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<document id="A97B342EDF66E17D21CB4F238348D6D8" ID-CLB-Dataset="6330" ID-DOI="10.5281/zenodo.4690007" ID-GBIF-Dataset="0baa733e-6e79-4600-a069-3a1a9f87fbec" ID-ISSN="1638-9387" ID-Zenodo-Dep="4690007" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618438256252" checkinUser="felipe" docAuthor="Monniot, Françoise &amp; Monniot, Claude" docDate="2008" docId="CF5AF0013B3F5C2BEB8DFE5BFCD0C57B" docLanguage="fr" docName="Zoosystema.30.4.799-872.pdf" docOrigin="Zoosystema 30 (4)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Pseudodistoma nigrum Monniot &amp; Monniot 2008, n. sp." docType="treatment" docVersion="5" lastPageNumber="836" masterDocId="336388793B1A5C0EEB52FFEEFFBAC617" masterDocTitle="Compléments sur la diversité des ascidies (Ascidiacea, Tunicata) de louest Pacifique tropical" masterLastPageNumber="872" masterPageNumber="799" pageNumber="836" updateTime="1698936655835" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0A100E717AF328F2E7A5741E7D8D1AA6">Compléments sur la diversité des ascidies (Ascidiacea, Tunicata) de louest Pacifique tropical</mods:title>
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<mods:namePart id="C92C8E4683FF806E8E7BF2F19A5EF193">Monniot, Françoise</mods:namePart>
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<mods:namePart id="218488CB0D53BF61EDA312D01D8A3156">Monniot, Claude</mods:namePart>
<mods:affiliation id="79D45B965AE9AA84DE9A5083D0341491">Muséum national dHistoire naturelle, Département Milieux et Peuplements aquatiques, case postale 51, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) monniot @ mnhn. fr</mods:affiliation>
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<mods:title id="1CB9AC639C1C328AC91B387BCC30B222">Zoosystema</mods:title>
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<mods:date id="6CEF12A6F938DAF7475931FE979F3F1B">2008</mods:date>
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<treatment id="CF5AF0013B3F5C2BEB8DFE5BFCD0C57B" ID-DOI="http://doi.org/10.5281/zenodo.4893689" ID-GBIF-Taxon="182485179" ID-Zenodo-Dep="4893689" LSID="urn:lsid:plazi:treatment:CF5AF0013B3F5C2BEB8DFE5BFCD0C57B" httpUri="http://treatment.plazi.org/id/CF5AF0013B3F5C2BEB8DFE5BFCD0C57B" lastPageNumber="836" pageId="37" pageNumber="836">
<subSubSection id="0FE9129C3B3F5C2BEB8DFE5BFDA3C7C6" box="[223,537,437,465]" pageId="37" pageNumber="836" type="nomenclature">
<paragraph id="474C41173B3F5C2BEB8DFE5BFDA3C7C6" blockId="37.[223,537,437,496]" box="[223,537,437,465]" pageId="37" pageNumber="836">
<heading id="1C04F67B3B3F5C2BEB8DFE5BFDA3C7C6" box="[223,537,437,465]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="836" reason="2">
<taxonomicName id="80F33A943B3F5C2BEB8DFE5BFE6DC7D8" authority="Monniot &amp; Monniot, 2008" authorityName="Monniot &amp; Monniot" authorityYear="2008" box="[223,471,437,464]" class="Ascidiacea" family="Pseudodistomidae" genus="Pseudodistoma" kingdom="Animalia" order="Enterogona" pageId="37" pageNumber="836" phylum="Chordata" rank="species" species="nigrum" status="sp. nov.">
<emphasis id="75879D053B3F5C2BEB8DFE5BFE6DC7D8" bold="true" box="[223,471,437,464]" italics="true" pageId="37" pageNumber="836">Pseudodistoma nigrum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EB4207E3B3F5C2BEA8CFE58FDA3C7C6" box="[478,537,438,465]" pageId="37" pageNumber="836" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="0FE9129C3B3F5C2BEA1FFE3BFCD0C57B" pageId="37" pageNumber="836" type="description">
<paragraph id="474C41173B3F5C2BEA1FFE3BFE11C7E7" blockId="37.[223,537,437,496]" box="[333,427,469,496]" pageId="37" pageNumber="836">
(
<figureCitation id="DFC85D923B3F5C2BEA07FE3BFE19C7E7" box="[341,419,469,496]" captionStart="FIG" captionStartId="38.[677,688,1494,1511]" captionTargetBox="[677,1210,220,1457]" captionTargetId="figure-19@38.[649,1231,201,1467]" captionTargetPageId="38" captionText="FIG. 49. — Pseudodistoma nigrum n. sp.: A, zoïde; B, C, zoïde en deux parties; D, détail de lestomac; E, larve. Échelles: A,2 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4690125" httpUri="https://zenodo.org/record/4690125/files/figure.png" pageId="37" pageNumber="836">Fig. 49</figureCitation>
)
</paragraph>
<paragraph id="474C41173B3F5C2BEB23FDFEFDAFC44C" blockId="37.[113,647,526,603]" pageId="37" pageNumber="836">
<typeStatus id="9848FFB53B3F5C2BEB23FDFEFF6CC432" box="[113,214,528,552]" pageId="37" pageNumber="836" type="syntype">SYNTYPES</typeStatus>
. —
<emphasis id="75879D053B3F5C2BEA5CFDE0FE52C431" bold="true" box="[270,488,526,550]" pageId="37" pageNumber="836">
<collectingCountry id="3FE401873B3F5C2BEA5CFDE0FE52C431" box="[270,488,526,550]" name="New Caledonia" pageId="37" pageNumber="836">Nouvelle-Calédonie</collectingCountry>
</emphasis>
.
<geoCoordinate id="22C727D03B3F5C2BEAA8FDE0FD3BC431" box="[506,641,526,550]" degrees="18" direction="south" minutes="07.013" orientation="latitude" pageId="37" pageNumber="836" precision="1" value="-18.116884">18°07.013S</geoCoordinate>
,
<geoCoordinate id="22C727D03B3F5C2BEB23FDC7FEB8C456" box="[113,258,553,577]" degrees="162" direction="east" minutes="52.926" orientation="longitude" pageId="37" pageNumber="836" precision="1" value="162.8821">162°52.926E</geoCoordinate>
,
<date id="334D67D73B3F5C2BEA5CFDC7FE2DC456" box="[270,407,553,577]" pageId="37" pageNumber="836" value="2003-08-16">16.VIII.2003</date>
(UA 406 st 736), coll. Laboute, 2 colonies (MNHN A1 PSE 74).
</paragraph>
<paragraph id="474C41173B3F5C2BEB23FD9AFE5EC49C" blockId="37.[113,484,627,652]" box="[113,484,627,652]" pageId="37" pageNumber="836">
ÉTYMOLOGIE. — Du latin
<emphasis id="75879D053B3F5C2BEA25FD9DFE1CC49D" box="[375,422,627,650]" italics="true" pageId="37" pageNumber="836">niger</emphasis>
: noir.
</paragraph>
<paragraph id="474C41173B3F5C2BEB23FD5AFEB8C4DB" blockId="37.[111,649,692,1700]" box="[113,258,692,718]" pageId="37" pageNumber="836">DESCRIPTION</paragraph>
<paragraph id="474C41173B3F5C2BEB23FD3DFF56C5DC" blockId="37.[111,649,692,1700]" pageId="37" pageNumber="836">
Les colonies forment des masses arrondies mamelonnées portées par un pédoncule court incrusté de sable. La plus grande colonie mesure 3,5 cm de haut. La partie supérieure est constituée de lobes soudés dont la surface est lisse sans incrustation. La couleur
<emphasis id="75879D053B3F5C2BEBBEFC9CFE89C59C" box="[236,307,882,907]" italics="true" pageId="37" pageNumber="836">in vivo</emphasis>
est crème et devient noire après fixation, mais la tunique du pédoncule nest pas pigmentée.
</paragraph>
<paragraph id="474C41173B3F5C2BEBDAFC3EFDEBC23C" blockId="37.[111,649,692,1700]" pageId="37" pageNumber="836">
Les zoïdes (
<figureCitation id="DFC85D923B3F5C2BEBAFFC3EFEE0C5FC" box="[253,346,976,1003]" captionStart="FIG" captionStartId="38.[677,688,1494,1511]" captionTargetBox="[677,1210,220,1457]" captionTargetId="figure-19@38.[649,1231,201,1467]" captionTargetPageId="38" captionText="FIG. 49. — Pseudodistoma nigrum n. sp.: A, zoïde; B, C, zoïde en deux parties; D, détail de lestomac; E, larve. Échelles: A,2 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4690125" httpUri="https://zenodo.org/record/4690125/files/figure.png" pageId="37" pageNumber="836">Fig. 49A</figureCitation>
) fixés sont incolores et mesurent jusquà
<quantity id="800BECF23B3F5C2BEBABFC1EFEF3C21C" box="[249,329,1008,1035]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="37" pageNumber="836" unit="mm" value="15.0">15 mm</quantity>
de long. Des prolongements vasculaires font suite au long post-abdomen.
</paragraph>
<paragraph id="474C41173B3F5C2BEBDAFBC1FE81C3B0" blockId="37.[111,649,692,1700]" pageId="37" pageNumber="836">
Les deux siphons sont courts à 6 lobes arrondis (
<figureCitation id="DFC85D923B3F5C2BEB28FBA1FF58C27D" box="[122,226,1103,1130]" captionStart="FIG" captionStartId="38.[677,688,1494,1511]" captionTargetBox="[677,1210,220,1457]" captionTargetId="figure-19@38.[649,1231,201,1467]" captionTargetPageId="38" captionText="FIG. 49. — Pseudodistoma nigrum n. sp.: A, zoïde; B, C, zoïde en deux parties; D, détail de lestomac; E, larve. Échelles: A,2 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4690125" httpUri="https://zenodo.org/record/4690125/files/figure.png" pageId="37" pageNumber="836">Fig. 49A</figureCitation>
). De chaque côté on compte 10 à 12 faisceaux musculaires longitudinaux sur le thorax qui se prolongent en restant très forts sur labdomen qui est généralement très contracté. La branchie a 3 rangs dune vingtaine de stigmates de chaque côté. Loesophage est long (
<figureCitation id="DFC85D923B3F5C2BEAC2FB03FE49C31F" box="[400,499,1261,1288]" captionStart="FIG" captionStartId="38.[677,688,1494,1511]" captionTargetBox="[677,1210,220,1457]" captionTargetId="figure-19@38.[649,1231,201,1467]" captionTargetPageId="38" captionText="FIG. 49. — Pseudodistoma nigrum n. sp.: A, zoïde; B, C, zoïde en deux parties; D, détail de lestomac; E, larve. Échelles: A,2 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4690125" httpUri="https://zenodo.org/record/4690125/files/figure.png" pageId="37" pageNumber="836">Fig. 49D</figureCitation>
) et lestomac, avec 4 côtes bien marquées se situe aux ⅔ de la longueur de labdomen. Le post-estomac est étroit suivi dun court segment peu renflé qui forme le fond de la boucle intestinale. Le rectum débute par un renflement net.
</paragraph>
<paragraph id="474C41173B3F5C2BEBDAFA45FDE0C072" blockId="37.[111,649,692,1700]" pageId="37" pageNumber="836">
Le post-abdomen est long et mince (
<figureCitation id="DFC85D923B3F5C2BEAAFFA45FDC3C3D1" box="[509,633,1451,1478]" captionStart="FIG" captionStartId="38.[677,688,1494,1511]" captionTargetBox="[677,1210,220,1457]" captionTargetId="figure-19@38.[649,1231,201,1467]" captionTargetPageId="38" captionText="FIG. 49. — Pseudodistoma nigrum n. sp.: A, zoïde; B, C, zoïde en deux parties; D, détail de lestomac; E, larve. Échelles: A,2 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4690125" httpUri="https://zenodo.org/record/4690125/files/figure.png" pageId="37" pageNumber="836">Fig. 49A, C</figureCitation>
). Lovaire est très éloigné de labdomen et suivi dune rangée irrégulière de vésicules testiculaires. Le coeur est terminal. Lextrémité du post-abdomen porte quelques protubérances arrondies entre lesquelles part un long appendice vasculaire (
<figureCitation id="DFC85D923B3F5C2BEABBF9A4FDF7C072" box="[489,589,1610,1637]" captionStart="FIG" captionStartId="38.[677,688,1494,1511]" captionTargetBox="[677,1210,220,1457]" captionTargetId="figure-19@38.[649,1231,201,1467]" captionTargetPageId="38" captionText="FIG. 49. — Pseudodistoma nigrum n. sp.: A, zoïde; B, C, zoïde en deux parties; D, détail de lestomac; E, larve. Échelles: A,2 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4690125" httpUri="https://zenodo.org/record/4690125/files/figure.png" pageId="37" pageNumber="836">Fig. 49C</figureCitation>
).
</paragraph>
<paragraph id="474C41173B3F5C2BEBDAF987FC0EC7E7" blockId="37.[111,649,692,1700]" lastBlockId="37.[675,1212,216,496]" pageId="37" pageNumber="836">
Une larve (
<figureCitation id="DFC85D923B3F5C2BEBAEF987FEE2C093" box="[252,344,1641,1668]" captionStart="FIG" captionStartId="38.[677,688,1494,1511]" captionTargetBox="[677,1210,220,1457]" captionTargetId="figure-19@38.[649,1231,201,1467]" captionTargetPageId="38" captionText="FIG. 49. — Pseudodistoma nigrum n. sp.: A, zoïde; B, C, zoïde en deux parties; D, détail de lestomac; E, larve. Échelles: A,2 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4690125" httpUri="https://zenodo.org/record/4690125/files/figure.png" pageId="37" pageNumber="836">Fig. 49E</figureCitation>
) est incubée dans une poche incubatrice qui se développe très tôt, à la jonction thorax abdomen. Le tronc mesure
<quantity id="800BECF23B3F5C2BEF78FF36FBCBC6E4" box="[1066,1137,216,243]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="37" pageNumber="836" unit="mm" value="1.0">1 mm</quantity>
. Les 3 vésicules adhésives sont divergentes, larges, sur des pédoncules étroits. Elles sont entourées de chaque côté du corps par 6 prolongements digitiformes disposés de la façon suivante: 1 dorsal, 1 ventral et 2 de chaque côté entre les vésicules adhésives. Deux petites ampoules sont accolées à la partie ventrale de la masse viscérale. Lune des larves montre de très nombreux stigmates.
</paragraph>
<paragraph id="474C41173B3F5C2BE9F7FDFBFC9DC43A" blockId="37.[674,1210,533,876]" box="[677,807,533,559]" pageId="37" pageNumber="836">REMARQUES</paragraph>
<paragraph id="474C41173B3F5C2BE9F7FDDAFCD0C57B" blockId="37.[674,1210,533,876]" pageId="37" pageNumber="836">
Par ses colonies nues sur un court pédondule ensablé,
<taxonomicName id="80F33A943B3F5C2BE986FDBAFC7BC479" authority="Monniot &amp; Monniot, 2008" authorityName="Monniot &amp; Monniot" authorityYear="2008" box="[724,961,596,622]" class="Ascidiacea" family="Pseudodistomidae" genus="Pseudodistoma" kingdom="Animalia" order="Enterogona" pageId="37" pageNumber="836" phylum="Chordata" rank="species" species="nigrum" status="sp. nov.">
<emphasis id="75879D053B3F5C2BE986FDBAFC7BC479" box="[724,961,596,622]" italics="true" pageId="37" pageNumber="836">Pseudodistoma nigrum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EB4207E3B3F5C2BE89BFDBAFBBCC478" box="[969,1030,596,623]" pageId="37" pageNumber="836" rank="species">n. sp.</taxonomicNameLabel>
se rapproche de
<taxonomicName id="80F33A943B3F5C2BE9F7FD9AFC2AC498" authority="Kott, 1992" authorityName="Kott" authorityYear="1992" box="[677,912,628,655]" class="Ascidiacea" family="Pseudodistomidae" genus="Pseudodistoma" kingdom="Animalia" order="Enterogona" pageId="37" pageNumber="836" phylum="Chordata" rank="species" species="pilatum">
<emphasis id="75879D053B3F5C2BE9F7FD9AFCAAC499" box="[677,784,628,654]" italics="true" pageId="37" pageNumber="836">P. pilatum</emphasis>
<bibRefCitation id="23623CE63B3F5C2BE84AFD9AFC2AC498" author="KOTT P." box="[792,912,628,655]" pageId="37" pageNumber="836" pagination="375 - 620" refId="ref36501" refString="KOTT P. 1992. - Th e Australian Ascidiacea. Part 3, Aplousobranchia (2). Memoirs of the Queensland Museum 32 (2): 375 - 620." type="journal article" year="1992">Kott, 1992</bibRefCitation>
</taxonomicName>
du sud de lAustralie. Il en diffère par la couleur noire de la tunique due à des cellules pigmentaires très denses et moins de zoïdes par lobe. Les larves de
<taxonomicName id="80F33A943B3F5C2BE8C8FD3DFBBFC4FA" authorityName="Kott" authorityYear="1992" box="[922,1029,723,749]" class="Ascidiacea" family="Pseudodistomidae" genus="Pseudodistoma" kingdom="Animalia" order="Enterogona" pageId="37" pageNumber="836" phylum="Chordata" rank="species" species="pilatum">
<emphasis id="75879D053B3F5C2BE8C8FD3DFBBFC4FA" box="[922,1029,723,749]" italics="true" pageId="37" pageNumber="836">P. pilatum</emphasis>
</taxonomicName>
sont inconnues, mais il nest pas fait mention dans la description de cette espèce de la dilatation de la paroi thoracique qui apparaît très tôt et qui formera la poche incubatrice de
<taxonomicName id="80F33A943B3F5C2BE991FCBCFC92C57C" authority="Monniot &amp; Monniot, 2008" authorityName="Monniot &amp; Monniot" authorityYear="2008" box="[707,808,850,875]" class="Ascidiacea" family="Pseudodistomidae" genus="Pseudodistoma" kingdom="Animalia" order="Enterogona" pageId="37" pageNumber="836" phylum="Chordata" rank="species" species="nigrum" status="sp. nov.">
<emphasis id="75879D053B3F5C2BE991FCBCFC92C57C" box="[707,808,850,875]" italics="true" pageId="37" pageNumber="836">P. nigrum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EB4207E3B3F5C2BE87DFCBFFCD0C57B" box="[815,874,849,876]" pageId="37" pageNumber="836" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>