361 lines
47 KiB
XML
361 lines
47 KiB
XML
<document id="51063C51B20097A0910353CD30DCAE55" ID-CLB-Dataset="82886" ID-DOI="10.1080/00222933.2021.2017499" ID-GBIF-Dataset="5cd01cfe-52bb-4092-97b2-804c3c7d145d" ID-ISSN="1464-5262" ID-Zenodo-Dep="6758313" ID-ZooBank="D766FC5E-CEC8-4184-BE90-C25350B90612" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656311158681" checkinUser="plazi" docAuthor="Costa, Leidiane O., Duarte, Paulo R. M., Iannuzzi, Luciana & Grossi, Paschoal C." docDate="2022" docId="CF46879EFFA9FF9CFE23747DFE6F2C02" docLanguage="en" docName="JNATHIST.56.1-4.191-225.pdf" docOrigin="Journal of Natural History 56 (1 - 4)" docSource="http://dx.doi.org/10.1080/00222933.2021.2017499" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Gibboryctes Endrodi 1974" docType="treatment" docVersion="7" lastPageNumber="202" masterDocId="337FFFE6FFAAFF90FFF57271FF852A10" masterDocTitle="Taxonomic revision and notes on natural history of the enigmatic beetle genus Gibboryctes Endrödi (Coleoptera: Melolonthidae: Dynastinae)" masterLastPageNumber="225" masterPageNumber="191" pageNumber="193" updateTime="1699352275692" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="D0A56E80CEFE6A12B2207FED94F22025" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="6A35A52073147CAAE4F0BC693B9FF191">
|
||
<mods:title id="F1B87406BA705D6E3935C9923DB9E389">Taxonomic revision and notes on natural history of the enigmatic beetle genus Gibboryctes Endrödi (Coleoptera: Melolonthidae: Dynastinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="EAC44C33AE7C670EE2E4BACE81B604F8" type="personal">
|
||
<mods:role id="65CFEF80FC6D66B7EC34675E6958F208">
|
||
<mods:roleTerm id="5144225766BCFC0E7C1570499305FDB9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="50347704CE39C973DCF596B182A2548B">Costa, Leidiane O.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="DE99278AEF7A65F949F58B0809C0764C" type="personal">
|
||
<mods:role id="C975F405F9C205B86AEC32DE936B5C4C">
|
||
<mods:roleTerm id="C746E504F4A9F86B51E6A0450BF4408C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4B9EA268821B7179BEEE9B03F97841A8">Duarte, Paulo R. M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="86D4C8A2EA8187E86958A9087E100E87" type="personal">
|
||
<mods:role id="E11647768929FFC40FE2F8F068E35C26">
|
||
<mods:roleTerm id="26635E939F333A021B4C6E762F7826F7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="07213D984BA19AFAB1565628A517A8AF">Iannuzzi, Luciana</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="8760A66BE75FD2E13035F2BF8A147A7E" type="personal">
|
||
<mods:role id="750531F04C41B6F72CF32BEF993E9302">
|
||
<mods:roleTerm id="035D007F9E5CC548394723938DEBAE87">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="38DAB0BD074F0031E7DC5B75B93F374E">Grossi, Paschoal C.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="F209709773328A10C01D633821BEF2EA">text</mods:typeOfResource>
|
||
<mods:relatedItem id="DAEEDD9CC7644CC814E9BB0BD439910E" type="host">
|
||
<mods:titleInfo id="E65632F33B65040DFBD1B52B01A8B736">
|
||
<mods:title id="78E1F497746E348CF4CCA571C1864C14">Journal of Natural History</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B3BAE0ACC47FF5732C6D9B57D4BE94B7">
|
||
<mods:date id="0954710B7602F4F3A518383D913F2CC3">2022</mods:date>
|
||
<mods:detail id="8C96A05D5D5CCBA6FDC2F0E232BC0232" type="pubDate">
|
||
<mods:number id="3948F75E01E6860DFF2C2E4B29474484">2022-05-17</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="CAC320279040468392AD4F8EA2ADA64B" type="volume">
|
||
<mods:number id="94185E906FED055F4801834877D9676D">56</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="3343560CC6ADC01B19AECCA9DB9B5DA9" type="issue">
|
||
<mods:number id="E71E76986C0ADBBCB1E14D55A498981A">1 - 4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="C6164392CFFC19DF0BF61A9AD04349CD" unit="page">
|
||
<mods:start id="11F668991FAF430DF0F879AFE9255EBA">191</mods:start>
|
||
<mods:end id="6B2E39C983BC944B77CA5BF84ECD7D94">225</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="B46ECDF1AB6C110E63D35980FA394054">
|
||
<mods:url id="0D8F5A9CF8D415F4F753353C3D871C77">http://dx.doi.org/10.1080/00222933.2021.2017499</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="6B0CDB0B89D7EBFF90BF2BB02A43CB3D">journal article</mods:classification>
|
||
<mods:identifier id="F58D54730A6DA71C34CEEA07BDA5017C" type="CLB-Dataset">82886</mods:identifier>
|
||
<mods:identifier id="8B473B1AB4B04E4BF90B10D6AE89EDC1" type="DOI">10.1080/00222933.2021.2017499</mods:identifier>
|
||
<mods:identifier id="73319535239626A91D2092F5BD0342D9" type="GBIF-Dataset">5cd01cfe-52bb-4092-97b2-804c3c7d145d</mods:identifier>
|
||
<mods:identifier id="90577998A551FE21832E163CAFCC8C01" type="ISSN">1464-5262</mods:identifier>
|
||
<mods:identifier id="29C071938ABB7DA021D4B90DBFEF274B" type="Zenodo-Dep">6758313</mods:identifier>
|
||
<mods:identifier id="A9B616A0895CDE00FCD544F10F758A06" type="ZooBank">D766FC5E-CEC8-4184-BE90-C25350B90612</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="CF46879EFFA9FF9CFE23747DFE6F2C02" ID-DOI="http://doi.org/10.5281/zenodo.6758315" ID-GBIF-Taxon="196400124" ID-Zenodo-Dep="6758315" LSID="urn:lsid:plazi:treatment:CF46879EFFA9FF9CFE23747DFE6F2C02" httpUri="http://treatment.plazi.org/id/CF46879EFFA9FF9CFE23747DFE6F2C02" lastPageId="12" lastPageNumber="202" pageId="3" pageNumber="193">
|
||
<subSubSection id="0FF56503FFA9FF93FE23747DFCC82C36" box="[470,845,1548,1574]" pageId="3" pageNumber="193" type="nomenclature">
|
||
<paragraph id="47503688FFA9FF93FE23747DFCC82C36" blockId="3.[470,845,1548,1574]" box="[470,845,1548,1574]" pageId="3" pageNumber="193">
|
||
<heading id="1C1881E4FFA9FF93FE23747DFCC82C36" bold="true" box="[470,845,1548,1574]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="193" reason="3">
|
||
<treatmentCitationGroup id="67FF11A6FFA9FF93FE23747DFCC82C36" box="[470,845,1548,1574]" pageId="3" pageNumber="193">
|
||
<treatmentCitation id="C64E1099FFA9FF93FE23747DFCC82C36" author="Endrodi S" box="[470,845,1548,1574]" page="13" pageId="3" pageNumber="193" year="1974">
|
||
<taxonomicName id="80EF4D0BFFA9FF93FE23747DFCC82C36" authority="Endrodi, 1974: 13" authorityName="Endrodi" authorityPageNumber="13" authorityYear="1974" box="[470,845,1548,1574]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="193" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="759BEA9AFFA9FF93FE23747DFCC82C36" bold="true" box="[470,845,1548,1574]" pageId="3" pageNumber="193">
|
||
Gibboryctes
|
||
<bibRefCitation id="237E4B79FFA9FF93FD80747DFCC82C36" author="Endrodi S" box="[629,845,1548,1574]" pageId="3" pageNumber="193" pagination="13 - 16" refId="ref13334" refString="Endrodi S. 1974. Gibboryctes szelenyii gen. nov. sp. nov. Folia Entomol Hung. 27: 13 - 16." type="journal article" year="1974">
|
||
<emphasis id="759BEA9AFFA9FF93FD80747DFCC82C36" bold="true" box="[629,845,1548,1574]" italics="true" pageId="3" pageNumber="193">Endrödi, 1974: 13</emphasis>
|
||
</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FF56503FFA9FF93FF557425FD532C81" pageId="3" pageNumber="193" type="type_taxon">
|
||
<paragraph id="47503688FFA9FF93FF557425FEB62C7E" blockId="3.[160,726,1620,1681]" box="[160,307,1620,1646]" pageId="3" pageNumber="193">
|
||
<emphasis id="759BEA9AFFA9FF93FF557425FEB62C7E" bold="true" box="[160,307,1620,1646]" italics="true" pageId="3" pageNumber="193">
|
||
<typeStatus id="9854882AFFA9FF93FF557425FF5D2C7E" box="[160,216,1620,1646]" pageId="3" pageNumber="193">Type</typeStatus>
|
||
species
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="47503688FFA9FF93FF557409FD532C81" blockId="3.[160,726,1620,1681]" box="[160,726,1656,1681]" pageId="3" pageNumber="193">
|
||
<taxonomicName id="80EF4D0BFFA9FF93FF557409FDA72C81" authority="Endrodi, 1974" authorityName="Endrodi" authorityYear="1974" box="[160,546,1656,1681]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="193" phylum="Arthropoda" rank="species" species="szelenyii">
|
||
<emphasis id="759BEA9AFFA9FF93FF557409FEFA2C81" box="[160,383,1656,1681]" italics="true" pageId="3" pageNumber="193">Gibboryctes szelenyii</emphasis>
|
||
<bibRefCitation id="237E4B79FFA9FF93FE727409FDA72C81" author="Endrodi S" box="[391,546,1656,1681]" pageId="3" pageNumber="193" pagination="13 - 16" refId="ref13334" refString="Endrodi S. 1974. Gibboryctes szelenyii gen. nov. sp. nov. Folia Entomol Hung. 27: 13 - 16." type="journal article" year="1974">Endrödi, 1974</bibRefCitation>
|
||
</taxonomicName>
|
||
(by monotypy).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="13906600FFAEFF94FF557727FDBD2FB8" ID-DOI="http://doi.org/10.5281/zenodo.6758317" ID-Zenodo-Dep="6758317" httpUri="https://zenodo.org/record/6758317/files/figure.png" pageId="4" pageNumber="194" startId="4.[160,227,1366,1389]" targetBox="[162,1148,150,1331]" targetPageId="4">
|
||
<paragraph id="47503688FFAEFF94FF557727FDBD2FB8" blockId="4.[160,1157,1366,1448]" pageId="4" pageNumber="194">
|
||
<emphasis id="759BEA9AFFAEFF94FF557727FE842F7D" bold="true" box="[160,257,1366,1389]" pageId="4" pageNumber="194">Figure 1.</emphasis>
|
||
<taxonomicName id="80EF4D0BFFAEFF94FEF87727FDF32F7D" authority="Endrodi, 1974" authorityName="Endrodi" authorityYear="1974" box="[269,630,1366,1389]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="194" phylum="Arthropoda" rank="species" species="szelenyii">
|
||
<emphasis id="759BEA9AFFAEFF94FEF87727FE5E2F7D" box="[269,475,1366,1389]" italics="true" pageId="4" pageNumber="194">Gibboryctes szelenyii</emphasis>
|
||
<bibRefCitation id="237E4B79FFAEFF94FE137727FDF32F7D" author="Endrodi S" box="[486,630,1366,1389]" pageId="4" pageNumber="194" pagination="13 - 16" refId="ref13334" refString="Endrodi S. 1974. Gibboryctes szelenyii gen. nov. sp. nov. Folia Entomol Hung. 27: 13 - 16." type="journal article" year="1974">Endrödi, 1974</bibRefCitation>
|
||
</taxonomicName>
|
||
. (a,b) Holotype male in dorsal and lateral views, respectively; (c) labels; (d,e) male habitus in dorsal and lateral views, respectively; (f) female habitus in dorsal view. (a,b) pictures out of scale.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0FF56503FFAEFF95FF557783FF652F20" lastPageId="5" lastPageNumber="195" pageId="4" pageNumber="194" type="diagnosis">
|
||
<paragraph id="47503688FFAEFF94FF557783FE912C1C" blockId="4.[159,1156,1522,1722]" box="[160,276,1522,1548]" pageId="4" pageNumber="194">
|
||
<emphasis id="759BEA9AFFAEFF94FF557783FE912C1C" bold="true" box="[160,276,1522,1548]" italics="true" pageId="4" pageNumber="194">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="47503688FFAEFF95FF557467FF652F20" blockId="4.[159,1156,1522,1722]" lastBlockId="5.[159,1155,1199,1328]" lastPageId="5" lastPageNumber="195" pageId="4" pageNumber="194">
|
||
<taxonomicName id="80EF4D0BFFAEFF94FF557467FE9F2C3F" box="[160,282,1558,1583]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="194" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="759BEA9AFFAEFF94FF557467FE9F2C3F" box="[160,282,1558,1583]" italics="true" pageId="4" pageNumber="194">Gibboryctes</emphasis>
|
||
</taxonomicName>
|
||
is separated from other Neotropical Oryctini by the following combination of characters: inconspicuous sexual dimorphism (
|
||
<figureCitation id="DFD42A0DFFAEFF94FD6E7448FD7F2C42" box="[667,762,1593,1618]" captionStart="Figure 1" captionStartId="4.[160,227,1366,1389]" captionTargetBox="[162,1148,150,1331]" captionTargetId="figure-77@4.[162,1154,142,1335]" captionTargetPageId="4" captionText="Figure 1. Gibboryctes szelenyii Endrödi, 1974. (a,b) Holotype male in dorsal and lateral views, respectively; (c) labels; (d,e) male habitus in dorsal and lateral views, respectively; (f) female habitus in dorsal view. (a,b) pictures out of scale." figureDoi="http://doi.org/10.5281/zenodo.6758317" httpUri="https://zenodo.org/record/6758317/files/figure.png" pageId="4" pageNumber="194">Figures 1</figureCitation>
|
||
(d–f) and 3(a–c)); transverse head, 2 times wider than long (
|
||
<figureCitation id="DFD42A0DFFAEFF94FE63742AFE682C64" box="[406,493,1627,1652]" captionStart="Figure 9" captionStartId="12.[160,225,1146,1169]" captionTargetBox="[184,1146,142,1069]" captionTargetId="figure-59@12.[166,1149,142,1114]" captionTargetPageId="12" captionText="Figure 9. Detail of head, pronotum and prothoracic legs of males in dorsal view: (a) Gibboryctes szelenyii (Endrödi); (b) Gibboryctes endroedii sp. nov.; (c) Heterogomphus gracilicornis Prell; (d) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758337" httpUri="https://zenodo.org/record/6758337/files/figure.png" pageId="4" pageNumber="194">Figure 9</figureCitation>
|
||
(a,b));both sexes with a transverse tubercle on head (
|
||
<figureCitation id="DFD42A0DFFAEFF94FBD7742AFB062C64" box="[1058,1155,1627,1652]" captionStart="Figure 9" captionStartId="12.[160,225,1146,1169]" captionTargetBox="[184,1146,142,1069]" captionTargetId="figure-59@12.[166,1149,142,1114]" captionTargetPageId="12" captionText="Figure 9. Detail of head, pronotum and prothoracic legs of males in dorsal view: (a) Gibboryctes szelenyii (Endrödi); (b) Gibboryctes endroedii sp. nov.; (c) Heterogomphus gracilicornis Prell; (d) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758337" httpUri="https://zenodo.org/record/6758337/files/figure.png" pageId="4" pageNumber="194">Figures 9</figureCitation>
|
||
(a,b) and 11(a–c)); bidentate mandibles with subapical tooth backward produced and larger than apical tooth (
|
||
<figureCitation id="DFD42A0DFFAEFF94FE6D74D0FD872CAA" box="[408,514,1697,1722]" captionStart="Figure 7" captionStartId="10.[160,225,1156,1179]" captionTargetBox="[169,1149,148,1090]" captionTargetId="figure-156@10.[166,1149,142,1125]" captionTargetPageId="10" captionText="Figure 7. Right mandible in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicate the apical tooth in (a–e), striate molar in (d)." figureDoi="http://doi.org/10.5281/zenodo.6758333" httpUri="https://zenodo.org/record/6758333/files/figure.png" pageId="4" pageNumber="194">Figure 7</figureCitation>
|
||
(a–c)); inner margin of galea bearing small teeth; pronotum with no horns,with only a small tubercle on anterior area combined with a shallow or absent concavity (
|
||
<figureCitation id="DFD42A0DFFAFFF95FE7276A3FE692EFB" box="[391,492,1234,1259]" captionStart="Figure 9" captionStartId="12.[160,225,1146,1169]" captionTargetBox="[184,1146,142,1069]" captionTargetId="figure-59@12.[166,1149,142,1114]" captionTargetPageId="12" captionText="Figure 9. Detail of head, pronotum and prothoracic legs of males in dorsal view: (a) Gibboryctes szelenyii (Endrödi); (b) Gibboryctes endroedii sp. nov.; (c) Heterogomphus gracilicornis Prell; (d) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758337" httpUri="https://zenodo.org/record/6758337/files/figure.png" pageId="5" pageNumber="195">Figures 9</figureCitation>
|
||
(a,b) and
|
||
<figureCitation id="DFD42A0DFFAFFF95FDA076A3FD412EFB" box="[597,708,1234,1259]" captionStart="Figure 11" captionStartId="14.[160,225,1441,1464]" captionTargetBox="[168,1146,144,1407]" captionTargetId="figure-42@14.[166,1149,142,1410]" captionTargetPageId="14" captionText="Figure 11. Detail of head, pronotum and prothoracic legs of females in dorsal view: (a) Gibboryctes szelenyii Endrödi;(b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedi sp.nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Arrow indicates punctate frons in (b)." figureDoi="http://doi.org/10.5281/zenodo.6758341" httpUri="https://zenodo.org/record/6758341/files/figure.png" pageId="5" pageNumber="195">Figures 11</figureCitation>
|
||
(a–c)); elytra with dense, large and deep punctures (
|
||
<figureCitation id="DFD42A0DFFAFFF95FEE87685FE022F1D" box="[285,391,1268,1293]" captionStart="Figure 12" captionStartId="15.[160,225,1189,1212]" captionTargetBox="[167,1149,146,1154]" captionTargetId="figure-100@15.[166,1149,142,1157]" captionTargetPageId="15" captionText="Figure 12. Females of Gibboryctes species with detail of head, pronotum and prothoracic legs in lateral view and left elytra in dorsal view, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes ebeninus sp. nov.; (e,f) Gibboryctes endroedii sp. nov. Jsi, juxtasutural interstria. Arrow indicates coalescent punctures in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758343" httpUri="https://zenodo.org/record/6758343/files/figure.png" pageId="5" pageNumber="195">Figure 12</figureCitation>
|
||
(b,d,f)); ventrite VI 2 times longer than ventrite V in female (
|
||
<figureCitation id="DFD42A0DFFAFFF95FBEE7685FB062F1D" box="[1051,1155,1268,1293]" captionStart="Figure 13" captionStartId="16.[160,225,1058,1081]" captionTargetBox="[166,1146,145,1023]" captionTargetId="figure-155@16.[166,1149,142,1026]" captionTargetPageId="16" captionText="Figure 13. Detail of ventrites I–VI of females. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov; (c) Gibboryctes endroedii sp. nov.; (d) Tetragus waldenfelsi (Endrödi). Red arrows indicate the proportions of ventrites V and VI." figureDoi="http://doi.org/10.5281/zenodo.6758345" httpUri="https://zenodo.org/record/6758345/files/figure.png" pageId="5" pageNumber="195">Figure 13</figureCitation>
|
||
(a–c)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="13906600FFAFFF95FF55766BFE9E2E5E" ID-DOI="http://doi.org/10.5281/zenodo.6758319" ID-Zenodo-Dep="6758319" httpUri="https://zenodo.org/record/6758319/files/figure.png" pageId="5" pageNumber="195" startId="5.[160,225,1050,1073]" targetBox="[170,1128,191,1006]" targetPageId="5">
|
||
<paragraph id="47503688FFAFFF95FF55766BFE9E2E5E" blockId="5.[160,1157,1050,1102]" pageId="5" pageNumber="195">
|
||
<emphasis id="759BEA9AFFAFFF95FF55766BFF7E2E21" bold="true" box="[160,251,1050,1073]" pageId="5" pageNumber="195">Figure 2.</emphasis>
|
||
<taxonomicName id="80EF4D0BFFAFFF95FEF6766BFE482E21" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[259,461,1050,1073]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="195" phylum="Arthropoda" rank="species" species="ebeninus" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFAFFF95FEF6766BFE482E21" box="[259,461,1050,1073]" italics="true" pageId="5" pageNumber="195">Gibboryctes ebeninus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFAFFF95FE20766BFDA62E21" box="[469,547,1050,1073]" pageId="5" pageNumber="195" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(a, b) Habitus of paratype female in dorsal and lateral views, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0FF56503FFAFFF9CFF55771DFE172FDD" lastPageId="12" lastPageNumber="202" pageId="5" pageNumber="195" type="description">
|
||
<paragraph id="47503688FFAFFF95FF55771DFE1E2F96" blockId="5.[160,1156,1388,1725]" box="[160,411,1388,1414]" pageId="5" pageNumber="195">
|
||
<emphasis id="759BEA9AFFAFFF95FF55771DFE1E2F96" bold="true" box="[160,411,1388,1414]" italics="true" pageId="5" pageNumber="195">Generic redescription</emphasis>
|
||
</paragraph>
|
||
<paragraph id="47503688FFAFFF9BFF5577FEFD8E2C06" blockId="5.[160,1156,1388,1725]" lastBlockId="11.[160,1155,1394,1558]" lastPageId="11" lastPageNumber="201" pageId="5" pageNumber="195">
|
||
<emphasis id="759BEA9AFFAFFF95FF5577FEFF762FB8" bold="true" box="[160,243,1423,1448]" pageId="5" pageNumber="195">Length</emphasis>
|
||
: 21.0–
|
||
<quantity id="80179B6DFFAFFF95FECF77FEFE252FB8" box="[314,416,1423,1448]" metricMagnitude="-2" metricUnit="m" metricValue="3.56" pageId="5" pageNumber="195" unit="mm" value="35.6">35.6 mm</quantity>
|
||
.
|
||
<emphasis id="759BEA9AFFAFFF95FE5977FEFE762FB8" bold="true" box="[428,499,1423,1448]" pageId="5" pageNumber="195">Width</emphasis>
|
||
: 11.0–
|
||
<quantity id="80179B6DFFAFFF95FDC977FEFD242FB8" box="[572,673,1423,1448]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="5" pageNumber="195" unit="mm" value="16.9">16.9 mm</quantity>
|
||
. Poorly apparent sexual dimorphism; body elongated, oval in dorsal view, convex in lateral view; dorsal surface nearly completely glabrous; ventral surface densely setose.
|
||
<emphasis id="759BEA9AFFAFFF95FD9377A5FD312FFD" bold="true" box="[614,692,1492,1517]" pageId="5" pageNumber="195">Colour</emphasis>
|
||
: From dark reddish brown to dark.
|
||
<emphasis id="759BEA9AFFAFFF95FBB577A5FBFB2FFD" bold="true" box="[1088,1150,1492,1517]" pageId="5" pageNumber="195">Head</emphasis>
|
||
: Transverse, about 2 times wider than long. Clypeus triangular, short, about 4 times wider than long, slightly narrow laterally, anteriorly acute; surface coarsely rugose, with scattered setae on each side. Frontoclypeal suture with a short, transverse, sometimes emarginate, subtrapezoidal tubercle. Frons from flattened to slightly concave; from rugopunctate to densely punctate. Ocular canthus rectangular, transverse, glabrous.
|
||
<emphasis id="759BEA9AFFAFFF95FC6A74F3FB972C8B" bold="true" box="[927,1042,1666,1691]" pageId="5" pageNumber="195">Antennae</emphasis>
|
||
: Provided with 10 segments; club short, globular, subequal in length to antennomeres II–VII combined; club decreasing in size from basal to apical lamella.
|
||
<emphasis id="759BEA9AFFA3FF99FCAE76A5FC602EFD" bold="true" box="[859,997,1236,1261]" pageId="9" pageNumber="199">Mouthparts</emphasis>
|
||
: Labrum from trapezoidal to parabolic in shape. Mandibles with 2 rounded teeth (1 apical, 1 subapical) on outer margin; teeth projected forward beyond clypeus in dorsal view, strongly reflexed above clypeus in lateral view; subapical tooth slightly backward produced, larger in size compared to apical tooth. Maxillae with small teeth at inner margin of galea; stipe produced laterally in triangular shape. Labium oval, densely covered with bristled punctures.
|
||
<emphasis id="759BEA9AFFA3FF99FF3E77D5FE982FAD" bold="true" box="[203,285,1444,1469]" pageId="9" pageNumber="199">Thorax</emphasis>
|
||
: Pronotum weakly convex in lateral view; horn absent, but with a small tubercle located at middle area contiguous to anterior margin; dorsal surface nearly completely covered with dense, large, deep, ocellated punctures; discal area covered with large, coalescent punctures, sometimes longitudinally scattered along of a middle line; anterior and lateral borders complete, posterior usually incomplete. Scutellar plate from subtriangular to parabolic, bearing irregularly scattered punctures. Elytral surface with 8 wellmarked striae longitudinally (4 discal, 4 lateral); striae covered with a row of large, ocellated, deep punctures; interstriae with mixed fine and large punctures.
|
||
<emphasis id="759BEA9AFFA3FF99FC9674E6FC1C2CA0" bold="true" box="[867,921,1687,1712]" pageId="9" pageNumber="199">Legs</emphasis>
|
||
: Protarsi with scytheshaped claws; tarsomere V clavate, elongated, 2 times longer than tarsomere IV; tarsomeres I–IV ‘beaker’-shaped, subequal in length. Protibial outer margins with 4 teeth (1 apical, 2 medial, 1 basal); apical tooth triangular, curved, pointed; medial and basal teeth triangular; basal tooth smaller than previous ones; dorsal surface weakly punctate, longitudinally covered with a double row of bristled punctures. Mesotarsi similar to metatarsi, with tarsomeres I–IV gradually decreasing in size. Meso- and metatibiae cylindrical in shape, with 2 transverse carinae on outer surface (1 basal, 1 medial); carinae and apex weakly crenulated, covered with stout spinules like-setae; inner surface moderately covered with long bristles.
|
||
<emphasis id="759BEA9AFFA0FF9AFEC2744EFE2E2C48" bold="true" box="[311,427,1599,1624]" pageId="10" pageNumber="200">Abdomen</emphasis>
|
||
: Tergite VII lacking stridulatory apparatus. Tergite VIII transverse, subtriangular, strongly convex in lateral view in male; surface glabrous, rugopunctate on sides, densely covered with large, deep punctures on discal area. Ventrites II–VI coarsely rugopunctate on sides, finely punctate and glabrous on discal area; ventrites III–V with an incomplete row of setigerous punctures confined on sides; posterior margin of ventrite VI emarginate, setose.
|
||
<emphasis id="759BEA9AFFA1FF9BFE7477E4FE702FBE" bold="true" box="[385,501,1429,1454]" pageId="11" pageNumber="201">Aedeagus</emphasis>
|
||
: Parameres, in dorsal view, symmetrical, basal half clearly wider than apical half, inner margin weakly bristled on apex. Parameres, in lateral view, 1.5 times shorter than phallobasis, dorsal surface convex, ventral surface with a weakly downward-produced carina at middle.
|
||
</paragraph>
|
||
<caption id="13906600FFACFF96FF5574BDFD662D10" ID-DOI="http://doi.org/10.5281/zenodo.6758321" ID-Zenodo-Dep="6758321" httpUri="https://zenodo.org/record/6758321/files/figure.png" pageId="6" pageNumber="196" startId="6.[160,225,1740,1763]" targetBox="[178,1132,148,1706]" targetPageId="6">
|
||
<paragraph id="47503688FFACFF96FF5574BDFD662D10" blockId="6.[160,1157,1740,1792]" pageId="6" pageNumber="196">
|
||
<emphasis id="759BEA9AFFACFF96FF5574BDFF792CF3" bold="true" box="[160,252,1740,1763]" pageId="6" pageNumber="196">Figure 3.</emphasis>
|
||
<taxonomicName id="80EF4D0BFFACFF96FEF074BDFE532CF3" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[261,470,1740,1763]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="196" phylum="Arthropoda" rank="species" species="endroedii" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFACFF96FEF074BDFE532CF3" box="[261,470,1740,1763]" italics="true" pageId="6" pageNumber="196">Gibboryctes endroedii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFACFF96FE2B74BDFDAB2CF3" box="[478,558,1740,1763]" pageId="6" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(a,b) Habitus of holotype male in dorsal and lateral views, respectively; (c) habitus of female paratype in dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="13906600FFADFF97FF5574A2FCF22D17" ID-DOI="http://doi.org/10.5281/zenodo.6758325" ID-Zenodo-Dep="6758325" httpUri="https://zenodo.org/record/6758325/files/figure.png" pageId="7" pageNumber="197" startId="7.[160,225,1747,1770]" targetBox="[171,1108,143,1711]" targetPageId="7">
|
||
<paragraph id="47503688FFADFF97FF5574A2FCF22D17" blockId="7.[160,1157,1747,1799]" pageId="7" pageNumber="197">
|
||
<emphasis id="759BEA9AFFADFF97FF5574A2FF7A2CFA" bold="true" box="[160,255,1747,1770]" pageId="7" pageNumber="197">Figure 4.</emphasis>
|
||
<taxonomicName id="80EF4D0BFFADFF97FEFF74A2FD102CFA" authority="Prell, 1912" authorityName="Prell" authorityYear="1912" box="[266,661,1747,1770]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="197" phylum="Arthropoda" rank="species" species="gracilicornis">
|
||
<emphasis id="759BEA9AFFADFF97FEFF74A2FD9A2CFA" box="[266,543,1747,1770]" italics="true" pageId="7" pageNumber="197">Heterogomphus gracilicornis</emphasis>
|
||
<bibRefCitation id="237E4B79FFADFF97FDDF74A2FD102CFA" author="Prell H" box="[554,661,1747,1770]" pageId="7" pageNumber="197" pagination="93 - 176" refId="ref13871" refString="Prell H. 1912. Revision des Dynastinen-Genus Heterogomphus Burm. Mem Soc Entomol Belg. 20: 93 - 176." type="journal article" year="1912">Prell, 1912</bibRefCitation>
|
||
</taxonomicName>
|
||
. (a,b) Male habitus in dorsal and lateral views, respectively; (c,d) female habitus in dorsal and lateral views, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="13906600FFA2FF98FF5574E4FCB22CDA" ID-DOI="http://doi.org/10.5281/zenodo.6758329" ID-Zenodo-Dep="6758329" httpUri="https://zenodo.org/record/6758329/files/figure.png" pageId="8" pageNumber="198" startId="8.[160,225,1685,1708]" targetBox="[221,1093,152,1648]" targetPageId="8">
|
||
<paragraph id="47503688FFA2FF98FF5574E4FCB22CDA" blockId="8.[160,1155,1685,1738]" pageId="8" pageNumber="198">
|
||
<emphasis id="759BEA9AFFA2FF98FF5574E4FF7E2CBC" bold="true" box="[160,251,1685,1708]" pageId="8" pageNumber="198">Figure 5.</emphasis>
|
||
<taxonomicName id="80EF4D0BFFA2FF98FEF774E4FDE02CBC" authority="(Endrodi, 1977)" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" baseAuthorityName="Endrodi" baseAuthorityYear="1977" box="[258,613,1685,1708]" class="Insecta" family="Scarabaeidae" genus="Tetragus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="198" phylum="Arthropoda" rank="species" species="waldenfelsi">
|
||
<emphasis id="759BEA9AFFA2FF98FEF774E4FE422CBC" box="[258,455,1685,1708]" italics="true" pageId="8" pageNumber="198">Tetragus waldenfelsi</emphasis>
|
||
(
|
||
<bibRefCitation id="237E4B79FFA2FF98FE2674E4FDD82CBC" author="Endrodi S" box="[467,605,1685,1708]" pageId="8" pageNumber="198" pagination="335 - 336" refId="ref13398" refString="Endrodi S. 1977. Strategus waldenfelsi sp. n. (Coleoptera, Dynastinae). Reichenbachia. 16: 335 - 336." type="journal article" year="1977">Endrödi, 1977</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. (a,b) Male habitus in dorsal and lateral views, respectively; (c,d) female habitus in dorsal and lateral views, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="13906600FFA3FF99FF557633FE842E84" ID-DOI="http://doi.org/10.5281/zenodo.6758331" ID-Zenodo-Dep="6758331" httpUri="https://zenodo.org/record/6758331/files/figure.png" pageId="9" pageNumber="199" startId="9.[160,225,1090,1113]" targetBox="[166,1145,146,1050]" targetPageId="9">
|
||
<paragraph id="47503688FFA3FF99FF557633FE842E84" blockId="9.[160,1156,1090,1172]" pageId="9" pageNumber="199">
|
||
<emphasis id="759BEA9AFFA3FF99FF557633FF792E49" bold="true" box="[160,252,1090,1113]" pageId="9" pageNumber="199">Figure 6.</emphasis>
|
||
Labrum in dorsal view. (a)
|
||
<taxonomicName id="80EF4D0BFFA3FF99FDE77633FCAA2E49" authority="Endrodi" authorityName="Endrodi" authorityYear="1974" box="[530,815,1090,1113]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="199" phylum="Arthropoda" rank="species" species="szelenyii">
|
||
<emphasis id="759BEA9AFFA3FF99FDE77633FD5D2E49" box="[530,728,1090,1113]" italics="true" pageId="9" pageNumber="199">Gibboryctes szelenyii</emphasis>
|
||
Endrödi
|
||
</taxonomicName>
|
||
; (b)
|
||
<taxonomicName id="80EF4D0BFFA3FF99FCA87633FBAD2E49" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[861,1064,1090,1113]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="199" phylum="Arthropoda" rank="species" species="ebeninus" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFA3FF99FCA87633FBAD2E49" box="[861,1064,1090,1113]" italics="true" pageId="9" pageNumber="199">Gibboryctes ebeninus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFA3FF99FBC57633FB042E49" box="[1072,1153,1090,1113]" pageId="9" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (c)
|
||
<taxonomicName id="80EF4D0BFFA3FF99FF31762EFE1E2E66" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[196,411,1119,1142]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="199" phylum="Arthropoda" rank="species" species="endroedii" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFA3FF99FF31762EFE1E2E66" box="[196,411,1119,1142]" italics="true" pageId="9" pageNumber="199">Gibboryctes endroedii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFA3FF99FE53762EFE792E66" box="[422,508,1119,1142]" pageId="9" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (d)
|
||
<taxonomicName id="80EF4D0BFFA3FF99FDC5762EFC012E66" authority="Prell" authorityName="Prell" box="[560,900,1119,1142]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="199" phylum="Arthropoda" rank="species" species="gracilicornis">
|
||
<emphasis id="759BEA9AFFA3FF99FDC5762EFCCF2E66" box="[560,842,1119,1142]" italics="true" pageId="9" pageNumber="199">Heterogomphus gracilicornis</emphasis>
|
||
Prell
|
||
</taxonomicName>
|
||
; (e)
|
||
<taxonomicName id="80EF4D0BFFA3FF99FC4D762EFF782E84" authority="(Endrodi)" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" baseAuthorityName="Endrodi" baseAuthorityYear="1977" class="Insecta" family="Scarabaeidae" genus="Tetragus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="199" phylum="Arthropoda" rank="species" species="waldenfelsi">
|
||
<emphasis id="759BEA9AFFA3FF99FC4D762EFB012E66" box="[952,1156,1119,1142]" italics="true" pageId="9" pageNumber="199">Tetragus waldenfelsi</emphasis>
|
||
(Endrödi)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="13906600FFA0FF9AFF5576F5FC0E2EC6" ID-DOI="http://doi.org/10.5281/zenodo.6758333" ID-Zenodo-Dep="6758333" httpUri="https://zenodo.org/record/6758333/files/figure.png" pageId="10" pageNumber="200" startId="10.[160,225,1156,1179]" targetBox="[169,1149,148,1090]" targetPageId="10">
|
||
<paragraph id="47503688FFA0FF9AFF5576F5FC0E2EC6" blockId="10.[160,1157,1156,1238]" pageId="10" pageNumber="200">
|
||
<emphasis id="759BEA9AFFA0FF9AFF5576F5FF7F2E8B" bold="true" box="[160,250,1156,1179]" pageId="10" pageNumber="200">Figure 7.</emphasis>
|
||
Right mandible in dorsal view. (a)
|
||
<taxonomicName id="80EF4D0BFFA0FF9AFDA676F5FCE92E8B" authority="Endrodi" authorityName="Endrodi" authorityYear="1974" box="[595,876,1156,1179]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="200" phylum="Arthropoda" rank="species" species="szelenyii">
|
||
<emphasis id="759BEA9AFFA0FF9AFDA676F5FC922E8B" box="[595,791,1156,1179]" italics="true" pageId="10" pageNumber="200">Gibboryctes szelenyii</emphasis>
|
||
Endrödi
|
||
</taxonomicName>
|
||
; (b)
|
||
<taxonomicName id="80EF4D0BFFA0FF9AFC6276F5FBE42E8B" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[919,1121,1156,1179]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="200" phylum="Arthropoda" rank="species" species="ebeninus" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFA0FF9AFC6276F5FBE42E8B" box="[919,1121,1156,1179]" italics="true" pageId="10" pageNumber="200">Gibboryctes ebeninus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFA0FF9AFB9D76F5FF482EA9" pageId="10" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (c)
|
||
<taxonomicName id="80EF4D0BFFA0FF9AFF0276D3FE422EA9" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[247,455,1186,1209]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="200" phylum="Arthropoda" rank="species" species="endroedii" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFA0FF9AFF0276D3FE422EA9" box="[247,455,1186,1209]" italics="true" pageId="10" pageNumber="200">Gibboryctes endroedii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFA0FF9AFE3B76D3FD9A2EA9" box="[462,543,1186,1209]" pageId="10" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (d)
|
||
<taxonomicName id="80EF4D0BFFA0FF9AFDB976D3FC162EA9" authority="Prell" authorityName="Prell" box="[588,915,1186,1209]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="200" phylum="Arthropoda" rank="species" species="gracilicornis">
|
||
<emphasis id="759BEA9AFFA0FF9AFDB976D3FCD82EA9" box="[588,861,1186,1209]" italics="true" pageId="10" pageNumber="200">Heterogomphus gracilicornis</emphasis>
|
||
Prell
|
||
</taxonomicName>
|
||
; (e)
|
||
<taxonomicName id="80EF4D0BFFA0FF9AFC4A76D3FF782EC6" authority="(Endrodi)" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" baseAuthorityName="Endrodi" baseAuthorityYear="1977" class="Insecta" family="Scarabaeidae" genus="Tetragus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="200" phylum="Arthropoda" rank="species" species="waldenfelsi">
|
||
<emphasis id="759BEA9AFFA0FF9AFC4A76D3FB012EA9" box="[959,1156,1186,1209]" italics="true" pageId="10" pageNumber="200">Tetragus waldenfelsi</emphasis>
|
||
(Endrödi)
|
||
</taxonomicName>
|
||
. Red arrows indicate the apical tooth in (a–e), striate molar in (d).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="13906600FFA1FF9BFF5576F8FD052F05" ID-DOI="http://doi.org/10.5281/zenodo.6758335" ID-Zenodo-Dep="6758335" httpUri="https://zenodo.org/record/6758335/files/figure.png" pageId="11" pageNumber="201" startId="11.[160,225,1161,1184]" targetBox="[167,1127,143,1126]" targetPageId="11">
|
||
<paragraph id="47503688FFA1FF9BFF5576F8FD052F05" blockId="11.[160,1156,1161,1301]" pageId="11" pageNumber="201">
|
||
<emphasis id="759BEA9AFFA1FF9BFF5576F8FF7C2EB0" bold="true" box="[160,249,1161,1184]" pageId="11" pageNumber="201">Figure 8.</emphasis>
|
||
Right maxilla in ventral view. (a)
|
||
<taxonomicName id="80EF4D0BFFA1FF9BFDC176F8FCC12EB0" authority="Endrodi" authorityName="Endrodi" authorityYear="1974" box="[564,836,1161,1184]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="201" phylum="Arthropoda" rank="species" species="szelenyii">
|
||
<emphasis id="759BEA9AFFA1FF9BFDC176F8FD772EB0" box="[564,754,1161,1184]" italics="true" pageId="11" pageNumber="201">Gibboryctes szelenyii</emphasis>
|
||
Endrödi
|
||
</taxonomicName>
|
||
; (b)
|
||
<taxonomicName id="80EF4D0BFFA1FF9BFC9E76F8FBB52EB0" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[875,1072,1161,1184]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="201" phylum="Arthropoda" rank="species" species="ebeninus" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFA1FF9BFC9E76F8FBB52EB0" box="[875,1072,1161,1184]" italics="true" pageId="11" pageNumber="201">Gibboryctes ebeninus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFA1FF9BFBC076F8FB042EB0" box="[1077,1153,1161,1184]" pageId="11" pageNumber="201" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (c)
|
||
<taxonomicName id="80EF4D0BFFA1FF9BFF4876D7FE032EAD" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[189,390,1190,1213]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="201" phylum="Arthropoda" rank="species" species="endroedii" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFA1FF9BFF4876D7FE032EAD" box="[189,390,1190,1213]" italics="true" pageId="11" pageNumber="201">Gibboryctes endroedii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFA1FF9BFE7E76D7FE5D2EAD" box="[395,472,1190,1213]" pageId="11" pageNumber="201" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (d)
|
||
<taxonomicName id="80EF4D0BFFA1FF9BFDF576D7FCBE2EAD" authority="Prell" authorityName="Prell" box="[512,827,1190,1213]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="201" phylum="Arthropoda" rank="species" species="gracilicornis">
|
||
<emphasis id="759BEA9AFFA1FF9BFDF576D7FC8C2EAD" box="[512,777,1190,1213]" italics="true" pageId="11" pageNumber="201">Heterogomphus gracilicornis</emphasis>
|
||
Prell
|
||
</taxonomicName>
|
||
; (e)
|
||
<taxonomicName id="80EF4D0BFFA1FF9BFC9776D7FB052EAD" authority="(Endrodi)" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" baseAuthorityName="Endrodi" baseAuthorityYear="1977" box="[866,1152,1190,1213]" class="Insecta" family="Scarabaeidae" genus="Tetragus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="201" phylum="Arthropoda" rank="species" species="waldenfelsi">
|
||
<emphasis id="759BEA9AFFA1FF9BFC9776D7FBA52EAD" box="[866,1056,1190,1213]" italics="true" pageId="11" pageNumber="201">Tetragus waldenfelsi</emphasis>
|
||
(Endrödi)
|
||
</taxonomicName>
|
||
. Red arrows indicating triangular galea in (a), elongate maxillary palpomere II in (b), lobed inner margin of galea and stipe with triangular lobe in (c), pointed and triangular teeth on galea in (d), scythe-shape d teeth and rounded inner margin of galea in (e).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="13906600FFA6FF9CFF55760BFEF72EDB" ID-DOI="http://doi.org/10.5281/zenodo.6758337" ID-Zenodo-Dep="6758337" httpUri="https://zenodo.org/record/6758337/files/figure.png" pageId="12" pageNumber="202" startId="12.[160,225,1146,1169]" targetBox="[184,1146,142,1069]" targetPageId="12">
|
||
<paragraph id="47503688FFA6FF9CFF55760BFEF72EDB" blockId="12.[160,1156,1146,1227]" pageId="12" pageNumber="202">
|
||
<emphasis id="759BEA9AFFA6FF9CFF55760BFF782E81" bold="true" box="[160,253,1146,1169]" pageId="12" pageNumber="202">Figure 9.</emphasis>
|
||
Detail of head, pronotum and prothoracic legs of males in dorsal view: (a)
|
||
<taxonomicName id="80EF4D0BFFA6FF9CFBE0760BFED72EBE" authority="(Endrodi)" baseAuthorityName="Endrodi" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="202" phylum="Arthropoda" rank="species" species="szelenyii">
|
||
<emphasis id="759BEA9AFFA6FF9CFBE0760BFF6A2EBE" italics="true" pageId="12" pageNumber="202">Gibboryctes szelenyii</emphasis>
|
||
(Endrödi)
|
||
</taxonomicName>
|
||
; (b)
|
||
<taxonomicName id="80EF4D0BFFA6FF9CFE8E76E6FDCC2EBE" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[379,585,1175,1198]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="202" phylum="Arthropoda" rank="species" species="endroedii" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFA6FF9CFE8E76E6FDCC2EBE" box="[379,585,1175,1198]" italics="true" pageId="12" pageNumber="202">Gibboryctes endroedii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFA6FF9CFDBA76E6FD1B2EBE" box="[591,670,1175,1198]" pageId="12" pageNumber="202" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; (c)
|
||
<taxonomicName id="80EF4D0BFFA6FF9CFD3176E6FB8D2EBE" authority="Prell" authorityName="Prell" box="[708,1032,1175,1198]" class="Insecta" family="Dynastidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="202" phylum="Arthropoda" rank="species" species="gracilicornis">
|
||
<emphasis id="759BEA9AFFA6FF9CFD3176E6FC512EBE" box="[708,980,1175,1198]" italics="true" pageId="12" pageNumber="202">Heterogomphus gracilicornis</emphasis>
|
||
Prell
|
||
</taxonomicName>
|
||
; (d)
|
||
<taxonomicName id="80EF4D0BFFA6FF9CFBC476E6FEEA2EDB" authority="(Endrodi)" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" baseAuthorityName="Endrodi" baseAuthorityYear="1977" class="Insecta" family="Scarabaeidae" genus="Tetragus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="202" phylum="Arthropoda" rank="species" species="waldenfelsi">
|
||
<emphasis id="759BEA9AFFA6FF9CFBC476E6FE8E2EDB" italics="true" pageId="12" pageNumber="202">Tetragus waldenfelsi</emphasis>
|
||
(Endrödi)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="47503688FFA6FF9CFF557684FE012F1F" blockId="12.[160,1155,1269,1399]" box="[160,388,1269,1295]" pageId="12" pageNumber="202">
|
||
<emphasis id="759BEA9AFFA6FF9CFF557684FE012F1F" bold="true" box="[160,388,1269,1295]" italics="true" pageId="12" pageNumber="202">Sexual dimorphism</emphasis>
|
||
</paragraph>
|
||
<paragraph id="47503688FFA6FF9CFF557768FCB42F67" blockId="12.[160,1155,1269,1399]" pageId="12" pageNumber="202">
|
||
The two sexes are largely identical, but the female can be distinguished in the following aspects:
|
||
<emphasis id="759BEA9AFFA6FF9CFF0B774DFEB92F45" bold="true" box="[254,316,1340,1365]" pageId="12" pageNumber="202">Head</emphasis>
|
||
: Anterior clypeal margin rounded.
|
||
<emphasis id="759BEA9AFFA6FF9CFD48774DFCB72F45" bold="true" box="[701,818,1340,1365]" pageId="12" pageNumber="202">Abdomen</emphasis>
|
||
: Tergite VIII flattened in lateral view; ventrite VI parabolic, lacking posterior emargination.
|
||
</paragraph>
|
||
<paragraph id="47503688FFA6FF9CFF5577C2FE172FDD" blockId="12.[160,1155,1459,1554]" box="[160,402,1459,1485]" pageId="12" pageNumber="202">
|
||
<emphasis id="759BEA9AFFA6FF9CFF5577C2FE172FDD" bold="true" box="[160,402,1459,1485]" italics="true" pageId="12" pageNumber="202">Specific composition</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FF56503FFA6FF9CFF5577A7FE6F2C02" pageId="12" pageNumber="202" type="discussion">
|
||
<paragraph id="47503688FFA6FF9CFF5577A7FE6F2C02" blockId="12.[160,1155,1459,1554]" pageId="12" pageNumber="202">
|
||
<taxonomicName id="80EF4D0BFFA6FF9CFF5577A7FEA72FFF" box="[160,290,1494,1519]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="759BEA9AFFA6FF9CFF5577A7FEA72FFF" box="[160,290,1494,1519]" italics="true" pageId="12" pageNumber="202">Gibboryctes</emphasis>
|
||
</taxonomicName>
|
||
comprises three species:
|
||
<taxonomicName id="80EF4D0BFFA6FF9CFDAB77A7FD5B2FFF" box="[606,734,1494,1519]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="202" phylum="Arthropoda" rank="species" species="szelenyii">
|
||
<emphasis id="759BEA9AFFA6FF9CFDAB77A7FD5B2FFF" box="[606,734,1494,1519]" italics="true" pageId="12" pageNumber="202">G. szelenyii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="80EF4D0BFFA6FF9CFD0577A7FC672FFF" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[752,994,1494,1519]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="202" phylum="Arthropoda" rank="species" species="ebeninus" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFA6FF9CFD0577A7FC672FFF" box="[752,994,1494,1519]" italics="true" pageId="12" pageNumber="202">Gibboryctes ebeninus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFA6FF9CFC1B77A7FBC82FFF" box="[1006,1101,1494,1519]" pageId="12" pageNumber="202" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="80EF4D0BFFA6FF9CFF557788FE0E2C02" authority="Costa & Duarte & Iannuzzi & Grossi, 2022" authorityName="Costa & Duarte & Iannuzzi & Grossi" authorityYear="2022" box="[160,395,1529,1554]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="202" phylum="Arthropoda" rank="species" species="endroedii" status="sp. nov.">
|
||
<emphasis id="759BEA9AFFA6FF9CFF557788FE0E2C02" box="[160,395,1529,1554]" italics="true" pageId="12" pageNumber="202">Gibboryctes endroedii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EA857E1FFA6FF9CFE677788FE6F2C02" box="[402,490,1529,1554]" pageId="12" pageNumber="202" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |