387 lines
45 KiB
XML
387 lines
45 KiB
XML
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<mods:title id="CEE175E388E0F4E6421F5D468EC13A19">Taxonomic revision and notes on natural history of the enigmatic beetle genus Gibboryctes Endrödi (Coleoptera: Melolonthidae: Dynastinae)</mods:title>
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<mods:namePart id="9621BB5752E2DC4A47A3C8667B150A43">Costa, Leidiane O.</mods:namePart>
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<mods:namePart id="8134F29654AFE92873936C8A4D5663D0">Duarte, Paulo R. M.</mods:namePart>
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<mods:namePart id="8239FA3812D1B52D438AEF50BF7E34AB">Iannuzzi, Luciana</mods:namePart>
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<mods:namePart id="8CFE790FFD71ED7B7AB1EB67256C3626">Grossi, Paschoal C.</mods:namePart>
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<mods:title id="78FDE5093727A610E1E54B0C8EDCCD0A">Journal of Natural History</mods:title>
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<mods:date id="E39879B20E0806DD1C4A638DB9FBBDF5">2022</mods:date>
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<treatment id="CF46879EFFB7FF8FFF0C731BFCDF2B23" ID-DOI="http://doi.org/10.5281/zenodo.6758327" ID-GBIF-Taxon="196400128" ID-Zenodo-Dep="6758327" LSID="urn:lsid:plazi:treatment:CF46879EFFB7FF8FFF0C731BFCDF2B23" httpUri="http://treatment.plazi.org/id/CF46879EFFB7FF8FFF0C731BFCDF2B23" lastPageId="31" lastPageNumber="221" pageId="29" pageNumber="219">
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<subSubSection id="0FF56503FFB7FF8DFF0C731BFBAF2B94" box="[249,1066,362,388]" pageId="29" pageNumber="219" type="nomenclature">
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<paragraph id="47503688FFB7FF8DFF0C731BFBAF2B94" blockId="29.[249,1066,362,388]" box="[249,1066,362,388]" pageId="29" pageNumber="219">
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<heading id="1C1881E4FFB7FF8DFF0C731BFBAF2B94" bold="true" box="[249,1066,362,388]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="219" reason="3">
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<treatmentCitationGroup id="67FF11A6FFB7FF8DFF0C731BFBAF2B94" box="[249,1066,362,388]" pageId="29" pageNumber="219">
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<emphasis id="759BEA9AFFB7FF8DFF0C731BFBAF2B94" bold="true" box="[249,1066,362,388]" pageId="29" pageNumber="219">
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<treatmentCitation id="C64E1099FFB7FF8DFF0C731BFCB92B94" author="Dechambre R-P" box="[249,828,362,388]" page="267" pageId="29" pageNumber="219" year="1981">
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<taxonomicName id="80EF4D0BFFB7FF8DFF0C731BFCB92B94" authority="Dechambre 1981: 267" authorityName="Dechambre" authorityPageNumber="267" authorityYear="1981" box="[249,828,362,388]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="219" phylum="Arthropoda" rank="species" species="waldenfelsi">
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Gibboryctes waldenfelsi
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<bibRefCitation id="237E4B79FFB7FF8DFDC5731BFCB92B94" author="Dechambre R-P" box="[560,828,362,388]" pageId="29" pageNumber="219" pagination="123 - 128" refId="ref13215" refString="Dechambre R-P. 1981. Nouvelles especes de Dynastidae de la region neotropicale (Coleoptera, Scarabaeoidea). Rev. Fr. Entomol. 3: 123 - 128." type="journal article" year="1981">
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<emphasis id="759BEA9AFFB7FF8DFDC5731BFCB92B94" bold="true" box="[560,828,362,388]" italics="true" pageId="29" pageNumber="219">Dechambre 1981: 267</emphasis>
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</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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<emphasis id="759BEA9AFFB7FF8DFCB1731BFBAF2B94" bold="true" box="[836,1066,362,388]" italics="true" pageId="29" pageNumber="219">(new combination)</emphasis>
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</emphasis>
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</treatmentCitationGroup>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0FF56503FFB7FF8DFF5573B3FC0D2854" pageId="29" pageNumber="219" type="diagnosis">
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<paragraph id="47503688FFB7FF8DFF5573B3FE912BCC" blockId="29.[160,1156,450,580]" box="[160,276,450,476]" pageId="29" pageNumber="219">
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<emphasis id="759BEA9AFFB7FF8DFF5573B3FE912BCC" bold="true" box="[160,276,450,476]" italics="true" pageId="29" pageNumber="219">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="47503688FFB7FF8DFF557397FC0D2854" blockId="29.[160,1156,450,580]" pageId="29" pageNumber="219">
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<taxonomicName id="80EF4D0BFFB7FF8DFF557397FE012BEF" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" baseAuthorityName="Endrodi" baseAuthorityYear="1977" box="[160,388,486,511]" class="Insecta" family="Scarabaeidae" genus="Tetragus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="219" phylum="Arthropoda" rank="species" species="waldenfelsi">
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<emphasis id="759BEA9AFFB7FF8DFF557397FE012BEF" box="[160,388,486,511]" italics="true" pageId="29" pageNumber="219">Tetragus waldenfelsi</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="80EF4D0BFFB7FF8DFDD67397FD4E2BEF" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" baseAuthorityName="Dupuis" baseAuthorityYear="2019" box="[547,715,486,511]" class="Insecta" family="Scarabaeidae" genus="Tetragus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="219" phylum="Arthropoda" rank="species" species="impunctatus">
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<emphasis id="759BEA9AFFB7FF8DFDD67397FD4E2BEF" box="[547,715,486,511]" italics="true" pageId="29" pageNumber="219">P. impunctatus</emphasis>
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</taxonomicName>
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in the following characters: pronotal surface punctate (punctures large on concavity, fine on other surfaces) in male; surface of tergite VIII flattened to slightly convex in lateral view in female.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="0FF56503FFB7FF8DFF557000FB01296E" pageId="29" pageNumber="219" type="materials_examined">
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<paragraph id="47503688FFB7FF8DFF557000FEC1289B" blockId="29.[160,1156,625,894]" box="[160,324,625,651]" pageId="29" pageNumber="219">
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<emphasis id="759BEA9AFFB7FF8DFF557000FEC1289B" bold="true" box="[160,324,625,651]" italics="true" pageId="29" pageNumber="219">
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<typeStatus id="9854882AFFB7FF8DFF557000FF5D289B" box="[160,216,625,651]" pageId="29" pageNumber="219">Type</typeStatus>
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material
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</emphasis>
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</paragraph>
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<paragraph id="47503688FFB7FF8DFF5570E4FB01296E" blockId="29.[160,1156,625,894]" pageId="29" pageNumber="219">
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<taxonomicName id="80EF4D0BFFB7FF8DFF5570E4FE0128BE" authorityName="Endrodi" authorityYear="1977" box="[160,388,661,686]" class="Insecta" family="Dynastidae" genus="Strategus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="219" phylum="Arthropoda" rank="species" species="waldenfelsi">
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||
<emphasis id="759BEA9AFFB7FF8DFF5570E4FE0128BE" box="[160,388,661,686]" italics="true" pageId="29" pageNumber="219">Strategus waldenfelsi</emphasis>
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</taxonomicName>
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,
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<typeStatus id="9854882AFFB7FF8DFE7A70E4FE7628BE" box="[399,499,661,686]" pageId="29" pageNumber="219" type="holotype">holotype</typeStatus>
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not examined.
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||
<taxonomicName id="80EF4D0BFFB7FF8DFD6B70E4FCEA28BE" authorityName="Dechambre" authorityYear="1981" box="[670,879,661,686]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="219" phylum="Arthropoda" rank="species" species="porioni">
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<emphasis id="759BEA9AFFB7FF8DFD6B70E4FCEA28BE" box="[670,879,661,686]" italics="true" pageId="29" pageNumber="219">Gibboryctes porioni</emphasis>
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</taxonomicName>
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,
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<typeStatus id="9854882AFFB7FF8DFC8F70E4FC5B28BE" box="[890,990,661,686]" pageId="29" pageNumber="219" type="holotype">holotype</typeStatus>
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male, labelled: (a) ‘trouve cadavre/1e 2
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<collectorName id="EA1A535EFFB7FF8DFE5170C6FE6828C0" box="[420,493,695,720]" pageId="29" pageNumber="219">Dmars</collectorName>
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1977/a SAÜL/
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<collectingCountry id="3FF87618FFB7FF8DFD7E70C6FD5B28C0" box="[651,734,695,720]" name="French Guiana" pageId="29" pageNumber="219">Guyane</collectingCountry>
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<collectorName id="EA1A535EFFB7FF8DFD1170C6FCD728C0" box="[740,850,695,720]" pageId="29" pageNumber="219">Francaise</collectorName>
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/
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<collectorName id="EA1A535EFFB7FF8DFCAF70C6FC0628C0" box="[858,899,695,720]" pageId="29" pageNumber="219">Coll</collectorName>
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. Th. PORION’ [old label, partially handwritten]; (b) ‘HOLOTYPE’ [red label]; (c) ‘gibboryctes/porioni
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<taxonomicNameLabel id="6EA857E1FFB7FF8DFBFD70ABFBB828E3" box="[1032,1085,730,755]" pageId="29" pageNumber="219" rank="species">n.sp.</taxonomicNameLabel>
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mihi/
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<typeStatus id="9854882AFFB7FF8DFF55708CFE9A2906" box="[160,287,765,790]" pageId="29" pageNumber="219" type="holotype">HOLOTYPE</typeStatus>
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/R. -P. Dechambre det. 1980’ [old white label, partially handwritten]; (d) ‘photographie 1978/R. -P.
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<location id="42306053FFB7FF8DFE8B716EFD832928" LSID="urn:lsid:plazi:treatment:CF46879EFFB7FF8FFF0C731BFCDF2B23:42306053FFB7FF8DFE8B716EFD832928" box="[382,518,799,824]" country="French Guiana" name="Dechambre'" pageId="29" pageNumber="219">Dechambre’</location>
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[old white label, partially handwritten]; (e) ‘gibboryctes/
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<taxonomicName id="80EF4D0BFFB7FF8DFF557133FE14294B" authority="(Endrodi)" baseAuthorityName="Endrodi" box="[160,401,834,859]" class="Insecta" family="Dynastidae" genus="Strategus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="219" phylum="Arthropoda" rank="species" species="waldenfelsi">waldenfelsi (Endrödi)</taxonomicName>
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/R. -P. Dechambre det. 1985’ [old white label, bordered in black border, partially handwritten]; (f) ‘MNHN/EC8408’ [white label/ bordered in black] (
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||
<collectionCode id="21FEAE4DFFB7FF8DFBDE7114FBFE296E" box="[1067,1147,869,894]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="29" pageNumber="219">MNHN</collectionCode>
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||
).
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="0FF56503FFB7FF8DFF5571DBFEFC29D4" box="[160,377,938,964]" pageId="29" pageNumber="219" type="description">
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<paragraph id="47503688FFB7FF8DFF5571DBFEFC29D4" blockId="29.[160,1156,938,1138]" box="[160,377,938,964]" pageId="29" pageNumber="219">
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||
<emphasis id="759BEA9AFFB7FF8DFF5571DBFEFC29D4" bold="true" box="[160,377,938,964]" italics="true" pageId="29" pageNumber="219">Non-type material</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FF56503FFB7FF8DFF5571BFFC392E62" pageId="29" pageNumber="219" type="materials_examined">
|
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<paragraph id="47503688FFB7FF8DFF5571BFFC392E62" blockId="29.[160,1156,938,1138]" pageId="29" pageNumber="219">
|
||
<materialsCitation id="F7873CD5FFB7FF8DFF5571BFFDAF29F7" ID-GBIF-Occurrence="3819562325" box="[160,554,974,999]" collectingDate="1979-06" collectingDateMax="1979-07-31" collectingDateMin="1979-06" country="Panama" location="Darien" pageId="29" pageNumber="219" specimenCount="1" stateProvince="Darien">
|
||
<collectingCountry id="3FF87618FFB7FF8DFF5571BFFE8C29F7" box="[160,265,974,999]" name="Panama" pageId="29" pageNumber="219">PANAMA</collectingCountry>
|
||
:
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||
<collectingRegion id="852BF86AFFB7FF8DFEE771BFFED829F7" box="[274,349,974,999]" country="Panama" name="Darien" pageId="29" pageNumber="219">Darién</collectingRegion>
|
||
,
|
||
<date id="33511048FFB7FF8DFE9371BFFD9529F7" box="[358,528,974,999]" pageId="29" pageNumber="219" value="1979-06" valueMax="1979-07-31" valueMin="1979-06-00">
|
||
<collectingDate id="2315E9A0FFB7FF8DFE9371BFFD9529F7" box="[358,528,974,999]" pageId="29" pageNumber="219" value="1979-06" valueMax="1979-07-31" valueMin="1979-06-00">June–July 1979</collectingDate>
|
||
</date>
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||
, R
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="F7873CD5FFB7FF8DFDC671BFFCAD29F7" ID-GBIF-Occurrence="3819562317" box="[563,808,974,999]" collectionCode="BCRC" country="Panama" location="Hinds" pageId="29" pageNumber="219" specimenCount="1" specimenCount-male="1">
|
||
<location id="42306053FFB7FF8DFDC671BFFDF729F7" LSID="urn:lsid:plazi:treatment:CF46879EFFB7FF8FFF0C731BFCDF2B23:42306053FFB7FF8DFDC671BFFDF729F7" box="[563,626,974,999]" country="Panama" name="Hinds" pageId="29" pageNumber="219">Hinds</location>
|
||
–
|
||
<specimenCount id="51E9FD01FFB7FF8DFD7971BFFD5329F7" box="[652,726,974,999]" pageId="29" pageNumber="219" type="male">1 male</specimenCount>
|
||
(
|
||
<collectionCode id="21FEAE4DFFB7FF8DFD1471BFFCA629F7" box="[737,803,974,999]" country="Taiwan" httpUri="http://grbio.org/cool/cv7g-ikqh" name="Bioresource Collection and Research Center" pageId="29" pageNumber="219">BCRC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="F7873CD5FFB7FF8DFCC471BFFE022E1A" ID-GBIF-Occurrence="3819562322" collectingDate="2008-01-10" country="French Guiana" location="Petit Savi" pageId="29" pageNumber="219" specimenCount="1">
|
||
<collectingCountry id="3FF87618FFB7FF8DFCC471BFFC6829F7" box="[817,1005,974,999]" name="French Guiana" pageId="29" pageNumber="219">FRENCH GUIANA</collectingCountry>
|
||
:
|
||
<location id="42306053FFB7FF8DFC0371BFFBDE29F7" LSID="urn:lsid:plazi:treatment:CF46879EFFB7FF8FFF0C731BFCDF2B23:42306053FFB7FF8DFC0371BFFBDE29F7" box="[1014,1115,974,999]" country="French Guiana" name="Petit Savi" pageId="29" pageNumber="219">Petit Savi</location>
|
||
PK,
|
||
<date id="33511048FFB7FF8DFF557180FEE22E1A" box="[160,359,1009,1034]" pageId="29" pageNumber="219" value="2008-01-10">
|
||
<collectingDate id="2315E9A0FFB7FF8DFF557180FEE22E1A" box="[160,359,1009,1034]" pageId="29" pageNumber="219" value="2008-01-10">10 January 2008</collectingDate>
|
||
</date>
|
||
, Y
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="F7873CD5FFB7FF8DFE627180FF342E3C" ID-GBIF-Occurrence="3819562319" collectingDate="2009-02" collectionCode="EPGC" country="French Guiana" location="Ponchel" pageId="29" pageNumber="219" specimenCode="RN1, PK70" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<location id="42306053FFB7FF8DFE627180FE772E1A" LSID="urn:lsid:plazi:treatment:CF46879EFFB7FF8FFF0C731BFCDF2B23:42306053FFB7FF8DFE627180FE772E1A" box="[407,498,1009,1034]" country="French Guiana" name="Ponchel" pageId="29" pageNumber="219">Ponchel</location>
|
||
–
|
||
<specimenCount id="51E9FD01FFB7FF8DFDEC7180FDEB2E1A" box="[537,622,1009,1034]" pageId="29" pageNumber="219" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="51E9FD01FFB7FF8DFD8B7180FD602E1A" box="[638,741,1009,1034]" pageId="29" pageNumber="219" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="21FEAE4DFFB7FF8DFD027180FCBE2E1A" box="[759,827,1009,1034]" pageId="29" pageNumber="219">EPGC</collectionCode>
|
||
);
|
||
<specimenCode id="17499EF3FFB7FF8DFCA57180FC432E1A" box="[848,966,1009,1034]" collectionCode="EPGC" pageId="29" pageNumber="219">RN1 PK70</specimenCode>
|
||
,
|
||
<date id="33511048FFB7FF8DFC237180FB042E1A" box="[982,1153,1009,1034]" pageId="29" pageNumber="219" value="2009-02">
|
||
<collectingDate id="2315E9A0FFB7FF8DFC237180FB042E1A" box="[982,1153,1009,1034]" pageId="29" pageNumber="219" value="2009-02">February 2009</collectingDate>
|
||
</date>
|
||
, Y
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="F7873CD5FFB7FF8DFF497662FDCA2E3C" ID-GBIF-Occurrence="3819562310" box="[188,591,1043,1068]" collectionCode="CERPE" country="French Guiana" location="Ponchel" pageId="29" pageNumber="219" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<location id="42306053FFB7FF8DFF497662FE912E3C" LSID="urn:lsid:plazi:treatment:CF46879EFFB7FF8FFF0C731BFCDF2B23:42306053FFB7FF8DFF497662FE912E3C" box="[188,276,1043,1068]" country="French Guiana" name="Ponchel" pageId="29" pageNumber="219">Ponchel</location>
|
||
–
|
||
<specimenCount id="51E9FD01FFB7FF8DFEC17662FE072E3C" box="[308,386,1043,1068]" pageId="29" pageNumber="219" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="51E9FD01FFB7FF8DFE7B7662FE752E3C" box="[398,496,1043,1068]" pageId="29" pageNumber="219" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="21FEAE4DFFB7FF8DFE087662FDCC2E3C" box="[509,585,1043,1068]" pageId="29" pageNumber="219">CERPE</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="F7873CD5FFB7FF8DFDAF7662FCC62E5F" ID-GBIF-Occurrence="3819562308" collectingDate="1965-06-10" collectingDateMax="1977-02-28" collectingDateMin="1965-06-10" collectionCode="BCRC" country="Brazil" county="Amazonas" location="Reserva Ducke" municipality="Maua" pageId="29" pageNumber="219" specimenCount="1" specimenCount-male="1">
|
||
<collectingCountry id="3FF87618FFB7FF8DFDAF7662FD2F2E3C" box="[602,682,1043,1068]" name="Brazil" pageId="29" pageNumber="219">BRAZIL</collectingCountry>
|
||
:
|
||
<collectingCounty id="AE314E04FFB7FF8DFD437662FCAF2E3C" box="[694,810,1043,1068]" pageId="29" pageNumber="219">Amazonas</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="A734ACF2FFB7FF8DFCC37662FCF02E3C" box="[822,885,1043,1068]" pageId="29" pageNumber="219">Maua</collectingMunicipality>
|
||
,
|
||
<date id="33511048FFB7FF8DFC747662FB922E3C" box="[897,1047,1043,1068]" pageId="29" pageNumber="219" value="1965-06-10">
|
||
<collectingDate id="2315E9A0FFB7FF8DFC747662FB922E3C" box="[897,1047,1043,1068]" pageId="29" pageNumber="219" value="1965-06-10">10 June 1965</collectingDate>
|
||
</date>
|
||
–
|
||
<specimenCount id="51E9FD01FFB7FF8DFBC37662FB062E3C" box="[1078,1155,1043,1068]" pageId="29" pageNumber="219" type="male">1 male</specimenCount>
|
||
(
|
||
<collectionCode id="21FEAE4DFFB7FF8DFF507647FF622E5F" box="[165,231,1078,1103]" country="Taiwan" httpUri="http://grbio.org/cool/cv7g-ikqh" name="Bioresource Collection and Research Center" pageId="29" pageNumber="219">BCRC</collectionCode>
|
||
);
|
||
<location id="42306053FFB7FF8DFF017647FE132E5F" LSID="urn:lsid:plazi:treatment:CF46879EFFB7FF8FFF0C731BFCDF2B23:42306053FFB7FF8DFF017647FE132E5F" box="[244,406,1078,1103]" country="Brazil" county="Amazonas" municipality="Maua" name="Reserva Ducke" pageId="29" pageNumber="219">Reserva Ducke</location>
|
||
,
|
||
<locationDeviation id="56E22FAAFFB7FF8DFE6B7647FDE62E5F" box="[414,611,1078,1103]" location="Manaus" pageId="29" pageNumber="219">
|
||
<quantity id="80179B6DFFB7FF8DFE6B7647FE642E5F" box="[414,481,1078,1103]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="219" unit="km" value="26.0">26 km</quantity>
|
||
NE Manaus
|
||
</locationDeviation>
|
||
,
|
||
<date id="33511048FFB7FF8DFD9E7647FCAF2E5F" box="[619,810,1078,1103]" pageId="29" pageNumber="219" value="1977-02-28">
|
||
<collectingDate id="2315E9A0FFB7FF8DFD9E7647FCAF2E5F" box="[619,810,1078,1103]" pageId="29" pageNumber="219" value="1977-02-28">28 February 1977</collectingDate>
|
||
</date>
|
||
, B
|
||
</materialsCitation>
|
||
.C.
|
||
<materialsCitation id="F7873CD5FFB7FF8DFC947647FB052E5F" ID-GBIF-Occurrence="3819562329" box="[865,1152,1078,1103]" collectionCode="BCRC" country="Brazil" location="Ratcliffe" pageId="29" pageNumber="219" specimenCount="1" specimenCount-female="1">
|
||
<location id="42306053FFB7FF8DFC947647FC3C2E5F" LSID="urn:lsid:plazi:treatment:CF46879EFFB7FF8FFF0C731BFCDF2B23:42306053FFB7FF8DFC947647FC3C2E5F" box="[865,953,1078,1103]" country="Brazil" name="Ratcliffe" pageId="29" pageNumber="219">Ratcliffe</location>
|
||
–
|
||
<specimenCount id="51E9FD01FFB7FF8DFC247647FBAA2E5F" box="[977,1071,1078,1103]" pageId="29" pageNumber="219" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="21FEAE4DFFB7FF8DFBCC7647FBFE2E5F" box="[1081,1147,1078,1103]" country="Taiwan" httpUri="http://grbio.org/cool/cv7g-ikqh" name="Bioresource Collection and Research Center" pageId="29" pageNumber="219">BCRC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="F7873CD5FFB7FF8DFF557628FDB82E62" ID-GBIF-Occurrence="3819562327" box="[160,573,1113,1138]" collectingDate="1954-10" country="Brazil" location="Xingu" pageId="29" pageNumber="219" specimenCount="1" stateProvince="Mato Grosso">
|
||
<collectingRegion id="852BF86AFFB7FF8DFF557628FEAB2E62" box="[160,302,1113,1138]" country="Brazil" name="Mato Grosso" pageId="29" pageNumber="219">Mato Grosso</collectingRegion>
|
||
,
|
||
<location id="42306053FFB7FF8DFEC27628FEFF2E62" LSID="urn:lsid:plazi:treatment:CF46879EFFB7FF8FFF0C731BFCDF2B23:42306053FFB7FF8DFEC27628FEFF2E62" box="[311,378,1113,1138]" country="Brazil" name="Xingu" pageId="29" pageNumber="219" stateProvince="Mato Grosso">Xingu</location>
|
||
,
|
||
<date id="33511048FFB7FF8DFE717628FD992E62" box="[388,540,1113,1138]" pageId="29" pageNumber="219" value="1954-10">
|
||
<collectingDate id="2315E9A0FFB7FF8DFE717628FD992E62" box="[388,540,1113,1138]" pageId="29" pageNumber="219" value="1954-10">October 1954</collectingDate>
|
||
</date>
|
||
, M
|
||
</materialsCitation>
|
||
<materialsCitation id="F7873CD5FFB7FF8DFDC87628FD672E62" ID-GBIF-Occurrence="3819562323" box="[573,738,1113,1138]" collectingDate="1954-10" collectorName="Alvarenga" country="Brazil" isTailEnumeration="true" location="Xingu" pageId="29" pageNumber="219" specimenCount="1" stateProvince="Mato Grosso">
|
||
.
|
||
<collectorName id="EA1A535EFFB7FF8DFDBD7628FD322E62" box="[584,695,1113,1138]" pageId="29" pageNumber="219">Alvarenga</collectorName>
|
||
leg
|
||
</materialsCitation>
|
||
<materialsCitation id="F7873CD5FFB7FF8DFD177628FC3D2E62" ID-GBIF-Occurrence="3819562305" box="[738,952,1113,1138]" collectionCode="CERPE" country="Brazil" pageId="29" pageNumber="219" specimenCount="1" specimenCount-female="1">
|
||
. -
|
||
<specimenCount id="51E9FD01FFB7FF8DFD0E7628FCDE2E62" box="[763,859,1113,1138]" pageId="29" pageNumber="219" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="21FEAE4DFFB7FF8DFC937628FC372E62" box="[870,946,1113,1138]" pageId="29" pageNumber="219">CERPE</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FF56503FFB7FF8EFF5576EFFC6D2CD0" lastPageId="30" lastPageNumber="220" pageId="29" pageNumber="219" type="description">
|
||
<paragraph id="47503688FFB7FF8DFF5576EFFE0F2EA8" blockId="29.[160,1157,1182,1728]" box="[160,394,1182,1208]" pageId="29" pageNumber="219">
|
||
<emphasis id="759BEA9AFFB7FF8DFF5576EFFE0F2EA8" bold="true" box="[160,394,1182,1208]" pageId="29" pageNumber="219">
|
||
Male
|
||
<emphasis id="759BEA9AFFB7FF8DFF1676EFFE0F2EA8" bold="true" box="[227,394,1182,1208]" italics="true" pageId="29" pageNumber="219">
|
||
(
|
||
<figureCitation id="DFD42A0DFFB7FF8DFF1E76EFFEC92EA8" box="[235,332,1182,1208]" captionStart="Figure 5" captionStartId="8.[160,225,1685,1708]" captionTargetBox="[221,1093,152,1648]" captionTargetId="figure-3@8.[219,1097,142,1654]" captionTargetPageId="8" captionText="Figure 5. Tetragus waldenfelsi (Endrödi, 1977). (a,b) Male habitus in dorsal and lateral views, respectively; (c,d) female habitus in dorsal and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.6758329" httpUri="https://zenodo.org/record/6758329/files/figure.png" pageId="29" pageNumber="219">Figure 5</figureCitation>
|
||
(a,b))
|
||
</emphasis>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="47503688FFB7FF8EFF5576B0FEBA2EA7" blockId="29.[160,1157,1182,1728]" lastBlockId="30.[160,1157,142,1207]" lastPageId="30" lastPageNumber="220" pageId="29" pageNumber="219">
|
||
<emphasis id="759BEA9AFFB7FF8DFF5576B0FEC32ECB" bold="true" box="[160,326,1217,1243]" italics="true" pageId="29" pageNumber="219">Redescription.</emphasis>
|
||
<emphasis id="759BEA9AFFB7FF8DFEA376B3FE2E2ECB" bold="true" box="[342,427,1218,1243]" pageId="29" pageNumber="219">Length</emphasis>
|
||
:
|
||
<quantity id="80179B6DFFB7FF8DFE4976B3FDE72ECB" box="[444,610,1218,1243]" metricMagnitude="-2" metricUnit="m" metricValue="2.525" metricValueMax="2.79" metricValueMin="2.26" pageId="29" pageNumber="219" unit="mm" value="25.25" valueMax="27.9" valueMin="22.6">22.6–27.9 mm</quantity>
|
||
.
|
||
<emphasis id="759BEA9AFFB7FF8DFD8476B3FD3F2ECB" bold="true" box="[625,698,1218,1243]" pageId="29" pageNumber="219">Width</emphasis>
|
||
:
|
||
<quantity id="80179B6DFFB7FF8DFD3F76B3FCF52ECB" box="[714,880,1218,1243]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" metricValueMax="1.24" metricValueMin="1.08" pageId="29" pageNumber="219" unit="mm" value="11.6" valueMax="12.4" valueMin="10.8">10.8–12.4 mm</quantity>
|
||
.
|
||
<emphasis id="759BEA9AFFB7FF8DFC8A76B3FC4A2ECB" bold="true" box="[895,975,1218,1243]" pageId="29" pageNumber="219">Colour</emphasis>
|
||
: Dorsal surface dark reddish; legs and ventral surface dark reddish brown.
|
||
<emphasis id="759BEA9AFFB7FF8DFCA97694FC1E2EEE" bold="true" box="[860,923,1253,1278]" pageId="29" pageNumber="219">Head</emphasis>
|
||
: Clypeus triangular, short (3.2 times wider than long), acuminate anteriorly, glabrous, finely punctate. Frontoclypeal suture well marked, provided with a short, conical tubercle at middle. Frons glabrous; sides with large, deep, C-shaped, coalescent punctures; longitudinal middle area from smooth to finely punctate; transversal posterior area smooth. Interocular width equals 3.5 times transverse eye diameter. Ocular canthus subrectangular, glabrous, rugopunctate.
|
||
<emphasis id="759BEA9AFFB7FF8DFDF577C4FD082FDE" bold="true" box="[512,653,1461,1486]" pageId="29" pageNumber="219">Mouthparts</emphasis>
|
||
: Labrum parabolic, covered with a fringe of bristles at apex (
|
||
<figureCitation id="DFD42A0DFFB7FF8DFE9377A6FE4D2FE0" box="[358,456,1495,1520]" captionStart="Figure 6" captionStartId="9.[160,225,1090,1113]" captionTargetBox="[166,1145,146,1050]" captionTargetId="figure-188@9.[166,1149,142,1058]" captionTargetPageId="9" captionText="Figure 6. Labrum in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758331" httpUri="https://zenodo.org/record/6758331/files/figure.png" pageId="29" pageNumber="219">Figure 6</figureCitation>
|
||
(e)). Mandibles with 2 teeth (1 apical, 1 subapical) at outer margin separated by a narrow notch; apical tooth truncated apically; subapical tooth rounded (
|
||
<figureCitation id="DFD42A0DFFB7FF8DFEE5746CFEE82C26" box="[272,365,1565,1590]" captionStart="Figure 7" captionStartId="10.[160,225,1156,1179]" captionTargetBox="[169,1149,148,1090]" captionTargetId="figure-156@10.[166,1149,142,1125]" captionTargetPageId="10" captionText="Figure 7. Right mandible in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicate the apical tooth in (a–e), striate molar in (d)." figureDoi="http://doi.org/10.5281/zenodo.6758333" httpUri="https://zenodo.org/record/6758333/files/figure.png" pageId="29" pageNumber="219">Figure 7</figureCitation>
|
||
(e)). Maxillae wider at middle than basis; outer margin rounded; inner margin with 6 teeth (3 apical, 2 medial, 1 basal); apical and medial teeth scythe-shaped; basal tooth triangular, smaller than previous ones (
|
||
<figureCitation id="DFD42A0DFFB7FF8DFD007413FCD12C6B" box="[757,852,1634,1659]" captionStart="Figure 8" captionStartId="11.[160,225,1161,1184]" captionTargetBox="[167,1127,143,1126]" captionTargetId="figure-61@11.[166,1149,142,1129]" captionTargetPageId="11" captionText="Figure 8. Right maxilla in ventral view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicating triangular galea in (a), elongate maxillary palpomere II in (b), lobed inner margin of galea and stipe with triangular lobe in (c), pointed and triangular teeth on galea in (d), scythe-shape d teeth and rounded inner margin of galea in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758335" httpUri="https://zenodo.org/record/6758335/files/figure.png" pageId="29" pageNumber="219">Figure 8</figureCitation>
|
||
(e)). Labium with rounded sides at basal half, convergent on apical half; apex wide, slightly rounded; surface surrounded with dense, setigerous punctures.
|
||
<emphasis id="759BEA9AFFB7FF8DFD4774D6FCA22CD0" bold="true" box="[690,807,1703,1728]" pageId="29" pageNumber="219">Antennae</emphasis>
|
||
: Club oval, subequal in length to antenomeres II–VII combined.
|
||
<emphasis id="759BEA9AFFB4FF8EFDDA72FFFD012AB7" bold="true" box="[559,644,142,167]" pageId="30" pageNumber="220">Thorax</emphasis>
|
||
: Pronotum with complete border; anterior margin with a horn at middle, followed by a wide, deep, U-shaped concavity, nearly reaching the posterior margin; horn, in dorsal view, from short (not covered the head) to long (covered the head), wide at basis, narrow laterally at middle, bifurcate at apex; horn, in lateral view, rarely wide dorsoventrally, obliquely produced regarding anteroposterior body axis; pronotal sides strongly convex in lateral view; surface of concavity with large, ocellate, irregularly scattered punctures (
|
||
<figureCitation id="DFD42A0DFFB4FF8EFD0F732FFCDF2B67" box="[762,858,350,375]" captionStart="Figure 9" captionStartId="12.[160,225,1146,1169]" captionTargetBox="[184,1146,142,1069]" captionTargetId="figure-59@12.[166,1149,142,1114]" captionTargetPageId="12" captionText="Figure 9. Detail of head, pronotum and prothoracic legs of males in dorsal view: (a) Gibboryctes szelenyii (Endrödi); (b) Gibboryctes endroedii sp. nov.; (c) Heterogomphus gracilicornis Prell; (d) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758337" httpUri="https://zenodo.org/record/6758337/files/figure.png" pageId="30" pageNumber="220">Figure 9</figureCitation>
|
||
(d)); sides usually smooth, posterior area finely punctate. Scutellar plate subtriangular, glabrous, irregularly punctate anteriorly, posteriorly smooth. Elytral with 10 well-marked striae (1 sutural, 5 discal, 4 lateral); punctures on striae ocellate, large, deep anteriorly, becoming shallow, small towards posterior area; first interstria covered with large, ocellate, oval punctures; other interstriae finely punctate.
|
||
<emphasis id="759BEA9AFFB4FF8EFE29707DFD962835" bold="true" box="[476,531,524,549]" pageId="30" pageNumber="220">Legs</emphasis>
|
||
: Protarsomere V clavate, elongated, subequal to tarsomeres II–IV combined; tarsomeres I–IV subequal in length, beaker-shaped. Protibiae with IV triangular, acute, short teeth, produced on outer margins. Mesofemora oval, scarcely setose ventrally. Mesotibiae cylindrical, with 2 carinae on outer surface (1 basal, 1 medial); carinae and apex covered with stout spinules like setae. Metafemora with ventral surface glabrous, from smooth to finely punctate at discal area. Metatibiae similar to mesotibiae. Meso- and metatarsi with tarsomeres I–IV gradually decreasing in size; outer apex of tarsomere I produced, acute, with 3 stout spinules; tarsomere V subequal in length to tarsomeres II–IV combined.
|
||
<emphasis id="759BEA9AFFB4FF8EFD1D7150FCDB292A" bold="true" box="[744,862,801,826]" pageId="30" pageNumber="220">Abdomen</emphasis>
|
||
: Tergite VII rugopunctate, with no stridulatory apparatus. Tergite VIII transverse strongly convex in lateral view; surface densely setose, rugopunctate on sides, finely punctate on disc. Ventrites II–V subequal in length, rugopunctate on sides, finely punctate on disc, with an incomplete row of setigerous punctures confined on sides; ventrite VI weakly rugopunctate on sides, finely punctate on disc, posterior margin setose, widely emarginate at middle.
|
||
<emphasis id="759BEA9AFFB4FF8EFF557180FF762E1A" bold="true" box="[160,243,1009,1034]" pageId="30" pageNumber="220">Venter</emphasis>
|
||
: Prosternal process conic, densely setose. Mesoventrite completely setose. Metaventrite nearly completely setose; discal area glabrous.
|
||
<emphasis id="759BEA9AFFB4FF8EFC947662FC5D2E3C" bold="true" box="[865,984,1043,1068]" pageId="30" pageNumber="220">Aedeagus</emphasis>
|
||
: Parameres, in dorsal view, wide at basal half, narrow at middle, becoming gradually wide at apical half, scarcely bristled lateroventrally at inner side of apex (
|
||
<figureCitation id="DFD42A0DFFB4FF8EFCF57628FCE92E62" box="[768,876,1113,1138]" captionStart="Figure 10" captionStartId="13.[160,226,1164,1187]" captionTargetBox="[169,1145,145,1132]" captionTargetId="figure-89@13.[166,1149,142,1133]" captionTargetPageId="13" captionText="Figure 10. Aedeagus with parameres in dorsal and lateral views, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes endroedii sp. nov.; (e,f) Heterogomphus gracilicornis Prell; (g,h) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758339" httpUri="https://zenodo.org/record/6758339/files/figure.png" pageId="30" pageNumber="220">Figure 10</figureCitation>
|
||
(g)). Parameres, in lateral view, sinuous dorsoventrally, ventrally with a strong carina at basis, apex narrow (
|
||
<figureCitation id="DFD42A0DFFB4FF8EFF5D76EFFE932EA7" box="[168,278,1182,1207]" captionStart="Figure 10" captionStartId="13.[160,226,1164,1187]" captionTargetBox="[169,1145,145,1132]" captionTargetId="figure-89@13.[166,1149,142,1133]" captionTargetPageId="13" captionText="Figure 10. Aedeagus with parameres in dorsal and lateral views, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes endroedii sp. nov.; (e,f) Heterogomphus gracilicornis Prell; (g,h) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758339" httpUri="https://zenodo.org/record/6758339/files/figure.png" pageId="30" pageNumber="220">Figure 10</figureCitation>
|
||
(h)).
|
||
</paragraph>
|
||
<paragraph id="47503688FFB4FF8EFF557692FE212EED" blockId="30.[160,1157,1251,1624]" box="[160,420,1251,1277]" pageId="30" pageNumber="220">
|
||
<emphasis id="759BEA9AFFB4FF8EFF557692FE212EED" bold="true" box="[160,420,1251,1277]" pageId="30" pageNumber="220">
|
||
Female
|
||
<emphasis id="759BEA9AFFB4FF8EFEF57692FE212EED" bold="true" box="[256,420,1251,1277]" italics="true" pageId="30" pageNumber="220">
|
||
(
|
||
<figureCitation id="DFD42A0DFFB4FF8EFEFC7692FEEC2EED" box="[265,361,1251,1277]" captionStart="Figure 5" captionStartId="8.[160,225,1685,1708]" captionTargetBox="[221,1093,152,1648]" captionTargetId="figure-3@8.[219,1097,142,1654]" captionTargetPageId="8" captionText="Figure 5. Tetragus waldenfelsi (Endrödi, 1977). (a,b) Male habitus in dorsal and lateral views, respectively; (c,d) female habitus in dorsal and lateral views, respectively." figureDoi="http://doi.org/10.5281/zenodo.6758329" httpUri="https://zenodo.org/record/6758329/files/figure.png" pageId="30" pageNumber="220">Figure 5</figureCitation>
|
||
(c,d))
|
||
</emphasis>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="47503688FFB4FF8EFF557777FEC72C48" blockId="30.[160,1157,1251,1624]" pageId="30" pageNumber="220">
|
||
<emphasis id="759BEA9AFFB4FF8EFF557777FEC32F30" bold="true" box="[160,326,1286,1312]" italics="true" pageId="30" pageNumber="220">Redescription.</emphasis>
|
||
<emphasis id="759BEA9AFFB4FF8EFEA37776FE292F30" bold="true" box="[342,428,1287,1312]" pageId="30" pageNumber="220">Length</emphasis>
|
||
:
|
||
<quantity id="80179B6DFFB4FF8EFE4B7776FDE02F30" box="[446,613,1287,1312]" metricMagnitude="-2" metricUnit="m" metricValue="2.9549999999999996" metricValueMax="3.2299999999999995" metricValueMin="2.68" pageId="30" pageNumber="220" unit="mm" value="29.549999999999997" valueMax="32.3" valueMin="26.8">26.8–32.3 mm</quantity>
|
||
;
|
||
<emphasis id="759BEA9AFFB4FF8EFD817776FD382F30" bold="true" box="[628,701,1287,1312]" pageId="30" pageNumber="220">Width</emphasis>
|
||
:
|
||
<quantity id="80179B6DFFB4FF8EFD3A7776FCF22F30" box="[719,887,1287,1312]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.41" metricValueMin="1.19" pageId="30" pageNumber="220" unit="mm" value="13.0" valueMax="14.1" valueMin="11.9">11.9–14.1 mm</quantity>
|
||
. Distinct from male in the following characters:
|
||
<emphasis id="759BEA9AFFB4FF8EFE257758FD8A2F52" bold="true" box="[464,527,1321,1346]" pageId="30" pageNumber="220">Head</emphasis>
|
||
: Clypeus from subtrapezoidal to subtriangular, truncated or slightly rounded at anterior margin, coarsely rugopunctate (
|
||
<figureCitation id="DFD42A0DFFB4FF8EFC12773DFBD22F75" box="[999,1111,1356,1381]" captionStart="Figure 11" captionStartId="14.[160,225,1441,1464]" captionTargetBox="[168,1146,144,1407]" captionTargetId="figure-42@14.[166,1149,142,1410]" captionTargetPageId="14" captionText="Figure 11. Detail of head, pronotum and prothoracic legs of females in dorsal view: (a) Gibboryctes szelenyii Endrödi;(b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedi sp.nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Arrow indicates punctate frons in (b)." figureDoi="http://doi.org/10.5281/zenodo.6758341" httpUri="https://zenodo.org/record/6758341/files/figure.png" pageId="30" pageNumber="220">Figure 11</figureCitation>
|
||
(e)); frons entirely, coarsely rugopunctate.
|
||
<emphasis id="759BEA9AFFB4FF8EFDAA771EFD312F98" bold="true" box="[607,692,1391,1416]" pageId="30" pageNumber="220">Thorax</emphasis>
|
||
: Pronotum with 2 flattened, transverse tubercles contiguous to middle of anterior margin; pronotal concavity V-shaped, transversely rugopunctate; concavity V-shaped, coarsely rugopunctate (
|
||
<figureCitation id="DFD42A0DFFB4FF8EFC1F77C5FBD22FDD" box="[1002,1111,1460,1485]" captionStart="Figure 11" captionStartId="14.[160,225,1441,1464]" captionTargetBox="[168,1146,144,1407]" captionTargetId="figure-42@14.[166,1149,142,1410]" captionTargetPageId="14" captionText="Figure 11. Detail of head, pronotum and prothoracic legs of females in dorsal view: (a) Gibboryctes szelenyii Endrödi;(b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedi sp.nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Arrow indicates punctate frons in (b)." figureDoi="http://doi.org/10.5281/zenodo.6758341" httpUri="https://zenodo.org/record/6758341/files/figure.png" pageId="30" pageNumber="220">Figure 11</figureCitation>
|
||
(e)); sides of pronotum weakly convex in lateral view.
|
||
<emphasis id="759BEA9AFFB4FF8EFD0577A6FCAD2FE0" bold="true" box="[752,808,1495,1520]" pageId="30" pageNumber="220">Legs</emphasis>
|
||
: Metafemora with large, oval punctures on ventral surface.
|
||
<emphasis id="759BEA9AFFB4FF8EFDFC7788FD052C02" bold="true" box="[521,640,1529,1554]" pageId="30" pageNumber="220">Abdomen</emphasis>
|
||
: Tergite VIII flattened to weakly convex in lateral view; ventrite V parabolic, moderately punctate, with no posterior emargination (
|
||
<figureCitation id="DFD42A0DFFB4FF8EFF5D744EFE9D2C48" box="[168,280,1599,1624]" captionStart="Figure 13" captionStartId="16.[160,225,1058,1081]" captionTargetBox="[166,1146,145,1023]" captionTargetId="figure-155@16.[166,1149,142,1026]" captionTargetPageId="16" captionText="Figure 13. Detail of ventrites I–VI of females. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov; (c) Gibboryctes endroedii sp. nov.; (d) Tetragus waldenfelsi (Endrödi). Red arrows indicate the proportions of ventrites V and VI." figureDoi="http://doi.org/10.5281/zenodo.6758345" httpUri="https://zenodo.org/record/6758345/files/figure.png" pageId="30" pageNumber="220">Figure 13</figureCitation>
|
||
(d)).
|
||
</paragraph>
|
||
<paragraph id="47503688FFB4FF8EFF5574F5FDE92C8E" blockId="30.[160,1000,1668,1728]" box="[160,620,1668,1694]" pageId="30" pageNumber="220">
|
||
<emphasis id="759BEA9AFFB4FF8EFF5574F5FDE92C8E" bold="true" box="[160,620,1668,1694]" pageId="30" pageNumber="220">
|
||
Geographic distribution
|
||
<emphasis id="759BEA9AFFB4FF8EFE3974F5FDE92C8E" bold="true" box="[460,620,1668,1694]" italics="true" pageId="30" pageNumber="220">
|
||
(
|
||
<figureCitation id="DFD42A0DFFB4FF8EFE2074F5FDC42C8E" box="[469,577,1668,1694]" captionStart="Figure 14" captionStartId="17.[160,227,1388,1411]" captionTargetBox="[168,1147,145,1355]" captionTargetId="figure-89@17.[166,1149,142,1357]" captionTargetPageId="17" captionText="Figure 14. Geographic distribution maps. (a) Tetragus species; (b) Gibboryctes species; (c) Heterogomphus gracilicornis Prell." figureDoi="http://doi.org/10.5281/zenodo.6758347" httpUri="https://zenodo.org/record/6758347/files/figure.png" pageId="30" pageNumber="220">Figure 14</figureCitation>
|
||
(a))
|
||
</emphasis>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="47503688FFB4FF8EFF5574D6FC6D2CD0" blockId="30.[160,1000,1668,1728]" box="[160,1000,1703,1728]" pageId="30" pageNumber="220">
|
||
<collectingCountry id="3FF87618FFB4FF8EFF5574D6FF782CD0" box="[160,253,1703,1728]" name="Panama" pageId="30" pageNumber="220">Panama</collectingCountry>
|
||
,
|
||
<collectingCountry id="3FF87618FFB4FF8EFEF274D6FE292CD0" box="[263,428,1703,1728]" name="French Guiana" pageId="30" pageNumber="220">French Guiana</collectingCountry>
|
||
,
|
||
<collectingCountry id="3FF87618FFB4FF8EFE4274D6FE762CD0" box="[439,499,1703,1728]" name="Brazil" pageId="30" pageNumber="220">Brazil</collectingCountry>
|
||
(Amazonas;
|
||
<collectingRegion id="852BF86AFFB4FF8EFD7774D6FC972CD0" box="[642,786,1703,1728]" country="Brazil" name="Mato Grosso" pageId="30" pageNumber="220">Mato Grosso</collectingRegion>
|
||
, new state record).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FF56503FFB5FF8FFF5572FFFCDF2B23" pageId="31" pageNumber="221" type="discussion">
|
||
<paragraph id="47503688FFB5FF8FFF5572FFFE832AB8" blockId="31.[160,1156,142,307]" box="[160,262,142,168]" pageId="31" pageNumber="221">
|
||
<emphasis id="759BEA9AFFB5FF8FFF5572FFFE832AB8" bold="true" box="[160,262,142,168]" italics="true" pageId="31" pageNumber="221">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="47503688FFB5FF8FFF5572C3FCDF2B23" blockId="31.[160,1156,142,307]" pageId="31" pageNumber="221">
|
||
Males of
|
||
<taxonomicName id="80EF4D0BFFB5FF8FFEF372C3FE1D2ADB" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" baseAuthorityName="Costa & Duarte & Iannuzzi & Grossi" baseAuthorityYear="1977" box="[262,408,178,203]" class="Insecta" family="Dynastidae" genus="Strategus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="221" phylum="Arthropoda" rank="species" species="waldenfelsi" status="comb. nov.">
|
||
<emphasis id="759BEA9AFFB5FF8FFEF372C3FE1D2ADB" box="[262,408,178,203]" italics="true" pageId="31" pageNumber="221">T. waldenfelsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFB5FF8FFE5572C3FD9B2ADB" box="[416,542,178,203]" pageId="31" pageNumber="221" rank="species">comb. nov.</taxonomicNameLabel>
|
||
have allometric development evidenced by the shape of male horn sometimes broader dorsoventrally at middle wider than basis in lateral view; longer (covering the head in dorsal view, surpassing the anterior margin of clypeus in lateral view) or shorter (not covering the head in dorsal view).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |