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<document id="C6E919F083AD1CFF979DB93C930E10A4" ID-CLB-Dataset="82886" ID-DOI="10.1080/00222933.2021.2017499" ID-GBIF-Dataset="5cd01cfe-52bb-4092-97b2-804c3c7d145d" ID-ISSN="1464-5262" ID-Zenodo-Dep="6758313" ID-ZooBank="D766FC5E-CEC8-4184-BE90-C25350B90612" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656311158681" checkinUser="plazi" docAuthor="Costa, Leidiane O., Duarte, Paulo R. M., Iannuzzi, Luciana &amp; Grossi, Paschoal C." docDate="2022" docId="CF46879EFFBEFF86FF55766EFE152BF0" docLanguage="en" docName="JNATHIST.56.1-4.191-225.pdf" docOrigin="Journal of Natural History 56 (1 - 4)" docSource="http://dx.doi.org/10.1080/00222933.2021.2017499" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Gibboryctes ebeninus Duarte and Grossi 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="212" masterDocId="337FFFE6FFAAFF90FFF57271FF852A10" masterDocTitle="Taxonomic revision and notes on natural history of the enigmatic beetle genus Gibboryctes Endrödi (Coleoptera: Melolonthidae: Dynastinae)" masterLastPageNumber="225" masterPageNumber="191" pageNumber="210" updateTime="1699352275692" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="61645598ACDB5937D21C61AC6E6C2F5E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1885DFFF53BBED9C7D9FB111D91A26CD">
<mods:title id="B3C76C06FDF6B970D22660EFA47630FC">Taxonomic revision and notes on natural history of the enigmatic beetle genus Gibboryctes Endrödi (Coleoptera: Melolonthidae: Dynastinae)</mods:title>
</mods:titleInfo>
<mods:name id="14474560077E4B08CBF1E5E272281CC8" type="personal">
<mods:role id="2D2BBA58813A51F30BB6EDC2BD0D7784">
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<mods:namePart id="79059A9FDA9D31A8185AC87B4758658A">Costa, Leidiane O.</mods:namePart>
</mods:name>
<mods:name id="EEF70691F724AA034459407B239D9757" type="personal">
<mods:role id="5546DCC77D9AB124043EF227E5692B03">
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<mods:namePart id="00A9A030F7D9D41E594E112912A00A7E">Duarte, Paulo R. M.</mods:namePart>
</mods:name>
<mods:name id="DE9A6723571C75E773C363B76ED26190" type="personal">
<mods:role id="18A80E236E1229D070336FE57C8D988B">
<mods:roleTerm id="589ED18FE690324F3F4C0EC503F124E4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="180FD212643FD15448444285974FEA5B">Iannuzzi, Luciana</mods:namePart>
</mods:name>
<mods:name id="EFEB8F924666C0856F74D073791AF5C5" type="personal">
<mods:role id="E7376CB62A0986EE674720CC5686E894">
<mods:roleTerm id="31DF24C09A341A8A8AA3D1D60FE2C02E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2EE7BEF7847FDF518227E0FE33BC3B52">Grossi, Paschoal C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5B8213D4F81CE0BAFFE629F086605F1A">text</mods:typeOfResource>
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<mods:titleInfo id="483707B959409B5892CA5A9E4AC1E5BA">
<mods:title id="98B7DF4505B8128E252709D89907DCC8">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="B59594EEDE05BE93CAAB170EFAE1DE58">
<mods:date id="F141440506A28D90C1699CED1CD7CA2B">2022</mods:date>
<mods:detail id="6D635033DF14F5FF0662E0D438E553FB" type="pubDate">
<mods:number id="B75C7A6F60C973DAE161142BF19021E4">2022-05-17</mods:number>
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<mods:detail id="26B9379484217C7C155148881DDBBBB2" type="volume">
<mods:number id="B356F83FCC16F370C51465D7DDC88DEB">56</mods:number>
</mods:detail>
<mods:detail id="2F41AAC2D976859876DCDF04D35E1298" type="issue">
<mods:number id="8E8D41526EA3D34B027936DD2863A91A">1 - 4</mods:number>
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<mods:start id="D4D1255FEF50FC4FC860493C39A3E7A2">191</mods:start>
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<mods:location id="405B1D888D995824F244B399D1ADC0B0">
<mods:url id="69F1A14AC092C7E9209E9512D25AFC5B">http://dx.doi.org/10.1080/00222933.2021.2017499</mods:url>
</mods:location>
<mods:classification id="417466B517A76B0FEE024ED83DA0CE47">journal article</mods:classification>
<mods:identifier id="FBB9F4408E652A0E6D6F731233591A24" type="CLB-Dataset">82886</mods:identifier>
<mods:identifier id="F2B6B5CBD7B64DBDB46706CD342BD84E" type="DOI">10.1080/00222933.2021.2017499</mods:identifier>
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<treatment id="CF46879EFFBEFF86FF55766EFE152BF0" ID-DOI="http://doi.org/10.5281/zenodo.6761926" ID-GBIF-Taxon="196400126" ID-Zenodo-Dep="6761926" LSID="urn:lsid:plazi:treatment:CF46879EFFBEFF86FF55766EFE152BF0" httpUri="http://treatment.plazi.org/id/CF46879EFFBEFF86FF55766EFE152BF0" lastPageId="22" lastPageNumber="212" pageId="20" pageNumber="210">
<subSubSection id="0FF56503FFBEFF84FF55766EFD772E29" box="[160,754,1055,1081]" pageId="20" pageNumber="210" type="nomenclature">
<paragraph id="47503688FFBEFF84FF55766EFD772E29" blockId="20.[160,1155,1055,1115]" box="[160,754,1055,1081]" pageId="20" pageNumber="210">
<taxonomicName id="80EF4D0BFFBEFF84FF55766EFD032E29" authority="Duarte and Grossi" authorityName="Duarte and Grossi" authorityYear="2022" box="[160,646,1055,1081]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="210" phylum="Arthropoda" rank="species" species="ebeninus" status="sp. nov.">Gibboryctes ebeninus Duarte and Grossi</taxonomicName>
,
<taxonomicNameLabel id="6EA857E1FFBEFF84FD66766EFD772E29" box="[659,754,1055,1081]" pageId="20" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBEFF84FD0C766EFC8B2E4B" pageId="20" pageNumber="210" type="description">
<paragraph id="47503688FFBEFF84FD0C766EFC8B2E4B" blockId="20.[160,1155,1055,1115]" pageId="20" pageNumber="210">
(
<figureCitation id="DFD42A0DFFBEFF84FCF7766EFCF42E29" box="[770,881,1055,1081]" captionStart="Figure 2" captionStartId="5.[160,225,1050,1073]" captionTargetBox="[170,1128,191,1006]" captionTargetId="figure-170@5.[162,1154,142,1018]" captionTargetPageId="5" captionText="Figure 2. Gibboryctes ebeninus sp. nov. (a, b) Habitus of paratype female in dorsal and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.6758319" httpUri="https://zenodo.org/record/6758319/files/figure.png" pageId="20" pageNumber="210">Figures 2</figureCitation>
;
<figureCitation id="DFD42A0DFFBEFF84FC75766EFC0B2E29" box="[896,910,1055,1081]" captionStart="Figure 6" captionStartId="9.[160,225,1090,1113]" captionTargetBox="[166,1145,146,1050]" captionTargetId="figure-188@9.[166,1149,142,1058]" captionTargetPageId="9" captionText="Figure 6. Labrum in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758331" httpUri="https://zenodo.org/record/6758331/files/figure.png" pageId="20" pageNumber="210">6</figureCitation>
(b); 7(b); 8(b); 11(b); 12(c,d); 13(b); 14(b))
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBEFF84FF5576D9FD832FA4" pageId="20" pageNumber="210" type="diagnosis">
<paragraph id="47503688FFBEFF84FF5576D9FE912ED2" blockId="20.[160,1155,1192,1460]" box="[160,276,1192,1218]" pageId="20" pageNumber="210">
<emphasis id="759BEA9AFFBEFF84FF5576D9FE912ED2" bold="true" box="[160,276,1192,1218]" italics="true" pageId="20" pageNumber="210">Diagnosis</emphasis>
</paragraph>
<paragraph id="47503688FFBEFF84FF5576BAFD832FA4" blockId="20.[160,1155,1192,1460]" pageId="20" pageNumber="210">
<taxonomicName id="80EF4D0BFFBEFF84FF5576BAFE032EF4" authorityName="Duarte and Grossi" authorityYear="2022" box="[160,390,1227,1252]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="210" phylum="Arthropoda" rank="species" species="ebeninus">
<emphasis id="759BEA9AFFBEFF84FF5576BAFE032EF4" box="[160,390,1227,1252]" italics="true" pageId="20" pageNumber="210">Gibboryctes ebeninus</emphasis>
</taxonomicName>
differs from other
<taxonomicName id="80EF4D0BFFBEFF84FDAC76BAFCAA2EF4" box="[601,815,1227,1252]" pageId="20" pageNumber="210">
<emphasis id="759BEA9AFFBEFF84FDAC76BAFD5D2EF4" box="[601,728,1227,1252]" italics="true" pageId="20" pageNumber="210">Gibboryctes</emphasis>
species
</taxonomicName>
by the following combination of characters: Body colouration black (
<figureCitation id="DFD42A0DFFBEFF84FDB2769FFD242F17" box="[583,673,1262,1287]" captionStart="Figure 2" captionStartId="5.[160,225,1050,1073]" captionTargetBox="[170,1128,191,1006]" captionTargetId="figure-170@5.[162,1154,142,1018]" captionTargetPageId="5" captionText="Figure 2. Gibboryctes ebeninus sp. nov. (a, b) Habitus of paratype female in dorsal and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.6758319" httpUri="https://zenodo.org/record/6758319/files/figure.png" pageId="20" pageNumber="210">Figure 2</figureCitation>
); labrum subtrapezoidal, bearing truncated apical margin (
<figureCitation id="DFD42A0DFFBEFF84FEB37760FE272F3A" box="[326,418,1297,1322]" captionStart="Figure 6" captionStartId="9.[160,225,1090,1113]" captionTargetBox="[166,1145,146,1050]" captionTargetId="figure-188@9.[166,1149,142,1058]" captionTargetPageId="9" captionText="Figure 6. Labrum in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758331" httpUri="https://zenodo.org/record/6758331/files/figure.png" pageId="20" pageNumber="210">Figure 6</figureCitation>
(b)); maxillary palpomere II long, about 2.8 times longer than wide at middle (
<figureCitation id="DFD42A0DFFBEFF84FEEC7742FEF32F5C" box="[281,374,1331,1356]" captionStart="Figure 8" captionStartId="11.[160,225,1161,1184]" captionTargetBox="[167,1127,143,1126]" captionTargetId="figure-61@11.[166,1149,142,1129]" captionTargetPageId="11" captionText="Figure 8. Right maxilla in ventral view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicating triangular galea in (a), elongate maxillary palpomere II in (b), lobed inner margin of galea and stipe with triangular lobe in (c), pointed and triangular teeth on galea in (d), scythe-shape d teeth and rounded inner margin of galea in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758335" httpUri="https://zenodo.org/record/6758335/files/figure.png" pageId="20" pageNumber="210">Figure 8</figureCitation>
(b)); head with punctate frons (
<figureCitation id="DFD42A0DFFBEFF84FD257742FCBD2F5C" box="[720,824,1331,1356]" captionStart="Figure 11" captionStartId="14.[160,225,1441,1464]" captionTargetBox="[168,1146,144,1407]" captionTargetId="figure-42@14.[166,1149,142,1410]" captionTargetPageId="14" captionText="Figure 11. Detail of head, pronotum and prothoracic legs of females in dorsal view: (a) Gibboryctes szelenyii Endrödi;(b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedi sp.nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Arrow indicates punctate frons in (b)." figureDoi="http://doi.org/10.5281/zenodo.6758341" httpUri="https://zenodo.org/record/6758341/files/figure.png" pageId="20" pageNumber="210">Figure 11</figureCitation>
(b)); pronotal anterior corners weakly punctate (punctures predominantly spaced above 1 diameter of punctures) (
<figureCitation id="DFD42A0DFFBEFF84FF5D7708FE972F82" box="[168,274,1401,1426]" captionStart="Figure 12" captionStartId="15.[160,225,1189,1212]" captionTargetBox="[167,1149,146,1154]" captionTargetId="figure-100@15.[166,1149,142,1157]" captionTargetPageId="15" captionText="Figure 12. Females of Gibboryctes species with detail of head, pronotum and prothoracic legs in lateral view and left elytra in dorsal view, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes ebeninus sp. nov.; (e,f) Gibboryctes endroedii sp. nov. Jsi, juxtasutural interstria. Arrow indicates coalescent punctures in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758343" httpUri="https://zenodo.org/record/6758343/files/figure.png" pageId="20" pageNumber="210">Figure 12</figureCitation>
(c)); juxtasutural interstriae with scarce and large punctures, confined on anterior area of elytra (
<figureCitation id="DFD42A0DFFBEFF84FE8477EAFE592FA4" box="[369,476,1435,1460]" captionStart="Figure 12" captionStartId="15.[160,225,1189,1212]" captionTargetBox="[167,1149,146,1154]" captionTargetId="figure-100@15.[166,1149,142,1157]" captionTargetPageId="15" captionText="Figure 12. Females of Gibboryctes species with detail of head, pronotum and prothoracic legs in lateral view and left elytra in dorsal view, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes ebeninus sp. nov.; (e,f) Gibboryctes endroedii sp. nov. Jsi, juxtasutural interstria. Arrow indicates coalescent punctures in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758343" httpUri="https://zenodo.org/record/6758343/files/figure.png" pageId="20" pageNumber="210">Figure 12</figureCitation>
(d)).
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBEFF84FF557781FEC12C1A" box="[160,324,1520,1546]" pageId="20" pageNumber="210" type="description">
<paragraph id="47503688FFBEFF84FF557781FEC12C1A" blockId="20.[160,1156,1520,1719]" box="[160,324,1520,1546]" pageId="20" pageNumber="210">
<emphasis id="759BEA9AFFBEFF84FF557781FEC12C1A" bold="true" box="[160,324,1520,1546]" italics="true" pageId="20" pageNumber="210">
<typeStatus id="9854882AFFBEFF84FF557781FF5D2C1A" box="[160,216,1520,1546]" pageId="20" pageNumber="210">Type</typeStatus>
material
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBEFF85FF557465FE132AB8" lastPageId="21" lastPageNumber="211" pageId="20" pageNumber="210" type="materials_examined">
<paragraph id="47503688FFBEFF84FF557465FE602CA7" blockId="20.[160,1156,1520,1719]" pageId="20" pageNumber="210">
<materialsCitation id="F7873CD5FFBEFF84FF557465FE3C2C62" ID-GBIF-Occurrence="3819562301" collectingDate="1991-11" collectionCode="CERPE" country="Brazil" location="Gibboryctes" municipality="Tres Marias" pageId="20" pageNumber="210" specimenCount="2" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="holotype">
<typeStatus id="9854882AFFBEFF84FF557465FE822C3D" box="[160,263,1556,1581]" pageId="20" pageNumber="210" type="holotype">Holotype</typeStatus>
<specimenCount id="51E9FD01FFBEFF84FEF87465FEDC2C3D" box="[269,345,1556,1581]" pageId="20" pageNumber="210" type="female">female</specimenCount>
dissected, labelled: (a) ‘
<collectingCountry id="3FF87618FFBEFF84FDAE7465FD192C3D" box="[603,668,1556,1581]" name="Brazil" pageId="20" pageNumber="210">Brasil</collectingCountry>
,
<collectingRegion id="852BF86AFFBEFF84FD517465FCB42C3D" box="[676,817,1556,1581]" country="Brazil" name="Minas Gerais" pageId="20" pageNumber="210">Minas Gerais</collectingRegion>
/
<collectingMunicipality id="A734ACF2FFBEFF84FCCD7465FC3D2C3D" box="[824,952,1556,1581]" pageId="20" pageNumber="210">Três Marias</collectingMunicipality>
,
<date id="33511048FFBEFF84FC347465FB922C3D" box="[961,1047,1556,1581]" pageId="20" pageNumber="210" value="1991-11">
<collectingDate id="2315E9A0FFBEFF84FC347465FB922C3D" box="[961,1047,1556,1581]" pageId="20" pageNumber="210" value="1991-11">xi-1991</collectingDate>
</date>
,/F. Z. Vazde-Mello’ [white label]; (b) ‘
<taxonomicName id="80EF4D0BFFBEFF84FE247447FD3D2C5F" authority="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi, 2022" authorityName="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi" authorityYear="2022" box="[465,696,1590,1615]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="210" phylum="Arthropoda" rank="species" species="ebeninus" status="sp. nov.">
<emphasis id="759BEA9AFFBEFF84FE247447FD3D2C5F" box="[465,696,1590,1615]" italics="true" pageId="20" pageNumber="210">
<location id="42306053FFBEFF84FE247447FDD52C5F" LSID="urn:lsid:plazi:treatment:CF46879EFFBEFF86FF55766EFE152BF0:42306053FFBEFF84FE247447FDD52C5F" box="[465,592,1590,1615]" country="Brazil" municipality="Tres Marias" name="Gibboryctes" pageId="20" pageNumber="210" stateProvince="Minas Gerais">Gibboryctes</location>
ebeninus
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EA857E1FFBEFF84FD4A7447FC8A2C5F" box="[703,783,1590,1615]" pageId="20" pageNumber="210" rank="species">
<specimenCount id="51E9FD01FFBEFF84FD4A7447FD5A2C5F" box="[703,735,1590,1615]" pageId="20" pageNumber="210" type="generic">sp.</specimenCount>
nov.
</taxonomicNameLabel>
/
<typeStatus id="9854882AFFBEFF84FCE07447FC182C5F" box="[789,925,1590,1615]" pageId="20" pageNumber="210" type="holotype">HOLOTYPE</typeStatus>
/
<location id="42306053FFBEFF84FC507447FB012C5F" LSID="urn:lsid:plazi:treatment:CF46879EFFBEFF86FF55766EFE152BF0:42306053FFBEFF84FC507447FB012C5F" box="[933,1156,1590,1615]" country="Brazil" municipality="Tres Marias" name="Duarte &amp; Grossi Det." pageId="20" pageNumber="210" stateProvince="Minas Gerais">
Duarte &amp;
<taxonomicName id="80EF4D0BFFBEFF84FBFE7447FB012C5F" authority="Det. 2021" authorityName="Det." authorityYear="2021" box="[1035,1156,1590,1615]" genus="Grossi" pageId="20" pageNumber="222" rank="genus" status="gen. nov.">Grossi Det.</taxonomicName>
</location>
2021’ [red label] (
<collectionCode id="21FEAE4DFFBEFF84FE977428FE352C62" box="[354,432,1625,1650]" collectionName="CERPE" pageId="20" pageNumber="210">CERPE</collectionCode>
).
</materialsCitation>
<materialsCitation id="F7873CD5FFBEFF84FE4A7428FE642CA7" ID-GBIF-Occurrence="3819562304" collectingDate="1989-10" collectionCode="CERPE" location="One" municipality="One" pageId="20" pageNumber="210" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<collectingMunicipality id="A734ACF2FFBEFF84FE4A7428FE6B2C62" box="[447,494,1625,1650]" pageId="20" pageNumber="210">One</collectingMunicipality>
<specimenCount id="51E9FD01FFBEFF84FE007428FDC42C62" box="[501,577,1625,1650]" pageId="20" pageNumber="210" type="female">female</specimenCount>
<typeStatus id="9854882AFFBEFF84FDBD7428FD292C62" box="[584,684,1625,1650]" pageId="20" pageNumber="210" type="paratype">paratype</typeStatus>
with same data as
<typeStatus id="9854882AFFBEFF84FC727428FC6E2C62" box="[903,1003,1625,1650]" pageId="20" pageNumber="210" type="holotype">holotype</typeStatus>
(
<collectionCode id="21FEAE4DFFBEFF84FC0D7428FBC32C62" box="[1016,1094,1625,1650]" collectionName="CERPE" pageId="20" pageNumber="210">CERPE</collectionCode>
).
<location id="42306053FFBEFF84FBA07428FB012C62" LSID="urn:lsid:plazi:treatment:CF46879EFFBEFF86FF55766EFE152BF0:42306053FFBEFF84FBA07428FB012C62" box="[1109,1156,1625,1650]" municipality="One" name="One" pageId="20" pageNumber="210">One</location>
<specimenCount id="51E9FD01FFBEFF84FF55740DFF692C85" box="[160,236,1660,1685]" pageId="20" pageNumber="210" type="female">female</specimenCount>
with same data as
<typeStatus id="9854882AFFBEFF84FE38740DFDB12C85" box="[461,564,1660,1685]" pageId="20" pageNumber="210" type="holotype">holotype</typeStatus>
, except: (a) ‘
<date id="33511048FFBEFF84FD35740DFC942C85" box="[704,785,1660,1685]" pageId="20" pageNumber="210" value="1989-10">
<collectingDate id="2315E9A0FFBEFF84FD35740DFC942C85" box="[704,785,1660,1685]" pageId="20" pageNumber="210" value="1989-10">x-1989</collectingDate>
</date>
’ [white label] (
<collectionCode id="21FEAE4DFFBEFF84FC4D740DFB832C85" box="[952,1030,1660,1685]" collectionName="CERPE" pageId="20" pageNumber="210">CERPE</collectionCode>
).
<typeStatus id="9854882AFFBEFF84FBE3740DFB062C85" box="[1046,1155,1660,1685]" pageId="20" pageNumber="210" type="paratype">Paratypes</typeStatus>
with a yellow
<typeStatus id="9854882AFFBEFF84FEC874EFFE242CA7" box="[317,417,1694,1719]" pageId="20" pageNumber="210" type="paratype">paratype</typeStatus>
label
</materialsCitation>
.
</paragraph>
<paragraph id="47503688FFBFFF85FF5572FFFE132AB8" blockId="21.[160,1157,142,1243]" box="[160,406,142,168]" pageId="21" pageNumber="211">
<emphasis id="759BEA9AFFBFFF85FF5572FFFE132AB8" bold="true" box="[160,406,142,168]" italics="true" pageId="21" pageNumber="211">
<typeStatus id="9854882AFFBFFF85FF5572FFFE8E2AB8" box="[160,267,142,168]" pageId="21" pageNumber="211" type="holotype">Holotype</typeStatus>
description
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBFFF85FF5572C0FE8B2F39" pageId="21" pageNumber="211" type="description">
<paragraph id="47503688FFBFFF85FF5572C0FEC22ECB" blockId="21.[160,1157,142,1243]" pageId="21" pageNumber="211">
<emphasis id="759BEA9AFFBFFF85FF5572C0FE672ADB" bold="true" box="[160,482,177,203]" pageId="21" pageNumber="211">
Female
<emphasis id="759BEA9AFFBFFF85FEF672C0FEF82ADB" bold="true" box="[259,381,177,203]" italics="true" pageId="21" pageNumber="211">
(
<figureCitation id="DFD42A0DFFBFFF85FEFE72C0FEE82ADB" box="[267,365,177,203]" captionStart="Figure 2" captionStartId="5.[160,225,1050,1073]" captionTargetBox="[170,1128,191,1006]" captionTargetId="figure-170@5.[162,1154,142,1018]" captionTargetPageId="5" captionText="Figure 2. Gibboryctes ebeninus sp. nov. (a, b) Habitus of paratype female in dorsal and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.6758319" httpUri="https://zenodo.org/record/6758319/files/figure.png" pageId="21" pageNumber="211">Figure 2</figureCitation>
).
</emphasis>
Length
</emphasis>
:
<quantity id="80179B6DFFBFFF85FE0672C3FDEC2ADB" box="[499,617,178,203]" metricMagnitude="-2" metricUnit="m" metricValue="2.654" pageId="21" pageNumber="211" unit="mm" value="26.54">26.54 mm</quantity>
.
<emphasis id="759BEA9AFFBFFF85FD8272C3FD452ADB" bold="true" box="[631,704,178,203]" pageId="21" pageNumber="211">Width</emphasis>
:
<quantity id="80179B6DFFBFFF85FD2472C3FCC22ADB" box="[721,839,178,203]" metricMagnitude="-2" metricUnit="m" metricValue="1.257" pageId="21" pageNumber="211" unit="mm" value="12.57">12.57 mm</quantity>
.
<emphasis id="759BEA9AFFBFFF85FCA372C3FC232ADB" bold="true" box="[854,934,178,203]" pageId="21" pageNumber="211">Colour</emphasis>
: Completely black.
<emphasis id="759BEA9AFFBFFF85FF5572A5FF5A2AFD" bold="true" box="[160,223,212,237]" pageId="21" pageNumber="211">Head</emphasis>
: Clypeus triangular, transverse (3 times wider than long), slightly constricted laterally, rounded apically; surface densely rugopunctate, with scattered setae on sides separated by a glabrous middle area. Frontoclypeal suture with transverse, short, flattened, weakly bilobed tubercle. Frons slightly concave, scarcely, irregularly punctate; interocular width equals 4.0 transverse eye diameters. Ocular canthus subrectangular, transverse, glabrous.
<emphasis id="759BEA9AFFBFFF85FE5773F3FDB72B8B" bold="true" box="[418,562,386,411]" pageId="21" pageNumber="211">Mouthparts</emphasis>
: Labrum trapezoidal, truncated at apical margin (
<figureCitation id="DFD42A0DFFBFFF85FF5D73D5FE892BAD" box="[168,268,420,445]" captionStart="Figure 6" captionStartId="9.[160,225,1090,1113]" captionTargetBox="[166,1145,146,1050]" captionTargetId="figure-188@9.[166,1149,142,1058]" captionTargetPageId="9" captionText="Figure 6. Labrum in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758331" httpUri="https://zenodo.org/record/6758331/files/figure.png" pageId="21" pageNumber="211">Figure 6</figureCitation>
(b)). Mandibles bidentate; teeth lobed, exposed laterally to clypeus; basal tooth larger in size compared to apical tooth (
<figureCitation id="DFD42A0DFFBFFF85FD4E73B6FC9E2BF0" box="[699,795,455,480]" captionStart="Figure 7" captionStartId="10.[160,225,1156,1179]" captionTargetBox="[169,1149,148,1090]" captionTargetId="figure-156@10.[166,1149,142,1125]" captionTargetPageId="10" captionText="Figure 7. Right mandible in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicate the apical tooth in (a–e), striate molar in (d)." figureDoi="http://doi.org/10.5281/zenodo.6758333" httpUri="https://zenodo.org/record/6758333/files/figure.png" pageId="21" pageNumber="211">Figure 7</figureCitation>
(b)). Maxillary galea rounded at apex; inner margin with 3 small, pointed teeth located at middle of galea; stipe weakly produced laterally (shorter in length than maxillary palpomere I) (
<figureCitation id="DFD42A0DFFBFFF85FC7B707DFC692835" box="[910,1004,524,549]" captionStart="Figure 8" captionStartId="11.[160,225,1161,1184]" captionTargetBox="[167,1127,143,1126]" captionTargetId="figure-61@11.[166,1149,142,1129]" captionTargetPageId="11" captionText="Figure 8. Right maxilla in ventral view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicating triangular galea in (a), elongate maxillary palpomere II in (b), lobed inner margin of galea and stipe with triangular lobe in (c), pointed and triangular teeth on galea in (d), scythe-shape d teeth and rounded inner margin of galea in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758335" httpUri="https://zenodo.org/record/6758335/files/figure.png" pageId="21" pageNumber="211">Figure 8</figureCitation>
(b)). Maxillary palpomere II beaker-shaped, elongate (2.8 times longer than width at middle) (
<figureCitation id="DFD42A0DFFBFFF85FBD3705EFB012858" box="[1062,1156,559,584]" captionStart="Figure 8" captionStartId="11.[160,225,1161,1184]" captionTargetBox="[167,1127,143,1126]" captionTargetId="figure-61@11.[166,1149,142,1129]" captionTargetPageId="11" captionText="Figure 8. Right maxilla in ventral view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicating triangular galea in (a), elongate maxillary palpomere II in (b), lobed inner margin of galea and stipe with triangular lobe in (c), pointed and triangular teeth on galea in (d), scythe-shape d teeth and rounded inner margin of galea in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758335" httpUri="https://zenodo.org/record/6758335/files/figure.png" pageId="21" pageNumber="211">Figure 8</figureCitation>
(b)). Labium suboval, with subapical sides constricted, apex rounded; surface surrounded with large, deep, coalescent, bristled punctures; disc scarcely punctate.
<emphasis id="759BEA9AFFBFFF85FF5570E6FF7128A0" bold="true" box="[160,244,663,688]" pageId="21" pageNumber="211">Thorax</emphasis>
: Pronotum rounded laterally in dorsal view, weakly convex in lateral view; anterior tubercle obsolete; posterior border interrupted at middle; anterior area densely covered with large, deep, C-shaped punctures; punctures gradually becoming small, shallow towards posterior area; anterior corners scarcely punctate (punctures predominantly spaced about 1 diameter of punctures (
<figureCitation id="DFD42A0DFFBFFF85FC7C7153FB83292B" box="[905,1030,802,827]" captionStart="Figure 11" captionStartId="14.[160,225,1441,1464]" captionTargetBox="[168,1146,144,1407]" captionTargetId="figure-42@14.[166,1149,142,1410]" captionTargetPageId="14" captionText="Figure 11. Detail of head, pronotum and prothoracic legs of females in dorsal view: (a) Gibboryctes szelenyii Endrödi;(b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedi sp.nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Arrow indicates punctate frons in (b)." figureDoi="http://doi.org/10.5281/zenodo.6758341" httpUri="https://zenodo.org/record/6758341/files/figure.png" pageId="21" pageNumber="211">Figures 11</figureCitation>
(c) and 12 (c)). Scutellar plate parabolic in shape, glabrous, scarcely punctate; punctures confined to anterior area, scattered about 1 diameter of punctures. Elytra longitudinally covered with 9 well-marked striae (1 sutural, 4 discal, 4 lateral); each stria with a row of large, ocellate, deep punctures; anterior area of first interstria with 9 large, deep, ocellated punctures; other interstriae finely punctate (
<figureCitation id="DFD42A0DFFBFFF85FC6671BEFB8F29F8" box="[915,1034,975,1000]" captionStart="Figure 12" captionStartId="15.[160,225,1189,1212]" captionTargetBox="[167,1149,146,1154]" captionTargetId="figure-100@15.[166,1149,142,1157]" captionTargetPageId="15" captionText="Figure 12. Females of Gibboryctes species with detail of head, pronotum and prothoracic legs in lateral view and left elytra in dorsal view, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes ebeninus sp. nov.; (e,f) Gibboryctes endroedii sp. nov. Jsi, juxtasutural interstria. Arrow indicates coalescent punctures in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758343" httpUri="https://zenodo.org/record/6758343/files/figure.png" pageId="21" pageNumber="211">Figure 12</figureCitation>
(d)).
<emphasis id="759BEA9AFFBFFF85FBB071BEFBF829F8" bold="true" box="[1093,1149,975,1000]" pageId="21" pageNumber="211">Legs</emphasis>
: Mesotibial outer carinae with 15 stout spinules like setae (6 on basal carina, 9 on medial carina). Metatibial outer carinae with short, stout spinules.
<emphasis id="759BEA9AFFBFFF85FBF67665FBFB2E3D" bold="true" box="[1027,1150,1044,1069]" pageId="21" pageNumber="211">Abdomen</emphasis>
: Tergite VIII rugopuctate on sides, punctate on disc. Ventrites II–V densely rugopunctate on sides, finely punctate on disc; ventrites II–V with a row of setigerous punctures confined to sides; ventrite VI parabolic in shape, surrounded with rugopunctures, smooth on discal area, covered with short setae on posterior margin (
<figureCitation id="DFD42A0DFFBFFF85FF5D76B3FE9E2ECB" box="[168,283,1218,1243]" captionStart="Figure 13" captionStartId="16.[160,225,1058,1081]" captionTargetBox="[166,1146,145,1023]" captionTargetId="figure-155@16.[166,1149,142,1026]" captionTargetPageId="16" captionText="Figure 13. Detail of ventrites I–VI of females. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov; (c) Gibboryctes endroedii sp. nov.; (d) Tetragus waldenfelsi (Endrödi). Red arrows indicate the proportions of ventrites V and VI." figureDoi="http://doi.org/10.5281/zenodo.6758345" httpUri="https://zenodo.org/record/6758345/files/figure.png" pageId="21" pageNumber="211">Figure 13</figureCitation>
(b)).
</paragraph>
<paragraph id="47503688FFBFFF85FF55777EFE8B2F39" blockId="21.[160,767,1295,1355]" box="[160,270,1295,1321]" pageId="21" pageNumber="211">
<emphasis id="759BEA9AFFBFFF85FF55777EFE8B2F39" bold="true" box="[160,270,1295,1321]" italics="true" pageId="21" pageNumber="211">Variation</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBFFF85FF557743FD7A2F5B" box="[160,767,1330,1355]" pageId="21" pageNumber="211" type="materials_examined">
<paragraph id="47503688FFBFFF85FF557743FD7A2F5B" blockId="21.[160,767,1295,1355]" box="[160,767,1330,1355]" pageId="21" pageNumber="211">
<typeStatus id="9854882AFFBFFF85FF557743FE882F5B" box="[160,269,1330,1355]" pageId="21" pageNumber="211" type="paratype">Paratypes</typeStatus>
do not vary significantly from the
<typeStatus id="9854882AFFBFFF85FD607743FD792F5B" box="[661,764,1330,1355]" pageId="21" pageNumber="211" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBFFF85FF5577F1FE962FAC" box="[160,275,1408,1468]" pageId="21" pageNumber="211" type="description">
<paragraph id="47503688FFBFFF85FF5577F1FF5F2F8A" blockId="21.[160,275,1408,1468]" box="[160,218,1408,1434]" pageId="21" pageNumber="211">
<emphasis id="759BEA9AFFBFFF85FF5577F1FF5F2F8A" bold="true" box="[160,218,1408,1434]" italics="true" pageId="21" pageNumber="211">Male</emphasis>
</paragraph>
<paragraph id="47503688FFBFFF85FF5577D2FE962FAC" blockId="21.[160,275,1408,1468]" box="[160,275,1443,1468]" pageId="21" pageNumber="211">Unknown.</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBFFF85FF557781FF652C40" pageId="21" pageNumber="211" type="etymology">
<paragraph id="47503688FFBFFF85FF557781FE982C1A" blockId="21.[160,1155,1520,1616]" box="[160,285,1520,1546]" pageId="21" pageNumber="211">
<emphasis id="759BEA9AFFBFFF85FF557781FE982C1A" bold="true" box="[160,285,1520,1546]" italics="true" pageId="21" pageNumber="211">Etymology</emphasis>
</paragraph>
<paragraph id="47503688FFBFFF85FF557465FF652C40" blockId="21.[160,1155,1520,1616]" pageId="21" pageNumber="211">
From latin ‘
<taxonomicName id="80EF4D0BFFBFFF85FED57465FE042C3D" authorityName="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi" authorityYear="2022" box="[288,385,1556,1581]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="211" phylum="Arthropoda" rank="species" species="ebeninus">
<emphasis id="759BEA9AFFBFFF85FED57465FE042C3D" box="[288,385,1556,1581]" italics="true" pageId="21" pageNumber="211">ebeninus</emphasis>
</taxonomicName>
’, meaning ‘ebony’, in reference to the black and shiny surface of the body.
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBFFF85FF5574F5FE0D2CD0" pageId="21" pageNumber="211" type="description">
<paragraph id="47503688FFBFFF85FF5574F5FDE82C8E" blockId="21.[160,621,1668,1728]" box="[160,621,1668,1694]" pageId="21" pageNumber="211">
<heading id="1C1881E4FFBFFF85FF5574F5FDE82C8E" bold="true" box="[160,621,1668,1694]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="211" reason="3">
<emphasis id="759BEA9AFFBFFF85FF5574F5FDE82C8E" bold="true" box="[160,621,1668,1694]" pageId="21" pageNumber="211">
Geographic distribution
<emphasis id="759BEA9AFFBFFF85FE3974F5FDE82C8E" bold="true" box="[460,621,1668,1694]" italics="true" pageId="21" pageNumber="211">
(
<figureCitation id="DFD42A0DFFBFFF85FE2074F5FDC42C8E" box="[469,577,1668,1694]" captionStart="Figure 14" captionStartId="17.[160,227,1388,1411]" captionTargetBox="[168,1147,145,1355]" captionTargetId="figure-89@17.[166,1149,142,1357]" captionTargetPageId="17" captionText="Figure 14. Geographic distribution maps. (a) Tetragus species; (b) Gibboryctes species; (c) Heterogomphus gracilicornis Prell." figureDoi="http://doi.org/10.5281/zenodo.6758347" httpUri="https://zenodo.org/record/6758347/files/figure.png" pageId="21" pageNumber="211">Figure 14</figureCitation>
(b))
</emphasis>
</emphasis>
</heading>
</paragraph>
<paragraph id="47503688FFBFFF85FF5574D6FE0D2CD0" blockId="21.[160,621,1668,1728]" box="[160,392,1703,1728]" pageId="21" pageNumber="211">
<collectingCountry id="3FF87618FFBFFF85FF5574D6FF592CD0" box="[160,220,1703,1728]" name="Brazil" pageId="21" pageNumber="211">Brazil</collectingCountry>
(
<collectingRegion id="852BF86AFFBFFF85FF1C74D6FEFB2CD0" box="[233,382,1703,1728]" country="Brazil" name="Minas Gerais" pageId="21" pageNumber="211">Minas Gerais</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBCFF86FF5572FFFE152BF0" pageId="22" pageNumber="212" type="discussion">
<paragraph id="47503688FFBCFF86FF5572FFFE832AB8" blockId="22.[160,1156,142,480]" box="[160,262,142,168]" pageId="22" pageNumber="212">
<emphasis id="759BEA9AFFBCFF86FF5572FFFE832AB8" bold="true" box="[160,262,142,168]" italics="true" pageId="22" pageNumber="212">Remarks</emphasis>
</paragraph>
<paragraph id="47503688FFBCFF86FF5572C3FE152BF0" blockId="22.[160,1156,142,480]" pageId="22" pageNumber="212">
<taxonomicName id="80EF4D0BFFBCFF86FF5572C3FE0D2ADB" authority="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi, 2022" authorityName="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi" authorityYear="2022" box="[160,392,178,203]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="212" phylum="Arthropoda" rank="species" species="ebeninus" status="sp. nov.">
<emphasis id="759BEA9AFFBCFF86FF5572C3FE0D2ADB" box="[160,392,178,203]" italics="true" pageId="22" pageNumber="212">Gibboryctes ebeninus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EA857E1FFBCFF86FE6472C3FE6F2ADB" box="[401,490,178,203]" pageId="22" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
is the most distinct species in the genus by virtue of four exclusive characters: body with dark colour; maxilla with short lateral stipe (shorter compared to maxillary palpomere I; maxillary palpomere II elongate (2.8 times longer than wide at middle); elytral interstriae predominantly covered with fine punctures, large punctures being scarce. These characters in the other two species are: body colouration reddish brown or dark reddish brown; frons coarsely rugopunctate; maxilla with stipe produced laterally (longer compared to maxillary palpomere I); maxillary palpomere II shorter (1.6 times longer than wide at middle); elytral interstriae almost entirely covered with large punctures.
</paragraph>
</subSubSection>
</treatment>
</document>
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