171 lines
19 KiB
XML
171 lines
19 KiB
XML
<document id="13AD75F8FA7F689C477539927A5EC6A3" ID-DOI="10.5281/zenodo.186920" ID-GBIF-Dataset="695e995e-3a9a-41b8-bafa-ed8366b37ef4" ID-ISSN="1175-5326" ID-Zenodo-Dep="186920" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461059560947" checkinUser="plazi" docAuthor="Breure, Abraham S. H." docDate="2009" docId="CF1A2655FFD6BD67E9A6FC871C69F9FE" docLanguage="en" docName="zt02065p050.pdf" docOrigin="Zootaxa 2065" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Drymaeus (Drymaeus) extraneus Haas 1955" docType="treatment" docVersion="8" lastPageNumber="40" masterDocId="33235E2DFFD9BD68E931FFE61949FF9C" masterDocTitle="New Orthalicidae (Mollusca, Gastropoda) from Venezuelan Guayana: unravelling secrets from the Lost World" masterLastPageNumber="50" masterPageNumber="25" pageNumber="40" updateTime="1698581906488" updateUser="plazi">
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<mods:mods id="D9579F66FD443A45CE11D3CF809DB3F2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="906474EFC9A310B1CBEDE62F8B445E7D">
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<mods:title id="1A970531190F7F35CFE87CCA00D9158A">New Orthalicidae (Mollusca, Gastropoda) from Venezuelan Guayana: unravelling secrets from the Lost World</mods:title>
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</mods:titleInfo>
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<mods:name id="01D9F20EE825DDB4F4718D8DDE4C90DB" type="personal">
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<mods:roleTerm id="F7E71C93C8A8038B88999E2064CAFC5A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CD5C1CFA768590A6B2FDF09463142E2D">Breure, Abraham S. H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="265917A12AEA81F705E4560A6B669E93">text</mods:typeOfResource>
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<mods:title id="BD56A5322615ABDDBE483E25C9444016">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="95E7580C1ED4DF125CAD80DAB0C74A67">
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<mods:date id="B51C6645E60A1CEF71036CCFB7702F7E">2009</mods:date>
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<mods:detail id="A555A699D5F81768FF0CD5AE6C9A6467" type="volume">
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<mods:number id="9E7C0958F4AB91DD96907DCAD0DD1E0F">2065</mods:number>
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</mods:detail>
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<mods:start id="E34A076C487B887474114A4391A5433A">25</mods:start>
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<mods:end id="0DD488D63D8AC47A981EE4B6495A9300">50</mods:end>
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<mods:classification id="4F4CF926CEA6B54AE237847057D2ACF3">journal article</mods:classification>
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<mods:identifier id="14E5371CDFA5C81BDCD54F1B346AFF0B" type="DOI">10.5281/zenodo.186920</mods:identifier>
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<mods:identifier id="1050EC0E1F3471387B1C87B58F64DD72" type="GBIF-Dataset">695e995e-3a9a-41b8-bafa-ed8366b37ef4</mods:identifier>
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<mods:identifier id="42FABCC1FE14D9243D07F4B6B654CAED" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="1D30C5B0F35AC15F2A47B0E42DAFA7C4" type="Zenodo-Dep">186920</mods:identifier>
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<treatment id="CF1A2655FFD6BD67E9A6FC871C69F9FE" ID-DOI="http://doi.org/10.5281/zenodo.5618377" ID-GBIF-Taxon="119613775" ID-Zenodo-Dep="5618377" LSID="urn:lsid:plazi:treatment:CF1A2655FFD6BD67E9A6FC871C69F9FE" httpUri="http://treatment.plazi.org/id/CF1A2655FFD6BD67E9A6FC871C69F9FE" lastPageNumber="40" pageId="15" pageNumber="40">
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<subSubSection id="0FA9C4C8FFD6BD67E9A6FC871865FC02" pageId="15" pageNumber="40" type="nomenclature">
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<paragraph id="470C9743FFD6BD67E9A6FC871B82FCE6" blockId="15.[151,715,864,926]" box="[151,715,864,891]" pageId="15" pageNumber="40">
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<heading id="1C44202FFFD6BD67E9A6FC871B82FCE6" bold="true" box="[151,715,864,891]" fontSize="11" level="1" pageId="15" pageNumber="40" reason="1">
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<taxonomicName id="80B3ECC0FFD6BD67E9A6FC871B82FCE6" authority="Haas, 1955" authorityName="Haas" authorityYear="1955" box="[151,715,864,891]" class="Gastropoda" family="Orthalicidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="40" phylum="Mollusca" rank="species" species="extraneus" subGenus="Drymaeus">
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<emphasis id="75C74B51FFD6BD67E9A6FC871B82FCE6" bold="true" box="[151,715,864,891]" pageId="15" pageNumber="40">
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<emphasis id="75C74B51FFD6BD67E9A6FC871B6EFCE7" bold="true" box="[151,551,865,891]" italics="true" pageId="15" pageNumber="40">Drymaeus (Drymaeus) extraneus</emphasis>
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(
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<bibRefCitation id="2322EAB2FFD6BD67EB06FC861B8BFCE6" author="Haas" box="[567,706,864,890]" pageId="15" pageNumber="49" refString="Haas, F. (1955) On some small collections of inland shells from South America. Fieldiana, Zoology, 34, 361 ─ 387." type="journal article" year="1955">Haas, 1955</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="470C9743FFD6BD67E9A6FC621865FC02" blockId="15.[151,715,864,926]" box="[151,300,900,926]" pageId="15" pageNumber="40">
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(
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<figureCitation id="DF888BC6FFD6BD67E9AEFC6219AAFC02" box="[159,227,900,926]" captionStart="FIGURE 8" captionStartId="16.[151,255,1462,1486]" captionTargetBox="[151,1434,274,1435]" captionTargetId="figure@16.[151,1435,274,1437]" captionTargetPageId="16" captionText="FIGURE 8. Shells in Drymaeus (Drymaeus). A ─ C. D. (D.) extraneus (Haas, 1955). A ─ B. Holotype FMNH 49736; C. FMNH 49734. D ─ F. D. (D.) steyermarki (Haas, 1955), holotype FMNH 49735. G ─ H. D. (D.) yapacanensis Breure & Eskens, 1981, holotype RMNH 55331. I ─ S. Drymaeus (Drymaeus) rex spec. nov. I ─ M, Q ─ S. Paratypes UF 284745, 284730, 284759, 284748, 284714, 284719, 284719 and 284715 respectively; N ─ P. Holotype UF 284726. Scale bar— 1 cm." httpUri="https://zenodo.org/record/186928/files/figure.png" pageId="15" pageNumber="40">Fig. 8</figureCitation>
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A─C)
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</paragraph>
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</subSubSection>
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<subSubSection id="0FA9C4C8FFD6BD67E9A6FC371C2FFBD4" pageId="15" pageNumber="40" type="reference_group">
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<paragraph id="470C9743FFD6BD67E9A6FC371D6AFB94" blockId="15.[151,1436,977,1096]" pageId="15" pageNumber="40">
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<treatmentCitationGroup id="67A3B06DFFD6BD67E9A6FC371D6AFB94" pageId="15" pageNumber="40">
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<taxonomicName id="80B3ECC0FFD6BD67E9A6FC371BFCFC74" ID-CoL="7QFPR" authority="Haas, 1955: 382" authorityName="Haas" authorityPageNumber="382" authorityYear="1955" box="[151,693,977,1000]" class="Gastropoda" family="Bulimulidae" genus="Bulimulus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="40" phylum="Mollusca" rank="species" species="extraneus" subGenus="Lissoacme">
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<emphasis id="75C74B51FFD6BD67E9A6FC3718B5FC74" box="[151,508,977,1000]" italics="true" pageId="15" pageNumber="40">Bulimulus (Lissoacme) extraneus</emphasis>
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<treatmentCitation id="C612B152FFD6BD67EB35FC371BFCFC74" author="Haas" box="[516,693,977,1000]" page="382" pageId="15" pageNumber="40" year="1955">
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<bibRefCitation id="2322EAB2FFD6BD67EB35FC371BC9FC74" author="Haas" box="[516,640,977,1000]" pageId="15" pageNumber="49" refString="Haas, F. (1955) On some small collections of inland shells from South America. Fieldiana, Zoology, 34, 361 ─ 387." type="journal article" year="1955">Haas, 1955</bibRefCitation>
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: 382
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</treatmentCitation>
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</taxonomicName>
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, fig. 82.
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<typeStatus id="980829E1FFD6BD67EA27FC371A02FC74" box="[790,843,977,1000]" pageId="15" pageNumber="40">Type</typeStatus>
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locality:
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<collectingCountry id="3FA4D7D3FFD6BD67EA80FC371D68FC74" box="[945,1057,977,1000]" name="Venezuela" pageId="15" pageNumber="40">Venezuela</collectingCountry>
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, State of Bolívar, northwest part of Chimantá massif, summit of Apacara tepui, ca.
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<quantity id="804B3AA6FFD6BD67EBF2FC171A5EFB94" box="[707,791,1009,1032]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="40" unit="m" value="2100.0">2100 m</quantity>
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;
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<typeStatus id="980829E1FFD6BD67EA10FC171A37FB94" box="[801,894,1009,1032]" pageId="15" pageNumber="40" type="holotype">holotype</typeStatus>
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FMNH 49736.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="470C9743FFD6BD67E9A6FBF71C2FFBD4" blockId="15.[151,1436,977,1096]" pageId="15" pageNumber="40">
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<treatmentCitationGroup id="67A3B06DFFD6BD67E9A6FBF71C2FFBD4" infered="true" pageId="15" pageNumber="40">
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<taxonomicName id="80B3ECC0FFD6BD67E9A6FBF71BD6FBB4" authority="Haas, 1955: 383" authorityName="Haas" authorityPageNumber="383" authorityYear="1955" box="[151,671,1041,1064]" class="Gastropoda" family="Orthalicidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="40" phylum="Mollusca" rank="species" species="griffini" subGenus="Drymaeus">
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<emphasis id="75C74B51FFD6BD67E9A6FBF718A9FBB4" box="[151,480,1041,1064]" italics="true" pageId="15" pageNumber="40">Drymaeus (Drymaeus) griffini</emphasis>
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<treatmentCitation id="C612B152FFD6BD67E8DBFBF71BD6FBB4" author="Haas" box="[490,671,1041,1064]" page="383" pageId="15" pageNumber="40" year="1955">
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<bibRefCitation id="2322EAB2FFD6BD67E8DBFBF71B21FBB4" author="Haas" box="[490,616,1041,1064]" pageId="15" pageNumber="49" refString="Haas, F. (1955) On some small collections of inland shells from South America. Fieldiana, Zoology, 34, 361 ─ 387." type="journal article" year="1955">Haas, 1955</bibRefCitation>
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: 383
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</treatmentCitation>
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</taxonomicName>
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, fig. 83.
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<typeStatus id="980829E1FFD6BD67EA34FBF71A73FBB4" box="[773,826,1041,1064]" pageId="15" pageNumber="40">Type</typeStatus>
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locality:
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<collectingCountry id="3FA4D7D3FFD6BD67EA95FBF71D5DFBB4" box="[932,1044,1041,1064]" name="Venezuela" pageId="15" pageNumber="40">Venezuela</collectingCountry>
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, State of Bolívar, Chimantá massif, western side of Abacapa tepui, drainage of Río Abacapa, ca.
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<quantity id="804B3AA6FFD6BD67EA63FBD71AEFFBD4" box="[850,934,1073,1096]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="15" pageNumber="40" unit="m" value="1300.0">1300 m</quantity>
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;
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<typeStatus id="980829E1FFD6BD67EA81FBD71D44FBD4" box="[944,1037,1073,1096]" pageId="15" pageNumber="40" type="holotype">holotype</typeStatus>
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FMNH 49734. New synonymy.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="0FA9C4C8FFD6BD67E9A6FB911B71FAB0" pageId="15" pageNumber="40" type="materials_examined">
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<paragraph id="470C9743FFD6BD67E9A6FB911B71FAB0" blockId="15.[151,1436,1143,1634]" pageId="15" pageNumber="40">
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<emphasis id="75C74B51FFD6BD67E9A6FB911842FB0D" bold="true" box="[151,267,1143,1169]" pageId="15" pageNumber="40">Material.</emphasis>
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<collectingCountry id="3FA4D7D3FFD6BD67E825FB9E18D9FB0E" box="[276,400,1144,1170]" name="Venezuela" pageId="15" pageNumber="40">Venezuela</collectingCountry>
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, Estado Bolívar, northwest part of Chimantá massif, summit of Apacará-tepui,
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<quantity id="804B3AA6FFD6BD67EC0AFB9E1CDEFB0E" box="[1339,1431,1144,1170]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="40" unit="m" value="2100.0">2100 m</quantity>
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, J.A. Steyermark leg., 20─
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<date id="330DB183FFD6BD67E8E0FB781B18FB24" box="[465,593,1182,1208]" pageId="15" pageNumber="40" value="1953-06-22">22.vi.1953</date>
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(
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<collectionCode id="21A20F86FFD6BD67EB53FB781BF3FB24" LSID="urn:lsid:biocol.org:col:34795" box="[610,698,1182,1208]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="15" pageNumber="40">FMNH</collectionCode>
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49736/
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<typeStatus id="980829E1FFD6BD67EA28FB781ACDFB24" box="[793,900,1182,1208]" pageId="15" pageNumber="40" type="holotype">holotype</typeStatus>
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); Ibidem, along trail between Río Tirica and lower summit camp,
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<quantity id="804B3AA6FFD6BD67E8BCFB2318AEFB43" box="[397,487,1221,1247]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="15" pageNumber="40" unit="m" value="1800.0">1800 m</quantity>
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, J.A. Steyermark & J.J. Wurdack leg.,
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<date id="330DB183FFD6BD67EA9BFB231D68FB43" box="[938,1057,1221,1247]" pageId="15" pageNumber="40" value="1955-02-26">26.ii.1955</date>
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(
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<collectionCode id="21A20F86FFD6BD67ED1EFB231DCFFB43" LSID="urn:lsid:biocol.org:col:34795" box="[1071,1158,1221,1247]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="15" pageNumber="40">FMNH</collectionCode>
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52444/
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<specimenCount id="51B55CCAFFD6BD67EDD5FB231C56FB43" box="[1252,1311,1221,1247]" pageId="15" pageNumber="40" type="generic">1 juv.</specimenCount>
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<typeStatus id="980829E1FFD6BD67EC17FB231CD9FB43" box="[1318,1424,1221,1247]" pageId="15" pageNumber="40" type="paratype">paratype</typeStatus>
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); Idem, west side of Abacapá-tepui,
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<quantity id="804B3AA6FFD6BD67EB1FFB0A1BC0FA9A" box="[558,649,1260,1286]" metricMagnitude="3" metricUnit="m" metricValue="1.189" pageId="15" pageNumber="40" unit="m" value="1189.0">1189 m</quantity>
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, J.A. Steyermark & C. Griffin leg.,
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<date id="330DB183FFD6BD67ED1AFB0A1DEEFA9A" box="[1067,1191,1260,1286]" pageId="15" pageNumber="40" value="1953-04-13">13.iv.1953</date>
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(
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<collectionCode id="21A20F86FFD6BD67ED87FB0A1C44FA9A" LSID="urn:lsid:biocol.org:col:34795" box="[1206,1293,1260,1286]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="15" pageNumber="40">FMNH</collectionCode>
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49734/1). Shell dimensions are given
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<tableCitation id="0A31A2F8FFD6BD67E8EBFAF41B7BFAB0" box="[474,562,1298,1324]" captionStart="TABLE 1" captionStartId="8.[151,240,267,291]" captionTargetBox="[151,1436,422,659]" captionTargetPageId="8" captionText="TABLE 1. Dimensions in selected species of Plekocheilus and Drymaeus in mm. Abbreviations: D, shell diameter; H, shell height; HA, height of aperture; LW, height of last whorl; M, mean; max, maximum; min, minimum; n, number of specimens measured; s, standard deviation; W, number of whorls; WA, width of aperture. For depositories of material, see section on material and methods." httpUri="http://table.plazi.org/id/13CCC7CBFFD1BD60E9A6FEED1B69FE17" pageId="15" pageNumber="40" tableUuid="13CCC7CBFFD1BD60E9A6FEED1B69FE17">Table 1</tableCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="0FA9C4C8FFD6BD67E9F4FADE1C69F9FE" pageId="15" pageNumber="40" type="discussion">
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<paragraph id="470C9743FFD6BD67E9F4FADE1C69F9FE" blockId="15.[151,1436,1143,1634]" pageId="15" pageNumber="40">
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<emphasis id="75C74B51FFD6BD67E9F4FADE1877FACE" bold="true" box="[197,318,1336,1362]" pageId="15" pageNumber="40">Remarks.</emphasis>
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Upon restudying the
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<typeStatus id="980829E1FFD6BD67EB0BFADF1B24FACF" box="[570,621,1337,1363]" pageId="15" pageNumber="40">type</typeStatus>
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material, also in the context of the material from Cerro de la Neblina, I have come to the conclusion that this species belongs to what I would like to call the
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<taxonomicName id="80B3ECC0FFD6BD67EDD3FA861868FA3C" class="Gastropoda" family="Orthalicidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="40" phylum="Mollusca" rank="species" species="steyermarki" subGenus="Drymaeus">
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<emphasis id="75C74B51FFD6BD67EDD3FA861868FA3C" italics="true" pageId="15" pageNumber="40">Drymaeus (D.) steyermarki</emphasis>
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</taxonomicName>
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-group; see discussion below.
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<taxonomicName id="80B3ECC0FFD6BD67EBB3FA611A06FA3C" box="[642,847,1415,1440]" class="Gastropoda" family="Bulimulidae" genus="Bulimulus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="40" phylum="Mollusca" rank="species" species="extraneus" subGenus="Drymaeus">
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<emphasis id="75C74B51FFD6BD67EBB3FA611A06FA3C" box="[642,847,1415,1440]" italics="true" pageId="15" pageNumber="40">D. (D.) extraneus</emphasis>
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</taxonomicName>
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resembles in shell shape most closely
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<taxonomicName id="80B3ECC0FFD6BD67EC24FA611CDDFA3C" box="[1301,1428,1415,1440]" class="Gastropoda" family="Orthalicidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="40" phylum="Mollusca" rank="species" species="rex" subGenus="Drymaeus">
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<emphasis id="75C74B51FFD6BD67EC24FA611CDDFA3C" box="[1301,1428,1415,1440]" italics="true" pageId="15" pageNumber="40">D. (D.) rex</emphasis>
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</taxonomicName>
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, but it is smaller, with the aperture less broad. The
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<typeStatus id="980829E1FFD6BD67EBDDFA4B1A56FA5B" box="[748,799,1453,1479]" pageId="15" pageNumber="40">type</typeStatus>
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specimens are unicoloured.
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<taxonomicName id="80B3ECC0FFD6BD67ED43FA4B1DA1FA5A" box="[1138,1256,1453,1478]" class="Gastropoda" family="Orthalicidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="40" phylum="Mollusca" rank="species" species="griffini">
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<emphasis id="75C74B51FFD6BD67ED43FA4B1DA1FA5A" box="[1138,1256,1453,1478]" italics="true" pageId="15" pageNumber="40">D. griffini</emphasis>
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</taxonomicName>
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—occurring on the same massif—proves to be synonymous; the name
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<taxonomicName id="80B3ECC0FFD6BD67EA2EFA321AD8FA71" box="[799,913,1492,1517]" class="Gastropoda" family="Bulimulidae" genus="Bulimulus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="40" phylum="Mollusca" rank="species" species="extraneus">
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<emphasis id="75C74B51FFD6BD67EA2EFA321AD8FA71" box="[799,913,1492,1517]" italics="true" pageId="15" pageNumber="40">extraneus</emphasis>
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</taxonomicName>
|
|
has page priority. The
|
|
<typeStatus id="980829E1FFD6BD67ED90FA321C41FA72" box="[1185,1288,1492,1518]" pageId="15" pageNumber="40" type="holotype">holotype</typeStatus>
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|
of
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|
<taxonomicName id="80B3ECC0FFD6BD67EC1EFA321CC9FA71" box="[1327,1408,1492,1517]" class="Gastropoda" family="Orthalicidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="40" phylum="Mollusca" rank="species" species="griffini">
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<emphasis id="75C74B51FFD6BD67EC1EFA321CC9FA71" box="[1327,1408,1492,1517]" italics="true" pageId="15" pageNumber="40">griffini</emphasis>
|
|
</taxonomicName>
|
|
is immature, but shows that also a banded colour pattern is possible in this taxon. The original label of
|
|
<taxonomicName id="80B3ECC0FFD6BD67EC16FA1D1CD2F988" box="[1319,1435,1531,1556]" class="Gastropoda" family="Orthalicidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="40" phylum="Mollusca" rank="species" species="griffini">
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|
<emphasis id="75C74B51FFD6BD67EC16FA1D1CD2F988" box="[1319,1435,1531,1556]" italics="true" pageId="15" pageNumber="40">D. griffini</emphasis>
|
|
</taxonomicName>
|
|
reads “
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|
<collectingCountry id="3FA4D7D3FFD6BD67E9D7F9C7182BF9A7" box="[230,354,1569,1595]" name="Venezuela" pageId="15" pageNumber="40">Venezuela</collectingCountry>
|
|
Botanical Expedition, 1953, Camp 33900, drainage of Rio Abacapá, western side Abacapátepui”. I have been unable to obtain more information from field notes made during the expedition.
|
|
</paragraph>
|
|
</subSubSection>
|
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</treatment>
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</document> |