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<mods:title id="FF0D22E7D646DF2D49B5143CD32D990E">A genus-level taxonomic review of primitively segmented spiders (Mesothelae, Liphistiidae)</mods:title>
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<mods:namePart id="444E9A452782A6BA98A4BB8A3CB5A902">Xu, Xin</mods:namePart>
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<mods:namePart id="3006912433A07AB9B175640FEE7609A2">Liu, Fengxiang</mods:namePart>
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<mods:namePart id="6C65091B4D48E864ACCEC674CB99FB78">Chen, Jian</mods:namePart>
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<mods:namePart id="72EA7C53481419EE64A0578E7CD0273C">Ono, Hirotsugu</mods:namePart>
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<mods:namePart id="1BD05D0DC24F8A2148BEF687543AB106">Li, Daiqin</mods:namePart>
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<mods:namePart id="CE82223277B98CFAE468F80BCA44708C">Kuntner, Matjaz</mods:namePart>
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<mods:title id="D30F60DA1D452D3D94E20E6E138CCE5C">ZooKeys</mods:title>
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<mods:date id="8785FC9ED9979404984B11D8A1A3DC21">2015</mods:date>
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<mods:number id="449872D345E2DF12F1068EDA4FBFC615">488</mods:number>
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<treatment id="D05DA3E9B8399DF494D4517D13E3A738" ID-GBIF-Taxon="152058740" LSID="urn:lsid:zoobank.org:act:BA103085-F5FB-4DF1-81C7-CA5CDE26E65E" httpUri="http://treatment.plazi.org/id/D05DA3E9B8399DF494D4517D13E3A738" lastPageId="9" lastPageNumber="130" pageId="8" pageNumber="129">
<subSubSection id="27C41AAD8F8135AFE5E4DDE59280FC53" pageId="8" pageNumber="129" type="multiple">
<paragraph id="C8D6F8E210D90CC847E2A7777D827827" pageId="8" pageNumber="129">Taxon classification Animalia Araneae Liphistiidae</paragraph>
</subSubSection>
<subSubSection id="D31B22B29B830F3B584B09CB07AEC71B" pageId="8" pageNumber="129" type="nomenclature">
<paragraph id="201CAF5FE80F2C67D94CDEE7E9929C9D" pageId="8" pageNumber="129">
Genus
<taxonomicName id="AEB64E8BE3741BBD5C3EE9DA29F65F21" ID-CoL="76B3" ID-ENA="1649364" LSID="http://zoobank.org/BA103085-F5FB-4DF1-81C7-CA5CDE26E65E" authority="Xu &amp; Kuntner" class="Arachnida" family="Heptathelidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela" order="Araneae" pageId="8" pageNumber="129" phylum="Arthropoda" rank="genus">Qiongthela Xu &amp; Kuntner</taxonomicName>
<taxonomicNameLabel id="06E0087A4502ACA3E6A97EDC5075FB96" pageId="8" pageNumber="129">gen. n.</taxonomicNameLabel>
Figures 22-27
</paragraph>
</subSubSection>
<subSubSection id="67776B8111AD1E4E88A6FB1367D30130" pageId="8" pageNumber="129" type="type species">
<paragraph id="73DA8D2251C8EBF2EB7797746D1E4A37" pageId="8" pageNumber="129">Type species.</paragraph>
<paragraph id="6E2C754B785FD94632CF8015A68FD3B5" pageId="8" pageNumber="129">
<taxonomicName id="7F296016A50B1960E36DFCDC70371F0F" class="Arachnida" family="Heptathelidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela baishensis" order="Araneae" pageId="8" pageNumber="129" phylum="Arthropoda" rank="species" species="baishensis">Qiongthela baishensis</taxonomicName>
sp. n.
</paragraph>
</subSubSection>
<subSubSection id="0538D4FCBDDC983B2479475D3E20E113" pageId="8" pageNumber="129" type="etymology">
<paragraph id="800EE998C4EF0731B65776D7AD72E239" pageId="8" pageNumber="129">Etymology.</paragraph>
<paragraph id="65F230436ED443796567869042B044F8" pageId="8" pageNumber="129">
The genera of heptathelines contain in their name the word
<normalizedToken id="5C0390307BC4320C0A4A217A3B1A3ED6" originalValue="‘thela’">'thela'</normalizedToken>
referring to spinnerets as the Greek word thele means nipple-like protuberance (
<bibRefCitation id="59BA4C45C94F3901BC963786BF34549C" author="Ono, H" journalOrPublisher="Memoirs of the National Science Museum" pageId="20" pageNumber="141" pagination="145 - 151" title="Zoogeographic and taxonomic notes on spiders of the subfamily Heptathelinae (Araneae, Mesothelae, Liphistiidae)." volume="33" year="2000">Ono 2000</bibRefCitation>
). We continue this tradition, but name the genus to start with Qiong-, referring to Hainan Province, China.
</paragraph>
</subSubSection>
<subSubSection id="F23DBB861A8C44AF465D6AFC52805AC8" pageId="8" pageNumber="129" type="diagnosis">
<paragraph id="867C262E257EE3C577A7506FEEF887CB" pageId="8" pageNumber="129">Diagnosis.</paragraph>
<paragraph id="947E2A2B802F485F57E635DA7B09CE7D" pageId="8" pageNumber="129">
<taxonomicName id="D7E13F7961053FFE9B603754E3CCC0BB" class="Arachnida" family="Heptathelidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela" order="Araneae" pageId="8" pageNumber="129" phylum="Arthropoda" rank="genus">Qiongthela</taxonomicName>
males differ from all other
<taxonomicName id="5E79F01028D4752A225B6637230BE140" lsidName="" pageId="8" pageNumber="129" rank="subFamily" subFamily="Heptathelinae">Heptathelinae</taxonomicName>
genera by the conductor with a narrow, blade-like, slightly hooked apex (Figures 22, 25-26), and by tegulum with two apophyses (Figures 23, 25-26).
<taxonomicName id="BDA6617CD5AFDF3990FA3E7A3A0322C9" class="Arachnida" family="Heptathelidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela" order="Araneae" pageId="8" pageNumber="129" phylum="Arthropoda" rank="genus">Qiongthela</taxonomicName>
females can be distinguished from all other
<taxonomicName id="2C61C5BE6F96990B857280F304A2B619" lsidName="" pageId="8" pageNumber="129" rank="subFamily" subFamily="Heptathelinae">Heptathelinae</taxonomicName>
by two paired receptacular clusters located at the anterior margin of the bursa copulatrix (Figure 27).
</paragraph>
</subSubSection>
<subSubSection id="A309032AAE31CBE514D8DB0C9120C802" pageId="8" pageNumber="129" type="description">
<paragraph id="7D4F12D8A6AB049BD8F03A5B8CC5565F" pageId="8" pageNumber="129">Description.</paragraph>
<paragraph id="B6285541763743EF752808D920A76B1D" pageId="8" pageNumber="129">Total length (excluding chelicerae) = 13-31 mm (N = 14); male palp with a distally free conductor, narrow, blade-like with slightly hook-like apex, (Figures 22, 25-26); tegulum with two margins, spinose paracymbium (Figures 23, 25-26); female genitalia with two paired receptacular clusters, all situated at the anterior margin of the bursa copulatrix with more or less distinct tubular stems (Figure 27).</paragraph>
</subSubSection>
<subSubSection id="A7F3DFD9B4AF1CA8E50D487C7CDE7E7F" pageId="8" pageNumber="129" type="species composition">
<paragraph id="9E42628A53DBC9AF74500C32C4FFE686" pageId="8" pageNumber="129">Species composition.</paragraph>
<paragraph id="0F8CDDC51E27BEA3BCB6EDB71C9A19E5" pageId="8" pageNumber="129">
<taxonomicName id="EC99476D7234C07E3C3AF8CF1B37E996" class="Arachnida" family="Heptathelidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela australis" order="Araneae" pageId="8" pageNumber="129" phylum="Arthropoda" rank="species" species="australis">Qiongthela australis</taxonomicName>
(Ono, 2002), comb. n.,
<taxonomicName id="B3F111CA8D5C0558FB022BE07FA93B66" class="Arachnida" family="Heptathelidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela nui" order="Araneae" pageId="8" pageNumber="129" phylum="Arthropoda" rank="species" species="nui">Qiongthela nui</taxonomicName>
(Schwendinger &amp; Ono, 2011), comb. n.,
<taxonomicName id="DDA5B7CA477DCB4ED03D11A410CF5FEF" class="Arachnida" family="Heptathelidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela baishensis" order="Araneae" pageId="8" pageNumber="129" phylum="Arthropoda" rank="species" species="baishensis">Qiongthela baishensis</taxonomicName>
sp. n. (3♂2♀), and three undescribed species (6♂8♀, 1♀ and 1♂1♀, respectively) from Hainan, China.
</paragraph>
</subSubSection>
<subSubSection id="FC8D102B406E6CEFCFFFCD4F878A9189" pageId="8" pageNumber="129" type="distribution">
<paragraph id="7893D0C47B6CB8326FF786D36E98B4CF" pageId="8" pageNumber="129">Distribution.</paragraph>
<paragraph id="AB8F9DE9BF393EFF1B8BE7FB9E7D2B65" pageId="8" pageNumber="129">Hainan (China) and Vietnam.</paragraph>
</subSubSection>
<subSubSection id="C8B264313B1C468BC4A892BAB587F6C8" pageId="9" pageNumber="130" type="remarks">
<paragraph id="0FC2D5D3E74E3FEA2EA92C76D2C3A0A1" pageId="9" pageNumber="130">
<pageBreakToken id="695C042782F086D4945738DA3F50C573" pageId="9" pageNumber="130" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph id="D3EA9AFA39F88B6F01127CEE4A1CD8D2" pageId="9" pageNumber="130">
Based on morphological descriptions, but not on phylogenetic analyses, we include two species from Vietnam in this genus, originally described as
<taxonomicName id="3649DAB9D80C543C9236A78EDB23B034" class="Arachnida" family="Heptathelidae" genus="Songthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Songthela australis" order="Araneae" pageId="9" pageNumber="130" phylum="Arthropoda" rank="species" species="australis">Songthela australis</taxonomicName>
Ono, 2002 and
<taxonomicName id="6B2F818F771F3605CA2C5133D32C1093" class="Arachnida" family="Heptathelidae" genus="Heptathela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptathela nui" order="Araneae" pageId="9" pageNumber="130" phylum="Arthropoda" rank="species" species="nui">Heptathela nui</taxonomicName>
Schwendinger &amp; Ono, 2011.
</paragraph>
</subSubSection>
</treatment>
</document>
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