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<document id="91D4DAE84BE26ECD3AAC3159423BC781" ID-CLB-Dataset="54423" ID-DOI="10.5252/geodiversitas2023v45a9" ID-GBIF-Dataset="11c1dda3-18f9-443b-b0fe-ffe6c4a60242" ID-ISSN="1638-9395" ID-Zenodo-Dep="8037926" ID-ZooBank="urn:lsid:zoobank.org:pub:1C430978-5EE6-49AE-AF7C-23C710161CB7" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1686745894455" checkinUser="felipe" docAuthor="Bronnert, Constance &amp; Métais, Grégoire" docDate="2023" docId="D066B24B520FB669FEBFFA42D9D4FD84" docLanguage="en" docName="Geodiversitas.45.9.277-326.pdf" docOrigin="Geodiversitas 45 (9)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Pliolophus quesnoyensis Bronnert, Gheerbrant, Godinot &amp; Metais 2018" docType="treatment" docVersion="7" lastPageNumber="286" masterDocId="2C5FCA335206B662FFF8FF96DB4AFF94" masterDocTitle="Early Eocene hippomorph perissodactyls (Mammalia) from the Paris Basin" masterLastPageNumber="326" masterPageNumber="277" pageNumber="284" updateTime="1701998466266" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="140DAC4C85BED8E34FE1D47557A6C7E2">Early Eocene hippomorph perissodactyls (Mammalia) from the Paris Basin</mods:title>
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<mods:namePart id="3060B209F39DB4B6B19D747FB036A551">Bronnert, Constance</mods:namePart>
<mods:affiliation id="4D1C97D4375F55FE860A66DB7FEDF0E7">Muséum national d’Histoire naturelle, Centre de Recherche en Paléontologie – Paris (CR 2 P, UMR 7207), CNRS / MNHN / Sorbonne Université, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:namePart id="98DB3687BF9759859ED5DDB55C918A7E">Métais, Grégoire</mods:namePart>
<mods:affiliation id="F76C47E455366ED44A61C376DA61B014">Muséum national d’Histoire naturelle, Centre de Recherche en Paléontologie – Paris (CR 2 P, UMR 7207), CNRS / MNHN / Sorbonne Université, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
<mods:nameIdentifier id="0A2C61AAA4DB0FAB3C0AB10F71AE16CD" type="email">gregoire.metais@mnhn.fr</mods:nameIdentifier>
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<mods:title id="26E4CFB34B021DE333F97155EA1DF1BE">Geodiversitas</mods:title>
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<mods:date id="D3BC2D2E313B7B87DF11DD5A00104BC2">2023</mods:date>
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<treatment id="D066B24B520FB669FEBFFA42D9D4FD84" ID-DOI="http://doi.org/10.5281/zenodo.8040621" ID-GBIF-Taxon="209690554" ID-Zenodo-Dep="8040621" LSID="urn:lsid:plazi:treatment:D066B24B520FB669FEBFFA42D9D4FD84" httpUri="http://treatment.plazi.org/id/D066B24B520FB669FEBFFA42D9D4FD84" lastPageId="11" lastPageNumber="286" pageId="9" pageNumber="284">
<subSubSection id="10D550D6520FB66BFEBFFA42D981F999" pageId="9" pageNumber="284" type="nomenclature">
<paragraph id="5870035D520FB66BFEBFFA42D981F999" blockId="9.[193,715,1491,1582]" pageId="9" pageNumber="284">
<taxonomicName id="9FCF78DE520FB66BFEBFFA42D981F999" authority="Bronnert, Gheerbrant, Godinot &amp; Metais, 2018" authorityName="Bronnert, Gheerbrant, Godinot &amp; Metais" authorityYear="2018" class="Mammalia" family="Equidae" genus="Pliolophus" kingdom="Animalia" order="Perissodactyla" pageId="9" pageNumber="284" phylum="Chordata" rank="species" species="quesnoyensis">
<heading id="0338B431520FB66BFEBFFA42D90EFA79" box="[327,580,1491,1518]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="284" reason="2">
<emphasis id="6ABBDF4F520FB66BFEBFFA42D90EFA79" bold="true" box="[327,580,1491,1518]" italics="true" pageId="9" pageNumber="284">Pliolophus quesnoyensis</emphasis>
</heading>
<bibRefCitation id="3C5E7EAC520FB66BFF39FA65D981F999" author="Bronnert, Gheerbrant, Godinot &amp; Metais" box="[193,715,1523,1549]" firstAuthor="Bronnert" pageId="9" pageNumber="284" pagination="237 - 250" refId="ref32707" refString="BRONNERT C., GHEERBRANT E., GODINOT M. &amp; METAIS G. 2018. - A primitive perissodactyl (Mammalia) from the early Eocene of Le Quesnoy (Mp 7, France). Historical Biology 30 (1 - 2): 237 - 250. https: // doi. org / 10.1080 / 08912963.2017.1341502" type="journal article" year="2018">Bronnert, Gheerbrant, Godinot &amp; Métais, 2018</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="10D550D6520FB66BFE66F985DA0FF9EA" pageId="9" pageNumber="284" type="description">
<paragraph id="5870035D520FB66BFE66F985DAA7F9B9" blockId="9.[193,715,1491,1582]" box="[414,493,1555,1582]" pageId="9" pageNumber="284">
(
<figureCitation id="C0F41FD8520FB66BFE5FF985DAAEF9BA" box="[423,484,1555,1582]" captionStart="FIG" captionStartId="10.[132,143,1667,1684]" captionTargetBox="[133,1454,215,1626]" captionTargetId="figure-142@10.[699,1267,778,1344]" captionTargetPageId="10" captionText="FIG. 5. — Teeth of Pliolophus quesnoyensisBronnert,Gheerbrant,Godinot &amp; Métais,2018 from Le Quesnoy,Try,Meudon and Sinceny:A, left p2 (MNHN-QNY2-3076); B, left p3 (MNHN-TRY16); C, left p4 (MNHN-TRY22-L); D, right M1/2 (MNHN-M-TRY23 [reversed]); E, left M3 (MNHN-M-TRY8); F, left m1 (MNHN-M-TRY5 [reversed]); G, left m1/2 (MNHN-ME16322 [reversed]); H, right DP2 (MNHN-SY3-Cn [reversed]); I, right P3 (MNHN-SY1-Cn [reversed]); J, left P4 (MNHN-SY5-DE); K, left M1/2 (MNHN-SY4-DE); L, right M1/2 (MNHN-ME16325 [reversed]); M, right DP3 (MNHN-M.TRY10 [reversed]); N, right DP4 (MNHN-M.TRY3 [reversed]); O-P, left mandible with p4, m2-m3 (coll. Phélizon); A-O, occlusal views; P, mesial view; A-N, coated with magnesium; H-K, casts. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8037940" httpUri="https://zenodo.org/record/8037940/files/figure.png" pageId="9" pageNumber="284">Fig. 5</figureCitation>
)
</paragraph>
<paragraph id="5870035D520FB66BFF7CF9DADA0FF9EA" blockId="9.[132,776,1612,1662]" pageId="9" pageNumber="284">
<taxonomicName id="9FCF78DE520FB66BFF7CF9DADBA6F9EA" authority="Bronnert, Gheerbrant, Godinot &amp; Metais, 2018: 239" authorityName="Bronnert, Gheerbrant, Godinot &amp; Metais" authorityPageNumber="239" authorityYear="2018" class="Mammalia" family="Equidae" genus="Pliolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="9" pageNumber="284" phylum="Chordata" rank="species" species="quesnoyensis">
<emphasis id="6ABBDF4F520FB66BFF7CF9DADA17F9F7" box="[132,349,1612,1636]" italics="true" pageId="9" pageNumber="284">Pliolophus quesnoyensis</emphasis>
<bibRefCitation id="3C5E7EAC520FB66BFE9DF9DADBA6F9EA" author="Bronnert, Gheerbrant, Godinot &amp; Metais" firstAuthor="Bronnert" page="239" pageId="9" pageNumber="284" pagination="237 - 250" refId="ref32707" refString="BRONNERT C., GHEERBRANT E., GODINOT M. &amp; METAIS G. 2018. - A primitive perissodactyl (Mammalia) from the early Eocene of Le Quesnoy (Mp 7, France). Historical Biology 30 (1 - 2): 237 - 250. https: // doi. org / 10.1080 / 08912963.2017.1341502" type="journal article" year="2018">Bronnert, Gheerbrant, Godinot &amp; Métais, 2018: 239</bibRefCitation>
</taxonomicName>
, figs 3-7.
</paragraph>
</subSubSection>
<subSubSection id="10D550D6520FB66BFF7CF900DB91F8C3" pageId="9" pageNumber="284" type="reference_group">
<paragraph id="5870035D520FB66BFF7CF900D97DF93A" blockId="9.[132,567,1686,1710]" box="[132,567,1686,1710]" pageId="9" pageNumber="284">
<taxonomicName id="9FCF78DE520FB66BFF7CF900DA79F939" box="[132,307,1686,1710]" class="Mammalia" family="Equidae" genus="Hyracotherium" kingdom="Animalia" order="Perissodactyla" pageId="9" pageNumber="284" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="6ABBDF4F520FB66BFF7CF900DA5BF93A" box="[132,273,1686,1710]" italics="true" pageId="9" pageNumber="284">Hyracotherium</emphasis>
sp.
</taxonomicName>
–
<bibRefCitation id="3C5E7EAC520FB66BFEB4F900D979F93A" author="Russell" box="[332,563,1686,1710]" etAl="et al." firstAuthor="Russell" page="432" pageId="9" pageNumber="284" pagination="429 - 433" refId="ref35960" refString="RUSSELL D. E., GALOYER A., LOUIS P. &amp; GINGERICH P. D. 1988. - Nouveaux vertebres sparnaciens du conglomerat de Meudon a Meudon, France. Compte-Rendus de l'Academie des Sciences de Paris, serie II, 307: 429 - 433. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 6327064 k / f 443. item" type="journal article" year="1988">
Russell
<emphasis id="6ABBDF4F520FB66BFE6EF901DA8EF93A" box="[406,452,1686,1710]" italics="true" pageId="9" pageNumber="284">et al.</emphasis>
1988: 432
</bibRefCitation>
.
</paragraph>
<paragraph id="5870035D520FB66BFF7CF953D9D0F94A" blockId="9.[132,666,1733,1758]" box="[132,666,1733,1758]" pageId="9" pageNumber="284">
<taxonomicName id="9FCF78DE520FB66BFF7CF953DA29F949" authorityName="Owen" authorityYear="1858" box="[132,355,1733,1757]" class="Mammalia" family="Equidae" genus="Pliolophus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Perissodactyla" pageId="9" pageNumber="284" phylum="Chordata" rank="species" species="vulpiceps">
<emphasis id="6ABBDF4F520FB66BFF7CF953DBAEF949" box="[132,228,1733,1757]" italics="true" pageId="9" pageNumber="284">Pliolophus</emphasis>
aff.
<emphasis id="6ABBDF4F520FB66BFEE8F953DA29F949" box="[272,355,1733,1757]" italics="true" pageId="9" pageNumber="284">vulpiceps</emphasis>
</taxonomicName>
–
<bibRefCitation id="3C5E7EAC520FB66BFE85F953D901F949" author="Hooker" box="[381,587,1733,1758]" firstAuthor="Hooker" page="53 - 54" pageId="9" pageNumber="284" pagination="29 - 63" refId="ref34268" refString="HOOKER J. J. 1994. - The beginning of the equoid radiation. Zoological Journal of the Linnean Society 112: 29 - 63. https: // doi. org / 10.1111 / j. 1096 - 3642.1994. tb 00311. x" type="journal article" year="1994">Hooker 1994: 53-54</bibRefCitation>
, fig. 17.
</paragraph>
<paragraph id="5870035D520FB66BFF7CF963D9C3F899" blockId="9.[132,649,1781,1805]" box="[132,649,1781,1805]" pageId="9" pageNumber="284">
“
<taxonomicName id="9FCF78DE520FB66BFF77F963DA7CF899" authorityName="Lemoine" authorityYear="1891" box="[143,310,1781,1805]" class="Mammalia" family="Palaeotheriidae" genus="Propachynolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="9" pageNumber="284" phylum="Chordata" rank="genus">
<emphasis id="6ABBDF4F520FB66BFF77F963DA7CF899" box="[143,310,1781,1805]" italics="true" pageId="9" pageNumber="284">Propachynolophus</emphasis>
</taxonomicName>
” sp. 1 –
<bibRefCitation id="3C5E7EAC520FB66BFE75F963D970F899" author="Hooker" box="[397,570,1781,1805]" firstAuthor="Hooker" page="60" pageId="9" pageNumber="284" pagination="29 - 63" refId="ref34268" refString="HOOKER J. J. 1994. - The beginning of the equoid radiation. Zoological Journal of the Linnean Society 112: 29 - 63. https: // doi. org / 10.1111 / j. 1096 - 3642.1994. tb 00311. x" type="journal article" year="1994">Hooker 1994: 60</bibRefCitation>
, fig. 26.
</paragraph>
<paragraph id="5870035D520FB66BFF7CF8B3DB91F8C3" blockId="9.[132,775,1829,1879]" pageId="9" pageNumber="284">
<taxonomicName id="9FCF78DE520FB66BFF7CF8B3DA54F8A9" box="[132,286,1829,1853]" class="Mammalia" family="Equidae" genus="Pliolophus" kingdom="Animalia" order="Perissodactyla" pageId="9" pageNumber="284" phylum="Chordata" rank="species" species="undefined-1">
<emphasis id="6ABBDF4F520FB66BFF7CF8B3DBAFF8A9" box="[132,229,1829,1853]" italics="true" pageId="9" pageNumber="284">Pliolophus</emphasis>
sp. 1
</taxonomicName>
–
<bibRefCitation id="3C5E7EAC520FB66BFEC4F8B3DAA2F8A9" author="Hooker" box="[316,488,1829,1853]" firstAuthor="Hooker" page="60" pageId="9" pageNumber="284" pagination="29 - 63" refId="ref34268" refString="HOOKER J. J. 1994. - The beginning of the equoid radiation. Zoological Journal of the Linnean Society 112: 29 - 63. https: // doi. org / 10.1111 / j. 1096 - 3642.1994. tb 00311. x" type="journal article" year="1994">Hooker 1994: 60</bibRefCitation>
(
<emphasis id="6ABBDF4F520FB66BFE01F8B3D970F8A8" box="[505,570,1829,1852]" italics="true" pageId="9" pageNumber="284">partim</emphasis>
), fig. 26; 2010: 117 (
<emphasis id="6ABBDF4F520FB66BFF74F8D6DB86F8C3" box="[140,204,1856,1879]" italics="true" pageId="9" pageNumber="284">partim</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="10D550D6520FB66BFF7CF8F9DE07FDB9" pageId="9" pageNumber="284" type="materials_examined">
<paragraph id="5870035D520FB66BFF7CF8F9D84DF813" blockId="9.[132,775,1903,1927]" box="[132,775,1903,1927]" pageId="9" pageNumber="284">
<materialsCitation id="E8A70900520FB66BFF7CF8F9D849F813" ID-GBIF-Occurrence="4130693351" box="[132,771,1903,1927]" collectionCode="MNHN-QNY" country="France" location="Le Quesnoy" municipality="Le Quesnoy" pageId="9" pageNumber="284" specimenCode="MNHN-QNY2-2696" specimenCount="1" stateProvince="Oise" typeStatus="holotype">
<smallCapsWord id="5E969581520FB66BFF7CF8F9DBA6F811" baselines="1920,1921" box="[132,236,1903,1926]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Holotype" pageId="9" pageNumber="284">
<typeStatus id="8774BDFF520FB66BFF7CF8F9DBA6F811" box="[132,236,1903,1926]" pageId="9" pageNumber="284" type="holotype">HOLOTYPE</typeStatus>
</smallCapsWord>
. —
<specimenCode id="0869AB26520FB66BFEEDF8F9DAA5F813" box="[277,495,1903,1927]" collectionCode="MNHN-QNY" pageId="9" pageNumber="284">MNHN-QNY2-2696</specimenCode>
, right mandible with m1-m3
</materialsCitation>
.
</paragraph>
<paragraph id="5870035D520FB66BFF7CF809DACEF87F" blockId="9.[131,775,1950,2027]" pageId="9" pageNumber="284">
<materialsCitation id="E8A70900520FB66BFF7CF809DACAF87F" ID-GBIF-Occurrence="4130693335" collectionCode="MNHN-QNY" pageId="9" pageNumber="284" specimenCode="MNHN-QNY2-3022, MNHN-QNY2-2700, MNHN-QNY2-2705" specimenCount="1" typeStatus="paratype">
<smallCapsWord id="5E969581520FB66BFF7CF809DBA6F821" baselines="1968,1969" box="[132,236,1951,1974]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Paratypes" pageId="9" pageNumber="284">
<typeStatus id="8774BDFF520FB66BFF7CF809DBA6F821" box="[132,236,1951,1974]" pageId="9" pageNumber="284" type="paratype">PARATYPES</typeStatus>
</smallCapsWord>
. —
<specimenCode id="0869AB26520FB66BFEEEF809DAB8F823" box="[278,498,1951,1975]" collectionCode="MNHN-QNY" pageId="9" pageNumber="284">MNHN-QNY2-3022</specimenCode>
, right mandible, P1 missing;
<specimenCode id="0869AB26520FB66BFF7BF82FDA22F845" box="[131,360,1977,2001]" collectionCode="MNHN-QNY" pageId="9" pageNumber="284">MNHN-QNY2-2700</specimenCode>
, right maxillary with P4-M3;
<specimenCode id="0869AB26520FB66BFD5FF82FDA4FF87F" collectionCode="MNHN-QNY" pageId="9" pageNumber="284">MNHN-QNY2-2705</specimenCode>
, right M1/2
</materialsCitation>
.
</paragraph>
<paragraph id="5870035D520FB66BFCD5FF4ED8F0FE9E" blockId="9.[811,1457,215,557]" pageId="9" pageNumber="284">
<materialsCitation id="E8A70900520FB66BFCD5FF4ED8FCFE9E" ID-GBIF-Occurrence="4130693314" pageId="9" pageNumber="284" specimenCount="1">
<smallCapsWord id="5E969581520FB66BFCD5FF4ED8C7FF7A" baselines="233,234" box="[813,909,216,239]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Material" pageId="9" pageNumber="284">MATERIAL</smallCapsWord>
. —
<emphasis id="6ABBDF4F520FB66BFC4BFF4EDF6CFF65" bold="true" box="[947,1062,216,241]" pageId="9" pageNumber="284">Le Quesnoy</emphasis>
: see
<bibRefCitation id="3C5E7EAC520FB66BFBA8FF4EDE46FF64" author="Bronnert" box="[1104,1292,215,240]" etAl="et al." firstAuthor="Bronnert" pageId="9" pageNumber="284" pagination="237 - 250" refId="ref32707" refString="BRONNERT C., GHEERBRANT E., GODINOT M. &amp; METAIS G. 2018. - A primitive perissodactyl (Mammalia) from the early Eocene of Le Quesnoy (Mp 7, France). Historical Biology 30 (1 - 2): 237 - 250. https: // doi. org / 10.1080 / 08912963.2017.1341502" type="journal article" year="2018">
Bronnert
<emphasis id="6ABBDF4F520FB66BFB54FF4EDF92FF7B" box="[1196,1240,215,239]" italics="true" pageId="9" pageNumber="284">et al.</emphasis>
2018
</bibRefCitation>
; p2 (R: MNHN-QNY2-3076)
</materialsCitation>
.
</paragraph>
<paragraph id="5870035D520FB66BFCD5FE9BD8EFFEE0" blockId="9.[811,1457,215,557]" pageId="9" pageNumber="284">
<materialsCitation id="E8A70900520FB66BFCD5FE9BD8EBFEE0" ID-GBIF-Occurrence="4130693331" pageId="9" pageNumber="284" specimenCount="1">
<emphasis id="6ABBDF4F520FB66BFCD5FE9BD8C8FEB1" bold="true" box="[813,898,269,293]" pageId="9" pageNumber="284">Meudon</emphasis>
: mandible with p4-m3 (L: coll. Phélizon); P3 (R: unnumbered [Marseille]); M1/2 (R: MNHN-ME-16325, unnumbered [Marseille]); m1/2 (L: MNHN-ME-16322); m3 (R: unnumbered [Marseille])
</materialsCitation>
.
</paragraph>
<paragraph id="5870035D520FB66BFCD5FEEEDE07FDB9" blockId="9.[811,1457,215,557]" pageId="9" pageNumber="284">
<materialsCitation id="E8A70900520FB66BFCD5FEEEDE03FDB9" ID-GBIF-Occurrence="4130693302" pageId="9" pageNumber="284" specimenCount="1">
<emphasis id="6ABBDF4F520FB66BFCD5FEEED832FE04" bold="true" box="[813,888,376,400]" pageId="9" pageNumber="284">Sinceny</emphasis>
: P2 (R: MNHN-SY3-CN); P3 (R: MNHN-SY1-CN; L: unnumbered MNHN coll. Cahuzac); P4 (L: MNHN-SY5-DE); M1/2 (L: MNHN-SY4-DE); M3 (R: unnumbered MNHN coll. Berton).
<emphasis id="6ABBDF4F520FB66BFCD5FE50D81AFE4A" bold="true" box="[813,848,454,478]" pageId="9" pageNumber="284">Try</emphasis>
: DP3 (R: MNHN-M-TRY10); DP4 (R: MNHN-M-TRY3); M1/2 (R: MNHN-M-TRY23, TRY17-L; L: MNHN-M-TRY9, M-TRY11); M3 (L: MNHN-M-TRY8); p3 (R: MNHN-M-TRY16); p4 (R: MNHN-TRY22-L); m1 (L: MNHN-M-TRY5)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="10D550D6520FB66BFCD5FDD3DE1DFD32" pageId="9" pageNumber="284" type="distribution">
<paragraph id="5870035D520FB66BFCD5FDD3DE72FDC8" blockId="9.[813,1336,581,605]" box="[813,1336,581,605]" pageId="9" pageNumber="284">
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<typeStatus id="8774BDFF520FB66BFCD5FDD3D817FDCE" box="[813,861,581,604]" pageId="9" pageNumber="284">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="5E969581520FB66BFC9CFDDCD8F5FDCF" baselines="599" box="[868,959,586,603]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="9" pageNumber="284">LOCALITY</smallCapsWord>
. — Le Quesnoy (Oise,
<collectingCountry id="20D843CD520FB66BFB51FDD3DFA4FDC9" box="[1193,1262,581,605]" name="France" pageId="9" pageNumber="284">France</collectingCountry>
), MP7.
</paragraph>
<paragraph id="5870035D520FB66BFCD5FDE3DE1DFD32" blockId="9.[811,1455,628,679]" pageId="9" pageNumber="284">
<smallCapsWord id="5E969581520FB66BFCD5FDE3D8FCFD1F" baselines="646,647" box="[813,950,629,652]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="9" pageNumber="284">DISTRIBUTION</smallCapsWord>
. — Le Quesnoy (Oise,
<collectingCountry id="20D843CD520FB66BFB68FDE2DF99FD18" box="[1168,1235,628,652]" name="France" pageId="9" pageNumber="284">France</collectingCountry>
), Try (Marne), Meudon (Hauts-de-Seine,
<collectingCountry id="20D843CD520FB66BFC2EFD19DF51FD33" box="[982,1051,655,679]" name="France" pageId="9" pageNumber="284">France</collectingCountry>
), Sinceny (Aisne,
<collectingCountry id="20D843CD520FB66BFB31FD19DE44FD33" box="[1225,1294,655,679]" name="France" pageId="9" pageNumber="284">France</collectingCountry>
), MP7.
</paragraph>
</subSubSection>
<subSubSection id="10D550D6520FB66BFCD5FD29DE0AFC2B" pageId="9" pageNumber="284" type="diagnosis">
<paragraph id="5870035D520FB66BFCD5FD29DEE5FCD4" blockId="9.[813,1456,702,832]" pageId="9" pageNumber="284">
<smallCapsWord id="5E969581520FB66BFCD5FD29D8DDFD41" baselines="720,721" box="[813,919,703,726]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="9" pageNumber="284">DIAGNOSIS</smallCapsWord>
(
<bibRefCitation id="3C5E7EAC520FB66BFC5FFD29DF3AFD42" author="Bronnert" box="[935,1136,702,726]" etAl="et al." firstAuthor="Bronnert" pageId="9" pageNumber="284" pagination="237 - 250" refId="ref32707" refString="BRONNERT C., GHEERBRANT E., GODINOT M. &amp; METAIS G. 2018. - A primitive perissodactyl (Mammalia) from the early Eocene of Le Quesnoy (Mp 7, France). Historical Biology 30 (1 - 2): 237 - 250. https: // doi. org / 10.1080 / 08912963.2017.1341502" type="journal article" year="2018">
Bronnert
<emphasis id="6ABBDF4F520FB66BFBFFFD29DF7FFD42" box="[1031,1077,702,726]" italics="true" pageId="9" pageNumber="284">et al.</emphasis>
2018
</bibRefCitation>
). — Small brachyodont perissodactyl; post-p1 diastema extremely short, almost absent; complete cheek tooth formula; sub-lophodont molars; paraconule close to protocone on P3; protocone on P2; straight centrocrista; enlarged crenulated cingulum on M3; hypoconulid lobe on m3 developed.
</paragraph>
<paragraph id="5870035D520FB66BFCD5FCCEDE0AFC2B" blockId="9.[811,1455,856,959]" pageId="9" pageNumber="284">
<smallCapsWord id="5E969581520FB66BFCD5FCCED8FFFCFA" baselines="873,874" box="[813,949,856,879]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Differential" pageId="9" pageNumber="284">DIFFERENTIAL</smallCapsWord>
<smallCapsWord id="5E969581520FB66BFC42FCCBDF6AFCFA" baselines="874" box="[954,1056,861,878]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="diagnosis" pageId="9" pageNumber="284">DIAGNOSIS</smallCapsWord>
(
<bibRefCitation id="3C5E7EAC520FB66BFBD5FCCEDFBAFCE4" author="Bronnert" box="[1069,1264,856,880]" etAl="et al." firstAuthor="Bronnert" pageId="9" pageNumber="284" pagination="237 - 250" refId="ref32707" refString="BRONNERT C., GHEERBRANT E., GODINOT M. &amp; METAIS G. 2018. - A primitive perissodactyl (Mammalia) from the early Eocene of Le Quesnoy (Mp 7, France). Historical Biology 30 (1 - 2): 237 - 250. https: // doi. org / 10.1080 / 08912963.2017.1341502" type="journal article" year="2018">
Bronnert
<emphasis id="6ABBDF4F520FB66BFB73FCCFDFFDFCE4" box="[1163,1207,856,880]" italics="true" pageId="9" pageNumber="284">et al.</emphasis>
2018
</bibRefCitation>
). — Species smaller than
<taxonomicName id="9FCF78DE520FB66BFCA5FCE5D889FC1E" authorityName="Owen" authorityYear="1858" box="[861,963,882,906]" class="Mammalia" family="Equidae" genus="Pliolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="9" pageNumber="284" phylum="Chordata" rank="species" species="vulpiceps">
<emphasis id="6ABBDF4F520FB66BFCA5FCE5D889FC1E" box="[861,963,882,906]" italics="true" pageId="9" pageNumber="284">P. vulpiceps</emphasis>
</taxonomicName>
and
<taxonomicName id="9FCF78DE520FB66BFC16FCE5DF0EFC1E" authorityName="Hooker" authorityYear="2010" box="[1006,1092,882,906]" class="Mammalia" family="Equidae" genus="Pliolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="9" pageNumber="284" phylum="Chordata" rank="species" species="barnesi">
<emphasis id="6ABBDF4F520FB66BFC16FCE5DF0EFC1E" box="[1006,1092,882,906]" italics="true" pageId="9" pageNumber="284">P. barnesi</emphasis>
</taxonomicName>
. Differs from the latters by a straighter centrocrista and a smaller metaconule on upper molars. Further differs from
<taxonomicName id="9FCF78DE520FB66BFC73FC3ED8BEFC2B" authorityName="Owen" authorityYear="1858" box="[907,1012,935,959]" class="Mammalia" family="Equidae" genus="Pliolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="9" pageNumber="284" phylum="Chordata" rank="species" species="vulpiceps">
<emphasis id="6ABBDF4F520FB66BFC73FC3ED8BEFC2B" box="[907,1012,935,959]" italics="true" pageId="9" pageNumber="284">P. vulpiceps</emphasis>
</taxonomicName>
by a shorter postcanine diastema.
</paragraph>
</subSubSection>
<subSubSection id="10D550D6520FB669FCD5FC63D9D4FD84" lastPageId="11" lastPageNumber="286" pageId="9" pageNumber="284" type="description">
<paragraph id="5870035D520FB66BFCD5FC63D8F7FB99" blockId="9.[813,1455,1013,1103]" box="[813,957,1013,1039]" pageId="9" pageNumber="284">
<smallCapsWord id="5E969581520FB66BFCD5FC63D8F7FB99" baselines="1032,1032" box="[813,957,1013,1039]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="9" pageNumber="284">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="5870035D520FB66BFCD5FB82DE47FBDA" blockId="9.[813,1455,1013,1103]" pageId="9" pageNumber="284">
See
<bibRefCitation id="3C5E7EAC520FB66BFCA0FB83DF0CFBBA" author="Bronnert" box="[856,1094,1044,1071]" etAl="et al." firstAuthor="Bronnert" pageId="9" pageNumber="284" pagination="237 - 250" refId="ref32707" refString="BRONNERT C., GHEERBRANT E., GODINOT M. &amp; METAIS G. 2018. - A primitive perissodactyl (Mammalia) from the early Eocene of Le Quesnoy (Mp 7, France). Historical Biology 30 (1 - 2): 237 - 250. https: // doi. org / 10.1080 / 08912963.2017.1341502" type="journal article" year="2018">
Bronnert
<emphasis id="6ABBDF4F520FB66BFC39FB83D8BFFBBA" box="[961,1013,1044,1070]" italics="true" pageId="9" pageNumber="284">et al.</emphasis>
(2018)
</bibRefCitation>
for a complete description of the species. Here is only described new material.
</paragraph>
<paragraph id="5870035D520FB66BFCD5FBE2D82EFB1A" blockId="9.[813,868,1140,1166]" box="[813,868,1140,1166]" pageId="9" pageNumber="284">
<heading id="0338B431520FB66BFCD5FBE2D82EFB1A" box="[813,868,1140,1166]" fontSize="11" level="2" pageId="9" pageNumber="284" reason="8">
<emphasis id="6ABBDF4F520FB66BFCD5FBE2D82EFB1A" box="[813,868,1140,1166]" italics="true" pageId="9" pageNumber="284">Teeth</emphasis>
</heading>
</paragraph>
<paragraph id="5870035D520FB668FCD5FB02DF12F87F" blockId="9.[811,1457,1172,2027]" lastBlockId="10.[811,1456,1841,2028]" lastPageId="10" lastPageNumber="285" pageId="9" pageNumber="284">
The P3 is nonmolariform (
<figureCitation id="C0F41FD8520FB66BFBA2FB02DFEDFB3A" box="[1114,1191,1172,1199]" captionStart="FIG" captionStartId="10.[132,143,1667,1684]" captionTargetBox="[133,1454,215,1626]" captionTargetId="figure-142@10.[699,1267,778,1344]" captionTargetPageId="10" captionText="FIG. 5. — Teeth of Pliolophus quesnoyensisBronnert,Gheerbrant,Godinot &amp; Métais,2018 from Le Quesnoy,Try,Meudon and Sinceny:A, left p2 (MNHN-QNY2-3076); B, left p3 (MNHN-TRY16); C, left p4 (MNHN-TRY22-L); D, right M1/2 (MNHN-M-TRY23 [reversed]); E, left M3 (MNHN-M-TRY8); F, left m1 (MNHN-M-TRY5 [reversed]); G, left m1/2 (MNHN-ME16322 [reversed]); H, right DP2 (MNHN-SY3-Cn [reversed]); I, right P3 (MNHN-SY1-Cn [reversed]); J, left P4 (MNHN-SY5-DE); K, left M1/2 (MNHN-SY4-DE); L, right M1/2 (MNHN-ME16325 [reversed]); M, right DP3 (MNHN-M.TRY10 [reversed]); N, right DP4 (MNHN-M.TRY3 [reversed]); O-P, left mandible with p4, m2-m3 (coll. Phélizon); A-O, occlusal views; P, mesial view; A-N, coated with magnesium; H-K, casts. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8037940" httpUri="https://zenodo.org/record/8037940/files/figure.png" pageId="9" pageNumber="284">Fig. 5I</figureCitation>
). The paracone and the metacone are close, and linked by a short centrocrista. A small paraconule is present. The postprotocrista is absent. The P4 is nonmolariform and more symmetrical than the P3.The cingulum is interrupted close to the protocone. The parastyle is small and crescentiform. The paraconule is developed and anterior to the paracone. The postprotocrista extends posteriolabially to the metacone. An endoprotocrista is present posterior to the postprotocrista. There is no mesostyle on upper molars (
<figureCitation id="C0F41FD8520FB66BFBDFFA25DFD6FA59" box="[1063,1180,1459,1486]" captionStart="FIG" captionStartId="10.[132,143,1667,1684]" captionTargetBox="[133,1454,215,1626]" captionTargetId="figure-142@10.[699,1267,778,1344]" captionTargetPageId="10" captionText="FIG. 5. — Teeth of Pliolophus quesnoyensisBronnert,Gheerbrant,Godinot &amp; Métais,2018 from Le Quesnoy,Try,Meudon and Sinceny:A, left p2 (MNHN-QNY2-3076); B, left p3 (MNHN-TRY16); C, left p4 (MNHN-TRY22-L); D, right M1/2 (MNHN-M-TRY23 [reversed]); E, left M3 (MNHN-M-TRY8); F, left m1 (MNHN-M-TRY5 [reversed]); G, left m1/2 (MNHN-ME16322 [reversed]); H, right DP2 (MNHN-SY3-Cn [reversed]); I, right P3 (MNHN-SY1-Cn [reversed]); J, left P4 (MNHN-SY5-DE); K, left M1/2 (MNHN-SY4-DE); L, right M1/2 (MNHN-ME16325 [reversed]); M, right DP3 (MNHN-M.TRY10 [reversed]); N, right DP4 (MNHN-M.TRY3 [reversed]); O-P, left mandible with p4, m2-m3 (coll. Phélizon); A-O, occlusal views; P, mesial view; A-N, coated with magnesium; H-K, casts. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8037940" httpUri="https://zenodo.org/record/8037940/files/figure.png" pageId="9" pageNumber="284">Fig. 5D, E</figureCitation>
, K-L). The centrocrista is straight. The paraconule is smaller than the protocone. The protoloph is slightly notched between the protocone and the paraconule. The metaconule is weak or absent. The metaloph ends by a metacone fold. The preparaconule crista joins the preparacrista. The ectocingulum is continuous. An accessory cusp is sometimes present on the cingulum between the metacone and the hypocone. The mandible is very worn and broken. The teeth are so worn that no enamel is present in occlusal view. The p2 has a single large cusp (
<figureCitation id="C0F41FD8520FB66BFAD9F944DE3CF978" box="[1313,1398,1746,1773]" captionStart="FIG" captionStartId="10.[132,143,1667,1684]" captionTargetBox="[133,1454,215,1626]" captionTargetId="figure-142@10.[699,1267,778,1344]" captionTargetPageId="10" captionText="FIG. 5. — Teeth of Pliolophus quesnoyensisBronnert,Gheerbrant,Godinot &amp; Métais,2018 from Le Quesnoy,Try,Meudon and Sinceny:A, left p2 (MNHN-QNY2-3076); B, left p3 (MNHN-TRY16); C, left p4 (MNHN-TRY22-L); D, right M1/2 (MNHN-M-TRY23 [reversed]); E, left M3 (MNHN-M-TRY8); F, left m1 (MNHN-M-TRY5 [reversed]); G, left m1/2 (MNHN-ME16322 [reversed]); H, right DP2 (MNHN-SY3-Cn [reversed]); I, right P3 (MNHN-SY1-Cn [reversed]); J, left P4 (MNHN-SY5-DE); K, left M1/2 (MNHN-SY4-DE); L, right M1/2 (MNHN-ME16325 [reversed]); M, right DP3 (MNHN-M.TRY10 [reversed]); N, right DP4 (MNHN-M.TRY3 [reversed]); O-P, left mandible with p4, m2-m3 (coll. Phélizon); A-O, occlusal views; P, mesial view; A-N, coated with magnesium; H-K, casts. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8037940" httpUri="https://zenodo.org/record/8037940/files/figure.png" pageId="9" pageNumber="284">Fig. 5A</figureCitation>
). On some unworn upper molars, denticules are present on the metaloph (
<figureCitation id="C0F41FD8520FB66BFC5CF884D8BFF8B8" box="[932,1013,1809,1836]" captionStart="FIG" captionStartId="10.[132,143,1667,1684]" captionTargetBox="[133,1454,215,1626]" captionTargetId="figure-142@10.[699,1267,778,1344]" captionTargetPageId="10" captionText="FIG. 5. — Teeth of Pliolophus quesnoyensisBronnert,Gheerbrant,Godinot &amp; Métais,2018 from Le Quesnoy,Try,Meudon and Sinceny:A, left p2 (MNHN-QNY2-3076); B, left p3 (MNHN-TRY16); C, left p4 (MNHN-TRY22-L); D, right M1/2 (MNHN-M-TRY23 [reversed]); E, left M3 (MNHN-M-TRY8); F, left m1 (MNHN-M-TRY5 [reversed]); G, left m1/2 (MNHN-ME16322 [reversed]); H, right DP2 (MNHN-SY3-Cn [reversed]); I, right P3 (MNHN-SY1-Cn [reversed]); J, left P4 (MNHN-SY5-DE); K, left M1/2 (MNHN-SY4-DE); L, right M1/2 (MNHN-ME16325 [reversed]); M, right DP3 (MNHN-M.TRY10 [reversed]); N, right DP4 (MNHN-M.TRY3 [reversed]); O-P, left mandible with p4, m2-m3 (coll. Phélizon); A-O, occlusal views; P, mesial view; A-N, coated with magnesium; H-K, casts. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8037940" httpUri="https://zenodo.org/record/8037940/files/figure.png" pageId="9" pageNumber="284">Fig. 5E</figureCitation>
), and we interpreted that as intraspecific variation. A small paraconid is present. Three crest extends posterior to the main cusp. A small hypoconid is present. The p3 display a protoconid larger than the metaconid. They are closely appressed. A cristid obliqua links the protolophid to the hypoconid. The paraconid is absent. The p4 is submolariform. The protoconid is larger than the metaconid, and they are closely appressed. The hypoconid is large and linked to the protolophid by the cristid obliqua. The m1 displays a cristid obliqua oriented slightly labially. The hypolophid is formed by two crests. A transversal crest linked to the entoconid extends toward the hypoconid and this crest suddenly becomes directed toward the hypoconulid, thus forminga right angle (
<figureCitation id="C0F41FD8520CB668FC8BF8A7D88AF8D8" box="[883,960,1841,1868]" captionStart="FIG" captionStartId="10.[132,143,1667,1684]" captionTargetBox="[133,1454,215,1626]" captionTargetId="figure-142@10.[699,1267,778,1344]" captionTargetPageId="10" captionText="FIG. 5. — Teeth of Pliolophus quesnoyensisBronnert,Gheerbrant,Godinot &amp; Métais,2018 from Le Quesnoy,Try,Meudon and Sinceny:A, left p2 (MNHN-QNY2-3076); B, left p3 (MNHN-TRY16); C, left p4 (MNHN-TRY22-L); D, right M1/2 (MNHN-M-TRY23 [reversed]); E, left M3 (MNHN-M-TRY8); F, left m1 (MNHN-M-TRY5 [reversed]); G, left m1/2 (MNHN-ME16322 [reversed]); H, right DP2 (MNHN-SY3-Cn [reversed]); I, right P3 (MNHN-SY1-Cn [reversed]); J, left P4 (MNHN-SY5-DE); K, left M1/2 (MNHN-SY4-DE); L, right M1/2 (MNHN-ME16325 [reversed]); M, right DP3 (MNHN-M.TRY10 [reversed]); N, right DP4 (MNHN-M.TRY3 [reversed]); O-P, left mandible with p4, m2-m3 (coll. Phélizon); A-O, occlusal views; P, mesial view; A-N, coated with magnesium; H-K, casts. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8037940" httpUri="https://zenodo.org/record/8037940/files/figure.png" pageId="10" pageNumber="285">Fig. 5F</figureCitation>
). A very faint crest extends from the hypoconid toward the transversal crest starting from the entoconid and reaches the point where the latter forms a right angle. This configuration of the hypolophid is unusual in
<taxonomicName id="9FCF78DE520CB668FAE0F804DEE5F83F" authorityName="Bronnert, Gheerbrant, Godinot &amp; Metais" authorityYear="2018" box="[1304,1455,1938,1963]" class="Mammalia" family="Equidae" genus="Pliolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="10" pageNumber="285" phylum="Chordata" rank="species" species="quesnoyensis">
<emphasis id="6ABBDF4F520CB668FAE0F804DEE5F83F" box="[1304,1455,1938,1963]" italics="true" pageId="10" pageNumber="285">P. quesnoyensis</emphasis>
</taxonomicName>
in which the hypolophid is generally formed by a transversal crest joining the hypoconid
</paragraph>
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. 5. — Teeth of
<taxonomicName id="9FCF78DE520CB668FEEDF915D804F900" authority="Bronnert, Gheerbrant, Godinot &amp; Metais, 2018" authorityName="Bronnert, Gheerbrant, Godinot &amp; Metais" authorityYear="2018" box="[277,846,1667,1684]" class="Mammalia" family="Equidae" genus="Pliolophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="10" pageNumber="285" phylum="Chordata" rank="species" species="quesnoyensis">
<emphasis id="6ABBDF4F520CB668FEEDF915DA91F900" box="[277,475,1667,1684]" italics="true" pageId="10" pageNumber="285">Pliolophus quesnoyensis</emphasis>
<bibRefCitation id="3C5E7EAC520CB668FE25F915D804F900" author="Bronnert, Gheerbrant, Godinot &amp; Metais" box="[477,846,1667,1684]" firstAuthor="Bronnert" pageId="10" pageNumber="285" pagination="237 - 250" refId="ref32707" refString="BRONNERT C., GHEERBRANT E., GODINOT M. &amp; METAIS G. 2018. - A primitive perissodactyl (Mammalia) from the early Eocene of Le Quesnoy (Mp 7, France). Historical Biology 30 (1 - 2): 237 - 250. https: // doi. org / 10.1080 / 08912963.2017.1341502" type="journal article" year="2018">Bronnert,Gheerbrant,Godinot &amp; Métais,2018</bibRefCitation>
</taxonomicName>
from Le Quesnoy,Try,Meudon and Sinceny:
<emphasis id="6ABBDF4F520CB668FB40F915DF8FF900" bold="true" box="[1208,1221,1667,1684]" pageId="10" pageNumber="285">A</emphasis>
, left p2 (MNHN-QNY2-3076);
<emphasis id="6ABBDF4F520CB668FF7CF90CDBDBF93F" bold="true" box="[132,145,1690,1707]" pageId="10" pageNumber="285">B</emphasis>
, left p3 (MNHN-TRY16);
<emphasis id="6ABBDF4F520CB668FE9AF90CDA3AF93F" bold="true" box="[354,368,1690,1707]" pageId="10" pageNumber="285">C</emphasis>
, left p4 (MNHN-TRY22-L);
<emphasis id="6ABBDF4F520CB668FDAAF90CD92AF93F" bold="true" box="[594,608,1690,1707]" pageId="10" pageNumber="285">D</emphasis>
, right M1/2 (MNHN-M-TRY23 [reversed]);
<emphasis id="6ABBDF4F520CB668FC38F90CD886F93F" bold="true" box="[960,972,1690,1707]" pageId="10" pageNumber="285">E</emphasis>
, left M3 (MNHN-M-TRY8);
<emphasis id="6ABBDF4F520CB668FB57F90CDFF0F93F" bold="true" box="[1199,1210,1690,1707]" pageId="10" pageNumber="285">F</emphasis>
, left m1 (MNHN-M-TRY5 [reversed]);
<emphasis id="6ABBDF4F520CB668FF28F927DB94F956" bold="true" box="[208,222,1713,1730]" pageId="10" pageNumber="285">G</emphasis>
, left m1/2 (MNHN-ME16322 [reversed]);
<emphasis id="6ABBDF4F520CB668FDCBF927D90BF956" bold="true" box="[563,577,1713,1730]" pageId="10" pageNumber="285">H</emphasis>
, right DP2 (MNHN-SY3-Cn [reversed]);
<emphasis id="6ABBDF4F520CB668FC73F927D8DBF956" bold="true" box="[907,913,1713,1730]" pageId="10" pageNumber="285">I</emphasis>
, right P3 (MNHN-SY1-Cn [reversed]);
<emphasis id="6ABBDF4F520CB668FB37F927DF93F956" bold="true" box="[1231,1241,1713,1730]" pageId="10" pageNumber="285">J</emphasis>
, left P4 (MNHN-SY5-DE);
<emphasis id="6ABBDF4F520CB668FF7CF95EDBDBF94D" bold="true" box="[132,145,1736,1753]" pageId="10" pageNumber="285">K</emphasis>
, left M1/2 (MNHN-SY4-DE);
<emphasis id="6ABBDF4F520CB668FE7AF95EDAC7F94D" bold="true" box="[386,397,1736,1753]" pageId="10" pageNumber="285">L</emphasis>
, right M1/2 (MNHN-ME16325 [reversed]);
<emphasis id="6ABBDF4F520CB668FD0AF95ED849F94D" bold="true" box="[754,771,1736,1753]" pageId="10" pageNumber="285">M</emphasis>
, right DP3 (MNHN-M.TRY10 [reversed]);
<emphasis id="6ABBDF4F520CB668FBA2F95EDF22F94D" bold="true" box="[1114,1128,1736,1753]" pageId="10" pageNumber="285">N</emphasis>
, right DP4 (MNHN-M.TRY3 [reversed]);
<emphasis id="6ABBDF4F520CB668FF7CF948DBECF97B" bold="true" box="[132,166,1758,1775]" pageId="10" pageNumber="285">O -P</emphasis>
, left mandible with p4, m2-m3 (coll. Phélizon);
<emphasis id="6ABBDF4F520CB668FDD3F948D904F97B" bold="true" box="[555,590,1758,1775]" pageId="10" pageNumber="285">A -O</emphasis>
, occlusal views;
<emphasis id="6ABBDF4F520CB668FD23F948D9ADF97B" bold="true" box="[731,743,1758,1775]" pageId="10" pageNumber="285">P</emphasis>
, mesial view;
<emphasis id="6ABBDF4F520CB668FCA5F948D835F97B" bold="true" box="[861,895,1758,1775]" pageId="10" pageNumber="285">A -N</emphasis>
, coated with magnesium;
<emphasis id="6ABBDF4F520CB668FBA3F948DF37F97B" bold="true" box="[1115,1149,1758,1775]" pageId="10" pageNumber="285">H -K</emphasis>
, casts. Scale bars: 5 mm.
</paragraph>
</caption>
<paragraph id="5870035D520DB669FF7CFF41DA6DFF65" blockId="11.[132,295,215,241]" box="[132,295,215,241]" pageId="11" pageNumber="286">
<heading id="0338B431520DB669FF7CFF41DA6DFF65" box="[132,295,215,241]" fontSize="11" level="2" pageId="11" pageNumber="286" reason="8">
<emphasis id="6ABBDF4F520DB669FF7CFF41DA6DFF65" box="[132,295,215,241]" italics="true" pageId="11" pageNumber="286">Deciduous teeth</emphasis>
</heading>
</paragraph>
<paragraph id="5870035D520DB669FF7CFF61D9D4FD84" blockId="11.[130,777,247,528]" pageId="11" pageNumber="286">
The DP2possesses a large paracone,much larger than the metacone (
<figureCitation id="C0F41FD8520DB669FF75FE81DB94FEA5" box="[141,222,279,306]" captionStart="FIG" captionStartId="10.[132,143,1667,1684]" captionTargetBox="[133,1454,215,1626]" captionTargetId="figure-142@10.[699,1267,778,1344]" captionTargetPageId="10" captionText="FIG. 5. — Teeth of Pliolophus quesnoyensisBronnert,Gheerbrant,Godinot &amp; Métais,2018 from Le Quesnoy,Try,Meudon and Sinceny:A, left p2 (MNHN-QNY2-3076); B, left p3 (MNHN-TRY16); C, left p4 (MNHN-TRY22-L); D, right M1/2 (MNHN-M-TRY23 [reversed]); E, left M3 (MNHN-M-TRY8); F, left m1 (MNHN-M-TRY5 [reversed]); G, left m1/2 (MNHN-ME16322 [reversed]); H, right DP2 (MNHN-SY3-Cn [reversed]); I, right P3 (MNHN-SY1-Cn [reversed]); J, left P4 (MNHN-SY5-DE); K, left M1/2 (MNHN-SY4-DE); L, right M1/2 (MNHN-ME16325 [reversed]); M, right DP3 (MNHN-M.TRY10 [reversed]); N, right DP4 (MNHN-M.TRY3 [reversed]); O-P, left mandible with p4, m2-m3 (coll. Phélizon); A-O, occlusal views; P, mesial view; A-N, coated with magnesium; H-K, casts. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8037940" httpUri="https://zenodo.org/record/8037940/files/figure.png" pageId="11" pageNumber="286">Fig. 5H</figureCitation>
). The parastyle is small.There is neither protocone nor hypocone, but a developed cingulum is present on the anteriolingual margin. The DP3 displays a larger parastyle (
<figureCitation id="C0F41FD8520DB669FD5BFEC1D9B1FEE5" box="[675,763,343,370]" captionStart="FIG" captionStartId="10.[132,143,1667,1684]" captionTargetBox="[133,1454,215,1626]" captionTargetId="figure-142@10.[699,1267,778,1344]" captionTargetPageId="10" captionText="FIG. 5. — Teeth of Pliolophus quesnoyensisBronnert,Gheerbrant,Godinot &amp; Métais,2018 from Le Quesnoy,Try,Meudon and Sinceny:A, left p2 (MNHN-QNY2-3076); B, left p3 (MNHN-TRY16); C, left p4 (MNHN-TRY22-L); D, right M1/2 (MNHN-M-TRY23 [reversed]); E, left M3 (MNHN-M-TRY8); F, left m1 (MNHN-M-TRY5 [reversed]); G, left m1/2 (MNHN-ME16322 [reversed]); H, right DP2 (MNHN-SY3-Cn [reversed]); I, right P3 (MNHN-SY1-Cn [reversed]); J, left P4 (MNHN-SY5-DE); K, left M1/2 (MNHN-SY4-DE); L, right M1/2 (MNHN-ME16325 [reversed]); M, right DP3 (MNHN-M.TRY10 [reversed]); N, right DP4 (MNHN-M.TRY3 [reversed]); O-P, left mandible with p4, m2-m3 (coll. Phélizon); A-O, occlusal views; P, mesial view; A-N, coated with magnesium; H-K, casts. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8037940" httpUri="https://zenodo.org/record/8037940/files/figure.png" pageId="11" pageNumber="286">Fig. 5M</figureCitation>
). There is no metaconule nor paraconule.The hypocone is larger than the protocone. The cingulum is interrupted close to the hypocone. The DP4 has a protocone larger than the hypocone (
<figureCitation id="C0F41FD8520DB669FF75FE40DB95FE64" box="[141,223,470,497]" captionStart="FIG" captionStartId="10.[132,143,1667,1684]" captionTargetBox="[133,1454,215,1626]" captionTargetId="figure-142@10.[699,1267,778,1344]" captionTargetPageId="10" captionText="FIG. 5. — Teeth of Pliolophus quesnoyensisBronnert,Gheerbrant,Godinot &amp; Métais,2018 from Le Quesnoy,Try,Meudon and Sinceny:A, left p2 (MNHN-QNY2-3076); B, left p3 (MNHN-TRY16); C, left p4 (MNHN-TRY22-L); D, right M1/2 (MNHN-M-TRY23 [reversed]); E, left M3 (MNHN-M-TRY8); F, left m1 (MNHN-M-TRY5 [reversed]); G, left m1/2 (MNHN-ME16322 [reversed]); H, right DP2 (MNHN-SY3-Cn [reversed]); I, right P3 (MNHN-SY1-Cn [reversed]); J, left P4 (MNHN-SY5-DE); K, left M1/2 (MNHN-SY4-DE); L, right M1/2 (MNHN-ME16325 [reversed]); M, right DP3 (MNHN-M.TRY10 [reversed]); N, right DP4 (MNHN-M.TRY3 [reversed]); O-P, left mandible with p4, m2-m3 (coll. Phélizon); A-O, occlusal views; P, mesial view; A-N, coated with magnesium; H-K, casts. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8037940" httpUri="https://zenodo.org/record/8037940/files/figure.png" pageId="11" pageNumber="286">Fig. 5N</figureCitation>
). A small paraconule and metaconule are present. The centrocrista is straight. The cingulum is continuous.
</paragraph>
</subSubSection>
</treatment>
</document>
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