689 lines
117 KiB
XML
689 lines
117 KiB
XML
<document id="AC209040B496FB5203F56BB106837A70" ID-DOI="10.3853/j.2201-4349.71.2019.1719" ID-ISSN="2201-4349" ID-Zenodo-Dep="4653485" ID-ZooBank="CB380F40-F8D8-471B-86B5-E567C8B8EEC9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617217195390" checkinUser="felipe" docAuthor="Smith, Graeme B. & Mitchell, Andrew" docDate="2019" docId="D026802BEE29A267FCAFA584FDEAFC7F" docLanguage="en" docName="RecAustMus.71.4.139-181.pdf" docOrigin="Records of the Australian Museum 71 (4)" docStyle="DocumentStyle:997BA24B490466A2F99594A367FA147F.3:RecAustMus.2019-.journal_article" docStyleId="997BA24B490466A2F99594A367FA147F" docStyleName="RecAustMus.2019-.journal_article" docStyleVersion="3" docTitle="Heterolepisma pyramidum Smith & Mitchell 2019, sp. nov." docType="treatment" docUuid="52FB2021-A86E-49BD-87EA-F77EC0F32755" docUuidSource="ZooBank" docVersion="4" lastPageNumber="164" masterDocId="2C1FF853EE3AA27EFFFBA404FFE5FF98" masterDocTitle="Species of Heterolepismatinae (Zygentoma: Lepismatidae) Found on some Remote Eastern Australian Islands" masterLastPageNumber="181" masterPageNumber="139" pageNumber="158" updateTime="1698898337455" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="3F66D89DE87ECAAE3B3F75AF3148A30C">
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<mods:title id="A3AD8AF6DD000D56DA3E4C681F5E7192">Species of Heterolepismatinae (Zygentoma: Lepismatidae) Found on some Remote Eastern Australian Islands</mods:title>
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<mods:namePart id="1F022BF299AAAB428D0FFBFC8EE13924">Smith, Graeme B.</mods:namePart>
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<mods:namePart id="DA7EB67FE47E0F2994F761607CC40488">Mitchell, Andrew</mods:namePart>
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<mods:typeOfResource id="90FE58A17FDE70878E650851ED72D361">text</mods:typeOfResource>
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<mods:title id="2DEDE969AE061477B014EA22EE2802D1">Records of the Australian Museum</mods:title>
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<mods:part id="86AC62E233AF449A4F5F452A90E3BF61">
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<mods:date id="76EC626BFF7E7A11CDBFF95275056C24">2019</mods:date>
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<mods:number id="13AAE66FBDCC09E022AECB572A552023">2019-07-24</mods:number>
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<mods:number id="9BE44CDC160F213EFD154B05779F36E9">71</mods:number>
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<mods:number id="A82C7BFF65EF85FB6565E583CDF95165">4</mods:number>
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<mods:start id="8F9121D400E12AAEA2D349C64AAB0647">139</mods:start>
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<mods:classification id="368F77FC0BBDECCC30564177035FA0F2">journal article</mods:classification>
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<mods:identifier id="FE72C7C6AFB1C0F381838FA18379F16A" type="DOI">10.3853/j.2201-4349.71.2019.1719</mods:identifier>
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<mods:identifier id="A42323C9ABA745852716C43106E9BB69" type="ISSN">2201-4349</mods:identifier>
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<mods:identifier id="A3436B6001A82DE2F7C96EF6939AE44A" type="Zenodo-Dep">4653485</mods:identifier>
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<mods:identifier id="521E837F0550265D634CA6335FA77C4D" type="ZooBank">CB380F40-F8D8-471B-86B5-E567C8B8EEC9</mods:identifier>
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</mods:mods>
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<treatment id="D026802BEE29A267FCAFA584FDEAFC7F" LSID="urn:lsid:zoobank.org:act:52FB2021-A86E-49BD-87EA-F77EC0F32755" httpUri="http://treatment.plazi.org/id/D026802BEE29A267FCAFA584FDEAFC7F" lastPageId="25" lastPageNumber="164" pageId="19" pageNumber="158">
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<subSubSection id="109562B6EE29A26DFCAFA584FAB5FE59" pageId="19" pageNumber="158" type="nomenclature">
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<paragraph id="5830313DEE29A26DFCAFA584FAB7FE07" blockId="19.[852,1362,384,415]" box="[852,1362,384,415]" pageId="19" pageNumber="158">
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<heading id="03788651EE29A26DFCAFA584FAB7FE07" bold="true" box="[852,1362,384,415]" centered="true" fontSize="12" level="2" pageId="19" pageNumber="158" reason="2">
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<emphasis id="6AFBED2FEE29A26DFCAFA584FAB7FE07" bold="true" box="[852,1362,384,415]" pageId="19" pageNumber="158">
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<taxonomicName id="9F8F4ABEEE29A26DFCAFA584FB3CFE07" authority="Smith & Mitchell, 2019" authorityName="Smith & Mitchell" authorityYear="2019" box="[852,1241,384,415]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="species" species="pyramidum" status="sp. nov.">
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<emphasis id="6AFBED2FEE29A26DFCAFA584FB3CFE07" bold="true" box="[852,1241,384,415]" italics="true" pageId="19" pageNumber="158">Heterolepisma pyramidum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="71C85054EE29A26DFB1AA584FAB7FE07" box="[1249,1362,384,415]" pageId="19" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="5830313DEE29A26DFCE1A5B6FAB5FE59" blockId="19.[794,1360,434,449]" box="[794,1360,434,449]" pageId="19" pageNumber="158">
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http://zoobank.org/NomenclaturalActs/
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<uuid id="2C290BE8EE29A26DFBE5A5B6FAB5FE59" box="[1054,1360,434,449]" pageId="19" pageNumber="158">52FB2021-A86E-49BD-87EA-F77EC0F32755</uuid>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="109562B6EE29A26DFBF2A5E5FB7BFE63" box="[1033,1182,481,507]" pageId="19" pageNumber="158" type="description">
|
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<paragraph id="5830313DEE29A26DFBF2A5E5FB7BFE63" blockId="19.[1033,1182,481,507]" box="[1033,1182,481,507]" pageId="19" pageNumber="158">
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<figureCitation id="C0B42DB8EE29A26DFBF2A5E5FB7BFE63" box="[1033,1182,481,507]" captionStart-0="Figures 86–98" captionStart-1="Figures 99–108" captionStart-2="Figures 109–122" captionStart-3="Figures 123–126" captionStartId-0="21.[121,196,1728,1750]" captionStartId-1="22.[166,243,1430,1452]" captionStartId-2="23.[121,198,1584,1606]" captionStartId-3="24.[166,243,781,803]" captionTargetBox-0="[121,1411,164,1712]" captionTargetBox-1="[173,1439,141,1416]" captionTargetBox-2="[129,1413,160,1568]" captionTargetBox-3="[183,1449,164,769]" captionTargetId-0="figure-47@21.[121,1421,151,1727]" captionTargetId-1="figure-8@22.[166,1466,130,1430]" captionTargetId-2="figure-10@23.[121,1421,151,1576]" captionTargetId-3="figure-8@24.[166,1466,151,771]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="24" captionText-0="Figures 86–98. Heterolepisma pyramidum sp.nov.holotype specimen K.261254 ♀ (86) habitus; (87) smooth,apically bifurcate macrochaeta of PI coxa; (88) carrot-shaped macrochaeta of PI tibia; (89) scale from mesonotum; (90) head; (91) antenna, pedicel and basal intervals of flagellum; (92) idem, most distal surviving interval showing basiconic sensilla type C (bsC); (93) mandible; (94) idem, incisor and molar regions; (95) maxilla, only more prominent setae of palp illustrated; (96) idem, lacinia and galea; (97) idem, apex of ultimate article of palp with its three branched papillae (bp); (98) labium. Scale bars = 0.1 mm unless otherwise indicated." captionText-1="Figures 99–108. Heterolepisma pyramidum sp. nov.holotype specimen K.261254 ♀ (99) ultimate article of labial palp showing basiconic sensillum (bsC); (100) pronotum, anterior and right margins; (101) idem, right anterior trichobothrial area; (102) idem, right posterior trichobothrial area; (103) idem, left posterior comb; (104) mesonotum, right side; (105) idem, trichobothrial areas, right side; (106) metanotum, right side; (107) idem, trichobothrial areas, right side; (108) presternum, prothoracic sternum and PI. Scale bars = 0.1 mm." captionText-2="Figures 109–122. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 or K.261255 ♀ (109) mesothoracic sternum; (110) metathoracic sternum and PIII; (111) PII; (112) urotergite III; (113) urotergite VII, left lateral comb; (114) idem, left sublateral lateral comb; (115) idem, left submedial comb; (116) urotergite IX, left infralateral comb; (117) urotergite X; (118) urosternite I; (119) idem, medial comb; (120) urosternite IV; (121) right comb of urosternite IV; (122) urosternite VII. Scale bars = 0.1 mm." captionText-3="Figures 123–126. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 ♀ (123) coxites VIII and IX, with styli and ovipositor; (124) apex of ovipositor valve; (125) base of cercus and medial dorsal appendage; (126) apex of cercus. Scale bars = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4653519" figureDoi-1="http://doi.org/10.5281/zenodo.4653523" figureDoi-2="http://doi.org/10.5281/zenodo.4653527" figureDoi-3="http://doi.org/10.5281/zenodo.4653529" httpUri-0="https://zenodo.org/record/4653519/files/figure.png" httpUri-1="https://zenodo.org/record/4653523/files/figure.png" httpUri-2="https://zenodo.org/record/4653527/files/figure.png" httpUri-3="https://zenodo.org/record/4653529/files/figure.png" pageId="19" pageNumber="158">Figs 86–126</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="109562B6EE29A26DFCE1A614FB78FD18" pageId="19" pageNumber="158" type="materials_examined">
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<paragraph id="5830313DEE29A26DFCE1A614FB78FD18" blockId="19.[794,1421,528,640]" pageId="19" pageNumber="158">
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<materialsCitation id="E8E73B60EE29A26DFCE1A614FB78FD18" collectingDate="2017-03-26" collectionCode="AM, LORD, HOWE, ISLAND" collectorName="Balls Pyramid & F. Koehler." latitude="-31.7575" longLatPrecision="6" longitude="159.255" pageId="19" pageNumber="158" specimenCode="HW 1.20" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<emphasis id="6AFBED2FEE29A26DFCE1A614FC62FDB0" bold="true" box="[794,903,528,552]" pageId="19" pageNumber="158">
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||
<typeStatus id="87348F9FEE29A26DFCE1A614FC62FDB0" box="[794,903,528,552]" pageId="19" pageNumber="158" type="holotype">Holotype</typeStatus>
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</emphasis>
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.
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<specimenCount id="4E89FAB4EE29A26DFC63A615FC48FDB0" box="[920,941,529,552]" pageId="19" pageNumber="158" type="female">♀</specimenCount>
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||
(
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<specimenCode id="08299946EE29A26DFC44A614FBC9FDB0" box="[959,1068,528,552]" collectionCode="HW" pageId="19" pageNumber="158">HW 1.20</specimenCode>
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||
) (
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||
<collectionCode id="3E9EA9F8EE29A26DFBBDA614FB91FDB0" box="[1094,1140,528,552]" country="Australia" name="Australian Museum" pageId="19" pageNumber="158" type="Museum">AM</collectionCode>
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K.261254 K.261255 on two slides)
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<collectionCode id="3E9EA9F8EE29A26DFC52A62AFB2AFDDE" box="[937,1231,558,582]" pageId="19" pageNumber="158">LORD HOWE ISLAND</collectionCode>
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||
:
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<collectorName id="F57A54EBEE29A26DFB26A62AFA6CFDDE" box="[1245,1417,558,582]" pageId="19" pageNumber="158">Balls Pyramid</collectorName>
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, rock platform at southern tip “Base Camp”,
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<geoCoordinate id="3DBB57FAEE29A26DFAE2A64FFA69FDFB" box="[1305,1420,587,611]" degrees="31.7575" direction="south" orientation="latitude" pageId="19" pageNumber="158" precision="5" value="-31.7575">31.7575°S</geoCoordinate>
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||
<geoCoordinate id="3DBB57FAEE29A26DFCE1A66CFC7EFD18" box="[794,923,616,640]" degrees="159.2550" direction="east" orientation="longitude" pageId="19" pageNumber="158" precision="5" value="159.255">159.2550°E</geoCoordinate>
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,
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<date id="2C3117FDEE29A26DFC5DA66CFBF9FD18" box="[934,1052,616,640]" pageId="19" pageNumber="158" value="2017-03-26">
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||
<collectingDate id="3C75EE15EE29A26DFC5DA66CFBF9FD18" box="[934,1052,616,640]" pageId="19" pageNumber="158" value="2017-03-26">26.iii.2017</collectingDate>
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</date>
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,
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<collectorName id="F57A54EBEE29A26DFBDCA66DFB78FD18" box="[1063,1181,616,640]" pageId="19" pageNumber="158">F. Koehler.</collectorName>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="109562B6EE29A26DFCE1A691FAEEFB38" pageId="19" pageNumber="158" type="diagnosis">
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<paragraph id="5830313DEE29A26DFCE1A691FAEEFB38" blockId="19.[794,1422,661,1184]" pageId="19" pageNumber="158">
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<emphasis id="6AFBED2FEE29A26DFCE1A691FC6DFD35" bold="true" box="[794,904,661,685]" pageId="19" pageNumber="158">Diagnosis</emphasis>
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. This species can be distinguished from many species of
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<taxonomicName id="9F8F4ABEEE29A26DFC75A6B7FBCFFD53" authorityName="Escherich. A" authorityYear="1905" box="[910,1066,691,715]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="genus">
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<emphasis id="6AFBED2FEE29A26DFC75A6B7FBCFFD53" box="[910,1066,691,715]" italics="true" pageId="19" pageNumber="158">Heterolepisma</emphasis>
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</taxonomicName>
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by the absence of macrochaetae along the anterior margin of the frons, by the presence of lanceolate scales on the clypeus, femora, tibia and terminal filaments and the presence of a medial comb on urosternite I. Within the remaining group of several species that definitely have these characters (
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<taxonomicName id="9F8F4ABEEE29A26DFB81A742FAE3FCC6" box="[1146,1286,838,862]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="species" species="bisetosum">
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<emphasis id="6AFBED2FEE29A26DFB81A742FAE3FCC6" box="[1146,1286,838,862]" italics="true" pageId="19" pageNumber="158">H. bisetosum</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="9F8F4ABEEE29A26DFAEFA742FA63FCC5" authorityName="Smith" authorityYear="2013" box="[1300,1414,838,861]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="species" species="parvum">
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<emphasis id="6AFBED2FEE29A26DFAEFA742FA63FCC5" box="[1300,1414,838,861]" italics="true" pageId="19" pageNumber="158">H. parvum</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="9F8F4ABEEE29A26DFCE1A760FC50FCE3" authority="Smith & Mitchell, 2019" authorityName="Smith & Mitchell" authorityYear="2019" box="[794,949,867,891]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="species" species="heraldense" status="sp. nov.">
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<emphasis id="6AFBED2FEE29A26DFCE1A760FC50FCE3" box="[794,949,867,891]" italics="true" pageId="19" pageNumber="158">H. heraldense</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="71C85054EE29A26DFC3BA760FBF2FCE3" box="[960,1047,868,891]" pageId="19" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="9F8F4ABEEE29A26DFBA9A760FB36FCE3" box="[1106,1235,868,891]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="species" species="serranoi">
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<emphasis id="6AFBED2FEE29A26DFBA9A760FB36FCE3" box="[1106,1235,868,891]" italics="true" pageId="19" pageNumber="158">H. serranoi</emphasis>
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</taxonomicName>
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) and others that probably have these characters (
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<taxonomicName id="9F8F4ABEEE29A26DFB92A785FB28FC00" box="[1129,1229,897,920]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="species" species="exacta">
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<emphasis id="6AFBED2FEE29A26DFB92A785FB28FC00" box="[1129,1229,897,920]" italics="true" pageId="19" pageNumber="158">H. exacta</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="9F8F4ABEEE29A26DFB22A785FA8DFC00" box="[1241,1384,897,920]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="species" species="japonicum">
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<emphasis id="6AFBED2FEE29A26DFB22A785FA8DFC00" box="[1241,1384,897,920]" italics="true" pageId="19" pageNumber="158">H. japonicum</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="9F8F4ABEEE29A26DFA88A785FC5AFC2E" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="species" species="mossambicense">
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<emphasis id="6AFBED2FEE29A26DFA88A785FC5AFC2E" italics="true" pageId="19" pageNumber="158">H. mossambicense</emphasis>
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||
</taxonomicName>
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,
|
||
<taxonomicName id="9F8F4ABEEE29A26DFC31A79AFBBEFC2E" box="[970,1115,926,950]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="species" species="primafrum">
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<emphasis id="6AFBED2FEE29A26DFC31A79AFBBEFC2E" box="[970,1115,926,950]" italics="true" pageId="19" pageNumber="158">H. primafrum</emphasis>
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||
</taxonomicName>
|
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and
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<taxonomicName id="9F8F4ABEEE29A26DFB70A79AFB04FC2D" box="[1163,1249,926,949]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="species" species="rouxi">
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<emphasis id="6AFBED2FEE29A26DFB70A79AFB04FC2D" box="[1163,1249,926,949]" italics="true" pageId="19" pageNumber="158">H. rouxi</emphasis>
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</taxonomicName>
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),
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<taxonomicName id="9F8F4ABEEE29A26DFB0EA79AFA68FC2E" authority="Smith & Mitchell, 2019" authorityName="Smith & Mitchell" authorityYear="2019" box="[1269,1421,926,950]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="species" species="pyramidum" status="sp. nov.">
|
||
<emphasis id="6AFBED2FEE29A26DFB0EA79AFA68FC2E" box="[1269,1421,926,950]" italics="true" pageId="19" pageNumber="158">H. pyramidum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71C85054EE29A26DFCE1A7B8FC88FC4B" box="[794,877,956,979]" pageId="19" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be distinguished from all by a combination of the number of macrochaetae in the posterior combs of the nota (one rather than two), the absence of a macrochaeta associated with the anterior trichobothrial area of the pronotum, the composition of the urosternal combs being of greater than a single macrochaeta, the arrangement of styli (three pair in the
|
||
<specimenCount id="4E89FAB4EE29A26DFC31A068FC3BFB1B" box="[970,990,1132,1155]" pageId="19" pageNumber="158" type="female">♀</specimenCount>
|
||
and only two pair in the
|
||
<specimenCount id="4E89FAB4EE29A26DFB25A06FFB17FB1B" box="[1246,1266,1131,1155]" pageId="19" pageNumber="158" type="male">♂</specimenCount>
|
||
) and the lesser number of combs on urotergite I (2+2 vs 3+3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="109562B6EE29A267FBF2A0C0FE9CFF54" lastPageId="25" lastPageNumber="164" pageId="19" pageNumber="158" type="description">
|
||
<paragraph id="5830313DEE29A26DFBF2A0C0FB78FB46" blockId="19.[1033,1181,1220,1246]" box="[1033,1181,1220,1246]" pageId="19" pageNumber="158">
|
||
<emphasis id="6AFBED2FEE29A26DFBF2A0C0FB78FB46" bold="true" box="[1033,1181,1220,1246]" pageId="19" pageNumber="158">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="5830313DEE29A26DFCE1A0F7FBF2FAFB" blockId="19.[794,1422,1267,1379]" pageId="19" pageNumber="158">
|
||
<emphasis id="6AFBED2FEE29A26DFCE1A0F7FC79FA92" box="[794,924,1267,1290]" italics="true" pageId="19" pageNumber="158">Appearance</emphasis>
|
||
: Medium sized silverfish, thorax about 15% wider than abdomen which only tapers slightly posteriorly from about the fifth abdominal segment; appearance when live unknown (
|
||
<figureCitation id="C0B42DB8EE29A26DFC41A14FFBEFFAFB" box="[954,1034,1355,1379]" captionStart="Figures 86–98" captionStartId="21.[121,196,1728,1750]" captionTargetBox="[121,1411,164,1712]" captionTargetId="figure-47@21.[121,1421,151,1727]" captionTargetPageId="21" captionText="Figures 86–98. Heterolepisma pyramidum sp.nov.holotype specimen K.261254 ♀ (86) habitus; (87) smooth,apically bifurcate macrochaeta of PI coxa; (88) carrot-shaped macrochaeta of PI tibia; (89) scale from mesonotum; (90) head; (91) antenna, pedicel and basal intervals of flagellum; (92) idem, most distal surviving interval showing basiconic sensilla type C (bsC); (93) mandible; (94) idem, incisor and molar regions; (95) maxilla, only more prominent setae of palp illustrated; (96) idem, lacinia and galea; (97) idem, apex of ultimate article of palp with its three branched papillae (bp); (98) labium. Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4653519" httpUri="https://zenodo.org/record/4653519/files/figure.png" pageId="19" pageNumber="158">Fig. 86</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5830313DEE29A26DFCE1A17CFA99F9BB" blockId="19.[794,1422,1400,1571]" pageId="19" pageNumber="158">
|
||
<emphasis id="6AFBED2FEE29A26DFCE1A17CFC78FA08" box="[794,925,1400,1424]" italics="true" pageId="19" pageNumber="158">Body length</emphasis>
|
||
: H+B
|
||
<quantity id="9F779CD8EE29A26DFC11A17CFBA7FA08" box="[1002,1090,1400,1424]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="19" pageNumber="158" unit="mm" value="8.9">8.9 mm</quantity>
|
||
; HW
|
||
<quantity id="9F779CD8EE29A26DFB7FA17CFB0CFA08" box="[1156,1257,1400,1424]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="158" unit="mm" value="1.2">1.20 mm</quantity>
|
||
; thorax: length
|
||
<quantity id="9F779CD8EE29A26DFCE1A192FC89FA35" box="[794,876,1430,1454]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="158" unit="mm" value="2.5">2.5 mm</quantity>
|
||
or 0.28 H+B; width up to
|
||
<quantity id="9F779CD8EE29A26DFB6FA192FB12FA35" box="[1172,1271,1430,1454]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="19" pageNumber="158" unit="mm" value="1.85">1.85 mm</quantity>
|
||
, widest at the mesonotum, pronotum and mesonotum slightly longer than the metanotum; antennae almost complete at
|
||
<quantity id="9F779CD8EE29A26DFB14A1D4FADBFA70" box="[1263,1342,1488,1512]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="19" pageNumber="158" unit="mm" value="5.3">5.3 mm</quantity>
|
||
or 0.59 H+B; cerci and median dorsal appendage almost complete at
|
||
<quantity id="9F779CD8EE29A26DFCCFA20FFC60F9BB" box="[820,901,1547,1571]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="19" pageNumber="158" unit="mm" value="5.2">5.2 mm</quantity>
|
||
and
|
||
<quantity id="9F779CD8EE29A26DFC42A20FFBEFF9BB" box="[953,1034,1547,1571]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="19" pageNumber="158" unit="mm" value="5.9">5.9 mm</quantity>
|
||
(0.58 and 0.67 H+B) respectively.
|
||
</paragraph>
|
||
<paragraph id="5830313DEE29A26DFCE1A23DFA6BF9F6" blockId="19.[794,1422,1592,1646]" pageId="19" pageNumber="158">
|
||
<emphasis id="6AFBED2FEE29A26DFCE1A23DFC4EF9C8" box="[794,939,1593,1616]" italics="true" pageId="19" pageNumber="158">Pigmentation</emphasis>
|
||
: Pigment brownish. Flagellum of antennae evenly light brown becoming slightly darker distally; pedicel
|
||
</paragraph>
|
||
<caption id="0CF061B5EE29A26DFF4CA2AEFAABF95A" box="[183,1358,1705,1730]" pageId="19" pageNumber="158">
|
||
<paragraph id="5830313DEE29A26DFF4CA2AEFAABF95A" blockId="19.[183,1358,1705,1759]" box="[183,1358,1705,1730]" pageId="19" pageNumber="158">
|
||
<emphasis id="6AFBED2FEE29A26DFF4CA2AEFEE2F959" bold="true" box="[183,263,1705,1730]" pageId="19" pageNumber="158">Table 6</emphasis>
|
||
. Selected measurement ratios showing a difference between
|
||
<taxonomicName id="9F8F4ABEEE29A26DFC87A2AEFB4AF95A" box="[892,1199,1706,1730]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="species" species="sclerophyllum">
|
||
<emphasis id="6AFBED2FEE29A26DFC87A2AEFB4AF95A" box="[892,1199,1706,1730]" italics="true" pageId="19" pageNumber="158">Heterolepisma sclerophyllum</emphasis>
|
||
</taxonomicName>
|
||
(all genotypes)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5830313DEE29A26AFF4CA2C3FDC0FD63" blockId="19.[183,1358,1705,1759]" lastBlockId="20.[166,795,152,763]" lastPageId="20" lastPageNumber="159" pageId="19" pageNumber="158">
|
||
and
|
||
<taxonomicName id="9F8F4ABEEE29A26DFF1EA2CCFE8DF947" authority="Smith & Mitchell, 2019" authorityName="Smith & Mitchell" authorityYear="2019" box="[229,360,1735,1759]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="species" species="milledgei" status="sp. nov.">
|
||
<emphasis id="6AFBED2FEE29A26DFF1EA2CCFE8DF947" box="[229,360,1735,1759]" italics="true" pageId="19" pageNumber="158">H. milledgei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71C85054EE29A26DFE94A2C3FE27F946" box="[367,450,1735,1758]" pageId="19" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Number of specimens within parentheses. without pigment, scape with small patch of pigment above as well as on its insertion pedestal. Terminal filaments annulated darker brown with only the portion of each major division distad of the rosette of larger macrochaetae unpigmented (e.g., the distal end of every eighth subdivision distally). Head with pigment around eyes and along lateral margins to the level of the perpendicular band of macrochaetae. Maxillary palp with light pigment on distal article, penultimate article and third article with even pigmentation, but much less on the second article. Ultimate article of labial palp with light pigment along lateral margins, penultimate article more pigmented, especially laterally and a small amount of pigment on external margin of second article. Precoxae with pigment and dark scales, coxae with light pigment only along lateral margins, trochanter without pigment, femur with small area of pigment distally on the leading edge as well as along the posterior margin, tibia with pigment over much of surface but darker on the dorsal margin, first tarsal article pigmented. Urotergite X and coxites IX pigmented. Styli IX with light pigment distributed evenly. Ovipositor white or perhaps slightly yellowish.
|
||
</paragraph>
|
||
<paragraph id="5830313DEE29A26DFDB1A2FEFB8AF852" pageId="19" pageNumber="158">
|
||
<table id="2A8FC39DEE295D81FF34A2FEFAD0F852" box="[207,1333,1786,1994]" gridcols="5" gridrows="6" pageId="19" pageNumber="158">
|
||
<tr id="E6BF337FEE295D81FF34A2FEFAD0F88A" box="[207,1333,1786,1810]" gridrow="0" pageId="19" pageNumber="158" rowspan-0="1">
|
||
<th id="A56E5A03EE295D81FDB1A2FEFCA2F88A" box="[586,839,1786,1810]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="19" pageNumber="158">
|
||
<taxonomicName id="9F8F4ABEEE29A26DFDB1A2FEFCA2F88A" authority="Smith" box="[586,839,1786,1810]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="species" species="sclerophyllum">
|
||
<emphasis id="6AFBED2FEE29A26DFDB1A2FEFCE4F88A" box="[586,769,1786,1810]" italics="true" pageId="19" pageNumber="158">H. sclerophyllum</emphasis>
|
||
Smith
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="A56E5A03EE295D81FC5CA2FEFAD0F88A" box="[935,1333,1786,1810]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="19" pageNumber="158">
|
||
<taxonomicName id="9F8F4ABEEE29A26DFC5CA2FEFBCEF88A" authority="Smith & Mitchell, 2019" authorityName="Smith & Mitchell" authorityYear="2019" box="[935,1067,1786,1810]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="19" pageNumber="158" phylum="Arthropoda" rank="species" species="milledgei" status="sp. nov.">
|
||
<emphasis id="6AFBED2FEE29A26DFC5CA2FEFBCEF88A" box="[935,1067,1786,1810]" italics="true" pageId="19" pageNumber="158">H. milledgei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71C85054EE29A26DFBC9A2FEFB60F889" box="[1074,1157,1786,1809]" pageId="19" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
</tr>
|
||
<tr id="E6BF337FEE295D81FF34A313FAD0F8B7" box="[207,1333,1815,1839]" gridrow="1" pageId="19" pageNumber="158" rowspan-0="1">
|
||
<td id="A56E5A03EE295D81FDB1A313FCA2F8B7" box="[586,839,1815,1839]" colspan="2" colspanRight="1" gridcol="1" gridrow="1" pageId="19" pageNumber="158">NSW all genotypes</td>
|
||
<td id="A56E5A03EE295D81FC5CA313FAD0F8B7" box="[935,1333,1815,1839]" colspan="2" colspanRight="1" gridcol="3" gridrow="1" pageId="19" pageNumber="158">Lord Howe Island and Balls Pyramid</td>
|
||
</tr>
|
||
<tr id="E6BF337FEE295D81FF34A34EFAD0F8FA" box="[207,1333,1866,1890]" gridrow="2" pageId="19" pageNumber="158">
|
||
<th id="A56E5A03EE295D81FF34A34EFDE9F8FA" box="[207,524,1866,1890]" gridcol="0" gridrow="2" pageId="19" pageNumber="158">measurement parameter</th>
|
||
<td id="A56E5A03EE295D81FDB1A34EFD49F8FA" box="[586,684,1866,1890]" gridcol="1" gridrow="2" pageId="19" pageNumber="158">mean</td>
|
||
<td id="A56E5A03EE295D81FD21A34EFCA2F8FA" box="[730,839,1866,1890]" gridcol="2" gridrow="2" pageId="19" pageNumber="158">range</td>
|
||
<td id="A56E5A03EE295D81FC5CA34EFC19F8FA" box="[935,1020,1866,1890]" gridcol="3" gridrow="2" pageId="19" pageNumber="158">mean</td>
|
||
<td id="A56E5A03EE295D81FBD6A34EFAD0F8FA" box="[1069,1333,1866,1890]" gridcol="4" gridrow="2" pageId="19" pageNumber="158">range</td>
|
||
</tr>
|
||
<tr id="E6BF337FEE295D81FF34A373FAD0F817" box="[207,1333,1911,1935]" gridrow="3" pageId="19" pageNumber="158">
|
||
<th id="A56E5A03EE295D81FF34A373FDE9F817" box="[207,524,1911,1935]" gridcol="0" gridrow="3" pageId="19" pageNumber="158">metasternum L gap/L combs</th>
|
||
<td id="A56E5A03EE295D81FDB1A373FD49F817" box="[586,684,1911,1935]" gridcol="1" gridrow="3" pageId="19" pageNumber="158">2.68 (44)</td>
|
||
<td id="A56E5A03EE295D81FD21A373FCA2F817" box="[730,839,1911,1935]" gridcol="2" gridrow="3" pageId="19" pageNumber="158">1.57–4.11</td>
|
||
<td id="A56E5A03EE295D81FC5CA373FC19F817" box="[935,1020,1911,1935]" gridcol="3" gridrow="3" pageId="19" pageNumber="158">3.80 (5)</td>
|
||
<td id="A56E5A03EE295D81FBD6A373FAD0F817" box="[1069,1333,1911,1935]" gridcol="4" gridrow="3" pageId="19" pageNumber="158">2.84–4.92</td>
|
||
</tr>
|
||
<tr id="E6BF337FEE295D81FF34A390FAD0F834" box="[207,1333,1940,1964]" gridrow="4" pageId="19" pageNumber="158">
|
||
<th id="A56E5A03EE295D81FF34A390FDE9F834" box="[207,524,1940,1964]" gridcol="0" gridrow="4" pageId="19" pageNumber="158">urotergite X (L/W)</th>
|
||
<td id="A56E5A03EE295D81FDB1A390FD49F834" box="[586,684,1940,1964]" gridcol="1" gridrow="4" pageId="19" pageNumber="158">0.57 (29)</td>
|
||
<td id="A56E5A03EE295D81FD21A390FCA2F834" box="[730,839,1940,1964]" gridcol="2" gridrow="4" pageId="19" pageNumber="158">0.43–0.73</td>
|
||
<td id="A56E5A03EE295D81FC5CA390FC19F834" box="[935,1020,1940,1964]" gridcol="3" gridrow="4" pageId="19" pageNumber="158">0.39 (5)</td>
|
||
<td id="A56E5A03EE295D81FBD6A390FAD0F834" box="[1069,1333,1940,1964]" gridcol="4" gridrow="4" pageId="19" pageNumber="158">0.36–0.47</td>
|
||
</tr>
|
||
<tr id="E6BF337FEE295D81FF34A3B6FAD0F852" box="[207,1333,1970,1994]" gridrow="5" pageId="19" pageNumber="158">
|
||
<th id="A56E5A03EE295D81FF34A3B6FDE9F852" box="[207,524,1970,1994]" gridcol="0" gridrow="5" pageId="19" pageNumber="158">ovipositor divisions (number)</th>
|
||
<td id="A56E5A03EE295D81FDB1A3B6FD49F852" box="[586,684,1970,1994]" gridcol="1" gridrow="5" pageId="19" pageNumber="158">37.3 (17)</td>
|
||
<td id="A56E5A03EE295D81FD21A3B6FCA2F852" box="[730,839,1970,1994]" gridcol="2" gridrow="5" pageId="19" pageNumber="158">32–41</td>
|
||
<td id="A56E5A03EE295D81FC5CA3B6FC19F852" box="[935,1020,1970,1994]" gridcol="3" gridrow="5" pageId="19" pageNumber="158">40.2 (2)</td>
|
||
<td id="A56E5A03EE295D81FBD6A3B6FAD0F852" box="[1069,1333,1970,1994]" gridcol="4" gridrow="5" pageId="19" pageNumber="158">39–42</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="5830313DEE2EA26AFF5DA70AFE47FCE6" blockId="20.[166,794,782,894]" pageId="20" pageNumber="159">
|
||
<emphasis id="6AFBED2FEE2EA26AFF5DA70AFEAEFCBE" box="[166,331,782,806]" italics="true" pageId="20" pageNumber="159">Macrochaetae</emphasis>
|
||
: Smooth, hyaline to straw-coloured to moderate brown, apically bifurcate with truncated tips to each bifurcation (
|
||
<figureCitation id="C0B42DB8EE2EA26AFEA0A74CFE42FCF8" box="[347,423,840,864]" captionStart="Figures 86–98" captionStartId="21.[121,196,1728,1750]" captionTargetBox="[121,1411,164,1712]" captionTargetId="figure-47@21.[121,1421,151,1727]" captionTargetPageId="21" captionText="Figures 86–98. Heterolepisma pyramidum sp.nov.holotype specimen K.261254 ♀ (86) habitus; (87) smooth,apically bifurcate macrochaeta of PI coxa; (88) carrot-shaped macrochaeta of PI tibia; (89) scale from mesonotum; (90) head; (91) antenna, pedicel and basal intervals of flagellum; (92) idem, most distal surviving interval showing basiconic sensilla type C (bsC); (93) mandible; (94) idem, incisor and molar regions; (95) maxilla, only more prominent setae of palp illustrated; (96) idem, lacinia and galea; (97) idem, apex of ultimate article of palp with its three branched papillae (bp); (98) labium. Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4653519" httpUri="https://zenodo.org/record/4653519/files/figure.png" pageId="20" pageNumber="159">Fig. 87</figureCitation>
|
||
). Some macrochaetae on tibia, stout carrot-shaped (
|
||
<figureCitation id="C0B42DB8EE2EA26AFEBDA762FE73FCE6" box="[326,406,870,894]" captionStart="Figures 86–98" captionStartId="21.[121,196,1728,1750]" captionTargetBox="[121,1411,164,1712]" captionTargetId="figure-47@21.[121,1421,151,1727]" captionTargetPageId="21" captionText="Figures 86–98. Heterolepisma pyramidum sp.nov.holotype specimen K.261254 ♀ (86) habitus; (87) smooth,apically bifurcate macrochaeta of PI coxa; (88) carrot-shaped macrochaeta of PI tibia; (89) scale from mesonotum; (90) head; (91) antenna, pedicel and basal intervals of flagellum; (92) idem, most distal surviving interval showing basiconic sensilla type C (bsC); (93) mandible; (94) idem, incisor and molar regions; (95) maxilla, only more prominent setae of palp illustrated; (96) idem, lacinia and galea; (97) idem, apex of ultimate article of palp with its three branched papillae (bp); (98) labium. Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4653519" httpUri="https://zenodo.org/record/4653519/files/figure.png" pageId="20" pageNumber="159">Fig. 88</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5830313DEE2EA26AFF5DA794FDA4FB56" blockId="20.[166,794,912,1230]" pageId="20" pageNumber="159">
|
||
<emphasis id="6AFBED2FEE2EA26AFF5DA794FF0FFC30" box="[166,234,912,936]" italics="true" pageId="20" pageNumber="159">Scales</emphasis>
|
||
: Quite broad, brown with numerous subparallel ribs that do not surpass the margin of the scale (
|
||
<figureCitation id="C0B42DB8EE2EA26AFD88A7AAFD26FC5E" box="[627,707,942,966]" captionStart="Figures 86–98" captionStartId="21.[121,196,1728,1750]" captionTargetBox="[121,1411,164,1712]" captionTargetId="figure-47@21.[121,1421,151,1727]" captionTargetPageId="21" captionText="Figures 86–98. Heterolepisma pyramidum sp.nov.holotype specimen K.261254 ♀ (86) habitus; (87) smooth,apically bifurcate macrochaeta of PI coxa; (88) carrot-shaped macrochaeta of PI tibia; (89) scale from mesonotum; (90) head; (91) antenna, pedicel and basal intervals of flagellum; (92) idem, most distal surviving interval showing basiconic sensilla type C (bsC); (93) mandible; (94) idem, incisor and molar regions; (95) maxilla, only more prominent setae of palp illustrated; (96) idem, lacinia and galea; (97) idem, apex of ultimate article of palp with its three branched papillae (bp); (98) labium. Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4653519" httpUri="https://zenodo.org/record/4653519/files/figure.png" pageId="20" pageNumber="159">Fig. 89</figureCitation>
|
||
). Scales found on top of head, some, either side of the tiny medial setae protruding well beyond the anterior margin; scales absent from clypeus and labrum as well as all cephalic appendages; dark scales present on all nota and abdominal tergites, clear scales on all thoracic and abdominal sterna, and coxae of legs, absent from styli. Lanceolate scales present on all femora and tibia; other leg articles without scales; lanceolate scales also present on basal third of the terminal filaments (except basal few divisions).
|
||
</paragraph>
|
||
<paragraph id="5830313DEE2EA26AFF5DA0E7FB12FB9B" blockId="20.[166,795,1251,2008]" lastBlockId="20.[839,1467,152,1027]" pageId="20" pageNumber="159">
|
||
<emphasis id="6AFBED2FEE2EA26AFF5DA0E7FF05FB63" box="[166,224,1251,1275]" italics="true" pageId="20" pageNumber="159">Head</emphasis>
|
||
: Wider than long (
|
||
<figureCitation id="C0B42DB8EE2EA26AFE4AA0E7FDE1FB63" box="[433,516,1251,1275]" captionStart="Figures 86–98" captionStartId="21.[121,196,1728,1750]" captionTargetBox="[121,1411,164,1712]" captionTargetId="figure-47@21.[121,1421,151,1727]" captionTargetPageId="21" captionText="Figures 86–98. Heterolepisma pyramidum sp.nov.holotype specimen K.261254 ♀ (86) habitus; (87) smooth,apically bifurcate macrochaeta of PI coxa; (88) carrot-shaped macrochaeta of PI tibia; (89) scale from mesonotum; (90) head; (91) antenna, pedicel and basal intervals of flagellum; (92) idem, most distal surviving interval showing basiconic sensilla type C (bsC); (93) mandible; (94) idem, incisor and molar regions; (95) maxilla, only more prominent setae of palp illustrated; (96) idem, lacinia and galea; (97) idem, apex of ultimate article of palp with its three branched papillae (bp); (98) labium. Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4653519" httpUri="https://zenodo.org/record/4653519/files/figure.png" pageId="20" pageNumber="159">Fig. 90</figureCitation>
|
||
), without distinct bushes. Anterior margin without macrochaetae except for four tiny setae in the middle; strong macrochaetae along lateral margins with peri-antennal groups of about seven macrochaetae which are contiguous laterally with the rows of macrochaetae along the margin, the marginal macrochaetae continue along the sides of the head behind the antennae with a line of four or five macrochaetae lying above the eyes. Clypeus with some longer macrochaetae and several setae extending across the face. Labrum with many setae across the proximal half and a line of six thinner setae anteriorly. — Scape quite short with a subdistal line of setae and smaller setae over the dorsal anterior face, pedicel (
|
||
<figureCitation id="C0B42DB8EE2EA26AFD3EA247FCF4F9C3" box="[709,785,1603,1627]" captionStart="Figures 86–98" captionStartId="21.[121,196,1728,1750]" captionTargetBox="[121,1411,164,1712]" captionTargetId="figure-47@21.[121,1421,151,1727]" captionTargetPageId="21" captionText="Figures 86–98. Heterolepisma pyramidum sp.nov.holotype specimen K.261254 ♀ (86) habitus; (87) smooth,apically bifurcate macrochaeta of PI coxa; (88) carrot-shaped macrochaeta of PI tibia; (89) scale from mesonotum; (90) head; (91) antenna, pedicel and basal intervals of flagellum; (92) idem, most distal surviving interval showing basiconic sensilla type C (bsC); (93) mandible; (94) idem, incisor and molar regions; (95) maxilla, only more prominent setae of palp illustrated; (96) idem, lacinia and galea; (97) idem, apex of ultimate article of palp with its three branched papillae (bp); (98) labium. Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4653519" httpUri="https://zenodo.org/record/4653519/files/figure.png" pageId="20" pageNumber="159">Fig. 91</figureCitation>
|
||
) short, also with a subapical rosette of small macrochaetae with setae scattered over the surface medially; first annulus of flagellum apparently glabrous, subsequent annuli with a rosette of small setae and some short trichobothria, the divisions between the annuli difficult to discern in the slide mounted material, the intervals split into two annuli by the 7th interval, into four by the 12th and into eight by the 17th. Most distal surviving intervals (
|
||
<figureCitation id="C0B42DB8EE2EA26AFDC0A32AFD6FF8DE" box="[571,650,1838,1862]" captionStart="Figures 86–98" captionStartId="21.[121,196,1728,1750]" captionTargetBox="[121,1411,164,1712]" captionTargetId="figure-47@21.[121,1421,151,1727]" captionTargetPageId="21" captionText="Figures 86–98. Heterolepisma pyramidum sp.nov.holotype specimen K.261254 ♀ (86) habitus; (87) smooth,apically bifurcate macrochaeta of PI coxa; (88) carrot-shaped macrochaeta of PI tibia; (89) scale from mesonotum; (90) head; (91) antenna, pedicel and basal intervals of flagellum; (92) idem, most distal surviving interval showing basiconic sensilla type C (bsC); (93) mandible; (94) idem, incisor and molar regions; (95) maxilla, only more prominent setae of palp illustrated; (96) idem, lacinia and galea; (97) idem, apex of ultimate article of palp with its three branched papillae (bp); (98) labium. Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4653519" httpUri="https://zenodo.org/record/4653519/files/figure.png" pageId="20" pageNumber="159">Fig. 92</figureCitation>
|
||
) divided into repeated patterns of eight annuli, each with two rosettes of setae, those basally long tapering, those distally thinner and almost parallel-sided, the apical article of each interval has at least one trichobothrium, the second and fourth annuli bear at least one long, sausage-shaped basiconic sensillum (
|
||
<typeStatus id="87348F9FEE2EA26AFCB5A49CFC60FF28" box="[846,901,152,176]" pageId="20" pageNumber="159">Type</typeStatus>
|
||
C of
|
||
<bibRefCitation id="3C1E4CCCEE2EA26AFC44A49CFBDFFF28" author="Adel, T." box="[959,1082,152,176]" pageId="20" pageNumber="159" pagination="191 - 217" refId="ref28396" refString="Adel, T. 1984. Sensilleninventar und sensillenmuster auf den Antennen von Thermobia domestica und Lepisma saccharina (Insecta: Zygentoma). Braunschweiger Naturkundliche Schriften 2: 191 - 217." type="journal article" year="1984">Adel, 1984</bibRefCitation>
|
||
). — Mandibles (
|
||
<figureCitation id="C0B42DB8EE2EA26AFB11A49CFA81FF28" box="[1258,1380,152,176]" captionStart="Figures 86–98" captionStartId="21.[121,196,1728,1750]" captionTargetBox="[121,1411,164,1712]" captionTargetId="figure-47@21.[121,1421,151,1727]" captionTargetPageId="21" captionText="Figures 86–98. Heterolepisma pyramidum sp.nov.holotype specimen K.261254 ♀ (86) habitus; (87) smooth,apically bifurcate macrochaeta of PI coxa; (88) carrot-shaped macrochaeta of PI tibia; (89) scale from mesonotum; (90) head; (91) antenna, pedicel and basal intervals of flagellum; (92) idem, most distal surviving interval showing basiconic sensilla type C (bsC); (93) mandible; (94) idem, incisor and molar regions; (95) maxilla, only more prominent setae of palp illustrated; (96) idem, lacinia and galea; (97) idem, apex of ultimate article of palp with its three branched papillae (bp); (98) labium. Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4653519" httpUri="https://zenodo.org/record/4653519/files/figure.png" pageId="20" pageNumber="159">Figs 93, 94</figureCitation>
|
||
) typical for
|
||
<taxonomicName id="9F8F4ABEEE2EA26AFC89A4B2FBAEFF56" authorityName="Mendes" authorityYear="1991" box="[882,1099,182,206]" class="Insecta" family="Lepismatidae" kingdom="Animalia" order="Zygentoma" pageId="20" pageNumber="159" phylum="Arthropoda" rank="subFamily" subFamily="Heterolepismatinae">Heterolepismatinae</taxonomicName>
|
||
with well-developed molar and incisor regions; a group of about six strong and short plus five thinner, apically bifurcated setae distally adjacent to the molar region and a bush of about 40 macrochaetae externally. — Maxilla (
|
||
<figureCitation id="C0B42DB8EE2EA26AFBC0A52FFB62FEDB" box="[1083,1159,299,323]" captionStart="Figures 86–98" captionStartId="21.[121,196,1728,1750]" captionTargetBox="[121,1411,164,1712]" captionTargetId="figure-47@21.[121,1421,151,1727]" captionTargetPageId="21" captionText="Figures 86–98. Heterolepisma pyramidum sp.nov.holotype specimen K.261254 ♀ (86) habitus; (87) smooth,apically bifurcate macrochaeta of PI coxa; (88) carrot-shaped macrochaeta of PI tibia; (89) scale from mesonotum; (90) head; (91) antenna, pedicel and basal intervals of flagellum; (92) idem, most distal surviving interval showing basiconic sensilla type C (bsC); (93) mandible; (94) idem, incisor and molar regions; (95) maxilla, only more prominent setae of palp illustrated; (96) idem, lacinia and galea; (97) idem, apex of ultimate article of palp with its three branched papillae (bp); (98) labium. Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4653519" httpUri="https://zenodo.org/record/4653519/files/figure.png" pageId="20" pageNumber="159">Fig. 95</figureCitation>
|
||
) with three apically bifurcate macrochaetae externally proximal to the palp; lacinia short and wide, with three strong teeth, one set further back than the other two, followed by about seven lamellate processes and a row of seven setae, galea with only short fine setulae (
|
||
<figureCitation id="C0B42DB8EE2EA26AFCB4A5BAFC7BFE4E" box="[847,926,446,470]" captionStart="Figures 86–98" captionStartId="21.[121,196,1728,1750]" captionTargetBox="[121,1411,164,1712]" captionTargetId="figure-47@21.[121,1421,151,1727]" captionTargetPageId="21" captionText="Figures 86–98. Heterolepisma pyramidum sp.nov.holotype specimen K.261254 ♀ (86) habitus; (87) smooth,apically bifurcate macrochaeta of PI coxa; (88) carrot-shaped macrochaeta of PI tibia; (89) scale from mesonotum; (90) head; (91) antenna, pedicel and basal intervals of flagellum; (92) idem, most distal surviving interval showing basiconic sensilla type C (bsC); (93) mandible; (94) idem, incisor and molar regions; (95) maxilla, only more prominent setae of palp illustrated; (96) idem, lacinia and galea; (97) idem, apex of ultimate article of palp with its three branched papillae (bp); (98) labium. Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4653519" httpUri="https://zenodo.org/record/4653519/files/figure.png" pageId="20" pageNumber="159">Fig. 96</figureCitation>
|
||
); apical article of maxillary palp (
|
||
<figureCitation id="C0B42DB8EE2EA26AFAFBA5BAFAA8FE4E" box="[1280,1357,446,470]" captionStart="Figures 86–98" captionStartId="21.[121,196,1728,1750]" captionTargetBox="[121,1411,164,1712]" captionTargetId="figure-47@21.[121,1421,151,1727]" captionTargetPageId="21" captionText="Figures 86–98. Heterolepisma pyramidum sp.nov.holotype specimen K.261254 ♀ (86) habitus; (87) smooth,apically bifurcate macrochaeta of PI coxa; (88) carrot-shaped macrochaeta of PI tibia; (89) scale from mesonotum; (90) head; (91) antenna, pedicel and basal intervals of flagellum; (92) idem, most distal surviving interval showing basiconic sensilla type C (bsC); (93) mandible; (94) idem, incisor and molar regions; (95) maxilla, only more prominent setae of palp illustrated; (96) idem, lacinia and galea; (97) idem, apex of ultimate article of palp with its three branched papillae (bp); (98) labium. Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4653519" httpUri="https://zenodo.org/record/4653519/files/figure.png" pageId="20" pageNumber="159">Fig. 97</figureCitation>
|
||
) 4.6 times longer than wide and 1.2 times longer than the penultimate article, the ultimate article with three branched papillae, other specialised sensilla (e.g., basiconic sensilla) not seen, last three articles of palp with fine setae only, two basal articles with subapical rosettes of thicker setae. — Labium (
|
||
<figureCitation id="C0B42DB8EE2EA26AFA9CA654FA56FDF0" box="[1383,1459,592,616]" captionStart="Figures 86–98" captionStartId="21.[121,196,1728,1750]" captionTargetBox="[121,1411,164,1712]" captionTargetId="figure-47@21.[121,1421,151,1727]" captionTargetPageId="21" captionText="Figures 86–98. Heterolepisma pyramidum sp.nov.holotype specimen K.261254 ♀ (86) habitus; (87) smooth,apically bifurcate macrochaeta of PI coxa; (88) carrot-shaped macrochaeta of PI tibia; (89) scale from mesonotum; (90) head; (91) antenna, pedicel and basal intervals of flagellum; (92) idem, most distal surviving interval showing basiconic sensilla type C (bsC); (93) mandible; (94) idem, incisor and molar regions; (95) maxilla, only more prominent setae of palp illustrated; (96) idem, lacinia and galea; (97) idem, apex of ultimate article of palp with its three branched papillae (bp); (98) labium. Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4653519" httpUri="https://zenodo.org/record/4653519/files/figure.png" pageId="20" pageNumber="159">Fig. 98</figureCitation>
|
||
) wider than long, postmentum with setae scattered in band across entire width including several small setae along and near the anterolateral margins, prementum with transverse and oblique groups of strong setae and with short setulae distally; apical article of labial palp (
|
||
<figureCitation id="C0B42DB8EE2EA26AFB32A6E7FAFDFD63" box="[1225,1304,739,763]" captionStart="Figures 99–108" captionStartId="22.[166,243,1430,1452]" captionTargetBox="[173,1439,141,1416]" captionTargetId="figure-8@22.[166,1466,130,1430]" captionTargetPageId="22" captionText="Figures 99–108. Heterolepisma pyramidum sp. nov.holotype specimen K.261254 ♀ (99) ultimate article of labial palp showing basiconic sensillum (bsC); (100) pronotum, anterior and right margins; (101) idem, right anterior trichobothrial area; (102) idem, right posterior trichobothrial area; (103) idem, left posterior comb; (104) mesonotum, right side; (105) idem, trichobothrial areas, right side; (106) metanotum, right side; (107) idem, trichobothrial areas, right side; (108) presternum, prothoracic sternum and PI. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653523" httpUri="https://zenodo.org/record/4653523/files/figure.png" pageId="20" pageNumber="159">Fig. 99</figureCitation>
|
||
), about as wide as long (L/
|
||
<geoCoordinate id="3DBB57FAEE2EA26AFC40A704FBEAFC80" box="[955,1039,768,792]" degrees="0.95" direction="west" orientation="longitude" pageId="20" pageNumber="159" precision="555" value="-0.95">W 0.95</geoCoordinate>
|
||
), with five papillae of the compact
|
||
<typeStatus id="87348F9FEE2EA26AFA76A705FA5EFC81" box="[1421,1467,769,793]" pageId="20" pageNumber="159">type</typeStatus>
|
||
arranged in a broad three plus two cluster arrangement, the distal three arranged in a curved line, a single basiconic sensillum (
|
||
<typeStatus id="87348F9FEE2EA26AFC41A75CFC14FCE8" box="[954,1009,856,880]" pageId="20" pageNumber="159">Type</typeStatus>
|
||
C) on the outer margin near the level of the papillae, covered with numerous fine short setae especially below; penultimate article with some stronger setae on the bulge medially; second article also with stronger setae along the inner margin and a subapical rosette; basal article with an apical line of stronger setae ventrally.
|
||
</paragraph>
|
||
<paragraph id="5830313DEE2EA26AFCBCA012FB41F90D" blockId="20.[839,1467,1046,2008]" pageId="20" pageNumber="159">
|
||
<emphasis id="6AFBED2FEE2EA26AFCBCA012FC71FBB6" box="[839,916,1046,1070]" italics="true" pageId="20" pageNumber="159">Thorax</emphasis>
|
||
: Pronotum (
|
||
<figureCitation id="C0B42DB8EE2EA26AFBE7A012FB9DFBB6" box="[1052,1144,1046,1070]" captionStart="Figures 99–108" captionStartId="22.[166,243,1430,1452]" captionTargetBox="[173,1439,141,1416]" captionTargetId="figure-8@22.[166,1466,130,1430]" captionTargetPageId="22" captionText="Figures 99–108. Heterolepisma pyramidum sp. nov.holotype specimen K.261254 ♀ (99) ultimate article of labial palp showing basiconic sensillum (bsC); (100) pronotum, anterior and right margins; (101) idem, right anterior trichobothrial area; (102) idem, right posterior trichobothrial area; (103) idem, left posterior comb; (104) mesonotum, right side; (105) idem, trichobothrial areas, right side; (106) metanotum, right side; (107) idem, trichobothrial areas, right side; (108) presternum, prothoracic sternum and PI. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653523" httpUri="https://zenodo.org/record/4653523/files/figure.png" pageId="20" pageNumber="159">Fig. 100</figureCitation>
|
||
) with complete setal collar of short macrochaetae and some cilia; setae of lateral margins lost, but consisting of some marginal setae as well as cilia and two submarginal macrochaetae, one about one quarter the distance along the margin and the other about seven-eighths along the margin associated with the posterior trichobothrial area. Anterior trichobothrium a little further than half way along the margin, with two to four setulae posterior to the trichobothrium, not associated with a macrochaeta (
|
||
<figureCitation id="C0B42DB8EE2EA26AFCABA11AFC52FAAE" box="[848,951,1310,1334]" captionStart="Figures 99–108" captionStartId="22.[166,243,1430,1452]" captionTargetBox="[173,1439,141,1416]" captionTargetId="figure-8@22.[166,1466,130,1430]" captionTargetPageId="22" captionText="Figures 99–108. Heterolepisma pyramidum sp. nov.holotype specimen K.261254 ♀ (99) ultimate article of labial palp showing basiconic sensillum (bsC); (100) pronotum, anterior and right margins; (101) idem, right anterior trichobothrial area; (102) idem, right posterior trichobothrial area; (103) idem, left posterior comb; (104) mesonotum, right side; (105) idem, trichobothrial areas, right side; (106) metanotum, right side; (107) idem, trichobothrial areas, right side; (108) presternum, prothoracic sternum and PI. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653523" httpUri="https://zenodo.org/record/4653523/files/figure.png" pageId="20" pageNumber="159">Fig. 101</figureCitation>
|
||
). The posterior trichobothrium mediad of a submarginal macrochaeta and its cilium, with two or three setulae posterior to this group (
|
||
<figureCitation id="C0B42DB8EE2EA26AFB6DA15CFB10FAE8" box="[1174,1269,1368,1392]" captionStart="Figures 99–108" captionStartId="22.[166,243,1430,1452]" captionTargetBox="[173,1439,141,1416]" captionTargetId="figure-8@22.[166,1466,130,1430]" captionTargetPageId="22" captionText="Figures 99–108. Heterolepisma pyramidum sp. nov.holotype specimen K.261254 ♀ (99) ultimate article of labial palp showing basiconic sensillum (bsC); (100) pronotum, anterior and right margins; (101) idem, right anterior trichobothrial area; (102) idem, right posterior trichobothrial area; (103) idem, left posterior comb; (104) mesonotum, right side; (105) idem, trichobothrial areas, right side; (106) metanotum, right side; (107) idem, trichobothrial areas, right side; (108) presternum, prothoracic sternum and PI. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653523" httpUri="https://zenodo.org/record/4653523/files/figure.png" pageId="20" pageNumber="159">Fig. 102</figureCitation>
|
||
). Posterior margin with 1+1 single macrochaetae each associated with two cilia (
|
||
<figureCitation id="C0B42DB8EE2EA26AFC7EA197FC06FA33" box="[901,995,1427,1451]" captionStart="Figures 99–108" captionStartId="22.[166,243,1430,1452]" captionTargetBox="[173,1439,141,1416]" captionTargetId="figure-8@22.[166,1466,130,1430]" captionTargetPageId="22" captionText="Figures 99–108. Heterolepisma pyramidum sp. nov.holotype specimen K.261254 ♀ (99) ultimate article of labial palp showing basiconic sensillum (bsC); (100) pronotum, anterior and right margins; (101) idem, right anterior trichobothrial area; (102) idem, right posterior trichobothrial area; (103) idem, left posterior comb; (104) mesonotum, right side; (105) idem, trichobothrial areas, right side; (106) metanotum, right side; (107) idem, trichobothrial areas, right side; (108) presternum, prothoracic sternum and PI. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653523" httpUri="https://zenodo.org/record/4653523/files/figure.png" pageId="20" pageNumber="159">Fig. 103</figureCitation>
|
||
). — Mesonotum (
|
||
<figureCitation id="C0B42DB8EE2EA26AFB51A197FAE3FA33" box="[1194,1286,1427,1451]" captionStart="Figures 99–108" captionStartId="22.[166,243,1430,1452]" captionTargetBox="[173,1439,141,1416]" captionTargetId="figure-8@22.[166,1466,130,1430]" captionTargetPageId="22" captionText="Figures 99–108. Heterolepisma pyramidum sp. nov.holotype specimen K.261254 ♀ (99) ultimate article of labial palp showing basiconic sensillum (bsC); (100) pronotum, anterior and right margins; (101) idem, right anterior trichobothrial area; (102) idem, right posterior trichobothrial area; (103) idem, left posterior comb; (104) mesonotum, right side; (105) idem, trichobothrial areas, right side; (106) metanotum, right side; (107) idem, trichobothrial areas, right side; (108) presternum, prothoracic sternum and PI. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653523" httpUri="https://zenodo.org/record/4653523/files/figure.png" pageId="20" pageNumber="159">Fig. 104</figureCitation>
|
||
) also with small setae along the margins as well as three combs each of two macrochaetae anterior to the anterior trichobothrial area, both trichobothrial areas (
|
||
<figureCitation id="C0B42DB8EE2EA26AFBA1A1EFFB50F99B" box="[1114,1205,1515,1539]" captionStart="Figures 99–108" captionStartId="22.[166,243,1430,1452]" captionTargetBox="[173,1439,141,1416]" captionTargetId="figure-8@22.[166,1466,130,1430]" captionTargetPageId="22" captionText="Figures 99–108. Heterolepisma pyramidum sp. nov.holotype specimen K.261254 ♀ (99) ultimate article of labial palp showing basiconic sensillum (bsC); (100) pronotum, anterior and right margins; (101) idem, right anterior trichobothrial area; (102) idem, right posterior trichobothrial area; (103) idem, left posterior comb; (104) mesonotum, right side; (105) idem, trichobothrial areas, right side; (106) metanotum, right side; (107) idem, trichobothrial areas, right side; (108) presternum, prothoracic sternum and PI. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653523" httpUri="https://zenodo.org/record/4653523/files/figure.png" pageId="20" pageNumber="159">Fig. 105</figureCitation>
|
||
) are more posterior than on the pronotum, the anterior trichobothrium located about 5/6 the distance along the margin and not associated with any macrochaeta and the posterior area as in the pronotum; 1+1 posterior macrochaetae as in pronotum. — Metanotum (
|
||
<figureCitation id="C0B42DB8EE2EA26AFA70A264FC40F90E" captionStart="Figures 99–108" captionStartId="22.[166,243,1430,1452]" captionTargetBox="[173,1439,141,1416]" captionTargetId="figure-8@22.[166,1466,130,1430]" captionTargetPageId="22" captionText="Figures 99–108. Heterolepisma pyramidum sp. nov.holotype specimen K.261254 ♀ (99) ultimate article of labial palp showing basiconic sensillum (bsC); (100) pronotum, anterior and right margins; (101) idem, right anterior trichobothrial area; (102) idem, right posterior trichobothrial area; (103) idem, left posterior comb; (104) mesonotum, right side; (105) idem, trichobothrial areas, right side; (106) metanotum, right side; (107) idem, trichobothrial areas, right side; (108) presternum, prothoracic sternum and PI. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653523" httpUri="https://zenodo.org/record/4653523/files/figure.png" pageId="20" pageNumber="159">Figs 106, 107</figureCitation>
|
||
) similar to mesonotum.
|
||
</paragraph>
|
||
<paragraph id="5830313DEE2EA26BFC9EA29FFBDAF803" blockId="20.[839,1467,1046,2008]" lastBlockId="21.[794,1422,1893,2005]" lastPageId="21" lastPageNumber="160" pageId="20" pageNumber="159">
|
||
Presternum with transverse row of strong macrochaetae (
|
||
<figureCitation id="C0B42DB8EE2EA26AFCB4A2BCFC4EF948" box="[847,939,1720,1744]" captionStart="Figures 99–108" captionStartId="22.[166,243,1430,1452]" captionTargetBox="[173,1439,141,1416]" captionTargetId="figure-8@22.[166,1466,130,1430]" captionTargetPageId="22" captionText="Figures 99–108. Heterolepisma pyramidum sp. nov.holotype specimen K.261254 ♀ (99) ultimate article of labial palp showing basiconic sensillum (bsC); (100) pronotum, anterior and right margins; (101) idem, right anterior trichobothrial area; (102) idem, right posterior trichobothrial area; (103) idem, left posterior comb; (104) mesonotum, right side; (105) idem, trichobothrial areas, right side; (106) metanotum, right side; (107) idem, trichobothrial areas, right side; (108) presternum, prothoracic sternum and PI. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653523" httpUri="https://zenodo.org/record/4653523/files/figure.png" pageId="20" pageNumber="159">Fig. 108</figureCitation>
|
||
) as well as some simple cilia. All thoracic sterna with hyaline scales. — Prothoracic sternum (
|
||
<figureCitation id="C0B42DB8EE2EA26AFAABA2D2FA57F976" box="[1360,1458,1750,1774]" captionStart="Figures 99–108" captionStartId="22.[166,243,1430,1452]" captionTargetBox="[173,1439,141,1416]" captionTargetId="figure-8@22.[166,1466,130,1430]" captionTargetPageId="22" captionText="Figures 99–108. Heterolepisma pyramidum sp. nov.holotype specimen K.261254 ♀ (99) ultimate article of labial palp showing basiconic sensillum (bsC); (100) pronotum, anterior and right margins; (101) idem, right anterior trichobothrial area; (102) idem, right posterior trichobothrial area; (103) idem, left posterior comb; (104) mesonotum, right side; (105) idem, trichobothrial areas, right side; (106) metanotum, right side; (107) idem, trichobothrial areas, right side; (108) presternum, prothoracic sternum and PI. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653523" httpUri="https://zenodo.org/record/4653523/files/figure.png" pageId="20" pageNumber="159">Fig. 108</figureCitation>
|
||
) cordiform, slightly wider than long (L/
|
||
<geoCoordinate id="3DBB57FAEE2EA26AFB28A2F7FAC7F893" box="[1235,1314,1779,1803]" degrees="0.91" direction="west" orientation="longitude" pageId="20" pageNumber="159" precision="555" value="-0.91">W 0.91</geoCoordinate>
|
||
), 1+1 irregular combs of eight to nine bifurcate macrochaetae running subparallel to the margin, as well as some marginal setae and cilia in distal one quarter, short posterior margin between combs straight. — Mesosternum (
|
||
<figureCitation id="C0B42DB8EE2EA26AFB4FA36CFAF6F818" box="[1204,1299,1896,1920]" captionStart="Figures 109–122" captionStartId="23.[121,198,1584,1606]" captionTargetBox="[129,1413,160,1568]" captionTargetId="figure-10@23.[121,1421,151,1576]" captionTargetPageId="23" captionText="Figures 109–122. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 or K.261255 ♀ (109) mesothoracic sternum; (110) metathoracic sternum and PIII; (111) PII; (112) urotergite III; (113) urotergite VII, left lateral comb; (114) idem, left sublateral lateral comb; (115) idem, left submedial comb; (116) urotergite IX, left infralateral comb; (117) urotergite X; (118) urosternite I; (119) idem, medial comb; (120) urosternite IV; (121) right comb of urosternite IV; (122) urosternite VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653527" httpUri="https://zenodo.org/record/4653527/files/figure.png" pageId="20" pageNumber="159">Figs 109</figureCitation>
|
||
) sub-parabolic, as long as wide at its base, (L/
|
||
<geoCoordinate id="3DBB57FAEE2EA26AFB5BA382FB11F806" box="[1184,1268,1926,1950]" degrees="1.01" direction="west" orientation="longitude" pageId="20" pageNumber="159" precision="555" value="-1.01">W 1.01</geoCoordinate>
|
||
) with about eight macrochaetae forming an irregular submarginal row along the posterior quarter of the margin, short posterior margin between combs straight. — Metasternum (
|
||
<figureCitation id="C0B42DB8EE2FA26BFDBEA361FD45F8E5" box="[581,672,1893,1917]" captionStart="Figures 109–122" captionStartId="23.[121,198,1584,1606]" captionTargetBox="[129,1413,160,1568]" captionTargetId="figure-10@23.[121,1421,151,1576]" captionTargetPageId="23" captionText="Figures 109–122. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 or K.261255 ♀ (109) mesothoracic sternum; (110) metathoracic sternum and PIII; (111) PII; (112) urotergite III; (113) urotergite VII, left lateral comb; (114) idem, left sublateral lateral comb; (115) idem, left submedial comb; (116) urotergite IX, left infralateral comb; (117) urotergite X; (118) urosternite I; (119) idem, medial comb; (120) urosternite IV; (121) right comb of urosternite IV; (122) urosternite VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653527" httpUri="https://zenodo.org/record/4653527/files/figure.png" pageId="21" pageNumber="160">Fig. 110</figureCitation>
|
||
) wider than long (L/
|
||
<geoCoordinate id="3DBB57FAEE2FA26BFEFDA387FEBCF803" box="[262,345,1923,1947]" degrees="0.74" direction="west" orientation="longitude" pageId="21" pageNumber="160" precision="555" value="-0.74">W 0.74</geoCoordinate>
|
||
), apically a little truncate, 1+1 apical combs of about five macrochaetae, the distance between the combs 2.3 times the average length of each comb (range 2.1–2.6), lateral margins with a few simple setae and cilia only adjacent to the combs.
|
||
</paragraph>
|
||
<caption id="0CF061B5EE2FA26BFF82A2C4FBBAF8D9" ID-DOI="http://doi.org/10.5281/zenodo.4653519" ID-Zenodo-Dep="4653519" httpUri="https://zenodo.org/record/4653519/files/figure.png" pageId="21" pageNumber="160" startId="21.[121,196,1728,1750]" targetBox="[121,1411,164,1712]" targetPageId="21">
|
||
<paragraph id="5830313DEE2FA26BFF82A2C4FBBAF8D9" blockId="21.[121,1422,1728,1857]" pageId="21" pageNumber="160">
|
||
<emphasis id="6AFBED2FEE2FA26BFF82A2C4FEE6F94D" bold="true" box="[121,259,1728,1750]" pageId="21" pageNumber="160">Figures 86–98</emphasis>
|
||
.
|
||
<taxonomicName id="9F8F4ABEEE2FA26BFEF6A2C4FDE0F94D" authority="Smith & Mitchell, 2019" authorityName="Smith & Mitchell" authorityYear="2019" box="[269,517,1728,1749]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="21" pageNumber="160" phylum="Arthropoda" rank="species" species="pyramidum" status="sp. nov.">
|
||
<emphasis id="6AFBED2FEE2FA26BFEF6A2C4FDE0F94D" box="[269,517,1728,1749]" italics="true" pageId="21" pageNumber="160">Heterolepisma pyramidum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71C85054EE2FA26BFDF2A2C4FDB4F94D" box="[521,593,1728,1749]" pageId="21" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
holotype specimen K.261254 ♀
|
||
<emphasis id="6AFBED2FEE2FA26BFC85A2C4FC40F94D" box="[894,933,1728,1749]" italics="true" pageId="21" pageNumber="160">(86)</emphasis>
|
||
habitus;
|
||
<emphasis id="6AFBED2FEE2FA26BFC0CA2C4FBFBF94D" box="[1015,1054,1728,1749]" italics="true" pageId="21" pageNumber="160">(87)</emphasis>
|
||
smooth, apically bifurcate macrochaeta of PI coxa;
|
||
<emphasis id="6AFBED2FEE2FA26BFF1EA2DFFEE8F968" box="[229,269,1755,1776]" italics="true" pageId="21" pageNumber="160">(88)</emphasis>
|
||
carrot-shaped macrochaeta of PI tibia;
|
||
<emphasis id="6AFBED2FEE2FA26BFD7AA2DFFD4CF968" box="[641,681,1755,1776]" italics="true" pageId="21" pageNumber="160">(89)</emphasis>
|
||
scale from mesonotum;
|
||
<emphasis id="6AFBED2FEE2FA26BFC69A2DFFC5CF968" box="[914,953,1755,1776]" italics="true" pageId="21" pageNumber="160">(90)</emphasis>
|
||
head;
|
||
<emphasis id="6AFBED2FEE2FA26BFC0CA2DFFBC5F968" box="[1015,1056,1755,1776]" italics="true" pageId="21" pageNumber="160">(91)</emphasis>
|
||
antenna, pedicel and basal intervals of flagellum;
|
||
<emphasis id="6AFBED2FEE2FA26BFF24A2F1FEE2F892" box="[223,263,1781,1802]" italics="true" pageId="21" pageNumber="160">(92)</emphasis>
|
||
idem, most distal surviving interval showing basiconic sensilla type C (
|
||
<emphasis id="6AFBED2FEE2FA26BFC4AA2F1FC3CF893" bold="true" box="[945,985,1781,1803]" pageId="21" pageNumber="160">bsC</emphasis>
|
||
);
|
||
<emphasis id="6AFBED2FEE2FA26BFC16A2F1FBF1F892" box="[1005,1044,1781,1802]" italics="true" pageId="21" pageNumber="160">(93)</emphasis>
|
||
mandible;
|
||
<emphasis id="6AFBED2FEE2FA26BFB84A2F1FB42F892" box="[1151,1191,1781,1802]" italics="true" pageId="21" pageNumber="160">(94)</emphasis>
|
||
idem, incisor and molar regions;
|
||
<emphasis id="6AFBED2FEE2FA26BFF36A314FF10F8BD" box="[205,245,1808,1829]" italics="true" pageId="21" pageNumber="160">(95)</emphasis>
|
||
maxilla, only more prominent setae of palp illustrated;
|
||
<emphasis id="6AFBED2FEE2FA26BFCF4A314FCD2F8BD" box="[783,823,1808,1829]" italics="true" pageId="21" pageNumber="160">(96)</emphasis>
|
||
idem, lacinia and galea;
|
||
<emphasis id="6AFBED2FEE2FA26BFBD2A314FBB4F8BD" box="[1065,1105,1808,1829]" italics="true" pageId="21" pageNumber="160">(97)</emphasis>
|
||
idem, apex of ultimate article of palp with its three branched papillae (
|
||
<emphasis id="6AFBED2FEE2FA26BFE1FA32FFE1AF8D8" bold="true" box="[484,511,1835,1856]" pageId="21" pageNumber="160">bp</emphasis>
|
||
);
|
||
<emphasis id="6AFBED2FEE2FA26BFDE8A32FFDDEF8D8" box="[531,571,1835,1856]" italics="true" pageId="21" pageNumber="160">(98)</emphasis>
|
||
labium. Scale bars = 0.1 mm unless otherwise indicated.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5830313DEE2FA269FCC3A3A4FDBFF81F" blockId="21.[794,1422,1893,2005]" lastBlockId="23.[121,749,1726,1927]" lastPageId="23" lastPageNumber="162" pageId="21" pageNumber="160">
|
||
Legs (
|
||
<figureCitation id="C0B42DB8EE2FA26BFC83A3A4FC32F820" box="[888,983,1952,1976]" captionStart="Figures 99–108" captionStartId="22.[166,243,1430,1452]" captionTargetBox="[173,1439,141,1416]" captionTargetId="figure-8@22.[166,1466,130,1430]" captionTargetPageId="22" captionText="Figures 99–108. Heterolepisma pyramidum sp. nov.holotype specimen K.261254 ♀ (99) ultimate article of labial palp showing basiconic sensillum (bsC); (100) pronotum, anterior and right margins; (101) idem, right anterior trichobothrial area; (102) idem, right posterior trichobothrial area; (103) idem, left posterior comb; (104) mesonotum, right side; (105) idem, trichobothrial areas, right side; (106) metanotum, right side; (107) idem, trichobothrial areas, right side; (108) presternum, prothoracic sternum and PI. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653523" httpUri="https://zenodo.org/record/4653523/files/figure.png" pageId="21" pageNumber="160">Figs 108</figureCitation>
|
||
,
|
||
<figureCitation id="C0B42DB8EE2FA26BFC1AA3A4FBDEF820" box="[993,1083,1952,1976]" captionStart="Figures 109–122" captionStartId="23.[121,198,1584,1606]" captionTargetBox="[129,1413,160,1568]" captionTargetId="figure-10@23.[121,1421,151,1576]" captionTargetPageId="23" captionText="Figures 109–122. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 or K.261255 ♀ (109) mesothoracic sternum; (110) metathoracic sternum and PIII; (111) PII; (112) urotergite III; (113) urotergite VII, left lateral comb; (114) idem, left sublateral lateral comb; (115) idem, left submedial comb; (116) urotergite IX, left infralateral comb; (117) urotergite X; (118) urosternite I; (119) idem, medial comb; (120) urosternite IV; (121) right comb of urosternite IV; (122) urosternite VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653527" httpUri="https://zenodo.org/record/4653527/files/figure.png" pageId="21" pageNumber="160">110, 111</figureCitation>
|
||
) becoming progressively longer and more slender with the tibia of PII being 1.3 times longer than that of PI and the tibia of PIII being 1.8 times longer than that of PI, tibia L/W ratio of legs PI 2.9, PII 3.2, PIII 3.6; tarsi L/W ratio PI 5.3, PII 8.0, PIII 9.2. Precoxa of PI with lateral combs of four macrochaetae. — Coxa of PI (
|
||
<figureCitation id="C0B42DB8EE2CA268FD14A287FF2AF921" captionStart="Figures 99–108" captionStartId="22.[166,243,1430,1452]" captionTargetBox="[173,1439,141,1416]" captionTargetId="figure-8@22.[166,1466,130,1430]" captionTargetPageId="22" captionText="Figures 99–108. Heterolepisma pyramidum sp. nov.holotype specimen K.261254 ♀ (99) ultimate article of labial palp showing basiconic sensillum (bsC); (100) pronotum, anterior and right margins; (101) idem, right anterior trichobothrial area; (102) idem, right posterior trichobothrial area; (103) idem, left posterior comb; (104) mesonotum, right side; (105) idem, trichobothrial areas, right side; (106) metanotum, right side; (107) idem, trichobothrial areas, right side; (108) presternum, prothoracic sternum and PI. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653523" httpUri="https://zenodo.org/record/4653523/files/figure.png" pageId="22" pageNumber="161">Fig. 108</figureCitation>
|
||
) without comb near the anterolateral corners, but many strong macrochaetae in approximately two rows along the external margin, becoming shorter distally; inner margin with five macrochaetae distally over the articulation and another seta subdistally near three long thin seta on the dorsal face. — Trochanter with small setae. — Femur ventrally with one large macrochaeta near the trochanter and three large bifurcate carrot-shaped macrochaetae plus two strong simple macrochaetae on the posterior bulge, distal anterior corner with two macrochaetae, lanceolate scales along the margin and extending onto the ventral face of the femur in the anterior half, rest of ventral surface with fine scattered setae. — Tibia with a few lanceolate scales on the ventral face proximally, rest of surface with setae, a strong carrot-shaped macrochaeta distally as well as a thinner pair subdistally, another pair midway along the margin and a smaller pair near the proximal end of the posterior margin, distal end of tibia with a row of stronger setae, anterior margin with two stout macrochaetae about one third and two thirds along the margin, as well as smaller setae; apical spine with several setae, some quite strong. — Tarsi of four articles, the basal tarsal article of PI not quite as long as the remaining articles together, bearing some stronger setae below and many other smaller setae over the surface; second and third articles the shortest, each with two stronger setae ventrally. Pretarsus with two long curved lateral claws and a shorter straight medial claw. PII (
|
||
<figureCitation id="C0B42DB8EE2CA269FA74A3C2FF45F94E" captionStart="Figures 109–122" captionStartId="23.[121,198,1584,1606]" captionTargetBox="[129,1413,160,1568]" captionTargetId="figure-10@23.[121,1421,151,1576]" captionTargetPageId="23" captionText="Figures 109–122. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 or K.261255 ♀ (109) mesothoracic sternum; (110) metathoracic sternum and PIII; (111) PII; (112) urotergite III; (113) urotergite VII, left lateral comb; (114) idem, left sublateral lateral comb; (115) idem, left submedial comb; (116) urotergite IX, left infralateral comb; (117) urotergite X; (118) urosternite I; (119) idem, medial comb; (120) urosternite IV; (121) right comb of urosternite IV; (122) urosternite VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653527" httpUri="https://zenodo.org/record/4653527/files/figure.png" lastPageId="23" lastPageNumber="162" pageId="22" pageNumber="161">Fig. 111</figureCitation>
|
||
) similar to PI except coxae with fewer macrochaetae along the outer margin, only becoming two macrochaetae wide in the distal half, with about seven stout setae over the articulation, the tibia with three stout macrochaetae along the dorsal margin and the first tarsal article is longer than the remaining three together, and PIII (
|
||
<figureCitation id="C0B42DB8EE2DA269FE0FA356FDAAF8F2" box="[500,591,1874,1898]" captionStart="Figures 109–122" captionStartId="23.[121,198,1584,1606]" captionTargetBox="[129,1413,160,1568]" captionTargetId="figure-10@23.[121,1421,151,1576]" captionTargetPageId="23" captionText="Figures 109–122. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 or K.261255 ♀ (109) mesothoracic sternum; (110) metathoracic sternum and PIII; (111) PII; (112) urotergite III; (113) urotergite VII, left lateral comb; (114) idem, left sublateral lateral comb; (115) idem, left submedial comb; (116) urotergite IX, left infralateral comb; (117) urotergite X; (118) urosternite I; (119) idem, medial comb; (120) urosternite IV; (121) right comb of urosternite IV; (122) urosternite VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653527" httpUri="https://zenodo.org/record/4653527/files/figure.png" pageId="23" pageNumber="162">Fig. 110</figureCitation>
|
||
) similar to PII but areas of lanceolate scales more extensive.
|
||
</paragraph>
|
||
<caption id="0CF061B5EE2CA268FF5DA192FA53FA63" ID-DOI="http://doi.org/10.5281/zenodo.4653523" ID-Zenodo-Dep="4653523" httpUri="https://zenodo.org/record/4653523/files/figure.png" pageId="22" pageNumber="161" startId="22.[166,243,1430,1452]" targetBox="[173,1439,141,1416]" targetPageId="22">
|
||
<paragraph id="5830313DEE2CA268FF5DA192FA53FA63" blockId="22.[166,1466,1430,1532]" pageId="22" pageNumber="161">
|
||
<emphasis id="6AFBED2FEE2CA268FF5DA192FEDAFA33" bold="true" box="[166,319,1430,1452]" pageId="22" pageNumber="161">Figures 99–108</emphasis>
|
||
.
|
||
<taxonomicName id="9F8F4ABEEE2CA268FEB1A192FDADFA33" authority="Smith & Mitchell, 2019" authorityName="Smith & Mitchell" authorityYear="2019" box="[330,584,1430,1451]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="22" pageNumber="161" phylum="Arthropoda" rank="species" species="pyramidum" status="sp. nov.">
|
||
<emphasis id="6AFBED2FEE2CA268FEB1A192FDADFA33" box="[330,584,1430,1451]" italics="true" pageId="22" pageNumber="161">Heterolepisma pyramidum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71C85054EE2CA268FDB7A192FD73FA33" box="[588,662,1430,1451]" pageId="22" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
|
||
holotype specimen K.261254 ♀
|
||
<emphasis id="6AFBED2FEE2CA268FC30A192FC17FA33" box="[971,1010,1430,1451]" italics="true" pageId="22" pageNumber="161">(99)</emphasis>
|
||
ultimate article of labial palp showing basiconic sensillum (
|
||
<emphasis id="6AFBED2FEE2CA268FEEAA1B4FEDCFA5E" bold="true" box="[273,313,1456,1478]" pageId="22" pageNumber="161">bsC</emphasis>
|
||
);
|
||
<emphasis id="6AFBED2FEE2CA268FEB4A1B5FE66FA5E" box="[335,387,1457,1478]" italics="true" pageId="22" pageNumber="161">(100)</emphasis>
|
||
pronotum, anterior and right margins;
|
||
<emphasis id="6AFBED2FEE2CA268FD06A1B5FCD4FA5E" box="[765,817,1457,1478]" italics="true" pageId="22" pageNumber="161">(101)</emphasis>
|
||
idem, right anterior trichobothrial area;
|
||
<emphasis id="6AFBED2FEE2CA268FB42A1B5FB08FA5E" box="[1209,1261,1457,1478]" italics="true" pageId="22" pageNumber="161">(102)</emphasis>
|
||
idem, right posterior trichobothrial area;
|
||
<emphasis id="6AFBED2FEE2CA268FE92A1CFFE79FA78" box="[361,412,1483,1504]" italics="true" pageId="22" pageNumber="161">(103)</emphasis>
|
||
idem, left posterior comb;
|
||
<emphasis id="6AFBED2FEE2CA268FD55A1CFFD07FA78" box="[686,738,1483,1504]" italics="true" pageId="22" pageNumber="161">(104)</emphasis>
|
||
mesonotum, right side;
|
||
<emphasis id="6AFBED2FEE2CA268FC2EA1CFFBECFA78" box="[981,1033,1483,1504]" italics="true" pageId="22" pageNumber="161">(105)</emphasis>
|
||
idem, trichobothrial areas, right side;
|
||
<emphasis id="6AFBED2FEE2CA268FA7DA1CFFA5FFA78" box="[1414,1466,1483,1504]" italics="true" pageId="22" pageNumber="161">(106)</emphasis>
|
||
metanotum, right side;
|
||
<emphasis id="6AFBED2FEE2CA268FE7EA1E2FE5CFA63" box="[389,441,1510,1531]" italics="true" pageId="22" pageNumber="161">(107)</emphasis>
|
||
idem, trichobothrial areas, right side;
|
||
<emphasis id="6AFBED2FEE2CA268FCDCA1E2FCBEFA63" box="[807,859,1510,1531]" italics="true" pageId="22" pageNumber="161">(108)</emphasis>
|
||
presternum, prothoracic sternum and PI. Scale bars = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0CF061B5EE2DA269FF82A234FB25F90D" ID-DOI="http://doi.org/10.5281/zenodo.4653527" ID-Zenodo-Dep="4653527" httpUri="https://zenodo.org/record/4653527/files/figure.png" pageId="23" pageNumber="162" startId="23.[121,198,1584,1606]" targetBox="[129,1413,160,1568]" targetPageId="23">
|
||
<paragraph id="5830313DEE2DA269FF82A234FB25F90D" blockId="23.[121,1422,1584,1686]" pageId="23" pageNumber="162">
|
||
<emphasis id="6AFBED2FEE2DA269FF82A234FEC4F9DD" bold="true" box="[121,289,1584,1606]" pageId="23" pageNumber="162">Figures 109–122</emphasis>
|
||
.
|
||
<taxonomicName id="9F8F4ABEEE2DA269FED6A234FDC8F9DD" authority="Smith & Mitchell, 2019" authorityName="Smith & Mitchell" authorityYear="2019" box="[301,557,1584,1605]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="23" pageNumber="162" phylum="Arthropoda" rank="species" species="pyramidum" status="sp. nov.">
|
||
<emphasis id="6AFBED2FEE2DA269FED6A234FDC8F9DD" box="[301,557,1584,1605]" italics="true" pageId="23" pageNumber="162">Heterolepisma pyramidum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71C85054EE2DA269FDCFA234FD9AF9DD" box="[564,639,1584,1605]" pageId="23" pageNumber="162" rank="species">sp. nov.</taxonomicNameLabel>
|
||
holotype specimen K.261254 or K.261255 ♀
|
||
<emphasis id="6AFBED2FEE2DA269FBC4A234FB96F9DD" box="[1087,1139,1584,1605]" italics="true" pageId="23" pageNumber="162">(109)</emphasis>
|
||
mesothoracic sternum;
|
||
<emphasis id="6AFBED2FEE2DA269FAA0A234FA68F9DD" box="[1371,1421,1584,1605]" italics="true" pageId="23" pageNumber="162">(110)</emphasis>
|
||
metathoracic sternum and PIII;
|
||
<emphasis id="6AFBED2FEE2DA269FE55A24FFE38F9F8" box="[430,477,1611,1632]" italics="true" pageId="23" pageNumber="162">(111)</emphasis>
|
||
PII;
|
||
<emphasis id="6AFBED2FEE2DA269FDEAA24FFDA6F9F8" box="[529,579,1611,1632]" italics="true" pageId="23" pageNumber="162">(112)</emphasis>
|
||
urotergite III;
|
||
<emphasis id="6AFBED2FEE2DA269FD2EA24FFCE2F9F8" box="[725,775,1611,1632]" italics="true" pageId="23" pageNumber="162">(113)</emphasis>
|
||
urotergite VII, left lateral comb;
|
||
<emphasis id="6AFBED2FEE2DA269FBB5A24FFB64F9F8" box="[1102,1153,1611,1632]" italics="true" pageId="23" pageNumber="162">(114)</emphasis>
|
||
idem, left sublateral lateral comb;
|
||
<emphasis id="6AFBED2FEE2DA269FF47A261FF0BF9E2" box="[188,238,1637,1658]" italics="true" pageId="23" pageNumber="162">(115)</emphasis>
|
||
idem, left submedial comb;
|
||
<emphasis id="6AFBED2FEE2DA269FDFDA261FDDCF9E2" box="[518,569,1637,1658]" italics="true" pageId="23" pageNumber="162">(116)</emphasis>
|
||
urotergite IX, left infralateral comb;
|
||
<emphasis id="6AFBED2FEE2DA269FC5EA261FC32F9E2" box="[933,983,1637,1658]" italics="true" pageId="23" pageNumber="162">(117)</emphasis>
|
||
urotergite X;
|
||
<emphasis id="6AFBED2FEE2DA269FB9AA261FB76F9E2" box="[1121,1171,1637,1658]" italics="true" pageId="23" pageNumber="162">(118)</emphasis>
|
||
urosternite I;
|
||
<emphasis id="6AFBED2FEE2DA269FAE5A261FAB5F9E2" box="[1310,1360,1637,1658]" italics="true" pageId="23" pageNumber="162">(119)</emphasis>
|
||
idem, medial comb;
|
||
<emphasis id="6AFBED2FEE2DA269FEF9A284FED3F90D" box="[258,310,1664,1685]" italics="true" pageId="23" pageNumber="162">(120)</emphasis>
|
||
urosternite IV;
|
||
<emphasis id="6AFBED2FEE2DA269FE36A284FDE4F90D" box="[461,513,1664,1685]" italics="true" pageId="23" pageNumber="162">(121)</emphasis>
|
||
right comb of urosternite IV;
|
||
<emphasis id="6AFBED2FEE2DA269FCDAA284FCB1F90D" box="[801,852,1664,1685]" italics="true" pageId="23" pageNumber="162">(122)</emphasis>
|
||
urosternite VII. Scale bars = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5830313DEE2DA266FF82A3A1FE8BFBDF" blockId="23.[121,748,1957,2010]" lastBlockId="24.[166,794,869,1586]" lastPageId="24" lastPageNumber="163" pageId="23" pageNumber="162">
|
||
<emphasis id="6AFBED2FEE2DA269FF82A3A1FF3BF825" box="[121,222,1957,1981]" italics="true" pageId="23" pageNumber="162">Abdomen</emphasis>
|
||
: Urotergite I with 2+2 combs, the lateral combs of two macrochaetae each associated with three marginal setae and a setula, the submedial combs with one or two marginal setae, 0–3 setulae and a cilium on the laterad end of each comb; urotergites II–VII (
|
||
<figureCitation id="C0B42DB8EE2DA269FB7FA2FCFB3AF888" box="[1156,1247,1784,1808]" captionStart="Figures 109–122" captionStartId="23.[121,198,1584,1606]" captionTargetBox="[129,1413,160,1568]" captionTargetId="figure-10@23.[121,1421,151,1576]" captionTargetPageId="23" captionText="Figures 109–122. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 or K.261255 ♀ (109) mesothoracic sternum; (110) metathoracic sternum and PIII; (111) PII; (112) urotergite III; (113) urotergite VII, left lateral comb; (114) idem, left sublateral lateral comb; (115) idem, left submedial comb; (116) urotergite IX, left infralateral comb; (117) urotergite X; (118) urosternite I; (119) idem, medial comb; (120) urosternite IV; (121) right comb of urosternite IV; (122) urosternite VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653527" httpUri="https://zenodo.org/record/4653527/files/figure.png" pageId="23" pageNumber="162">Fig. 112</figureCitation>
|
||
) with 3+3 small combs as shown in table 7, each lateral comb associated with 1–3 marginal setae, 1–3 setulae, usually a cilium at the mediad end of each comb and sometimes one between, and anterior to, the two most mediad macrochaetae, each sublateral comb with 0–2 marginal setae and 1–3 setulae, the submedial combs usually with a cilium on either side of the single macrochaeta and rarely a setula between the macrochaeta and the margin (
|
||
<figureCitation id="C0B42DB8EE22A266FE20A762FD8BFCE5" box="[475,622,869,894]" captionStart="Figures 109–122" captionStartId="23.[121,198,1584,1606]" captionTargetBox="[129,1413,160,1568]" captionTargetId="figure-10@23.[121,1421,151,1576]" captionTargetPageId="23" captionText="Figures 109–122. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 or K.261255 ♀ (109) mesothoracic sternum; (110) metathoracic sternum and PIII; (111) PII; (112) urotergite III; (113) urotergite VII, left lateral comb; (114) idem, left sublateral lateral comb; (115) idem, left submedial comb; (116) urotergite IX, left infralateral comb; (117) urotergite X; (118) urosternite I; (119) idem, medial comb; (120) urosternite IV; (121) right comb of urosternite IV; (122) urosternite VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653527" httpUri="https://zenodo.org/record/4653527/files/figure.png" pageId="24" pageNumber="163">Figs 113–115</figureCitation>
|
||
); urotergite VIII with 2+2 combs, lacking the sublateral comb; urotergite IX without combs but with two delicate marginal setae and two cilia in each infralateral corner (
|
||
<figureCitation id="C0B42DB8EE22A266FDC3A7B8FD72FC4C" box="[568,663,956,980]" captionStart="Figures 109–122" captionStartId="23.[121,198,1584,1606]" captionTargetBox="[129,1413,160,1568]" captionTargetId="figure-10@23.[121,1421,151,1576]" captionTargetPageId="23" captionText="Figures 109–122. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 or K.261255 ♀ (109) mesothoracic sternum; (110) metathoracic sternum and PIII; (111) PII; (112) urotergite III; (113) urotergite VII, left lateral comb; (114) idem, left sublateral lateral comb; (115) idem, left submedial comb; (116) urotergite IX, left infralateral comb; (117) urotergite X; (118) urosternite I; (119) idem, medial comb; (120) urosternite IV; (121) right comb of urosternite IV; (122) urosternite VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653527" httpUri="https://zenodo.org/record/4653527/files/figure.png" pageId="24" pageNumber="163">Fig. 116</figureCitation>
|
||
). Urotergite X (
|
||
<figureCitation id="C0B42DB8EE22A266FF30A7DDFECFFC69" box="[203,298,985,1009]" captionStart="Figures 109–122" captionStartId="23.[121,198,1584,1606]" captionTargetBox="[129,1413,160,1568]" captionTargetId="figure-10@23.[121,1421,151,1576]" captionTargetPageId="23" captionText="Figures 109–122. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 or K.261255 ♀ (109) mesothoracic sternum; (110) metathoracic sternum and PIII; (111) PII; (112) urotergite III; (113) urotergite VII, left lateral comb; (114) idem, left sublateral lateral comb; (115) idem, left submedial comb; (116) urotergite IX, left infralateral comb; (117) urotergite X; (118) urosternite I; (119) idem, medial comb; (120) urosternite IV; (121) right comb of urosternite IV; (122) urosternite VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653527" httpUri="https://zenodo.org/record/4653527/files/figure.png" pageId="24" pageNumber="163">Fig. 117</figureCitation>
|
||
) short and distinctly trapezoidal (L/W about 0.35), with 1+1 macrochaetae in the posterolateral corners, each associated with cilia on each side and numerous setae along the margins.
|
||
</paragraph>
|
||
<caption id="0CF061B5EE22A266FF5DA709FABBFCA6" ID-DOI="http://doi.org/10.5281/zenodo.4653529" ID-Zenodo-Dep="4653529" httpUri="https://zenodo.org/record/4653529/files/figure.png" pageId="24" pageNumber="163" startId="24.[166,243,781,803]" targetBox="[183,1449,164,769]" targetPageId="24">
|
||
<paragraph id="5830313DEE22A266FF5DA709FABBFCA6" blockId="24.[166,1467,781,830]" pageId="24" pageNumber="163">
|
||
<emphasis id="6AFBED2FEE22A266FF5DA709FEA9FCBA" bold="true" box="[166,332,781,803]" pageId="24" pageNumber="163">Figures 123–126</emphasis>
|
||
.
|
||
<taxonomicName id="9F8F4ABEEE22A266FEADA709FDB6FCBA" authority="Smith & Mitchell, 2019" authorityName="Smith & Mitchell" authorityYear="2019" box="[342,595,781,802]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="24" pageNumber="163" phylum="Arthropoda" rank="species" species="pyramidum" status="sp. nov.">
|
||
<emphasis id="6AFBED2FEE22A266FEADA709FDB6FCBA" box="[342,595,781,802]" italics="true" pageId="24" pageNumber="163">Heterolepisma pyramidum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71C85054EE22A266FDA3A70AFD44FCBB" box="[600,673,782,803]" pageId="24" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
|
||
holotype specimen K.261254 ♀
|
||
<emphasis id="6AFBED2FEE22A266FC2DA70AFBEFFCBB" box="[982,1034,782,803]" italics="true" pageId="24" pageNumber="163">(123)</emphasis>
|
||
coxites VIII and IX, with styli and ovipositor;
|
||
<emphasis id="6AFBED2FEE22A266FF5DA72CFF3FFCA5" box="[166,218,808,829]" italics="true" pageId="24" pageNumber="163">(124)</emphasis>
|
||
apex of ovipositor valve;
|
||
<emphasis id="6AFBED2FEE22A266FE2DA72CFDECFCA5" box="[470,521,808,829]" italics="true" pageId="24" pageNumber="163">(125)</emphasis>
|
||
base of cercus and medial dorsal appendage;
|
||
<emphasis id="6AFBED2FEE22A266FC3AA72CFC10FCA5" box="[961,1013,808,829]" italics="true" pageId="24" pageNumber="163">(126)</emphasis>
|
||
apex of cercus. Scale bars = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5830313DEE22A266FF3EA048FB14FCE5" blockId="24.[166,794,869,1586]" lastBlockId="24.[839,1467,870,1539]" pageId="24" pageNumber="163">
|
||
Urosternite I with a medial comb of three macrochaetae and two setulae on a non-protruding region (
|
||
<figureCitation id="C0B42DB8EE22A266FD80A06DFCE8FB19" box="[635,781,1129,1153]" captionStart="Figures 109–122" captionStartId="23.[121,198,1584,1606]" captionTargetBox="[129,1413,160,1568]" captionTargetId="figure-10@23.[121,1421,151,1576]" captionTargetPageId="23" captionText="Figures 109–122. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 or K.261255 ♀ (109) mesothoracic sternum; (110) metathoracic sternum and PIII; (111) PII; (112) urotergite III; (113) urotergite VII, left lateral comb; (114) idem, left sublateral lateral comb; (115) idem, left submedial comb; (116) urotergite IX, left infralateral comb; (117) urotergite X; (118) urosternite I; (119) idem, medial comb; (120) urosternite IV; (121) right comb of urosternite IV; (122) urosternite VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653527" httpUri="https://zenodo.org/record/4653527/files/figure.png" pageId="24" pageNumber="163">Figs 118, 119</figureCitation>
|
||
), urosternite II–VI (
|
||
<figureCitation id="C0B42DB8EE22A266FE94A082FDECFB06" box="[367,521,1158,1182]" captionStart="Figures 109–122" captionStartId="23.[121,198,1584,1606]" captionTargetBox="[129,1413,160,1568]" captionTargetId="figure-10@23.[121,1421,151,1576]" captionTargetPageId="23" captionText="Figures 109–122. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 or K.261255 ♀ (109) mesothoracic sternum; (110) metathoracic sternum and PIII; (111) PII; (112) urotergite III; (113) urotergite VII, left lateral comb; (114) idem, left sublateral lateral comb; (115) idem, left submedial comb; (116) urotergite IX, left infralateral comb; (117) urotergite X; (118) urosternite I; (119) idem, medial comb; (120) urosternite IV; (121) right comb of urosternite IV; (122) urosternite VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653527" httpUri="https://zenodo.org/record/4653527/files/figure.png" pageId="24" pageNumber="163">Figs 120, 121</figureCitation>
|
||
) with 1+1 lateral combs each of four or five macrochaetae and two marginal setulae plus a cilium at the laterad end of each comb, the distance between each comb is about nine times the average length of the combs on the segment (range 8.7–10.2), urosternite VII (
|
||
<figureCitation id="C0B42DB8EE22A266FF55A112FEECFAB6" box="[174,265,1302,1326]" captionStart="Figures 109–122" captionStartId="23.[121,198,1584,1606]" captionTargetBox="[129,1413,160,1568]" captionTargetId="figure-10@23.[121,1421,151,1576]" captionTargetPageId="23" captionText="Figures 109–122. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 or K.261255 ♀ (109) mesothoracic sternum; (110) metathoracic sternum and PIII; (111) PII; (112) urotergite III; (113) urotergite VII, left lateral comb; (114) idem, left sublateral lateral comb; (115) idem, left submedial comb; (116) urotergite IX, left infralateral comb; (117) urotergite X; (118) urosternite I; (119) idem, medial comb; (120) urosternite IV; (121) right comb of urosternite IV; (122) urosternite VII. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653527" httpUri="https://zenodo.org/record/4653527/files/figure.png" pageId="24" pageNumber="163">Fig. 122</figureCitation>
|
||
) with styli, 1+1 combs each of two macrochaetae located mediad of each stylus, associated with two marginal setulae, laterad of the stylus is a cilium associated with two marginal setulae. Urosternite VIII in
|
||
<specimenCount id="4E89FAB4EE22A266FDDCA169FDDEFA1C" box="[551,571,1389,1412]" pageId="24" pageNumber="163" type="female">♀</specimenCount>
|
||
divided into separate coxites shaped as shown in figure 123, each bearing a small stylus which is apically armed with some strong setae, one or two macrochaetae mediad of the stylus insertion associated with a few small marginal setae, with two or three setae and a cilium on the outer side of the stylus. Styli present in three pairs, those on IX about twice as long as those on VIII which are just a little longer than those on VII.
|
||
</paragraph>
|
||
<paragraph id="5830313DEE22A266FC9EA787FC23FB46" blockId="24.[839,1467,870,1539]" pageId="24" pageNumber="163">
|
||
Coxite IX of
|
||
<specimenCount id="4E89FAB4EE22A266FC0BA787FBE1FC02" box="[1008,1028,899,922]" pageId="24" pageNumber="163" type="female">♀</specimenCount>
|
||
(
|
||
<figureCitation id="C0B42DB8EE22A266FBE9A787FB89FC03" box="[1042,1132,899,923]" captionStart="Figures 123–126" captionStartId="24.[166,243,781,803]" captionTargetBox="[183,1449,164,769]" captionTargetId="figure-8@24.[166,1466,151,771]" captionTargetPageId="24" captionText="Figures 123–126. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 ♀ (123) coxites VIII and IX, with styli and ovipositor; (124) apex of ovipositor valve; (125) base of cercus and medial dorsal appendage; (126) apex of cercus. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653529" httpUri="https://zenodo.org/record/4653529/files/figure.png" pageId="24" pageNumber="163">Fig. 123</figureCitation>
|
||
) with the internal process acute apically, about three times longer than the external process and 1.3 times as long as wide at its base, reaching to about one third the length of the stylus; external and internal margins of internal process and external margin with a few moderately strong setae, apex of outer process with three or four strong setae. — Ovipositor (
|
||
<figureCitation id="C0B42DB8EE22A266FB33A037FAC6FBD3" box="[1224,1315,1075,1099]" captionStart="Figures 123–126" captionStartId="24.[166,243,781,803]" captionTargetBox="[183,1449,164,769]" captionTargetId="figure-8@24.[166,1466,151,771]" captionTargetPageId="24" captionText="Figures 123–126. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 ♀ (123) coxites VIII and IX, with styli and ovipositor; (124) apex of ovipositor valve; (125) base of cercus and medial dorsal appendage; (126) apex of cercus. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653529" httpUri="https://zenodo.org/record/4653529/files/figure.png" pageId="24" pageNumber="163">Fig. 123</figureCitation>
|
||
) probably not yet fully developed as its only reaches the end of the styli (1.20 HW), composed of about 45 divisions (range 44–47). Distal divisions of gonapophyses difficult to see adequately in prepared material (
|
||
<figureCitation id="C0B42DB8EE22A266FBCEA0ACFB77FB58" box="[1077,1170,1192,1216]" captionStart="Figures 123–126" captionStartId="24.[166,243,781,803]" captionTargetBox="[183,1449,164,769]" captionTargetId="figure-8@24.[166,1466,151,771]" captionTargetPageId="24" captionText="Figures 123–126. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 ♀ (123) coxites VIII and IX, with styli and ovipositor; (124) apex of ovipositor valve; (125) base of cercus and medial dorsal appendage; (126) apex of cercus. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653529" httpUri="https://zenodo.org/record/4653529/files/figure.png" pageId="24" pageNumber="163">Fig. 124</figureCitation>
|
||
) with only short fine setae and setulae.
|
||
</paragraph>
|
||
<paragraph id="5830313DEE22A266FC9EA0E7FAC4F99B" blockId="24.[839,1467,870,1539]" pageId="24" pageNumber="163">
|
||
Cerci (
|
||
<figureCitation id="C0B42DB8EE22A266FC50A0E7FBD9FB63" box="[939,1084,1251,1275]" captionStart="Figures 123–126" captionStartId="24.[166,243,781,803]" captionTargetBox="[183,1449,164,769]" captionTargetId="figure-8@24.[166,1466,151,771]" captionTargetPageId="24" captionText="Figures 123–126. Heterolepisma pyramidum sp. nov. holotype specimen K.261254 ♀ (123) coxites VIII and IX, with styli and ovipositor; (124) apex of ovipositor valve; (125) base of cercus and medial dorsal appendage; (126) apex of cercus. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4653529" httpUri="https://zenodo.org/record/4653529/files/figure.png" pageId="24" pageNumber="163">Figs 125, 126</figureCitation>
|
||
) with basal divisions as long as wide (or perhaps it is several divisions with indistinguishable sutures), divisions two to four shorter than wide, fifth division as long as wide consisting of two annuli and first lanceolate scales seen, following division s consisting of four annuli eventually increasing to eight annuli with the setae, macrochaetae and trichobothria becoming increasingly stronger and longer, lanceolate scales not visible in distal two thirds of cerci and not seen on median dorsal appendage.. — Median dorsal appendage (
|
||
<figureCitation id="C0B42DB8EE22A266FB93A1EFFB27F99B" box="[1128,1218,1515,1539]" captionStart="Figures 127–136" captionStartId="26.[166,243,1700,1722]" captionTargetBox="[170,1457,142,1687]" captionTargetId="figure-8@26.[166,1466,127,1695]" captionTargetPageId="26" captionText="Figures 127–136. Heterolepisma heraldense sp. nov. holotype specimen K.261309 ♀ (127) habitus; (128) scale from urotergite VII; (129) head (cross-hatched area obscured by eye pigment); (130) antenna, scape, pedicel and basal intervals of flagellum; (131) idem, most distal surviving interval showing basiconic sensilla (bsB); (132) mandible (poorly orientated); (133) maxilla, only more prominent setae of palp illustrated; (134) idem, lacinia; (135) idem, ultimate article of palp with its three branched papillae (bp); (136) labium. Scale bars = 0.1 mm unless otherwise indicated." figureDoi="http://doi.org/10.5281/zenodo.4653531" httpUri="https://zenodo.org/record/4653531/files/figure.png" pageId="24" pageNumber="163">Fig. 128</figureCitation>
|
||
) similar.
|
||
</paragraph>
|
||
<paragraph id="5830313DEE22A266FCBCA21FFC18F9AB" blockId="24.[839,1021,1563,1587]" box="[839,1021,1563,1587]" pageId="24" pageNumber="163">
|
||
<emphasis id="6AFBED2FEE22A266FCBCA21FFC9BF9AB" box="[839,894,1563,1587]" italics="true" pageId="24" pageNumber="163">Male</emphasis>
|
||
: Unknown.
|
||
</paragraph>
|
||
<caption id="0CF061B5EE22A266FEF2A259FB24F9EC" box="[265,1217,1629,1653]" pageId="24" pageNumber="163">
|
||
<paragraph id="5830313DEE22A266FEF2A259FB24F9EC" blockId="24.[265,1217,1629,1653]" box="[265,1217,1629,1653]" pageId="24" pageNumber="163">
|
||
<emphasis id="6AFBED2FEE22A266FEF2A259FEBEF9ED" bold="true" box="[265,347,1629,1653]" pageId="24" pageNumber="163">Table 7</emphasis>
|
||
. Number of macrochaetae per bristle comb—
|
||
<taxonomicName id="9F8F4ABEEE22A266FCB0A259FB82F9ED" authority="Smith & Mitchell, 2019" authorityName="Smith & Mitchell" authorityYear="2019" box="[843,1127,1629,1653]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="24" pageNumber="163" phylum="Arthropoda" rank="species" species="pyramidum" status="sp. nov.">
|
||
<emphasis id="6AFBED2FEE22A266FCB0A259FB82F9ED" box="[843,1127,1629,1653]" italics="true" pageId="24" pageNumber="163">Heterolepisma pyramidum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71C85054EE22A266FB95A259FB24F9EC" box="[1134,1217,1629,1652]" pageId="24" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5830313DEE22A266FE48A28BFB31F852" pageId="24" pageNumber="163">
|
||
<table id="2A8FC39DEE225D81FEDAA28BFB0BF852" box="[289,1262,1679,1994]" colsContinueIn="30.[289,1262,1678,1993]" gridcols="6" gridrows="10" pageId="24" pageNumber="163">
|
||
<tr id="E6BF337FEE225D81FEDAA28BFB0BF93F" box="[289,1262,1679,1703]" gridrow="0" pageId="24" pageNumber="163" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-5="1">
|
||
<th id="A56E5A03EE225D81FE48A28BFDFFF93F" box="[435,538,1679,1703]" gridcol="1" gridrow="0" pageId="24" pageNumber="163">urotergite</th>
|
||
<th id="A56E5A03EE225D81FBE8A28BFB60F93F" box="[1043,1157,1679,1703]" gridcol="4" gridrow="0" pageId="24" pageNumber="163">urosternite</th>
|
||
</tr>
|
||
<tr id="E6BF337FEE225D81FEDAA2B3FB0BF948" box="[289,1262,1719,1744]" gridrow="1" pageId="24" pageNumber="163">
|
||
<th id="A56E5A03EE225D81FEDAA2B3FE9FF948" box="[289,378,1719,1744]" gridcol="0" gridrow="1" pageId="24" pageNumber="163">segment</th>
|
||
<td id="A56E5A03EE225D81FE48A2B3FDFFF948" box="[435,538,1719,1744]" gridcol="1" gridrow="1" pageId="24" pageNumber="163">lateral</td>
|
||
<td id="A56E5A03EE225D81FDB0A2B3FD56F948" box="[587,691,1719,1744]" gridcol="2" gridrow="1" pageId="24" pageNumber="163">sublateral</td>
|
||
<td id="A56E5A03EE225D81FD18A2B3FCB4F948" box="[739,849,1719,1744]" gridcol="3" gridrow="1" pageId="24" pageNumber="163">submedial</td>
|
||
<td id="A56E5A03EE225D81FBE8A2B3FB60F948" box="[1043,1157,1719,1744]" gridcol="4" gridrow="1" pageId="24" pageNumber="163">medial</td>
|
||
<td id="A56E5A03EE225D81FB50A2B3FB0BF948" box="[1195,1262,1719,1744]" gridcol="5" gridrow="1" pageId="24" pageNumber="163">lateral</td>
|
||
</tr>
|
||
<tr id="E6BF337FEE225D81FEDAA2E1FB0BF965" box="[289,1262,1765,1789]" gridrow="2" pageId="24" pageNumber="163">
|
||
<th id="A56E5A03EE225D81FEDAA2E1FE9FF965" box="[289,378,1765,1789]" gridcol="0" gridrow="2" pageId="24" pageNumber="163">I</th>
|
||
<td id="A56E5A03EE225D81FE48A2E1FDFFF965" box="[435,538,1765,1789]" gridcol="1" gridrow="2" pageId="24" pageNumber="163">2</td>
|
||
<td id="A56E5A03EE225D81FDB0A2E1FD56F965" box="[587,691,1765,1789]" gridcol="2" gridrow="2" pageId="24" pageNumber="163">1–2</td>
|
||
<td id="A56E5A03EE225D81FD18A2E1FCB4F965" box="[739,849,1765,1789]" gridcol="3" gridrow="2" pageId="24" pageNumber="163">—</td>
|
||
<td id="A56E5A03EE225D81FBE8A2E1FB60F965" box="[1043,1157,1765,1789]" gridcol="4" gridrow="2" pageId="24" pageNumber="163">3</td>
|
||
<td id="A56E5A03EE225D81FB50A2E1FB0BF965" box="[1195,1262,1765,1789]" gridcol="5" gridrow="2" pageId="24" pageNumber="163">—</td>
|
||
</tr>
|
||
<tr id="E6BF337FEE225D81FEDAA306FB0BF882" box="[289,1262,1794,1818]" gridrow="3" pageId="24" pageNumber="163">
|
||
<th id="A56E5A03EE225D81FEDAA306FE9FF882" box="[289,378,1794,1818]" gridcol="0" gridrow="3" pageId="24" pageNumber="163">II</th>
|
||
<td id="A56E5A03EE225D81FE48A306FDFFF882" box="[435,538,1794,1818]" gridcol="1" gridrow="3" pageId="24" pageNumber="163">3</td>
|
||
<td id="A56E5A03EE225D81FDB0A306FD56F882" box="[587,691,1794,1818]" gridcol="2" gridrow="3" pageId="24" pageNumber="163">3</td>
|
||
<td id="A56E5A03EE225D81FD18A306FCB4F882" box="[739,849,1794,1818]" gridcol="3" gridrow="3" pageId="24" pageNumber="163">1</td>
|
||
<td id="A56E5A03EE225D81FBE8A306FB60F882" box="[1043,1157,1794,1818]" gridcol="4" gridrow="3" pageId="24" pageNumber="163">—</td>
|
||
<td id="A56E5A03EE225D81FB50A306FB0BF882" box="[1195,1262,1794,1818]" gridcol="5" gridrow="3" pageId="24" pageNumber="163">4</td>
|
||
</tr>
|
||
<tr id="E6BF337FEE225D81FEDAA31BFB0BF8AF" box="[289,1262,1823,1847]" gridrow="4" pageId="24" pageNumber="163">
|
||
<th id="A56E5A03EE225D81FEDAA31BFE9FF8AF" box="[289,378,1823,1847]" gridcol="0" gridrow="4" pageId="24" pageNumber="163">III</th>
|
||
<td id="A56E5A03EE225D81FE48A31BFDFFF8AF" box="[435,538,1823,1847]" gridcol="1" gridrow="4" pageId="24" pageNumber="163">3</td>
|
||
<td id="A56E5A03EE225D81FDB0A31BFD56F8AF" box="[587,691,1823,1847]" gridcol="2" gridrow="4" pageId="24" pageNumber="163">3</td>
|
||
<td id="A56E5A03EE225D81FD18A31BFCB4F8AF" box="[739,849,1823,1847]" gridcol="3" gridrow="4" pageId="24" pageNumber="163">1</td>
|
||
<td id="A56E5A03EE225D81FBE8A31BFB60F8AF" box="[1043,1157,1823,1847]" gridcol="4" gridrow="4" pageId="24" pageNumber="163">—</td>
|
||
<td id="A56E5A03EE225D81FB50A31BFB0BF8AF" box="[1195,1262,1823,1847]" gridcol="5" gridrow="4" pageId="24" pageNumber="163">4–5</td>
|
||
</tr>
|
||
<tr id="E6BF337FEE225D81FEDAA339FB0BF8CD" box="[289,1262,1853,1877]" gridrow="5" pageId="24" pageNumber="163">
|
||
<th id="A56E5A03EE225D81FEDAA339FE9FF8CD" box="[289,378,1853,1877]" gridcol="0" gridrow="5" pageId="24" pageNumber="163">IV</th>
|
||
<td id="A56E5A03EE225D81FE48A339FDFFF8CD" box="[435,538,1853,1877]" gridcol="1" gridrow="5" pageId="24" pageNumber="163">3–4</td>
|
||
<td id="A56E5A03EE225D81FDB0A339FD56F8CD" box="[587,691,1853,1877]" gridcol="2" gridrow="5" pageId="24" pageNumber="163">3</td>
|
||
<td id="A56E5A03EE225D81FD18A339FCB4F8CD" box="[739,849,1853,1877]" gridcol="3" gridrow="5" pageId="24" pageNumber="163">1</td>
|
||
<td id="A56E5A03EE225D81FBE8A339FB60F8CD" box="[1043,1157,1853,1877]" gridcol="4" gridrow="5" pageId="24" pageNumber="163">—</td>
|
||
<td id="A56E5A03EE225D81FB50A339FB0BF8CD" box="[1195,1262,1853,1877]" gridcol="5" gridrow="5" pageId="24" pageNumber="163">4–5</td>
|
||
</tr>
|
||
<tr id="E6BF337FEE225D81FEDAA35EFB0BF8EA" box="[289,1262,1882,1906]" gridrow="6" pageId="24" pageNumber="163">
|
||
<th id="A56E5A03EE225D81FEDAA35EFE9FF8EA" box="[289,378,1882,1906]" gridcol="0" gridrow="6" pageId="24" pageNumber="163">V</th>
|
||
<td id="A56E5A03EE225D81FE48A35EFDFFF8EA" box="[435,538,1882,1906]" gridcol="1" gridrow="6" pageId="24" pageNumber="163">4</td>
|
||
<td id="A56E5A03EE225D81FDB0A35EFD56F8EA" box="[587,691,1882,1906]" gridcol="2" gridrow="6" pageId="24" pageNumber="163">3</td>
|
||
<td id="A56E5A03EE225D81FD18A35EFCB4F8EA" box="[739,849,1882,1906]" gridcol="3" gridrow="6" pageId="24" pageNumber="163">1</td>
|
||
<td id="A56E5A03EE225D81FBE8A35EFB60F8EA" box="[1043,1157,1882,1906]" gridcol="4" gridrow="6" pageId="24" pageNumber="163">—</td>
|
||
<td id="A56E5A03EE225D81FB50A35EFB0BF8EA" box="[1195,1262,1882,1906]" gridcol="5" gridrow="6" pageId="24" pageNumber="163">4–5</td>
|
||
</tr>
|
||
<tr id="E6BF337FEE225D81FEDAA373FB0BF817" box="[289,1262,1911,1935]" gridrow="7" pageId="24" pageNumber="163">
|
||
<th id="A56E5A03EE225D81FEDAA373FE9FF817" box="[289,378,1911,1935]" gridcol="0" gridrow="7" pageId="24" pageNumber="163">VI</th>
|
||
<td id="A56E5A03EE225D81FE48A373FDFFF817" box="[435,538,1911,1935]" gridcol="1" gridrow="7" pageId="24" pageNumber="163">4</td>
|
||
<td id="A56E5A03EE225D81FDB0A373FD56F817" box="[587,691,1911,1935]" gridcol="2" gridrow="7" pageId="24" pageNumber="163">3</td>
|
||
<td id="A56E5A03EE225D81FD18A373FCB4F817" box="[739,849,1911,1935]" gridcol="3" gridrow="7" pageId="24" pageNumber="163">1</td>
|
||
<td id="A56E5A03EE225D81FBE8A373FB60F817" box="[1043,1157,1911,1935]" gridcol="4" gridrow="7" pageId="24" pageNumber="163">—</td>
|
||
<td id="A56E5A03EE225D81FB50A373FB0BF817" box="[1195,1262,1911,1935]" gridcol="5" gridrow="7" pageId="24" pageNumber="163">5</td>
|
||
</tr>
|
||
<tr id="E6BF337FEE225D81FEDAA391FB0BF835" box="[289,1262,1941,1965]" gridrow="8" pageId="24" pageNumber="163">
|
||
<th id="A56E5A03EE225D81FEDAA391FE9FF835" box="[289,378,1941,1965]" gridcol="0" gridrow="8" pageId="24" pageNumber="163">VII</th>
|
||
<td id="A56E5A03EE225D81FE48A391FDFFF835" box="[435,538,1941,1965]" gridcol="1" gridrow="8" pageId="24" pageNumber="163">3–4</td>
|
||
<td id="A56E5A03EE225D81FDB0A391FD56F835" box="[587,691,1941,1965]" gridcol="2" gridrow="8" pageId="24" pageNumber="163">3</td>
|
||
<td id="A56E5A03EE225D81FD18A391FCB4F835" box="[739,849,1941,1965]" gridcol="3" gridrow="8" pageId="24" pageNumber="163">1</td>
|
||
<td id="A56E5A03EE225D81FBE8A391FB60F835" box="[1043,1157,1941,1965]" gridcol="4" gridrow="8" pageId="24" pageNumber="163">—</td>
|
||
<td id="A56E5A03EE225D81FB50A391FB0BF835" box="[1195,1262,1941,1965]" gridcol="5" gridrow="8" pageId="24" pageNumber="163">2</td>
|
||
</tr>
|
||
<tr id="E6BF337FEE225D81FEDAA3B6FB0BF852" box="[289,1262,1970,1994]" gridrow="9" pageId="24" pageNumber="163">
|
||
<th id="A56E5A03EE225D81FEDAA3B6FE9FF852" box="[289,378,1970,1994]" gridcol="0" gridrow="9" pageId="24" pageNumber="163">VIII</th>
|
||
<td id="A56E5A03EE225D81FE48A3B6FDFFF852" box="[435,538,1970,1994]" gridcol="1" gridrow="9" pageId="24" pageNumber="163">3</td>
|
||
<td id="A56E5A03EE225D81FDB0A3B6FD56F852" box="[587,691,1970,1994]" gridcol="2" gridrow="9" pageId="24" pageNumber="163">—</td>
|
||
<td id="A56E5A03EE225D81FD18A3B6FCB4F852" box="[739,849,1970,1994]" gridcol="3" gridrow="9" pageId="24" pageNumber="163">1</td>
|
||
<td id="A56E5A03EE225D81FBE8A3B6FB60F852" box="[1043,1157,1970,1994]" gridcol="4" gridrow="9" pageId="24" pageNumber="163">—</td>
|
||
<td id="A56E5A03EE225D81FB50A3B6FB0BF852" box="[1195,1262,1970,1994]" gridcol="5" gridrow="9" pageId="24" pageNumber="163">1–2</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="5830313DEE23A267FF82A493FE9CFF54" blockId="25.[121,748,151,205]" pageId="25" pageNumber="164">
|
||
<emphasis id="6AFBED2FEE23A267FF82A493FF35FF37" bold="true" box="[121,208,151,175]" pageId="25" pageNumber="164">Habitat</emphasis>
|
||
. Collected on a remote rock pyramid, the remnant of an old volcanic cone.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="109562B6EE23A267FF82A4E6FDBFFE8F" pageId="25" pageNumber="164" type="etymology">
|
||
<paragraph id="5830313DEE23A267FF82A4E6FDBFFE8F" blockId="25.[121,749,226,279]" pageId="25" pageNumber="164">
|
||
<emphasis id="6AFBED2FEE23A267FF82A4E6FF11FF62" bold="true" box="[121,244,226,250]" pageId="25" pageNumber="164">Etymology</emphasis>
|
||
. Named after the dramatic appearance of the island from which this species was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="109562B6EE23A267FE82A53DFDEAFC7F" pageId="25" pageNumber="164" type="discussion">
|
||
<paragraph id="5830313DEE23A267FE82A53DFE08FECB" blockId="25.[377,493,313,339]" box="[377,493,313,339]" pageId="25" pageNumber="164">
|
||
<emphasis id="6AFBED2FEE23A267FE82A53DFE08FECB" bold="true" box="[377,493,313,339]" pageId="25" pageNumber="164">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="5830313DEE23A267FF82A56CFD08FD47" blockId="25.[121,749,359,1000]" pageId="25" pageNumber="164">
|
||
Even though only a single subadult female specimen was available, the species is described as new because it has a clear morphological difference to the published description of its probable nearest neighbour (
|
||
<taxonomicName id="9F8F4ABEEE23A267FDF6A5C4FD88FE4F" box="[525,621,448,471]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="164" phylum="Arthropoda" rank="species" species="rouxi">
|
||
<emphasis id="6AFBED2FEE23A267FDF6A5C4FD88FE4F" box="[525,621,448,471]" italics="true" pageId="25" pageNumber="164">H. rouxi</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="209871ADEE23A267FD42A5C4FF02FE6D" name="New Caledonia" pageId="25" pageNumber="164">New Caledonia</collectingCountry>
|
||
) and good molecular sequence data are available for it.
|
||
<taxonomicName id="9F8F4ABEEE23A267FF39A5FEFE7AFD8A" box="[194,415,506,530]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="164" phylum="Arthropoda" rank="species" species="rouxi">
|
||
<emphasis id="6AFBED2FEE23A267FF39A5FEFE7AFD8A" box="[194,415,506,530]" italics="true" pageId="25" pageNumber="164">Heterolepisma rouxi</emphasis>
|
||
</taxonomicName>
|
||
is in need of redescription as characters such as the chaetotaxy of the anterior margin of the frons, the position of the trichobothrial areas of the nota and the presence or absence of lanceolate scales were not reported.
|
||
<taxonomicName id="9F8F4ABEEE23A267FEF3A66BFDC5FD1F" authority="Smith & Mitchell, 2019" authorityName="Smith & Mitchell" authorityYear="2019" box="[264,544,623,647]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="164" phylum="Arthropoda" rank="species" species="heraldense" status="sp. nov.">
|
||
<emphasis id="6AFBED2FEE23A267FEF3A66BFDC5FD1F" box="[264,544,623,647]" italics="true" pageId="25" pageNumber="164">Heterolepisma heraldense</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71C85054EE23A267FDDCA674FD99FD1F" box="[551,636,624,647]" pageId="25" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, described below, while distinct from
|
||
<taxonomicName id="9F8F4ABEEE23A267FE50A68AFDA9FD3D" authority="Smith & Mitchell, 2019" authorityName="Smith & Mitchell" authorityYear="2019" box="[427,588,653,677]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="164" phylum="Arthropoda" rank="species" species="pyramidum" status="sp. nov.">
|
||
<emphasis id="6AFBED2FEE23A267FE50A68AFDA9FD3D" box="[427,588,653,677]" italics="true" pageId="25" pageNumber="164">H. pyramidum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71C85054EE23A267FDADA689FD55FD3C" box="[598,688,653,676]" pageId="25" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, may eventually be shown to be conspecific with
|
||
<taxonomicName id="9F8F4ABEEE23A267FDA9A6AFFD4EFD5A" box="[594,683,683,706]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="164" phylum="Arthropoda" rank="species" species="rouxi">
|
||
<emphasis id="6AFBED2FEE23A267FDA9A6AFFD4EFD5A" box="[594,683,683,706]" italics="true" pageId="25" pageNumber="164">H. rouxi</emphasis>
|
||
</taxonomicName>
|
||
when the latter species is redescribed according to current criteria.
|
||
</paragraph>
|
||
<paragraph id="5830313DEE23A267FF6CA6E1FDEAFC7F" blockId="25.[121,749,359,1000]" pageId="25" pageNumber="164">
|
||
The character analysis places
|
||
<taxonomicName id="9F8F4ABEEE23A267FE10A6E2FD6EFD65" authority="Smith & Mitchell, 2019" authorityName="Smith & Mitchell" authorityYear="2019" box="[491,651,741,765]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="164" phylum="Arthropoda" rank="species" species="pyramidum" status="sp. nov.">
|
||
<emphasis id="6AFBED2FEE23A267FE10A6E2FD6EFD65" box="[491,651,741,765]" italics="true" pageId="25" pageNumber="164">H. pyramidum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71C85054EE23A267FD6EA6E1FD08FD64" box="[661,749,741,764]" pageId="25" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in a clade with
|
||
<taxonomicName id="9F8F4ABEEE23A267FEDFA707FE72FC82" authorityName="Smith" authorityYear="2013" box="[292,407,771,794]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="164" phylum="Arthropoda" rank="species" species="parvum">
|
||
<emphasis id="6AFBED2FEE23A267FEDFA707FE72FC82" box="[292,407,771,794]" italics="true" pageId="25" pageNumber="164">H. parvum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="9F8F4ABEEE23A267FE5CA707FDDBFC82" authority="Smith & Mitchell, 2019" authorityName="Smith & Mitchell" authorityYear="2019" box="[423,574,770,794]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="164" phylum="Arthropoda" rank="species" species="heraldense" status="sp. nov.">
|
||
<emphasis id="6AFBED2FEE23A267FE5CA707FDDBFC82" box="[423,574,770,794]" italics="true" pageId="25" pageNumber="164">H. heraldense</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71C85054EE23A267FDBCA707FD79FC82" box="[583,668,771,794]" pageId="25" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="9F8F4ABEEE23A267FD29A707FF36FCAF" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="164" phylum="Arthropoda" rank="species" species="serranoi">
|
||
<emphasis id="6AFBED2FEE23A267FD29A707FF36FCAF" italics="true" pageId="25" pageNumber="164">H. serranoi</emphasis>
|
||
</taxonomicName>
|
||
which all have a glabrous anterior margin to the frons, lanceolate scales, a medial comb on urosternite I and sublateral combs on urosternites II-VII consisting of more than one macrochaeta. The limited molecular data available for
|
||
<taxonomicName id="9F8F4ABEEE23A267FF66A792FEE9FC35" authorityName="Smith" authorityYear="2013" box="[157,268,918,941]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="164" phylum="Arthropoda" rank="species" species="parvum">
|
||
<emphasis id="6AFBED2FEE23A267FF66A792FEE9FC35" box="[157,268,918,941]" italics="true" pageId="25" pageNumber="164">H. parvum</emphasis>
|
||
</taxonomicName>
|
||
(28S of three specimens) and one specimen of
|
||
<taxonomicName id="9F8F4ABEEE23A267FF82A7B7FEEAFC52" authority="Smith & Mitchell, 2019" authorityName="Smith & Mitchell" authorityYear="2019" box="[121,271,946,970]" class="Insecta" family="Lepismatidae" genus="Heterolepisma" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="164" phylum="Arthropoda" rank="species" species="pyramidum" status="sp. nov.">
|
||
<emphasis id="6AFBED2FEE23A267FF82A7B7FEEAFC52" box="[121,271,946,970]" italics="true" pageId="25" pageNumber="164">H. pyramidum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="71C85054EE23A267FEEFA7B7FE81FC52" box="[276,356,947,970]" pageId="25" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
offers no support for this arrangement (see supplementary data on Figshare).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |