber="303" phylum="Chordata" rank="species" species="parisiensis">
<emphasis id="6ABBDF4F521AB67EFEA1F958D95BF972" box="[345,529,1742,1766]" italics="true" pageId="28" pageNumber="303">Hallensia parisiensis</emphasis>
,
<bibRefCitation id="3C5E7EAC521AB67EFDE5F959D9EDF972" author="Franzen" box="[541,679,1742,1766]" firstAuthor="Franzen" pageId="28" pageNumber="303" pagination="175 - 201" refId="ref33404" refString="FRANZEN J. L. 1990. - Hallensia (Mammalia, Perissodactyla) aus Messel und dem Pariser Becken sowie Nachtrage aus dem Geiseltal. Bulletin de l'Institut royal des Sciences naturelles de Belgique, Sciences de la Terre 60: 175 - 201. http: // pascal-francis. inist. fr / vibad / index. php? action = getRecordDetail &amp; idt = 19633641" type="journal article" year="1990">Franzen, 1990</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="9FCF78DE521AB67EFD56F958DA05F895" authority=", Hooker, 1994" authorityName=", Hooker" authorityYear="1994" class="Mammalia" family="Lophialetidae" genus="Hallensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="28" pageNumber="303" phylum="Chordata" rank="species" species="louisi">
<emphasis id="6ABBDF4F521AB67EFD56F958DBFCF895" italics="true" pageId="28" pageNumber="303">Hallensia louisi</emphasis>
,
<bibRefCitation id="3C5E7EAC521AB67EFF3BF97FDA05F895" author="Hooker" box="[195,335,1769,1793]" firstAuthor="Hooker" pageId="28" pageNumber="303" pagination="29 - 63" refId="ref34268" refString="HOOKER J. J. 1994. - The beginning of the equoid radiation. Zoological Journal of the Linnean Society 112: 29 - 63. https: // doi. org / 10.1111 / j. 1096 - 3642.1994. tb 00311. x" type="journal article" year="1994">Hooker, 1994</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<subSubSection id="10D550D6521AB67EFF7CF88FDA53F87F" pageId="28" pageNumber="303" type="diagnosis">
<paragraph id="5870035D521AB67EFF7CF88FDA53F87F" blockId="28.[131,777,1817,2027]" pageId="28" pageNumber="303">
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<smallCapsWord id="5E969581521AB67EFF12F888DA05F8BB" baselines="1835" box="[234,335,1822,1839]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="diagnosis" pageId="28" pageNumber="303">DIAGNOSIS</smallCapsWord>
. — Perissodactyl of medium size (basicranium length:
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) with brachyodont and bunolophodont dentition. Dental formula 3.1.4.3/3.1.4.3. Nasal incisure extending at the posterior border of the I3 alveolus. Upper molars with or without mesostyle and without postprotocrista. Strong cingula on upper molars. M3 with strong hypocone. Upper and lower M3 larger than M2. Lower molars without paraconid. Hypoconulid of m3 relatively large.
</paragraph>
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<subSubSection id="10D550D6521AB67EFCD5FF41D8A3FE65" pageId="28" pageNumber="303" type="discussion">
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<taxonomicName id="9FCF78DE521AB67EFCD5FF61D8C4FE85" authorityName="Franzen &amp; Haubold" authorityYear="1986" box="[813,910,247,273]" class="Mammalia" family="Lophialetidae" genus="Hallensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="28" pageNumber="303" phylum="Chordata" rank="genus">
<emphasis id="6ABBDF4F521AB67EFCD5FF61D8C4FE85" box="[813,910,247,273]" italics="true" pageId="28" pageNumber="303">Hallensia</emphasis>
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is a perissodactyl with a very characteristic morphology. This primitive morphology pushed
<bibRefCitation id="3C5E7EAC521AB67EFB2DFE8ED83FFEC5" author="Franzen &amp; Haubold" firstAuthor="Franzen" pageId="28" pageNumber="303" pagination="159 - 170" refId="ref33472" refString="FRANZEN J. L. &amp; HAUBOLD H. 1986. - The middle Eocene of European mammalian stratigraphy. Definition of the Geiseltalian. Modern Geology 10 (2 - 3): 159 - 170." type="journal article" year="1986">Franzen &amp; Haubold (1986)</bibRefCitation>
to classify it within the phenacodontid condylarths. The discovery of a complete skeleton of
<taxonomicName id="9FCF78DE521AB67EFB23FECEDE1CFEE5" authorityName="Franzen &amp; Haubold" authorityYear="1986" box="[1243,1366,343,369]" class="Mammalia" family="Lophialetidae" genus="Hallensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="28" pageNumber="303" phylum="Chordata" rank="species" species="matthesi">
<emphasis id="6ABBDF4F521AB67EFB23FECEDE1CFEE5" box="[1243,1366,343,369]" italics="true" pageId="28" pageNumber="303">H. matthesi</emphasis>
</taxonomicName>
in Messel (
<collectingCountry id="20D843CD521AB67EFCA1FEE1D889FE05" box="[857,963,375,401]" name="Germany" pageId="28" pageNumber="303">Germany</collectingCountry>
) indicates that it is a perissodactyl (
<bibRefCitation id="3C5E7EAC521AB67EFAA0FEE1D82CFE25" author="Franzen" firstAuthor="Franzen" pageId="28" pageNumber="303" pagination="175 - 201" refId="ref33404" refString="FRANZEN J. L. 1990. - Hallensia (Mammalia, Perissodactyla) aus Messel und dem Pariser Becken sowie Nachtrage aus dem Geiseltal. Bulletin de l'Institut royal des Sciences naturelles de Belgique, Sciences de la Terre 60: 175 - 201. http: // pascal-francis. inist. fr / vibad / index. php? action = getRecordDetail &amp; idt = 19633641" type="journal article" year="1990">Franzen 1990</bibRefCitation>
), mostly because of the saddle-shaped navicular facet of its astragalus, a synapomorphic feature of perissodactyls (
<bibRefCitation id="3C5E7EAC521AB67EFCCEFE40D890FE64" author="Radinsky" box="[822,986,470,496]" firstAuthor="Radinsky" pageId="28" pageNumber="303" pagination="408 - 417" refId="ref35676" refString="RADINSKY L. B. 1966. - The adaptive radiation of the phenacodontid condylarths and the origin of the Perissodactyla. Evolution 20 (3): 408 - 417. https: // doi. org / 10.2307 / 2406639" type="journal article" year="1966">Radinsky 1966</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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