258 lines
31 KiB
XML
258 lines
31 KiB
XML
<document id="58E2EDE3843E2A112C6C6C7E634809A9" ID-DOI="http://dx.doi.org/10.3897/BDJ.1.e957" ID-PMC="PMC3964709" ID-Pensoft-Pub="1314-2828--957" ID-PubMed="24723759" ID-ZBK="CABF7796A0F44526917B5286F2C7CA0C" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1314-2828--e957" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="The millipede family Paradoxosomatidae in the Philippines, with a description of Eustrongylosomapenevi sp.n., and notes on Anoplodesmusanthracinus Pocock, 1895, recorded in Malaysia and Sri Lanka for the first time (Diplopoda, Polydesmida)" checkinTime="1553126941720" checkinUser="pensoft" docAuthor="Golovatch, Sergei & Stoev, Pavel" docDate="2013" docId="D001B89A0EA0E1AA03E0540221E6D1F5" docLanguage="en" docName="BiodivDatJour 1: e957" docOrigin="Biodiversity Data Journal 1" docSource="http://dx.doi.org/10.3897/BDJ.1.e957" docTitle="Anoplodesmus anthracinus Pocock 1895" docType="treatment" docVersion="4" lastPageNumber="957" masterDocId="F979FFB6721F7E1F78569D4FFF85FFB0" masterDocTitle="The millipede family Paradoxosomatidae in the Philippines, with a description of Eustrongylosomapenevi sp. n., and notes on Anoplodesmusanthracinus Pocock, 1895, recorded in Malaysia and Sri Lanka for the first time (Diplopoda, Polydesmida)" masterLastPageNumber="957" masterPageNumber="957" pageNumber="957" updateTime="1668122260419" updateUser="ExternalLinkService">
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<mods:titleInfo id="93AF065AFC8B30E2850231002468C1E1">
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<mods:title id="B059E8146912637BD98BFD0421715788">The millipede family Paradoxosomatidae in the Philippines, with a description of Eustrongylosomapenevi sp. n., and notes on Anoplodesmusanthracinus Pocock, 1895, recorded in Malaysia and Sri Lanka for the first time (Diplopoda, Polydesmida)</mods:title>
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<mods:namePart id="C4EE48E200C883FA614B81A9300A88AC">Golovatch, Sergei</mods:namePart>
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<mods:namePart id="E25C0230A7F37108D605485DD24472C8">Stoev, Pavel</mods:namePart>
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<mods:typeOfResource id="A1124328D97CCAF825A066680C38FE61">text</mods:typeOfResource>
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<mods:title id="105AA5B6A6C3222C9DFC1C5911E35E77">Biodiversity Data Journal</mods:title>
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<mods:part id="5C5A1A073198690B466062602037A461">
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<mods:date id="7849F372534EBC4CAA4322B45AD6A6B8">2013</mods:date>
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<mods:detail id="2487AC0CCD4EC7CDF2494BD7130A561D" type="volume">
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<mods:number id="FB1E71B64BBD2AB07A35AEA94C438135">1</mods:number>
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<mods:start id="6054EEE60812260FBA5458658FC8B64A">957</mods:start>
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<mods:end id="0EE167D3ECD48A71C7E6553D21183822">957</mods:end>
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<mods:classification id="F8D695D1A28B17227D395154F0C5B18F">journal article</mods:classification>
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<mods:identifier id="549728D18E1DC8FB1264E64133F60106" type="DOI">http://dx.doi.org/10.3897/BDJ.1.e957</mods:identifier>
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<mods:identifier id="CEB60A027FA1AAC4436EC56F0623780B" type="Pensoft-Pub">1314-2828--957</mods:identifier>
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<treatment id="D001B89A0EA0E1AA03E0540221E6D1F5" LSID="urn:lsid:plazi:treatment:D001B89A0EA0E1AA03E0540221E6D1F5" httpUri="http://treatment.plazi.org/id/D001B89A0EA0E1AA03E0540221E6D1F5" lastPageNumber="957" pageId="0" pageNumber="957">
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<subSubSection id="102E3B1AA38ABB42DCCF856251FBBCEA" pageId="0" pageNumber="957" type="nomenclature">
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<paragraph id="09BB8D22588A85AAA3954F814C974F09" pageId="0" pageNumber="957">
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<taxonomicName id="24377C4279EC272DB13BC15C74AAFED7" ID-CoL="678WD" authority="Pocock, 1895" authorityName="Pocock" authorityYear="1895" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodesmus anthracinus" order="Polydesmida" pageId="0" pageNumber="957" phylum="Arthropoda" rank="species" species="anthracinus">Anoplodesmus anthracinus Pocock, 1895</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="D915A800EC01599B09DC6DFB28C7149A" pageId="0" pageNumber="957" type="materials_examined">
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<paragraph id="5D7521D92D947B39C81E85B4147532E8" pageId="0" pageNumber="957">Materials</paragraph>
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<paragraph id="30930A730CFB0A29F38ACDA9D9EEEAD4" pageId="0" pageNumber="957">
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<materialsCitation id="EAEE587907CA7E7B8E37E5702CFF7F6C" collectingDate="2011-06-15" collectionCode="NMNHS" collectorName="P. Stoev & L. Penev" country="Malaysia" latitude="5.54" location="Malaysia" longLatPrecision="21" longitude="100.46972" pageId="0" pageNumber="957" specimenCount="1" specimenCount-male="1">
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Occurrence: recordedBy:
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<collectorName id="5283068AEA32B827EA3BBFBA2DF0624A" pageId="0" pageNumber="957">P. Stoev & L. Penev</collectorName>
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; individualCount:
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<specimenCount id="65698DAA749CED407D155B9F5A843442" pageId="0" pageNumber="957">1</specimenCount>
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; sex:
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<specimenType id="85421EA2DB5AD2629A67A75F079392C7" pageId="0" pageNumber="957">male</specimenType>
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; behavior: copulation observed; Location: higherGeography: Malay Peninsula; country:
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<collectingCountry id="A07F3F61CD61F24A9305D196B0384706" pageId="0" pageNumber="957">Malaysia</collectingCountry>
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; stateProvince: State Pulau Penang; verbatimLocality: Station MARDI Seberang Perai; locationRemarks: agriculture area, in close proximity to experimental rice fields, under wooden board; verbatimLatitude:
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<geoCoordinate id="97168F90FB9357292A339ED8A982F53F" direction="north" orientation="latitude" precision="15" value="5.54">5°32'24''N</geoCoordinate>
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; verbatimLongitude:
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<geoCoordinate id="17124A5B3AF6F21F9854ED020523EB8C" direction="east" orientation="longitude" precision="15" value="100.46972">100°28'11''E</geoCoordinate>
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; Event: eventDate:
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<collectingDate id="E101839CD4D3B7F3F833D0428B23922F" pageId="0" pageNumber="957" value="2011-06-15">15 June 2011</collectingDate>
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; Record Level: institutionCode:
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<collectionCode id="E93F4A079BB19B4F838534D5BD749704" pageId="0" pageNumber="957">NMNHS</collectionCode>
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</materialsCitation>
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<materialsCitation id="27A4C8D00A7FF4310828E9EE300F6CB3" collectingDate="2011-06-15" collectionCode="NMNHS" collectorName="P. Stoev & L. Penev" country="Malaysia" latitude="5.54" location="Malaysia" longLatPrecision="21" longitude="100.46972" pageId="0" pageNumber="957" specimenCount="2" specimenCount-female="2">
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Occurrence: recordedBy:
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<collectorName id="86838066F06A6B91CAAF68DD60887513" pageId="0" pageNumber="957">P. Stoev & L. Penev</collectorName>
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; individualCount:
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<specimenCount id="6512267DD0C5F3916378DC623D149B0A" pageId="0" pageNumber="957">2</specimenCount>
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; sex:
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<specimenType id="C34C970376680CBA94569984F457834B" pageId="0" pageNumber="957">females</specimenType>
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; behavior: copulation observed; Location: higherGeography: Malay Peninsula; country:
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<collectingCountry id="70AB36146328176C2799B3D58FC69BBF" pageId="0" pageNumber="957">Malaysia</collectingCountry>
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; stateProvince: State Pulau Penang; verbatimLocality: Station MARDI Seberang Perai; locationRemarks: agriculture area, in close proximity to experimental rice fields, under wooden board; verbatimLatitude:
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<geoCoordinate id="4370190E41BDB9F3E6FC2E84CF5682DE" direction="north" orientation="latitude" precision="15" value="5.54">5°32'24''N</geoCoordinate>
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; verbatimLongitude:
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<geoCoordinate id="61F0A35442B4C92C7F39292670233BA2" direction="east" orientation="longitude" precision="15" value="100.46972">100°28'11''E</geoCoordinate>
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; Event: eventDate:
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<collectingDate id="0F3BE52B6A037A69BA793E5DD1BBEF17" pageId="0" pageNumber="957" value="2011-06-15">15 June 2011</collectingDate>
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; Record Level: institutionCode:
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<collectionCode id="CF880788F6B5BE1CF3C55F3923F482D0" pageId="0" pageNumber="957">NMNHS</collectionCode>
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</materialsCitation>
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<materialsCitation id="040AC3959D0601DE929126D2C1468D7A" collectingDate="2012-12-19" collectionCode="ZMUM" collectorName="I. Melnik" country="Sri Lanka" latitude="7.277778" location="Sri Lanka" longLatPrecision="21" longitude="80.386665" pageId="0" pageNumber="957" specimenCount="1" specimenCount-male="1">
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Occurrence: recordedBy:
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<collectorName id="12BE223F17BF54ECEFA6C13C34370583" pageId="0" pageNumber="957">I. Melnik</collectorName>
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; individualCount:
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<specimenCount id="CB5F60CE3B0C1A2B0E904BBF06ECD63A" pageId="0" pageNumber="957">1</specimenCount>
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; sex:
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<specimenType id="C0BB0E6C98962F3E5B39F8B58628D8D3" pageId="0" pageNumber="957">male</specimenType>
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; Location: country:
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<collectingCountry id="E9E3E7AA94A93CC955E43BEE83DB692D" pageId="0" pageNumber="957">Sri Lanka</collectingCountry>
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; stateProvince: Sabaragamuwa Prov.; verbatimLocality: Millenium Foundation Orphanage; verbatimElevation:
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<elevation id="282CD4F35974E4F06A7904782653EB45" pageId="0" pageNumber="957">90 m</elevation>
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; verbatimLatitude:
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<geoCoordinate id="3230AFFCB8F0C994C9D8400FF68EF4C2" direction="north" orientation="latitude" precision="15" value="7.277778">7°16'40"N</geoCoordinate>
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; verbatimLongitude:
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<geoCoordinate id="D1F9C893D21D6E9B62734C48B44DCE87" direction="east" orientation="longitude" precision="15" value="80.386665">80°23'12"E</geoCoordinate>
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; Event: eventDate:
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<collectingDate id="4241E1756C7E425A838A382DCCEBF7CC" pageId="0" pageNumber="957" value="2012-12-19" valueMax="2012-12-22" valueMin="2012-12-19">19-22.XII.2012</collectingDate>
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; Record Level: institutionCode:
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<collectionCode id="D40F33B0097F82128D168A746F269546" pageId="0" pageNumber="957">ZMUM</collectionCode>
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</materialsCitation>
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<materialsCitation id="45A444AFFB51D84D68C9D1440FBAF842" collectingDate="2012-12-19" collectionCode="ZMUM" collectorName="I. Melnik" country="Sri Lanka" latitude="7.277778" location="Sri Lanka" longLatPrecision="21" longitude="80.386665" pageId="0" pageNumber="957" specimenCount="2" specimenCount-female="2">
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Occurrence: recordedBy:
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<collectorName id="48583F003DE2418131DE0494F3AD2558" pageId="0" pageNumber="957">I. Melnik</collectorName>
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; individualCount:
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<specimenCount id="2CFBE4F3C7E62AAB136EC8FD82A18719" pageId="0" pageNumber="957">2</specimenCount>
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; sex:
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<specimenType id="A477DAAEC2AF05640E5DC9214EA2ED29" pageId="0" pageNumber="957">females</specimenType>
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; Location: country:
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<collectingCountry id="2A44E035B52257364F968993DD7DB902" pageId="0" pageNumber="957">Sri Lanka</collectingCountry>
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; stateProvince: Sabaragamuwa Prov.; verbatimLocality: Millenium Foundation Orphanage; verbatimElevation:
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<elevation id="35B1E0698EBE7918D39F3C2AC0B30D74" pageId="0" pageNumber="957">90 m</elevation>
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; verbatimLatitude:
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<geoCoordinate id="8E5F48033ED895F27DE0CAA56E0EC62A" direction="north" orientation="latitude" precision="15" value="7.277778">7°16'40"N</geoCoordinate>
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; verbatimLongitude:
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<geoCoordinate id="7EB779131625035900F2A42E6D559F1F" direction="east" orientation="longitude" precision="15" value="80.386665">80°23'12"E</geoCoordinate>
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; Event: eventDate:
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<collectingDate id="02966A46000C4BC4AC2C554265DA534D" pageId="0" pageNumber="957" value="2012-12-19" valueMax="2012-12-22" valueMin="2012-12-19">19-22.XII.2012</collectingDate>
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; Record Level: institutionCode:
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<collectionCode id="39C1A137474DCA899C46B6B9EDA83A34" pageId="0" pageNumber="957">ZMUM</collectionCode>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="ED9F697F31612CD0EFF0DF5BF9133CEB" pageId="0" pageNumber="957" type="description">
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<paragraph id="4BE53552B3311740DCBFBE32ECD958CF" pageId="0" pageNumber="957">Description</paragraph>
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<paragraph id="9A4BF95C3D9C6831DA15CBA13D2848A8" pageId="0" pageNumber="957">Length ca 33 mm, width of pro- and metazona 2.8 and 3.8 mm, respectively (♂), or 25, 3.0 and 4.0 mm, respectively (♀). Colour pattern highly vivid (Fig. 5), shiny blackish to dark brown, with contrasting yellowish paraterga and the immediately adjacent regions. Paraterga very well-developed, set rather high (about 1/4th of midbody height) (Fig. 6a), callus wide (Fig. 6b), thicker in pore-bearing paraterga. Pleurosternal carinae longitudinally arched ribs, increasingly poorly developed towards telson to totally disappear in segment 15. Legs only slightly enlarged in male, rather long and slender, about 1.3 (♂) or 0.9 (♀) times as long as midbody height. ♂ legs 5 (Fig. 7a) and 6 with large femoral humps, ♂ femur 7 with a humped process even greater than in leg 6 (Fig. 7b). Epiproct subtruncate (Fig. 6c). Hypoproct roundly subtrapeziform (Fig. 6d). Sternal lamina between ♂ coxae 4 semi-circular (Fig. 6e).</paragraph>
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<paragraph id="B27617611D13ECCD43568C5CCE33BDC5" pageId="0" pageNumber="957">Gonopods very simple (Fig. 6f): coxite with a few strong setae distodorsally, prefemoral part prominent, only slightly shorter than acropodite; femorite with a strong ventral tooth (a), solenophore bipartite, with two apical lobes (b, c), lobe c supporting a short solenomere (sl).</paragraph>
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</subSubSection>
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<subSubSection id="89B76ACE4E5BE3692C6E00D4D85058CC" pageId="0" pageNumber="957" type="notes">
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<paragraph id="6199DC3B35570009A5AF06175CB3B73F" pageId="0" pageNumber="957">Notes</paragraph>
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<paragraph id="7E4820E9E2E9E826EBF8ED1F77B62AB3" pageId="0" pageNumber="957">
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This species was originally described from Yangon (= Rangoon), Myanmar (
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<bibRefCitation id="7FC98B2DD6EC5A9B4242CD9E1DC1EEBF" author="Pocock, RI" journalOrPublisher="Annali del Museo Civico di Storia Naturale " Giacomo Doria ", Genova" pageId="0" pageNumber="957" pagination="787 - 834" title="The Myriapoda of Burma, Pt. IV. Report upon the Polydesmoidea collected by Sig. L. Fea, Mr. E. W. Oates and others." volume="2" year="1895">Pocock 1895</bibRefCitation>
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).
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<bibRefCitation id="BB0610D06B3978CD716D59380FB3F299" author="Attems, C" journalOrPublisher="Das Tierreich" pageId="0" pageNumber="957" pagination="1 - 300" title="Myriapoda 3. Polydesmoidea I. Fam. Strongylosomidae." volume="68" year="1937">Attems (1937)</bibRefCitation>
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synonymized it with
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<taxonomicName id="2FAFEC6754C7E6052B73343113F38B6D" class="Diplopoda" family="Paradoxosomatidae" genus="Jonespeltis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Jonespeltis splendidus" order="Polydesmida" pageId="0" pageNumber="957" phylum="Arthropoda" rank="species" species="splendidus">Jonespeltis splendidus</taxonomicName>
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Verhoeff, 1936, from southern India, but
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<bibRefCitation id="8B61774A0F00B9250A611683E311696A" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="957" pagination="95 - 144" title="A revision of the Burmese Paradoxosomatidae (Diplopoda, Polydesmida) in the Museo Civico di Storia Naturale at Genoa (Part I)." volume="108" year="1965">Jeekel (1965)</bibRefCitation>
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revalidated the latter species and returned
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<taxonomicName id="AA446C65BD0371E0668383916F47CDE2" lsidName="A. anthracinus" pageId="0" pageNumber="957" rank="species" species="anthracinus">A. anthracinus</taxonomicName>
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to its original scope. Furthermore, Jeekel provided very useful illustrations and a detailed redescription of the species, based on a part of the type series.
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<bibRefCitation id="91F9301B4263652A29FDCCF92856DB90" author="Hoffman, RL" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="957" pagination="361 - 389" title="Descriptions and allocation of new or poorly known genera and species of Paradoxomatidae from south-eastern Asia (Diplopoda: Polydesmida)." volume="7" year="1973">Hoffman (1973)</bibRefCitation>
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gave more illustrations of the gonopods, based on a paratype of
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<taxonomicName id="5BA1962B610D832A8965ACDA0D4172AD" lsidName="A. kathanus" pageId="0" pageNumber="957" rank="species" species="kathanus">A. kathanus</taxonomicName>
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(Chamberlin, 1921), from Katha, north of Yangon, Myanmar, and synonymized it with
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<taxonomicName id="0E96F07F1F0173622D798C5C7EBF4FD2" lsidName="A. anthracinus" pageId="0" pageNumber="957" rank="species" species="anthracinus">A. anthracinus</taxonomicName>
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.
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</paragraph>
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<paragraph id="BFEDAE45664024B8E39C6F961C861C8E" pageId="0" pageNumber="957">
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Our record of
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<taxonomicName id="44D87E4B9EA691641FCB3537C6241EB0" lsidName="A. anthracinus" pageId="0" pageNumber="957" rank="species" species="anthracinus">A. anthracinus</taxonomicName>
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in the State of Pulau Penang, Malaysia considerably extends the range of this species to the south. The studied sample agrees well with the description provided by
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<bibRefCitation id="AEA73200E6C9F7A2BF1FAA6EFDF784F8" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="957" pagination="95 - 144" title="A revision of the Burmese Paradoxosomatidae (Diplopoda, Polydesmida) in the Museo Civico di Storia Naturale at Genoa (Part I)." volume="108" year="1965">Jeekel (1965)</bibRefCitation>
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and
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<bibRefCitation id="177CDD30AA8EE5157CEC0E13D62D9B9E" author="Hoffman, RL" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="957" pagination="361 - 389" title="Descriptions and allocation of new or poorly known genera and species of Paradoxomatidae from south-eastern Asia (Diplopoda: Polydesmida)." volume="7" year="1973">Hoffman (1973)</bibRefCitation>
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in most characters (Figs 6f, 7), including humps in ♂ femora 5 and 6, as well as a process surmounting a hump in ♂ femur 7. Only slight variations have been noticed in the shapes of paraterga and sternal lobe between ♂ coxae 4. The same can be said about the samples from Sri Lanka which are also identified as
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<taxonomicName id="77D95B717288F479958164805D6E26E7" lsidName="A. anthracinus" pageId="0" pageNumber="957" rank="species" species="anthracinus">A. anthracinus</taxonomicName>
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.
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</paragraph>
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<paragraph id="E394BBFC6A9F0DEB351798BF759BFE4B" pageId="0" pageNumber="957">These are the first formal records of the species in Malaysia and Sri Lanka (Figs 8, 10). However, actually they might well represent introductions. In fact, in Malaysia the species was observed and collected in a highly agricultural and urbanized area, in close proximity to experimental rice fields (Fig. 9), while in Sri Lanka, the collecting locality is a human settlement.</paragraph>
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<paragraph id="1017D431DF4ABC363C0C931E1B58D801" pageId="0" pageNumber="957">
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It is noteworthy that Sri Lanka hosts several formal species of
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<taxonomicName id="BCDC34557C841D54CF280CB3029C68AD" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodesmus" order="Polydesmida" pageId="0" pageNumber="957" phylum="Arthropoda" rank="genus">Anoplodesmus</taxonomicName>
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, nearly all very similar to one another:
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</paragraph>
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<paragraph id="9BD021D6D4AE6629F005A78D91828BDB" pageId="0" pageNumber="957">
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<taxonomicName id="C8ACFB9CBCEDB9F90FCFAA3D4EEF406F" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodesmus saussurii" order="Polydesmida" pageId="0" pageNumber="957" phylum="Arthropoda" rank="species" species="saussurii">Anoplodesmus saussurii</taxonomicName>
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(Humbert, 1865), originally described from Sri Lanka, later recorded also in Fiji and Mauritius (
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<bibRefCitation id="23B45B827BBD84239034B292B48C002A" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="957" pagination="95 - 144" title="A revision of the Burmese Paradoxosomatidae (Diplopoda, Polydesmida) in the Museo Civico di Storia Naturale at Genoa (Part I)." volume="108" year="1965">Jeekel 1965</bibRefCitation>
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,
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<bibRefCitation id="3D18466FBBA140CE73F9E234FA656BAD" author="Jeekel, CAW" journalOrPublisher="Beaufortia" pageId="0" pageNumber="957" pagination="1 - 5" title="The " endemic " paradoxosomatids (Diplopoda, Polydesmida) of the Fiji Islands." volume="20" year="1972">Jeekel 1972</bibRefCitation>
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,
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<bibRefCitation id="792021DE8140422CF798B82E5CB3309B" author="Jeekel, CAW" journalOrPublisher="Entomologische Berichten" pageId="0" pageNumber="957" pagination="122 - 127" title="Records of Diplopoda of the order Polydesmida from the Fiji Islands." volume="40" year="1980">Jeekel 1980a</bibRefCitation>
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). The only meaningful difference from
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<taxonomicName id="B12F52EAB2A2EE0E5A55B9EAF36565EB" lsidName="A. anthracinus" pageId="0" pageNumber="957" rank="species" species="anthracinus">A. anthracinus</taxonomicName>
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is said to lie in the absence of a ventral hump in ♂ femur 5. However, given considerable variation in the presence or absence of this character, when such a hump in
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<taxonomicName id="26CA9590A18D30518816B5537ADE0439" lsidName="A. anthracinus" pageId="0" pageNumber="957" rank="species" species="anthracinus">A. anthracinus</taxonomicName>
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can either be present in or absent from ♂ femur 4 (
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<bibRefCitation id="CB3CB755A76BB52A519346EA102B2F2C" author="Attems, C" journalOrPublisher="Das Tierreich" pageId="0" pageNumber="957" pagination="1 - 300" title="Myriapoda 3. Polydesmoidea I. Fam. Strongylosomidae." volume="68" year="1937">Attems 1937</bibRefCitation>
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,
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<bibRefCitation id="70F8BDF64598667A21ED3B0F925741D1" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="957" pagination="95 - 144" title="A revision of the Burmese Paradoxosomatidae (Diplopoda, Polydesmida) in the Museo Civico di Storia Naturale at Genoa (Part I)." volume="108" year="1965">Jeekel 1965</bibRefCitation>
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), its status versus the older name
|
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<taxonomicName id="C875EABB668045D014F36792B8F7A367" lsidName="A. saussurii" pageId="0" pageNumber="957" rank="species" species="saussurii">A. saussurii</taxonomicName>
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is to be questioned.
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</paragraph>
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<paragraph id="00C193CB32993307AAD570F84C698D98" pageId="0" pageNumber="957">
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<taxonomicName id="AC41C02299F2103B3AD581222C7ABC07" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodesmus luctuosus" order="Polydesmida" pageId="0" pageNumber="957" phylum="Arthropoda" rank="species" species="luctuosus">Anoplodesmus luctuosus</taxonomicName>
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(Peters, 1864), from Rambodde;
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<taxonomicName id="82B6390567979E0809E8DC0B8097941C" lsidName="A. inornatus" pageId="0" pageNumber="957" rank="species" species="inornatus">A. inornatus</taxonomicName>
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(Humbert, 1865),
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<taxonomicName id="14E47729EE5590BEDD6C6378A1A82176" lsidName="A. layardi" pageId="0" pageNumber="957" rank="species" species="layardi">A. layardi</taxonomicName>
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(Humbert, 1865),
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|
<taxonomicName id="FF532DC76EC09D2D48E0EAE5E45148F5" lsidName="A. thwaitesii" pageId="0" pageNumber="957" rank="species" species="thwaitesii">A. thwaitesii</taxonomicName>
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|
(Humbert, 1865) and
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|
<taxonomicName id="72F1EF7A60F2FB4414514031D9BF9DD9" lsidName="A. humberti" pageId="0" pageNumber="957" rank="species" species="humberti">A. humberti</taxonomicName>
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(Carl, 1902), all from Paradeniya; and
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|
<taxonomicName id="4398595E2FF781E92C1B2524023204E7" lsidName="A. sabulosus" pageId="0" pageNumber="957" rank="species" species="sabulosus">A. sabulosus</taxonomicName>
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Attems, 1898, from Kandy. All of them have been described from Sri Lanka, still known only from that island. Some of these taxa are however dubious, being based on female or even juvenile material, but most could be included into a key (
|
|
<bibRefCitation id="904950F8F632AEE92CDE62502B00322C" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="957" pagination="95 - 144" title="A revision of the Burmese Paradoxosomatidae (Diplopoda, Polydesmida) in the Museo Civico di Storia Naturale at Genoa (Part I)." volume="108" year="1965">Jeekel 1965</bibRefCitation>
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|
). Regrettably, the first couplet in the key is purely geographic, separating the species from Myanmar and Sumatra from those described from Sri Lanka and India (
|
|
<bibRefCitation id="EB18182334E6C4CCD8376A3353C69E67" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="957" pagination="95 - 144" title="A revision of the Burmese Paradoxosomatidae (Diplopoda, Polydesmida) in the Museo Civico di Storia Naturale at Genoa (Part I)." volume="108" year="1965">Jeekel 1965</bibRefCitation>
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|
). As one can see from the presently known distributions of
|
|
<taxonomicName id="FF0FFDDE02F182C1F3F5455E4781FAC2" lsidName="A. saussurii" pageId="0" pageNumber="957" rank="species" species="saussurii">A. saussurii</taxonomicName>
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|
and
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|
<taxonomicName id="F16F8A5295B2BB784473FDE7B7DC923F" lsidName="A. anthracinus" pageId="0" pageNumber="957" rank="species" species="anthracinus">A. anthracinus</taxonomicName>
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|
, this distinction does not hold, also strongly suggesting several introductions through human agency. The only feasible solution lies in collecting new and/or spotting topotypic museum samples of the still enigmatic
|
|
<taxonomicName id="2BAB2504EDA58E2BEEFFCF0B8C7870CC" lsidName="A. inornatus" pageId="0" pageNumber="957" rank="species" species="inornatus">A. inornatus</taxonomicName>
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|
and
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|
<taxonomicName id="DDD26D945918BAF0F10B87A4FF5430A9" lsidName="A. layardi" pageId="0" pageNumber="957" rank="species" species="layardi">A. layardi</taxonomicName>
|
|
from Paradeniya, and of
|
|
<taxonomicName id="0DC5F2E9E1CE050267473F5C00AD398D" lsidName="A. sabulosus" pageId="0" pageNumber="957" rank="species" species="sabulosus">A. sabulosus</taxonomicName>
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|
from Kandy, to properly compare them to their type material. In addition, bar-coding could help tracing genetic relationships. Last, but not least, a few congeneric species, most of which also very similar to
|
|
<taxonomicName id="4C6F9C7E6A223FD34A182467D1404AF3" lsidName="A. anthracinus" pageId="0" pageNumber="957" rank="species" species="anthracinus">A. anthracinus</taxonomicName>
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|
, are known to occur in southern India as well.
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|
</paragraph>
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|
<paragraph id="28ADEA852F0F4C226A3DA68D2BF46512" pageId="0" pageNumber="957">
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Since
|
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<taxonomicName id="7530F66118FE606F9CADF5E5AF3F39BF" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodesmus" order="Polydesmida" pageId="0" pageNumber="957" phylum="Arthropoda" rank="genus">Anoplodesmus</taxonomicName>
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|
is a senior synonym of
|
|
<taxonomicName id="9EEBE3D1DE3A4C33CCAAB6910DFD3755" class="Diplopoda" family="Paradoxosomatidae" genus="Paranedyopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paranedyopus" order="Polydesmida" pageId="0" pageNumber="957" phylum="Arthropoda" rank="genus">Paranedyopus</taxonomicName>
|
|
(
|
|
<bibRefCitation id="B7CB71FDCC26A7C82355C1B41AD91E81" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="957" pagination="215 - 220" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda, Polydesmida), VII." volume="8" year="2000">Golovatch 2000</bibRefCitation>
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|
,
|
|
<bibRefCitation id="0478C8A7AEEA9614C49D10DD6C737B6F" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="957" pagination="1 - 31" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda, Polydesmida), XIII." volume="22" year="2013">Golovatch 2013</bibRefCitation>
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|
), the sole erstwhile component species of the latter genus from Sri Lanka,
|
|
<taxonomicName id="72EA4795CA57F0CEF0352249993BA7A2" lsidName="A. simplex" pageId="0" pageNumber="957" rank="species" species="simplex">A. simplex</taxonomicName>
|
|
(Humbert, 1865), from Pundaloya (
|
|
<bibRefCitation id="2313B956C80747E77BD9095C7873201C" author="Jeekel, CAW" journalOrPublisher="Beaufortia" pageId="0" pageNumber="957" pagination="163 - 178" title="On some little known Paradoxosomatidae from India and Ceylon, with the description of four new genera (Diplopoda, Polydesmida)." volume="30" year="1980">Jeekel 1980c</bibRefCitation>
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|
), must be considered as well. However, like any former
|
|
<taxonomicName id="1E6881D63F9CA476B36ED743A98549C5" class="Diplopoda" family="Paradoxosomatidae" genus="Paranedyopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paranedyopus" order="Polydesmida" pageId="0" pageNumber="957" phylum="Arthropoda" rank="genus">Paranedyopus</taxonomicName>
|
|
species,
|
|
<taxonomicName id="3A265E9ED583C7B8285FAFA5A9476C74" lsidName="A. simplex" pageId="0" pageNumber="957" rank="species" species="simplex">A. simplex</taxonomicName>
|
|
shows reduced paraterga and more elaborate gonopods (
|
|
<bibRefCitation id="D55E494E606B2AD1A00B74D2BD61C499" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="957" pagination="1 - 31" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda, Polydesmida), XIII." volume="22" year="2013">Golovatch 2013</bibRefCitation>
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|
). In other words,
|
|
<taxonomicName id="C71C1998000F85CF9CB4A1B01790DA83" lsidName="A. simplex" pageId="0" pageNumber="957" rank="species" species="simplex">A. simplex</taxonomicName>
|
|
is quite distinct from the above congeners from Sri Lanka which all have strongly developed paraterga and highly simple gonopods. In contrast, it seems to be more similar to
|
|
<taxonomicName id="C59C7FCBC8D21568A1AA759F338FFB97" lsidName="A. rufocinctus" pageId="0" pageNumber="957" rank="species" species="rufocinctus">A. rufocinctus</taxonomicName>
|
|
(Carl, 1932) and
|
|
<taxonomicName id="81D71285569971ABD26B7D4134A14B4F" lsidName="A. subcylindricus" pageId="0" pageNumber="957" rank="species" species="subcylindricus">A. subcylindricus</taxonomicName>
|
|
(Carl, 1932), both latter taxa from southern India (
|
|
<bibRefCitation id="78E678EC1AABF80BACB1E465CA473EEA" author="Jeekel, CAW" journalOrPublisher="Beaufortia" pageId="0" pageNumber="957" pagination="163 - 178" title="On some little known Paradoxosomatidae from India and Ceylon, with the description of four new genera (Diplopoda, Polydesmida)." volume="30" year="1980">Jeekel 1980c</bibRefCitation>
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).
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|
</paragraph>
|
|
<paragraph id="150EFB419C1F5B4182DDB11A72769BFA" pageId="0" pageNumber="957">
|
|
<taxonomicName id="8A71E5BD3D2B1764656D0205D80EA8BE" class="Diplopoda" family="Paradoxosomatidae" genus="Anoplodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoplodesmus anthracinus" order="Polydesmida" pageId="0" pageNumber="957" phylum="Arthropoda" rank="species" species="anthracinus">Anoplodesmus anthracinus</taxonomicName>
|
|
, new to the fauna of Sri Lanka.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |