327 lines
36 KiB
XML
327 lines
36 KiB
XML
<document id="87B0EA4C3F834B44B536C1DCD5092438" ID-DOI="10.2478/if-2019-0024" ID-ISSN="2533-4069" ID-Zenodo-Dep="5382362" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630294818620" checkinUser="diego" docAuthor="Qiu, Zhuding & Ni, Xijun" docDate="2019" docId="D035C10BFF9BFF82FBF6F9C504615A81" docLanguage="en" docName="07_Qiu-1.pdf" docOrigin="Fossil Imprint 75 (3 - 4)" docSource="http://dx.doi.org/10.2478/if-2019-0024" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Typhlomys storchi Qiu & Ni 2019, n. sp." docType="treatment" docVersion="3" lastPageNumber="394" masterDocId="2C0CB973FF9CFF89FFF2FF90060D5B70" masterDocTitle="Platacanthomyids (Rodentia, Mammalia) From The Late Miocene Yuanmou Hominoid Locality Of Yunnan, China" masterLastPageNumber="396" masterPageNumber="383" pageNumber="390" updateTime="1699218172350" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="894B0E0921C26EC8E5FC9CBF877CC401">Platacanthomyids (Rodentia, Mammalia) From The Late Miocene Yuanmou Hominoid Locality Of Yunnan, China</mods:title>
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<mods:namePart id="5588114B3E7F475CF8BB55F4E5A0A523">Qiu, Zhuding</mods:namePart>
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<mods:namePart id="437416AF7A64A60E01EC87DE77A76400">Ni, Xijun</mods:namePart>
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<mods:title id="D7DE1D7B09C615A24BDAD0FFB13DCC0E">Fossil Imprint</mods:title>
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<mods:part id="7BA8921640D074ACFB60D9A023937726">
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<mods:date id="6CCF9FD80E8F9D44F3B969A943C06234">2019</mods:date>
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<mods:number id="7D453FB8BEC8313270395DF73D58CE12">2019-12-30</mods:number>
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<mods:number id="70D434A2FFE998EFDAC1E11F9363BF04">3 - 4</mods:number>
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<treatment id="D035C10BFF9BFF82FBF6F9C504615A81" LSID="urn:lsid:plazi:treatment:D035C10BFF9BFF82FBF6F9C504615A81" httpUri="http://treatment.plazi.org/id/D035C10BFF9BFF82FBF6F9C504615A81" lastPageId="11" lastPageNumber="394" pageId="7" pageNumber="390">
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<subSubSection id="10862396FF9BFF8EFBF6F9C5031D5D1E" box="[1028,1296,1621,1646]" pageId="7" pageNumber="390" type="nomenclature">
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<paragraph id="5823701DFF9BFF8EFBF6F9C5031D5D1E" blockId="7.[1028,1296,1621,1646]" box="[1028,1296,1621,1646]" pageId="7" pageNumber="390">
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<heading id="036BC771FF9BFF8EFBF6F9C5031D5D1E" box="[1028,1296,1621,1646]" centered="true" fontSize="8" level="2" pageId="7" pageNumber="390" reason="2">
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<emphasis id="6AE8AC0FFF9BFF8EFBF6F9C5031D5D1E" bold="true" box="[1028,1296,1621,1646]" pageId="7" pageNumber="390">
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<taxonomicName id="9F9C0B9EFF9BFF8EFBF6F9C502C35D1D" authority="Qiu & Ni, 2019" authorityName="Qiu & Ni" authorityYear="2019" box="[1028,1230,1621,1645]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="390" phylum="Chordata" rank="species" species="storchi" status="sp. nov.">
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<emphasis id="6AE8AC0FFF9BFF8EFBF6F9C502C35D1D" bold="true" box="[1028,1230,1621,1645]" italics="true" pageId="7" pageNumber="390">Typhlomys storchi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="71DB1174FF9BFF8EFB26F9C6031D5D1E" box="[1236,1296,1622,1646]" pageId="7" pageNumber="390" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="10862396FF9BFF82FBB6F9E604615A81" lastPageId="11" lastPageNumber="394" pageId="7" pageNumber="390" type="description">
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<paragraph id="5823701DFF9BFF8EFBB6F9E602DD5DFE" blockId="7.[1092,1232,1653,1678]" box="[1092,1232,1653,1678]" pageId="7" pageNumber="390">
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<figureCitation id="C0A76C98FF9BFF8EFBB6F9E602B55DFD" box="[1092,1208,1653,1678]" captionStart="Text-fig" captionStartId="8.[123,204,1932,1953]" captionTargetBox="[126,1437,148,1894]" captionTargetId="figure-107@8.[499,876,981,1261]" captionTargetPageId="8" captionText="Text-fig. 6. Scatter diagrams (length and width) of Typhlomys aff. T. primitivus measurements from Leilao, Yuanmou in comparison with those of T. hipparionum and T. storchi from Leilao, and of T. primitivus and T. hipparionum from Shihuiba, Lufeng." figureDoi="http://doi.org/10.5281/zenodo.5382375" httpUri="https://zenodo.org/record/5382375/files/figure.png" pageId="7" pageNumber="390">Text-figs 6</figureCitation>
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,
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<figureCitation id="C0A76C98FF9BFF8EFB31F9E602DD5DFE" box="[1219,1232,1654,1678]" captionStart="Text-fig" captionStartId="10.[125,206,1012,1033]" captionTargetBox="[122,1444,149,1004]" captionTargetId="figure-555@10.[122,1444,149,1004]" captionTargetPageId="10" captionText="Text-fig. 8. Typhlomys storchi n. sp. from Leilao, Yuanmou. a) left M1 (IVPP V 25887.1, paratype), b) right M3 (IVPP V 25887.2, reversed), c) right M1 (IVPP V 25886, holotype, reversed), d) right M2 (IVPP V 25887.3, reversed)." figureDoi="http://doi.org/10.5281/zenodo.5382379" httpUri="https://zenodo.org/record/5382379/files/figure.png" pageId="7" pageNumber="390">8</figureCitation>
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</paragraph>
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||
<paragraph id="5823701DFF9BFF8EFC83F93203C75DC9" blockId="7.[843,1482,1697,1753]" box="[881,1482,1697,1724]" pageId="7" pageNumber="390">
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H o l o t y p e. Right M
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<subScript id="C4187258FF9BFF8EFB9CF93E027B5DCC" attach="left" box="[1134,1142,1710,1724]" fontSize="6" pageId="7" pageNumber="390">1</subScript>
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(IVPP
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<accessionNumber id="47CFEDFEFF9BFF8EFB3FF932033F5DC9" box="[1229,1330,1697,1721]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V25886" pageId="7" pageNumber="390" type="EnaNcbi">V 25886</accessionNumber>
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); dimensions:
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</paragraph>
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<paragraph id="5823701DFF9BFF8EFCB9F95102BD5DA9" blockId="7.[843,1482,1697,1753]" box="[843,1200,1729,1753]" pageId="7" pageNumber="390">
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2.75 ×
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<quantity id="9F64DDF8FF9BFF8EFC64F95105F95DA9" box="[918,1012,1729,1753]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="390" unit="mm" value="1.3">1.30 mm</quantity>
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(length × width).
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</paragraph>
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<paragraph id="5823701DFF9BFF8EFC83F97D03C45C75" blockId="7.[843,1481,1773,1829]" box="[881,1481,1773,1797]" pageId="7" pageNumber="390">E t y m o l o g y. This new species is dedicated to Gerhard</paragraph>
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<paragraph id="5823701DFF9BFF8EFCB9F89D05955C55" blockId="7.[843,1481,1773,1829]" box="[843,920,1805,1829]" pageId="7" pageNumber="390">Storch.</paragraph>
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<paragraph id="5823701DFF9BFF8EFC83F8A903C75C20" blockId="7.[843,1482,1848,1905]" box="[881,1482,1848,1873]" pageId="7" pageNumber="390">T y p e l o c a l i t y. Loc. 9905, Leilao, Yuanmou,</paragraph>
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<paragraph id="5823701DFF9BFF8EFCB9F8C9025B5C01" blockId="7.[843,1482,1848,1905]" box="[843,1110,1881,1905]" pageId="7" pageNumber="390">
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<collectingRegion id="9A58BEFFFF9BFF8EFCB9F8C902085C01" box="[843,1029,1881,1905]" country="China" name="Yunnan" pageId="7" pageNumber="390">Yunnan Province</collectingRegion>
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,
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<collectingCountry id="208B308DFF9BFF8EFBFDF8C9025F5C01" box="[1039,1106,1881,1905]" name="China" pageId="7" pageNumber="390">China</collectingCountry>
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.
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</paragraph>
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<paragraph id="5823701DFF9BFF8EFC83F81502815CCC" blockId="7.[843,1482,1924,1980]" pageId="7" pageNumber="390">
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T y p e s t r a t u m.
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<quantity id="9F64DDF8FF9BFF8EFBB8F81402CF5CEC" box="[1098,1218,1924,1948]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" metricValueMax="15.0" metricValueMin="3.0" pageId="7" pageNumber="390" unit="cm" value="90.0" valueMax="150.0" valueMin="30.0">30–150 cm</quantity>
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grayish bands of pebbly sandstone, Xiaohe Formation.
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</paragraph>
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<paragraph id="5823701DFF9BFF8EFC83F84002585C98" blockId="7.[881,1109,2000,2024]" box="[881,1109,2000,2024]" pageId="7" pageNumber="390">A g e. Late Miocene.</paragraph>
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<paragraph id="5823701DFF9BFF8EFC83F86C03C75363" blockId="7.[843,1482,2042,2100]" box="[881,1482,2042,2068]" pageId="7" pageNumber="390">
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P a r a t y p e. 1 left M
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||
<superScript id="AFE9DD55FF9BFF8EFB9FF86A02785378" attach="left" box="[1133,1141,2042,2056]" fontSize="6" pageId="7" pageNumber="390">1</superScript>
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(IVPP V 25887.1) from Loc.
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</paragraph>
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<paragraph id="5823701DFF9BFF8EFCB9F78C03765344" blockId="7.[843,1482,2042,2100]" box="[843,1403,2076,2100]" pageId="7" pageNumber="390">
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||
9905; dimensions: 2.05 ×
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<quantity id="9F64DDF8FF9BFF8EFB93F78C02B25343" box="[1121,1215,2076,2100]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="390" unit="mm" value="1.3">1.30 mm</quantity>
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(length × width).
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||
</paragraph>
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||
<caption id="0CE32095FF94FF81FF89F81C03315CCE" ID-DOI="http://doi.org/10.5281/zenodo.5382375" ID-Zenodo-Dep="5382375" httpUri="https://zenodo.org/record/5382375/files/figure.png" pageId="8" pageNumber="391" startId="8.[123,204,1932,1953]" targetBox="[126,1437,148,1894]" targetPageId="8">
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||
<paragraph id="5823701DFF94FF81FF89F81C03315CCE" blockId="8.[123,1439,1869,1982]" pageId="8" pageNumber="391">
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||
Text-fig. 6. Scatter diagrams (length and width) of
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||
<taxonomicName id="9F9C0B9EFF94FF81FD84F81C04EC5CD1" authorityName="Milne-Edwards" authorityYear="1877" box="[630,737,1932,1953]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="391" phylum="Chordata" rank="genus">
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<emphasis id="6AE8AC0FFF94FF81FD84F81C04EC5CD1" bold="true" box="[630,737,1932,1953]" italics="true" pageId="8" pageNumber="391">Typhlomys</emphasis>
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</taxonomicName>
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aff.
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<taxonomicName id="9F9C0B9EFF94FF81FCFFF81C05855CD1" authorityName="QIU" authorityYear="1989" box="[781,904,1932,1953]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="391" phylum="Chordata" rank="species" species="primitivus">
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<emphasis id="6AE8AC0FFF94FF81FCFFF81C05855CD1" bold="true" box="[781,904,1932,1953]" italics="true" pageId="8" pageNumber="391">T. primitivus</emphasis>
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</taxonomicName>
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measurements from Leilao, Yuanmou in comparison with those of
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<taxonomicName id="9F9C0B9EFF94FF81FEF6F83907935CCE" authorityName="QIU" authorityYear="1989" box="[260,414,1961,1982]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="391" phylum="Chordata" rank="species" species="hipparionum">
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<emphasis id="6AE8AC0FFF94FF81FEF6F83907935CCE" bold="true" box="[260,414,1961,1982]" italics="true" pageId="8" pageNumber="391">T. hipparionum</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="9F9C0B9EFF94FF81FE23F83904205CCE" authorityName="Qiu & Ni" authorityYear="2019" box="[465,557,1961,1982]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="391" phylum="Chordata" rank="species" species="storchi">
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<emphasis id="6AE8AC0FFF94FF81FE23F83904205CCE" bold="true" box="[465,557,1961,1982]" italics="true" pageId="8" pageNumber="391">T. storchi</emphasis>
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</taxonomicName>
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from Leilao, and of
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<taxonomicName id="9F9C0B9EFF94FF81FD0CF83905775CCE" authorityName="QIU" authorityYear="1989" box="[766,890,1961,1982]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="391" phylum="Chordata" rank="species" species="primitivus">
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<emphasis id="6AE8AC0FFF94FF81FD0CF83905775CCE" bold="true" box="[766,890,1961,1982]" italics="true" pageId="8" pageNumber="391">T. primitivus</emphasis>
|
||
</taxonomicName>
|
||
and
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<taxonomicName id="9F9C0B9EFF94FF81FC5FF839024B5CCE" authorityName="QIU" authorityYear="1989" box="[941,1094,1961,1982]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="391" phylum="Chordata" rank="species" species="hipparionum">
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<emphasis id="6AE8AC0FFF94FF81FC5FF839024B5CCE" bold="true" box="[941,1094,1961,1982]" italics="true" pageId="8" pageNumber="391">T. hipparionum</emphasis>
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</taxonomicName>
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from Shihuiba, Lufeng.
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</paragraph>
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</caption>
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<paragraph id="5823701DFF94FF81FF62F86704E4537E" blockId="8.[106,745,2037,2097]" box="[144,745,2037,2062]" pageId="8" pageNumber="391">
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R e f e r r e d m a t e r i a l. 2 isolated teeth (1 M
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<superScript id="AFE9DD55FF94FF81FD51F86504A65373" attach="left" box="[675,683,2037,2051]" fontSize="6" pageId="8" pageNumber="391">3</superScript>
|
||
from
|
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</paragraph>
|
||
<paragraph id="5823701DFF94FF81FF98F78704A1535E" blockId="8.[106,745,2037,2097]" box="[106,684,2070,2097]" pageId="8" pageNumber="391">
|
||
Loc. 9906, 1 M
|
||
<subScript id="C4187258FF94FF81FEFDF7B3071A5341" attach="left" box="[271,279,2083,2097]" fontSize="6" pageId="8" pageNumber="391">2</subScript>
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from Loc. 9903; IVPP V 25887.2–3).
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</paragraph>
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<paragraph id="5823701DFF94FF81FCA8F86703BE537E" blockId="8.[820,1459,2038,2097]" box="[858,1459,2038,2062]" pageId="8" pageNumber="391">M e a s u r e m e n t s f o r r e f e r r e d m a t e r i a l.</paragraph>
|
||
<paragraph id="5823701DFF94FF81FCC6F78703A0535E" blockId="8.[820,1459,2038,2097]" box="[820,1453,2069,2097]" pageId="8" pageNumber="391">
|
||
M
|
||
<superScript id="AFE9DD55FF94FF81FCBEF78505595353" attach="left" box="[844,852,2069,2083]" fontSize="6" pageId="8" pageNumber="391">3</superScript>
|
||
: 1.60 ×
|
||
<quantity id="9F64DDF8FF94FF81FC5FF7860202535E" box="[941,1039,2070,2094]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="8" pageNumber="391" unit="mm" value="1.25">1.25 mm</quantity>
|
||
; M
|
||
<subScript id="C4187258FF94FF81FBC3F7B302345341" attach="left" box="[1073,1081,2083,2097]" fontSize="6" pageId="8" pageNumber="391">2</subScript>
|
||
: 2.00 ×
|
||
<quantity id="9F64DDF8FF94FF81FB60F78602FC535E" box="[1170,1265,2070,2094]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="8" pageNumber="391" unit="mm" value="1.35">1.35 mm</quantity>
|
||
(length × width).
|
||
</paragraph>
|
||
<caption id="0CE32095FF95FF80FF61FA5304DA5D40" ID-DOI="http://doi.org/10.5281/zenodo.5382377" ID-Zenodo-Dep="5382377" httpUri="https://zenodo.org/record/5382377/files/figure.png" pageId="9" pageNumber="392" startId="9.[147,228,1475,1496]" targetBox="[147,1462,151,1455]" targetPageId="9">
|
||
<paragraph id="5823701DFF95FF80FF61FA5304DA5D40" blockId="9.[147,1463,1473,1587]" pageId="9" pageNumber="392">
|
||
<emphasis id="6AE8AC0FFF95FF80FF61FA5307A75D43" bold="true" pageId="9" pageNumber="392">
|
||
Text-fig. 7.
|
||
<taxonomicName id="9F9C0B9EFF95FF80FEFAFA5307F75EA8" authorityName="QIU" authorityYear="1989" box="[264,506,1475,1496]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="392" phylum="Chordata" rank="species" species="hipparionum">
|
||
<emphasis id="6AE8AC0FFF95FF80FEFAFA5307F75EA8" bold="true" box="[264,506,1475,1496]" italics="true" pageId="9" pageNumber="392">Typhlomys hipparionum</emphasis>
|
||
</taxonomicName>
|
||
Q, 1989 from Leilao, Yuanmou. a) left M
|
||
<superScript id="AFE9DD55FF95FF80FC4AFA5105B25EBE" attach="left" box="[952,959,1473,1486]" fontSize="5" pageId="9" pageNumber="392">1</superScript>
|
||
(IVPP V 25885.4), b) right M
|
||
<superScript id="AFE9DD55FF95FF80FB02FA5102FA5EBE" attach="left" box="[1264,1271,1473,1486]" fontSize="5" pageId="9" pageNumber="392">1</superScript>
|
||
(IVPP V 25885.5), IU c) left M
|
||
<superScript id="AFE9DD55FF95FF80FF1CFA4F06F85E9C" attach="left" box="[238,245,1503,1516]" fontSize="5" pageId="9" pageNumber="392">2</superScript>
|
||
(IVPP V 25885.6), d) left M
|
||
<superScript id="AFE9DD55FF95FF80FDECFA4F04285E9C" attach="left" box="[542,549,1503,1516]" fontSize="5" pageId="9" pageNumber="392">2</superScript>
|
||
(IVPP V 25885.7, reversed), e) left M
|
||
<superScript id="AFE9DD55FF95FF80FC42FA4F05BA5E9C" attach="left" box="[944,951,1503,1516]" fontSize="5" pageId="9" pageNumber="392">3</superScript>
|
||
(IVPP V 25885.8), f) right M
|
||
<superScript id="AFE9DD55FF95FF80FB1EFA4F02FE5E9C" attach="left" box="[1260,1267,1503,1516]" fontSize="5" pageId="9" pageNumber="392">3</superScript>
|
||
(IVPP V 25885.9), g) left M
|
||
<subScript id="C4187258FF95FF80FF18F99906FC5D66" attach="left" box="[234,241,1545,1558]" fontSize="5" pageId="9" pageNumber="392">1</subScript>
|
||
(IVPP V 25885.10), h) right M
|
||
<subScript id="C4187258FF95FF80FDD5F99904235D66" attach="left" box="[551,558,1545,1558]" fontSize="5" pageId="9" pageNumber="392">1</subScript>
|
||
(IVPP V 25885.11), i) left M
|
||
<subScript id="C4187258FF95FF80FCBEF999055E5D66" attach="left" box="[844,851,1545,1558]" fontSize="5" pageId="9" pageNumber="392">2</subScript>
|
||
(IVPP V 25885.12), j) right M
|
||
<subScript id="C4187258FF95FF80FB76F99902865D66" attach="left" box="[1156,1163,1545,1558]" fontSize="5" pageId="9" pageNumber="392">2</subScript>
|
||
(IVPP V 25885.13), k) left M
|
||
<subScript id="C4187258FF95FF80FA42F99903BA5D66" attach="left" box="[1456,1463,1545,1558]" fontSize="5" pageId="9" pageNumber="392">3</subScript>
|
||
(IVPP V 25885.1), l) left M
|
||
<subScript id="C4187258FF95FF80FE51F9B607A75D43" attach="left" box="[419,426,1574,1587]" fontSize="5" pageId="9" pageNumber="392">3</subScript>
|
||
</emphasis>
|
||
(IVPP V 25885.14, reversed).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5823701DFF95FF80FF54F9E707D75C22" blockId="9.[129,768,1655,1877]" pageId="9" pageNumber="392">
|
||
D i a g n o s i s. Large-sized species of
|
||
<taxonomicName id="9F9C0B9EFF95FF80FD75F9E704F45DFF" authorityName="Milne-Edwards" authorityYear="1877" box="[647,761,1655,1679]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="392" phylum="Chordata" rank="genus">
|
||
<emphasis id="6AE8AC0FFF95FF80FD75F9E704F45DFF" box="[647,761,1655,1679]" italics="true" pageId="9" pageNumber="392">Typhlomys</emphasis>
|
||
</taxonomicName>
|
||
. Molars with concave grinding surface, six inclined diagonal ridges separated by five synclines, continuous endoloph(id)s and ectoloph(id)s incorporated with the anteroloph(id)s and posteroloph(id)s to circle the occlusal surface without buccal opening of synclines II and III. Anterior portion of M
|
||
<subScript id="C4187258FF95FF80FF6AF8D706AD5C25" attach="left" box="[152,160,1863,1877]" fontSize="6" pageId="9" pageNumber="392">1</subScript>
|
||
distinctly extending forward.
|
||
</paragraph>
|
||
<paragraph id="5823701DFF95FF80FF54F8FA048B5CD2" blockId="9.[128,767,1898,1955]" pageId="9" pageNumber="392">
|
||
S t r a t i g r a p h i c r a n g e.
|
||
<taxonomicName id="9F9C0B9EFF95FF80FE17F8FA04AB5CF2" authority="Qiu & Ni, 2019" authorityName="Qiu & Ni" authorityYear="2019" box="[485,678,1898,1922]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="392" phylum="Chordata" rank="species" species="storchi" status="sp. nov.">
|
||
<emphasis id="6AE8AC0FFF95FF80FE17F8FA04AB5CF2" box="[485,678,1898,1922]" italics="true" pageId="9" pageNumber="392">Typhlomys storchi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71DB1174FF95FF80FD5FF8FA04EA5CF1" box="[685,743,1898,1921]" pageId="9" pageNumber="392" rank="species">n. sp.</taxonomicNameLabel>
|
||
is known so far only from Leilao, the
|
||
<typeStatus id="8727CEBFFF95FF80FE0EF81B04275CD3" box="[508,554,1931,1955]" pageId="9" pageNumber="392">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph id="5823701DFF95FF80FF54F82B036D5DDF" blockId="9.[128,768,1977,2100]" lastBlockId="9.[843,1483,1655,2100]" pageId="9" pageNumber="392">
|
||
D e s c r i p t i o n. The available molars (M
|
||
<superScript id="AFE9DD55FF95FF80FD79F829049E5CB7" attach="left" box="[651,659,1977,1991]" fontSize="6" pageId="9" pageNumber="392">1</superScript>
|
||
, M
|
||
<superScript id="AFE9DD55FF95FF80FD4CF82904CB5CB7" attach="left" box="[702,710,1977,1991]" fontSize="6" pageId="9" pageNumber="392">3</superScript>
|
||
, M
|
||
<subScript id="C4187258FF95FF80FD03F85704F45CA5" attach="left" box="[753,761,1991,2005]" fontSize="6" pageId="9" pageNumber="392">1</subScript>
|
||
, M
|
||
<subScript id="C4187258FF95FF80FF6AF87706AD5C85" attach="left" box="[152,160,2023,2037]" fontSize="6" pageId="9" pageNumber="392">2</subScript>
|
||
) are lophodont, with concave grinding surface, slight unilateral hypsodonty of the upper molars, and have six diagonal/transversal ridges separated by five synclines. The endoloph and the endolophid are rather developed, and the synclines are laterally closed in all stages of wear.
|
||
</paragraph>
|
||
<paragraph id="5823701DFF95FF80FC83F928021F5344" blockId="9.[843,1483,1655,2100]" pageId="9" pageNumber="392">
|
||
The M
|
||
<superScript id="AFE9DD55FF95FF80FC49F92605CE5DB4" attach="left" box="[955,963,1718,1732]" fontSize="6" pageId="9" pageNumber="392">1</superScript>
|
||
is subrectangular in occlusal view, with nearly parallel lateral margins. The anteroloph is pronounced and anteriorly curved. The anterior extra ridge is prominent, enclosing a crescent syncline Ia with the anterolophid. The protoloph and mesoloph are long and strong, roughly parallel to the anterior extra ridge in direction. The mesoloph has a mesial angle to the longitudinal axis of 53°. The metaloph is short, but longer than the anterior extra ridge. The posteroloph is nearly transverse. Synclines II and III are marked, and in parallel arrangement; there is no trace of a buccal opening of the synclines. Syncline IV is close to syncline Ia in size.
|
||
</paragraph>
|
||
<caption id="0CE32095FF96FF83FF8FFC64026F5F57" ID-DOI="http://doi.org/10.5281/zenodo.5382379" ID-Zenodo-Dep="5382379" httpUri="https://zenodo.org/record/5382379/files/figure.png" pageId="10" pageNumber="393" startId="10.[125,206,1012,1033]" targetBox="[122,1444,149,1004]" targetPageId="10">
|
||
<paragraph id="5823701DFF96FF83FF8FFC64026F5F57" blockId="10.[125,1440,1011,1066]" pageId="10" pageNumber="393">
|
||
<emphasis id="6AE8AC0FFF96FF83FF8FFC64054F5F5A" bold="true" pageId="10" pageNumber="393">
|
||
Text-fig. 8.
|
||
<taxonomicName id="9F9C0B9EFF96FF83FF00FC6507A55F7A" authority="Qiu & Ni, 2019" authorityName="Qiu & Ni" authorityYear="2019" box="[242,424,1013,1034]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="393" phylum="Chordata" rank="species" species="storchi" status="sp. nov.">
|
||
<emphasis id="6AE8AC0FFF96FF83FF00FC6507A55F7A" bold="true" box="[242,424,1013,1034]" italics="true" pageId="10" pageNumber="393">Typhlomys storchi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71DB1174FF96FF83FE5DFC6407EB5F79" box="[431,486,1012,1033]" pageId="10" pageNumber="393" rank="species">n. sp.</taxonomicNameLabel>
|
||
from Leilao, Yuanmou. a) left M
|
||
<superScript id="AFE9DD55FF96FF83FCCFFC6305495F70" attach="left" box="[829,836,1011,1024]" fontSize="5" pageId="10" pageNumber="393">1</superScript>
|
||
(IVPP V 25887.1, paratype), b) right M
|
||
<superScript id="AFE9DD55FF96FF83FB13FC6302E55F70" attach="left" box="[1249,1256,1011,1024]" fontSize="5" pageId="10" pageNumber="393">3</superScript>
|
||
(IVPP V 25887.2, reversed), c) right M
|
||
<subScript id="C4187258FF96FF83FEA2FB8D075A5F5A" attach="left" box="[336,343,1053,1066]" fontSize="5" pageId="10" pageNumber="393">1</subScript>
|
||
(IVPP
|
||
<accessionNumber id="47CFEDFEFF96FF83FE50FB8207FB5F57" box="[418,502,1042,1063]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V25886" pageId="10" pageNumber="393" type="EnaNcbi">V 25886</accessionNumber>
|
||
, holotype, reversed), d) right M
|
||
<subScript id="C4187258FF96FF83FCC9FB8D054F5F5A" attach="left" box="[827,834,1053,1066]" fontSize="5" pageId="10" pageNumber="393">2</subScript>
|
||
</emphasis>
|
||
(IVPP V 25887.3, reversed).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5823701DFF96FF83FF62FBFE04465EDA" blockId="10.[106,746,1132,2100]" pageId="10" pageNumber="393">
|
||
The M
|
||
<superScript id="AFE9DD55FF96FF83FF29FBFC06EE5F0A" attach="left" box="[219,227,1132,1146]" fontSize="6" pageId="10" pageNumber="393">3</superScript>
|
||
is an elongated-triangle in occlusal view, with the lateral margins being slightly convergent posteriorly. It has a reduced posterior part and relatively transverse ridges. The anterior extra ridge is fused with the thick anteroloph lingually. The protoloph and mesoloph are conspicuous and nearly transverse in direction. The metaloph is short and slightly curved backward. A trace of syncline Ia is discernible at the anterobuccal corner of the tooth. Syncline II is the longest furrow on the tooth. Syncline IV is more distinct than syncline Ia. The M3 has 3 roots.
|
||
</paragraph>
|
||
<paragraph id="5823701DFF96FF83FF62FA2304E45C82" blockId="10.[106,746,1132,2100]" pageId="10" pageNumber="393">
|
||
The M
|
||
<subScript id="C4187258FF96FF83FF2DFA2F06EA5EBD" attach="left" box="[223,231,1471,1485]" fontSize="6" pageId="10" pageNumber="393">1</subScript>
|
||
is much longer than wide, with the lateral margins converging and distinctly extended anteriorly. The tooth is in very early wear, but shows continuous endolophid and ectolophid. No deep buccal opening of synclines II and III is visible as seen in other
|
||
<taxonomicName id="9F9C0B9EFF96FF83FE13F9A5045E5D3D" authorityName="Milne-Edwards" authorityYear="1877" box="[481,595,1589,1613]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="393" phylum="Chordata" rank="genus">
|
||
<emphasis id="6AE8AC0FFF96FF83FE13F9A5045E5D3D" box="[481,595,1589,1613]" italics="true" pageId="10" pageNumber="393">Typhlomys</emphasis>
|
||
</taxonomicName>
|
||
. The anterior extra ridge is long and pronounced, but does not touch the anterolophid in this stage of wear. The paraconid-metaconid connection is the longest diagonal/transverse ridge on the tooth, with a mesial angle to the longitudinal axis of about 40°. The mesolophid is nearly transverse in the lingual area, and then turns anterobuccally at the middle of the tooth width. The hypolophid is short and transverse, joining with the posterolophid to form a continuous loop posteriorly. Syncline Ia is long but narrow, due to the thickening of the endolophid anteriorly. Syncline I is the longest syncline on the tooth. Synclines II and III are similar in shape and parallel in arrangement, but the former is longer than the latter. Syncline IV is short and transverse. M
|
||
<subScript id="C4187258FF96FF83FDB7F87704405C85" attach="left" box="[581,589,2023,2037]" fontSize="6" pageId="10" pageNumber="393">1</subScript>
|
||
has two roots.
|
||
</paragraph>
|
||
<paragraph id="5823701DFF96FF83FF62F86B05AD5D63" blockId="10.[106,746,1132,2100]" lastBlockId="10.[820,1459,1134,1555]" pageId="10" pageNumber="393">
|
||
The M
|
||
<subScript id="C4187258FF96FF83FF2CF79806EB5366" attach="left" box="[222,230,2056,2070]" fontSize="6" pageId="10" pageNumber="393">2</subScript>
|
||
is subrectangular in occlusal outline, with a straight anterior margin and slightly curved posterior margin. The tooth is in early wear, and shows continuous endolophid and ectolophid, which are incorporated with the anterolophid and posterolophid to circle the occlusal surface. As in the M
|
||
<subScript id="C4187258FF96FF83FCBEFB4E05595F9C" attach="left" box="[844,852,1246,1260]" fontSize="6" pageId="10" pageNumber="393">1</subScript>
|
||
, there is no trace of a buccal opening of synclines II and III. The anterior extra ridge is short, joining the endolophid and the anterolophid to enclose an elongated oval syncline Ia. The paraconid-metaconid connection, the mesolophid and the hypolophid are roughly parallel in arrangement, but successively reduced in length. The mesolophid has a ~60° mesial angle to the longitudinal axis. Synclines I, II and III are similar in shape and orientation. Syncline IV is short, but longer than syncline Ia, slightly curved backward. M
|
||
<subScript id="C4187258FF96FF83FA8CFA77038B5E85" attach="left" box="[1406,1414,1511,1525]" fontSize="6" pageId="10" pageNumber="393">2</subScript>
|
||
has two roots.
|
||
</paragraph>
|
||
<paragraph id="5823701DFF96FF83FCA8F9BC02915C5A" blockId="10.[820,1459,1579,2100]" pageId="10" pageNumber="393">
|
||
D i s c u s s i o n. The described specimens are referred to the genus
|
||
<taxonomicName id="9F9C0B9EFF96FF83FC31F9DC02385D14" authorityName="Milne-Edwards" authorityYear="1877" box="[963,1077,1612,1636]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="393" phylum="Chordata" rank="genus">
|
||
<emphasis id="6AE8AC0FFF96FF83FC31F9DC02385D14" box="[963,1077,1612,1636]" italics="true" pageId="10" pageNumber="393">Typhlomys</emphasis>
|
||
</taxonomicName>
|
||
, due to the lophodont molars with concave grinding surface, six diagonal/transverse ridges and five synclines, mid-ridges/synclines less inclined to the longitudinal axis, developed endoloph in the upper molars and continuous endolophid in the lower molars. In these characters, they resemble
|
||
<taxonomicName id="9F9C0B9EFF96FF83FB7DF962030C5C7A" authorityName="Milne-Edwards" authorityYear="1877" box="[1167,1281,1778,1802]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="393" phylum="Chordata" rank="genus">
|
||
<emphasis id="6AE8AC0FFF96FF83FB7DF962030C5C7A" box="[1167,1281,1778,1802]" italics="true" pageId="10" pageNumber="393">Typhlomys</emphasis>
|
||
</taxonomicName>
|
||
, but differ from
|
||
<taxonomicName id="9F9C0B9EFF96FF83FCC6F88405B85C5B" authorityName="SCHAUB et ZAPFE" authorityYear="1953" box="[820,949,1812,1835]" class="Mammalia" family="Muridae" genus="Neocometes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="393" phylum="Chordata" rank="genus">
|
||
<emphasis id="6AE8AC0FFF96FF83FCC6F88405B85C5B" box="[820,949,1812,1835]" italics="true" pageId="10" pageNumber="393">Neocometes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9F9C0B9EFF96FF83FC1BF88302985C5B" authorityName="BLYTH" authorityYear="1859" box="[1001,1173,1811,1835]" class="Mammalia" family="Platacanthomyidae" genus="Platacanthomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="393" phylum="Chordata" rank="genus">
|
||
<emphasis id="6AE8AC0FFF96FF83FC1BF88302985C5B" box="[1001,1173,1811,1835]" italics="true" pageId="10" pageNumber="393">Platacanthomys</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5823701DFF96FF82FCA8F8A4041C5A7F" blockId="10.[820,1459,1579,2100]" lastBlockId="11.[129,768,151,497]" lastPageId="11" lastPageNumber="394" pageId="10" pageNumber="393">
|
||
The huge dimensions (
|
||
<figureCitation id="C0A76C98FF96FF83FBBAF8A402B55C3C" box="[1096,1208,1844,1868]" captionStart="Text-fig" captionStartId="8.[123,204,1932,1953]" captionTargetBox="[126,1437,148,1894]" captionTargetId="figure-107@8.[499,876,981,1261]" captionTargetPageId="8" captionText="Text-fig. 6. Scatter diagrams (length and width) of Typhlomys aff. T. primitivus measurements from Leilao, Yuanmou in comparison with those of T. hipparionum and T. storchi from Leilao, and of T. primitivus and T. hipparionum from Shihuiba, Lufeng." figureDoi="http://doi.org/10.5281/zenodo.5382375" httpUri="https://zenodo.org/record/5382375/files/figure.png" pageId="10" pageNumber="393">Text-fig. 6</figureCitation>
|
||
) of these specimens are striking, even larger than those of
|
||
<taxonomicName id="9F9C0B9EFF96FF83FB6EF8C5038B5C1D" baseAuthorityName="Zheng" baseAuthorityYear="1993" box="[1180,1414,1877,1901]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="393" phylum="Chordata" rank="species" species="macrourus">
|
||
<emphasis id="6AE8AC0FFF96FF83FB6EF8C5038B5C1D" box="[1180,1414,1877,1901]" italics="true" pageId="10" pageNumber="393">Typhlomys macrourus</emphasis>
|
||
</taxonomicName>
|
||
, the currently largest known form of the genus from Longgupo of Wushan,
|
||
<collectingRegion id="9A58BEFFFF96FF83FC45F80702235CDF" box="[951,1070,1943,1967]" country="China" name="Chongqing" pageId="10" pageNumber="393">Chongqing</collectingRegion>
|
||
(
|
||
<bibRefCitation id="3C0D0DECFF96FF83FBCEF80702CE5CDF" author="Zheng, S. - H." box="[1084,1219,1943,1967]" pageId="10" pageNumber="393" refId="ref10129" refString="Zheng, S. - H. (1993): Quaternary rodents of Sichuan - Guizhou area, China. - Science Press, Beijing, 270 pp. (in Chinese with English summary)" type="book" year="1993">Zheng 1993</bibRefCitation>
|
||
). The relative anterior extension of M
|
||
<subScript id="C4187258FF96FF83FC2FF85505E85CA3" attach="left" box="[989,997,1989,2003]" fontSize="6" pageId="10" pageNumber="393">1</subScript>
|
||
is also unique in
|
||
<taxonomicName id="9F9C0B9EFF96FF83FB40F82803295CA0" authorityName="Milne-Edwards" authorityYear="1877" box="[1202,1316,1976,2000]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="393" phylum="Chordata" rank="genus">
|
||
<emphasis id="6AE8AC0FFF96FF83FB40F82803295CA0" box="[1202,1316,1976,2000]" italics="true" pageId="10" pageNumber="393">Typhlomys</emphasis>
|
||
</taxonomicName>
|
||
. In addition, the specimens possess morphological features different from all known species of
|
||
<taxonomicName id="9F9C0B9EFF96FF83FBEEF86B02835363" authorityName="Milne-Edwards" authorityYear="1877" box="[1052,1166,2043,2067]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="393" phylum="Chordata" rank="genus">
|
||
<emphasis id="6AE8AC0FFF96FF83FBEEF86B02835363" box="[1052,1166,2043,2067]" italics="true" pageId="10" pageNumber="393">Typhlomys</emphasis>
|
||
</taxonomicName>
|
||
, for example, the presence of continuous endoloph(id)s and ectoloph(id)s, and the lack of buccal opening of synclines II and III. The four molars from Leilao obviously represent a new platacanthomyid. Therefore, it is assigned to a new species of
|
||
<taxonomicName id="9F9C0B9EFF97FF82FD75FF4804F45B80" authorityName="Milne-Edwards" authorityYear="1877" box="[647,761,216,240]" class="Mammalia" family="Platacanthomyidae" genus="Typhlomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="394" phylum="Chordata" rank="genus">
|
||
<emphasis id="6AE8AC0FFF97FF82FD75FF4804F45B80" box="[647,761,216,240]" italics="true" pageId="11" pageNumber="394">Typhlomys</emphasis>
|
||
</taxonomicName>
|
||
, although the material is rather scarce.
|
||
</paragraph>
|
||
<paragraph id="5823701DFF97FF82FF54FE8804615A81" blockId="11.[129,768,151,497]" pageId="11" pageNumber="394">
|
||
A possibility of the species described here belonging to a new genus of the
|
||
<taxonomicName id="9F9C0B9EFF97FF82FEA2FEA804165A20" authorityName="ALSTON" authorityYear="1876" box="[336,539,312,336]" class="Mammalia" family="Platacanthomyidae" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="394" phylum="Chordata" rank="family">Platacanthomyidae</taxonomicName>
|
||
cannot be precluded, due to the huge size, the presence of continuous endoloph(id)s and ectoloph(id)s, the distinct extension of anterior portion on M
|
||
<subScript id="C4187258FF97FF82FF4BFE3506CC5AC3" attach="left" box="[185,193,421,435]" fontSize="6" pageId="11" pageNumber="394">1</subScript>
|
||
, and the lack of buccal opening of synclines II and I in molars. Nevertheless, it is considered inadvisable to create a new genus in view of the inadequate material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |