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<mods:title id="1DD06710BD5BD5FEBAB250B845546F6F">Revision Of The Asian Catfish Genus Hemibagrus Bleeker, 1862 (Teleostei: Siluriformes: Bagridae)</mods:title>
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<emphasis id="75C56466FFC8FF96FF44F56BFE2E8197" bold="true" box="[170,430,637,662]" italics="true" pageId="33" pageNumber="238">Hemibagrus filamentus</emphasis>
(Fang &amp;
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)
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(
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)
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<taxonomicName id="80B1C3F7FFC8FF96FF9FF5CBFE4E81F3" authority="Fang &amp; Chaux" authorityName="Fang &amp; Chaux" authorityYear="1949" box="[113,462,732,754]" class="Insecta" family="Cerambycidae" genus="Macrones" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="238" phylum="Arthropoda" rank="species" species="filamentus">
<emphasis id="75C56466FFC8FF96FF9FF5CBFEB981F0" box="[113,313,733,753]" italics="true" pageId="33" pageNumber="238">Macrones filamentus</emphasis>
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, in
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&amp;
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, 1949: 200,
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(
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locality:
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<emphasis id="75C56466FFC8FF96FF9FF401FEA2802A" box="[113,290,791,811]" italics="true" pageId="33" pageNumber="238">Mystus filamentus</emphasis>
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-
<treatmentCitation id="C6109E65FFC8FF96FED3F400FD97802A" author="Lim, P &amp; Lek, S" box="[317,535,790,812]" page="382" pageId="33" pageNumber="238" year="1996">
<bibRefCitation id="2320C585FFC8FF96FED3F400FD97802A" author="Lim, P &amp; Lek, S" box="[317,535,790,812]" pageId="33" pageNumber="238" pagination="379 - 386" refId="ref72604" refString="Lim, P., S. Lek, S. T. Touch, S. - O. Mao &amp; B. Chhouk, 1996. Diversity and spatial distribution of freshwater fish in Great Lake and Tonle Sap river (Cambodia, Southeast Asia). Aquatic Living Resources, 12: 379 - 386." type="journal article" year="1996">Lim et al., 1996: 382</bibRefCitation>
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;
<treatmentCitation id="C6109E65FFC8FF96FDCAF400FD6C802A" author="Rainboth, W" box="[548,748,790,812]" page="144" pageId="33" pageNumber="238" year="1996">
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, Pl. 19 Fig. 147.
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<treatmentCitationGroup id="67A19F5AFFC8FF96FF9FF446FE2F80BA" pageId="33" pageNumber="238">
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<emphasis id="75C56466FFC8FF96FF9FF446FEE28065" box="[113,354,848,868]" italics="true" pageId="33" pageNumber="238">Hemibagrus filamentus</emphasis>
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- Ng &amp; Rainboth, 1999: 555,
<figureCitation id="DF8AA4F1FFC8FF96FD45F459FD6A8064" box="[683,746,847,869]" captionStart="Fig" captionStartId="2.[137,172,1975,1997]" captionTargetBox="[136,1471,759,1944]" captionTargetId="figure-78@2.[135,1471,755,1944]" captionTargetPageId="2" captionText="Fig. 1. Method of measurements used in this study: 1, Standard length; 2, Predorsal length; 3, Preanal length; 4, Prepelvic length; 5, Prepectoral length; 6, Dorsal-spine length; 7, Dorsal-fin length; 8, Length of dorsal-fin base; 9, Length of anal-fin base; 10, Pelvic-fin length; 11, Pectoral-fin length; 12, Pectoral-spine length; 13, Caudal-fin length; 14, Length of adipose-fin base; 15, Maximum height of adipose fin; 16, Dorsal to adipose distance; 17, Post-adipose distance; 18, Length of caudal peduncle; 19, Depth of caudal peduncle; 20, Body depth at anus; 21, Head length; 22, Head width; 23, Head depth; 24, Snout length; 25, Interorbital distance; 26, Eye diameter." figureDoi="http://doi.org/10.5281/zenodo.5351790" httpUri="https://zenodo.org/record/5351790/files/figure.png" pageId="33" pageNumber="238">Fig. 1</figureCitation>
;
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;
<treatmentCitation id="C6109E65FFC8FF96FE57F47AFD088080" author="Kottelat, M" box="[441,648,876,898]" page="122" pageId="33" pageNumber="238" year="2001">
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, Fig. 327;
<treatmentCitation id="C6109E65FFC8FF96FF79F49FFECC809F" author="Ferraris, C" box="[151,332,905,927]" page="88" pageId="33" pageNumber="238" year="2007">
<bibRefCitation id="2320C585FFC8FF96FF79F49FFECC809F" author="Ferraris, C" box="[151,332,905,927]" pageId="33" pageNumber="238" pagination="1 - 628" refId="ref68483" refString="Ferraris, C. J., 2007. Checklist of catfishes, recent and fossil (Osteichthyes: Siluriformes), and catalogue of siluriform primary types. Zootaxa, 1418: 1 - 628." type="journal article" year="2007">Ferraris, 2007: 88</bibRefCitation>
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;
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</treatmentCitation>
;
<treatmentCitation id="C6109E65FFC8FF96FDB9F49FFF6F80BA" author="Rainboth, W" page="72" pageId="33" pageNumber="238" year="2012">
<bibRefCitation id="2320C585FFC8FF96FDB9F49FFF6F80BA" author="Rainboth, W" pageId="33" pageNumber="238" pagination="1 - 172" refId="ref75936" refString="Rainboth, W. J., C. Vidthayanon &amp; D. Y. Mai, 2012. Fishes of the greater Mekong ecosystem with species list and photographic atlas. Miscellaneous Publications. Museum of Zoology, University of Michigan, 201: i - xvi + 1 - 172, 121 pls." type="book chapter" year="2012">Rainboth et al., 2012: 72</bibRefCitation>
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, Plate 45, Fig. 943.
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: confluence of Nam Leuk and Nam Ngong,
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SL,
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, a tributary of
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;
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<collectionCode id="21A020B1FFC8FF96FEB0F345FE158769" box="[350,405,1107,1128]" pageId="33" pageNumber="238">CMK</collectionCode>
13393,
<specimenCount id="51B773FDFFC8FF96FE07F345FD9E8769" box="[489,542,1107,1128]" pageId="33" pageNumber="238" type="generic">9 ex.</specimenCount>
,
<quantity id="80491591FFC8FF96FDC4F345FD458769" box="[554,709,1107,1129]" metricMagnitude="-2" metricUnit="m" metricValue="9.625" metricValueMax="12.93" metricValueMin="6.32" pageId="33" pageNumber="238" unit="mm" value="96.25" valueMax="129.3" valueMin="63.2">63.2129.3 mm</quantity>
SL,
<collectingRegion id="85757696FFC8FF96FF9FF379FEA38784" box="[113,291,1135,1157]" country="Laos" name="Viangchan" pageId="33" pageNumber="238">Vientiane province</collectingRegion>
,
<collectingMunicipality id="A76A220EFFC8FF96FEC4F366FE008784" box="[298,384,1136,1157]" pageId="33" pageNumber="238">Nam San</collectingMunicipality>
downstream of
<location id="426EEEAFFFC8FF96FDFCF366FD068784" LSID="urn:lsid:plazi:treatment:CF180962FFC8FF95FF44F56BFE3A85F0:426EEEAFFFC8FF96FDFCF366FD068784" box="[530,646,1136,1157]" country="Laos" latitude="18.534166" longLatPrecision="21" longitude="102.93" municipality="Nam San" name="Ban Pa Man" pageId="33" pageNumber="238" stateProvince="Viangchan">Ban Pa Man</location>
,
<geoCoordinate id="2285DEB3FFC8FF96FD60F366FD708784" box="[654,752,1136,1157]" degrees="18" direction="north" minutes="32" orientation="latitude" pageId="33" pageNumber="238" precision="15" seconds="3" value="18.534166">18°32'3&quot;N</geoCoordinate>
<geoCoordinate id="2285DEB3FFC8FF96FF9FF39AFF6C87A3" box="[113,236,1164,1186]" degrees="102" direction="east" minutes="55" orientation="longitude" pageId="33" pageNumber="238" precision="15" seconds="48" value="102.93">102°55'48&quot;E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="F7D9B229FFC8FF96FF1BF39BFE8787DD" collectionCode="UMMZ" country="Laos" location="Nam Ngum" municipality="Nam Tou" pageId="33" pageNumber="238" specimenCode="UMMZ 238567" specimenCount="30" stateProvince="Viangchan">
<specimenCode id="1717100FFFC8FF96FF1BF39BFE0C87A3" box="[245,396,1164,1186]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="33" pageNumber="238" type="University or college">UMMZ 238567</specimenCode>
,
<specimenCount id="51B773FDFFC8FF96FE7BF39AFE5087A3" box="[405,464,1164,1186]" pageId="33" pageNumber="238" type="generic">30 ex.</specimenCount>
, 62.0138.0 mm SL,
<collectingRegion id="85757696FFC8FF96FD78F39AFF4A87BE" country="Laos" name="Viangchan" pageId="33" pageNumber="238">Vientiane province</collectingRegion>
:
<collectingMunicipality id="A76A220EFFC8FF96FF38F3BFFEB687BF" box="[214,310,1193,1214]" pageId="33" pageNumber="238">Nam Tou</collectingMunicipality>
,
<quantity id="80491591FFC8FF96FEADF3BFFEF787BE" box="[323,375,1193,1215]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="33" pageNumber="238" unit="km" value="8.0">8 km</quantity>
upstream from confluence with
<location id="426EEEAFFFC8FF96FD2FF3BFFF2C87DA" LSID="urn:lsid:plazi:treatment:CF180962FFC8FF95FF44F56BFE3A85F0:426EEEAFFFC8FF96FD2FF3BFFF2C87DA" country="Laos" municipality="Nam Tou" name="Nam Ngum" pageId="33" pageNumber="238" stateProvince="Viangchan">Nam Ngum</location>
reservoir
</materialsCitation>
;
<materialsCitation id="F7D9B229FFC8FF96FEE1F3D0FD1987F8" collectionCode="UMMZ" country="Laos" location="Nam Ngum" municipality="Nam Ngao" pageId="33" pageNumber="238" specimenCode="UMMZ 238677" specimenCount="8" stateProvince="Viangchan">
<specimenCode id="1717100FFFC8FF96FEE1F3D0FE2687DD" box="[271,422,1222,1244]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="33" pageNumber="238" type="University or college">UMMZ 238677</specimenCode>
,
<specimenCount id="51B773FDFFC8FF96FE40F3D0FE5D87DA" box="[430,477,1222,1243]" pageId="33" pageNumber="238" type="generic">8 ex.</specimenCount>
, 44.066.0 mm SL,
<collectingRegion id="85757696FFC8FF96FD78F3D0FF4987F8" country="Laos" name="Viangchan" pageId="33" pageNumber="238">Vientiane province</collectingRegion>
:
<collectingMunicipality id="A76A220EFFC8FF96FF3BF3F5FEC187F9" box="[213,321,1251,1272]" pageId="33" pageNumber="238">Nam Ngao</collectingMunicipality>
at S end of
<location id="426EEEAFFFC8FF96FE2DF3F5FDB787F9" LSID="urn:lsid:plazi:treatment:CF180962FFC8FF95FF44F56BFE3A85F0:426EEEAFFFC8FF96FE2DF3F5FDB787F9" box="[451,567,1251,1272]" country="Laos" municipality="Nam Ngao" name="Nam Ngum" pageId="33" pageNumber="238" stateProvince="Viangchan">Nam Ngum</location>
reservoir
</materialsCitation>
;
<materialsCitation id="F7D9B229FFC8FF96FD4BF3F5FDD28633" collectionCode="UMMZ" country="Laos" elevation="100" location="Nam Ngum" municipality="Nam Ti" pageId="33" pageNumber="238" specimenCode="UMMZ 240634" specimenCount="12" stateProvince="Viangchan">
<specimenCode id="1717100FFFC8FF96FD4BF3F5FF3B8614" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="33" pageNumber="238" type="University or college">UMMZ 240634</specimenCode>
,
<specimenCount id="51B773FDFFC8FF96FF26F216FE878614" box="[200,263,1280,1301]" pageId="33" pageNumber="238" type="generic">12 ex.</specimenCount>
, 50.0
<quantity id="80491591FFC8FF96FEA4F3E9FE2C8614" box="[330,428,1279,1301]" metricMagnitude="-1" metricUnit="m" metricValue="1.4449999999999998" pageId="33" pageNumber="238" unit="mm" value="144.5">144.5 mm</quantity>
SL,
<collectingRegion id="85757696FFC8FF96FE30F3E9FD1A8614" box="[478,666,1279,1301]" country="Laos" name="Viangchan" pageId="33" pageNumber="238">Vientiane province</collectingRegion>
:
<collectingMunicipality id="A76A220EFFC8FF96FD4BF216FD708614" box="[677,752,1279,1301]" pageId="33" pageNumber="238">Nam Ti</collectingMunicipality>
at junction with
<location id="426EEEAFFFC8FF96FEFFF20BFE038633" LSID="urn:lsid:plazi:treatment:CF180962FFC8FF95FF44F56BFE3A85F0:426EEEAFFFC8FF96FEFFF20BFE038633" box="[273,387,1309,1330]" country="Laos" municipality="Nam Ti" name="Nam Ngum" pageId="33" pageNumber="238" stateProvince="Viangchan">Nam Ngum</location>
and upstream
<quantity id="80491591FFC8FF96FDFCF20AFDD28633" box="[530,594,1308,1330]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="238" unit="m" value="100.0">
<elevation id="CC9C5F47FFC8FF96FDFCF20AFDD28633" box="[530,594,1308,1330]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="238" unit="m" value="100.0">100 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="F7D9B229FFC8FF96FDB3F20BFD2E864E" accessionNumber="NRM 15013" collectionCode="NRM" country="Laos" latitude="14.15" location="Pakse" longLatPrecision="1289" longitude="105.833336" municipality="Sedone River" pageId="33" pageNumber="238" specimenCount="1">
<collectionCode id="21A020B1FFC8FF96FDB3F20BFD148633" box="[605,660,1309,1330]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="33" pageNumber="238" type="Museum">NRM</collectionCode>
15013,
<specimenCount id="51B773FDFFC8FF96FD0AF20AFF10864F" pageId="33" pageNumber="238" type="generic">1 ex.</specimenCount>
,
<quantity id="80491591FFC8FF96FF72F22FFF7F864F" box="[156,255,1337,1359]" metricMagnitude="-1" metricUnit="m" metricValue="2.368" pageId="33" pageNumber="238" unit="mm" value="236.8">236.8 mm</quantity>
SL,
<collectingMunicipality id="A76A220EFFC8FF96FEDFF22FFE36864E" box="[305,438,1337,1359]" pageId="33" pageNumber="238">Sedone River</collectingMunicipality>
,
<location id="426EEEAFFFC8FF96FE2FF22FFE7B864E" LSID="urn:lsid:plazi:treatment:CF180962FFC8FF95FF44F56BFE3A85F0:426EEEAFFFC8FF96FE2FF22FFE7B864E" box="[449,507,1337,1359]" country="Laos" latitude="14.15" longLatPrecision="1289" longitude="105.833336" municipality="Sedone River" name="Pakse" pageId="33" pageNumber="238">Pakse</location>
,
<geoCoordinate id="2285DEB3FFC8FF96FDE9F22FFDCB864F" box="[519,587,1337,1358]" degrees="14" direction="north" minutes="9" orientation="latitude" pageId="33" pageNumber="238" precision="925" value="14.15">14°9'N</geoCoordinate>
<geoCoordinate id="2285DEB3FFC8FF96FDBCF22FFD2E864E" box="[594,686,1337,1359]" degrees="105" direction="east" minutes="50" orientation="longitude" pageId="33" pageNumber="238" precision="925" value="105.833336">105°50'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="F7D9B229FFC8FF96FD57F22FFE09866A" accessionNumber="NRM 15019" collectionCode="NRM" country="Laos" pageId="33" pageNumber="238" specimenCount="1">
<collectionCode id="21A020B1FFC8FF96FD57F22FFD70864F" box="[697,752,1337,1358]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="33" pageNumber="238" type="Museum">NRM</collectionCode>
15019,
<specimenCount id="51B773FDFFC8FF96FF52F240FF70866A" box="[188,240,1366,1387]" pageId="33" pageNumber="238" type="generic">1 ex.</specimenCount>
,
<quantity id="80491591FFC8FF96FF13F240FEE1866A" box="[253,353,1366,1388]" metricMagnitude="-1" metricUnit="m" metricValue="1.985" pageId="33" pageNumber="238" unit="mm" value="198.5">198.5 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="F7D9B229FFC8FF96FE7BF240FD6C866A" accessionNumber="NRM 15020" box="[405,748,1366,1388]" collectionCode="NRM" country="Laos" pageId="33" pageNumber="238" specimenCount="1">
<collectionCode id="21A020B1FFC8FF96FE7BF240FE4C866A" box="[405,460,1366,1387]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="33" pageNumber="238" type="Museum">NRM</collectionCode>
15020,
<specimenCount id="51B773FDFFC8FF96FDCEF240FDD4866A" box="[544,596,1366,1387]" pageId="33" pageNumber="238" type="generic">1 ex.</specimenCount>
,
<quantity id="80491591FFC8FF96FD8EF240FD44866A" box="[608,708,1366,1387]" metricMagnitude="-1" metricUnit="m" metricValue="1.919" pageId="33" pageNumber="238" unit="mm" value="191.9">191.9 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="F7D9B229FFC8FF96FF9FF265FE7A8689" accessionNumber="NRM 15022" box="[113,506,1394,1416]" collectionCode="NRM" country="Laos" location="Laos" pageId="33" pageNumber="238" specimenCount="1">
<collectionCode id="21A020B1FFC8FF96FF9FF265FF288689" box="[113,168,1395,1416]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="33" pageNumber="238" type="Museum">NRM</collectionCode>
15022,
<specimenCount id="51B773FDFFC8FF96FF16F265FEAB8689" box="[248,299,1395,1416]" pageId="33" pageNumber="238" type="generic">1 ex.</specimenCount>
,
<quantity id="80491591FFC8FF96FEDBF264FE188689" box="[309,408,1394,1416]" metricMagnitude="-1" metricUnit="m" metricValue="2.516" pageId="33" pageNumber="238" unit="mm" value="251.6">251.6 mm</quantity>
SL,
<location id="426EEEAFFFC8FF96FE26F265FE7A8689" LSID="urn:lsid:plazi:treatment:CF180962FFC8FF95FF44F56BFE3A85F0:426EEEAFFFC8FF96FE26F265FE7A8689" box="[456,506,1395,1416]" country="Laos" name="Laos" pageId="33" pageNumber="238">Laos</location>
</materialsCitation>
.
</paragraph>
<paragraph id="470EB874FFC8FF96FF9FF2BAFEA2858D" blockId="33.[113,753,1452,1676]" pageId="33" pageNumber="238">
<materialsCitation id="F7D9B229FFC8FF96FF9FF2BAFF5786DE" collectionCode="MNHN" country="Cambodia" location="Cambodia" pageId="33" pageNumber="238" specimenCode="MNHN 1966-728" specimenCount="1" typeStatus="holotype">
<collectingCountry id="3FA6F8E4FFC8FF96FF9FF2BAFE8186C0" box="[113,257,1452,1473]" name="Cambodia" pageId="33" pageNumber="238">CAMBODIA</collectingCountry>
:
<specimenCode id="1717100FFFC8FF96FEFEF2BBFE5586C3" box="[272,469,1452,1474]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="238">MNHN 1966728</specimenCode>
,
<typeStatus id="980A06D6FFC8FF96FE0AF2BAFDC486C3" box="[484,580,1452,1474]" pageId="33" pageNumber="238" type="holotype">holotype</typeStatus>
,
<quantity id="80491591FFC8FF96FDBDF2BAFD4186C3" box="[595,705,1452,1474]" metricMagnitude="-1" metricUnit="m" metricValue="4.198" pageId="33" pageNumber="238" unit="mm" value="419.8">419.8 mm</quantity>
SL,
<location id="426EEEAFFFC8FF96FF9FF2DFFF5786DE" LSID="urn:lsid:plazi:treatment:CF180962FFC8FF95FF44F56BFE3A85F0:426EEEAFFFC8FF96FF9FF2DFFF5786DE" box="[113,215,1481,1503]" country="Cambodia" name="Cambodia" pageId="33" pageNumber="238">Cambodia</location>
</materialsCitation>
;
<materialsCitation id="F7D9B229FFC8FF96FF0EF2DFFEB48518" collectionCode="UMMZ" country="Cambodia" latitude="13.266666" location="Mekong" longLatPrecision="1291" longitude="103.8" municipality="River" pageId="33" pageNumber="238" specimenCode="UMMZ 232423" specimenCount="3" stateProvince="Siem Reap" typeStatus="holotype">
<specimenCode id="1717100FFFC8FF96FF0EF2DFFEFA86DF" box="[224,378,1481,1502]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="33" pageNumber="238" type="University or college">UMMZ 232423</specimenCode>
,
<specimenCount id="51B773FDFFC8FF96FE6AF2DFFE3486DF" box="[388,436,1481,1502]" pageId="33" pageNumber="238" type="generic">3 ex.</specimenCount>
,
<quantity id="80491591FFC8FF96FE53F2DFFDD486DF" box="[445,596,1481,1503]" metricMagnitude="-2" metricUnit="m" metricValue="8.54" metricValueMax="10.209999999999999" metricValueMin="6.87" pageId="33" pageNumber="238" unit="mm" value="85.4" valueMax="102.1" valueMin="68.7">68.7102.1 mm</quantity>
SL,
<collectingRegion id="85757696FFC8FF96FD6FF2DFFD6D86DF" box="[641,749,1481,1503]" country="Cambodia" name="Siem Reap" pageId="33" pageNumber="238">Siem Reap</collectingRegion>
: floating village at mouth of
<collectingRegion id="85757696FFC8FF96FE62F2F0FE7786FA" box="[396,503,1510,1532]" country="Cambodia" name="Siem Reap" pageId="33" pageNumber="238">Siem Reap</collectingRegion>
<collectingMunicipality id="A76A220EFFC8FF96FDEFF2F0FDB886FD" box="[513,568,1510,1532]" pageId="33" pageNumber="238">River</collectingMunicipality>
,
<location id="426EEEAFFFC8FF96FDAAF2F0FD1486FD" LSID="urn:lsid:plazi:treatment:CF180962FFC8FF95FF44F56BFE3A85F0:426EEEAFFFC8FF96FDAAF2F0FD1486FD" box="[580,660,1510,1532]" country="Cambodia" latitude="13.266666" longLatPrecision="1291" longitude="103.8" municipality="River" name="Mekong" pageId="33" pageNumber="238" stateProvince="Siem Reap">Mekong</location>
drainage (
<geoCoordinate id="2285DEB3FFC8FF96FF97F115FF488518" box="[121,200,1539,1561]" degrees="13" direction="north" minutes="16" orientation="latitude" pageId="33" pageNumber="238" precision="925" value="13.266666">13°16'N</geoCoordinate>
<geoCoordinate id="2285DEB3FFC8FF96FF3FF115FEAD8518" box="[209,301,1539,1561]" degrees="103" direction="east" minutes="48" orientation="longitude" pageId="33" pageNumber="238" precision="925" value="103.8">103°48'E</geoCoordinate>
)
</materialsCitation>
;
<materialsCitation id="F7D9B229FFC8FF96FEAEF115FDAA8553" collectionCode="UMMZ" country="Cambodia" latitude="13.266666" location="Mekong" longLatPrecision="1291" longitude="103.76667" municipality="River" pageId="33" pageNumber="238" specimenCode="UMMZ 232462" specimenCount="1" stateProvince="Siem Reap" typeStatus="holotype">
<specimenCode id="1717100FFFC8FF96FEAEF115FE5D8518" box="[320,477,1539,1561]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="33" pageNumber="238" type="University or college">UMMZ 232462</specimenCode>
,
<specimenCount id="51B773FDFFC8FF96FE07F115FD9D8519" box="[489,541,1539,1560]" pageId="33" pageNumber="238" type="generic">1 ex.</specimenCount>
,
<quantity id="80491591FFC8FF96FDC7F114FD0C8519" box="[553,652,1538,1560]" metricMagnitude="-1" metricUnit="m" metricValue="1.5059999999999998" pageId="33" pageNumber="238" unit="mm" value="150.6">150.6 mm</quantity>
SL,
<collectingRegion id="85757696FFC8FF96FD51F115FF288534" country="Cambodia" name="Siem Reap" pageId="33" pageNumber="238">Siem Reap</collectingRegion>
:
<collectingMethod id="9EF0C063FFC8FF96FF58F109FE8C8534" box="[182,268,1567,1589]" pageId="33" pageNumber="238">fish trap</collectingMethod>
W of floating village at mouth of
<collectingRegion id="85757696FFC8FF96FD6EF109FD6F8534" box="[640,751,1567,1589]" country="Cambodia" name="Siem Reap" pageId="33" pageNumber="238">Siem Reap</collectingRegion>
<collectingMunicipality id="A76A220EFFC8FF96FF9FF12AFF298553" box="[113,169,1596,1618]" pageId="33" pageNumber="238">River</collectingMunicipality>
,
<location id="426EEEAFFFC8FF96FF58F12AFE878553" LSID="urn:lsid:plazi:treatment:CF180962FFC8FF95FF44F56BFE3A85F0:426EEEAFFFC8FF96FF58F12AFE878553" box="[182,263,1596,1618]" country="Cambodia" latitude="13.266666" longLatPrecision="1291" longitude="103.76667" municipality="River" name="Mekong" pageId="33" pageNumber="238" stateProvince="Siem Reap">Mekong</location>
drainage,
<geoCoordinate id="2285DEB3FFC8FF96FE9AF12AFE438553" box="[372,451,1596,1618]" degrees="13" direction="north" minutes="16" orientation="latitude" pageId="33" pageNumber="238" precision="925" value="13.266666">13°16'N</geoCoordinate>
<geoCoordinate id="2285DEB3FFC8FF96FE23F12AFDAA8553" box="[461,554,1596,1618]" degrees="103" direction="east" minutes="46" orientation="longitude" pageId="33" pageNumber="238" precision="925" value="103.76667">103°46'E</geoCoordinate>
</materialsCitation>
;
<materialsCitation id="F7D9B229FFC8FF96FDD9F12BFE9F858D" collectionCode="UMMZ" country="Cambodia" location="Mekong" municipality="Kompong Chhnang" pageId="33" pageNumber="238" specimenCode="UMMZ 232698" specimenCount="5" typeStatus="holotype">
<specimenCode id="1717100FFFC8FF96FDD9F12BFD578553" box="[567,727,1596,1618]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="33" pageNumber="238" type="University or college">UMMZ 232698</specimenCode>
,
<specimenCount id="51B773FDFFC8FF96FD0AF12AFF11856F" pageId="33" pageNumber="238" type="generic">5 ex.</specimenCount>
,
<quantity id="80491591FFC8FF96FF70F14FFEC8856F" box="[158,328,1625,1647]" metricMagnitude="-1" metricUnit="m" metricValue="1.6344999999999998" metricValueMax="1.751" metricValueMin="1.518" pageId="33" pageNumber="238" unit="mm" value="163.45" valueMax="175.1" valueMin="151.8">151.8175.1 mm</quantity>
SL,
<collectingMunicipality id="A76A220EFFC8FF96FE93F14FFDC0856E" box="[381,576,1625,1647]" pageId="33" pageNumber="238">Kompong Chhnang</collectingMunicipality>
morning market,
<location id="426EEEAFFFC8FF96FF9FF160FF41858D" LSID="urn:lsid:plazi:treatment:CF180962FFC8FF95FF44F56BFE3A85F0:426EEEAFFFC8FF96FF9FF160FF41858D" box="[113,193,1654,1676]" country="Cambodia" municipality="Kompong Chhnang" name="Mekong" pageId="33" pageNumber="238">Mekong</location>
drainage
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0FABEBFFFFC8FF96FF9FF1A2FAB88177" pageId="33" pageNumber="238" type="diagnosis">
<paragraph id="470EB874FFC8FF96FF9FF1A2FAB88177" blockId="33.[113,753,1716,2092]" lastBlockId="33.[813,1452,190,630]" pageId="33" pageNumber="238">
<emphasis id="75C56466FFC8FF96FF9FF1A2FF6685CD" bold="true" box="[113,230,1716,1740]" pageId="33" pageNumber="238">
<emphasis id="75C56466FFC8FF96FF9FF1A2FF5F85CD" bold="true" box="[113,223,1716,1740]" italics="true" pageId="33" pageNumber="238">Diagnosis</emphasis>
.
</emphasis>
<taxonomicName id="80B1C3F7FFC8FF96FEE0F1A3FD8785CD" baseAuthorityName="Fang &amp; Chaux" baseAuthorityYear="1949" box="[270,519,1717,1740]" class="Actinopterygii" family="Bagridae" genus="Hemibagrus" kingdom="Animalia" order="Siluriformes" pageId="33" pageNumber="238" phylum="Chordata" rank="species" species="filamentus">
<emphasis id="75C56466FFC8FF96FEE0F1A3FD8785CD" box="[270,519,1717,1740]" italics="true" pageId="33" pageNumber="238">Hemibagrus filamentus</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="80B1C3F7FFC8FF96FF9FF1C3FF7D85ED" authorityName="Popta" authorityYear="1904" box="[113,253,1749,1772]" class="Insecta" family="Cerambycidae" genus="Macrones" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="238" phylum="Arthropoda" rank="species" species="capitulum">
<emphasis id="75C56466FFC8FF96FF9FF1C3FF7D85ED" box="[113,253,1749,1772]" italics="true" pageId="33" pageNumber="238">H. capitulum</emphasis>
</taxonomicName>
in having a longer, narrower head (head width 5560% HL vs. 5970), from
<taxonomicName id="80B1C3F7FFC8FF96FE55F1E3FD95840D" box="[443,533,1781,1804]" class="Insecta" family="Cerambycidae" genus="Macrones" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="238" phylum="Arthropoda" rank="species" species="fortis">
<emphasis id="75C56466FFC8FF96FE55F1E3FD95840D" box="[443,533,1781,1804]" italics="true" pageId="33" pageNumber="238">H. fortis</emphasis>
</taxonomicName>
in having a shorter, deeper adipose fin (maximum height of adipose fin 2.33.2 times in the length of its base vs. 2.84.9) and the anteriormost branch of dorsal-fin rays longer than other branches (vs. branches of the dorsal-fin rays of almost equal length.) giving the dorsal fin a jagged (vs. rounded) distal margin (
<figureCitation id="DF8AA4F1FFC8FF96FF97F0A2FF4B84CD" box="[121,203,1972,1996]" captionStart="Fig" captionStartId="25.[814,849,1975,1997]" captionTargetBox="[811,1452,1048,1964]" captionTargetId="figure-8@25.[808,1462,1030,1964]" captionTargetPageId="25" captionText="Fig. 20. Schematic illustrations of fully extended dorsal fins of members of the H. nemurus species group: a, H. fortis, ZRC 40522, 126.9 mm SL; b, other members of the H. nemurus species group (H. nemurus, ZRC 39181, 132.4 mm SL, illustrated). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5351826" httpUri="https://zenodo.org/record/5351826/files/figure.png" pageId="33" pageNumber="238">Fig. 20</figureCitation>
), and from
<taxonomicName id="80B1C3F7FFC8FF96FEA5F0A3FE4884CD" authorityName="Bleeker" authorityYear="1846" box="[331,456,1973,1996]" class="Actinopterygii" family="Bagridae" genus="Bagrus" kingdom="Animalia" order="Siluriformes" pageId="33" pageNumber="238" phylum="Chordata" rank="species" species="hoevenii">
<emphasis id="75C56466FFC8FF96FEA5F0A3FE4884CD" box="[331,456,1973,1996]" italics="true" pageId="33" pageNumber="238">H. hoevenii</emphasis>
</taxonomicName>
in having the premaxillary tooth band not (vs. partially) exposed when the mouth is closed, rounded (vs. tapering) caudal-fin lobes, absence (vs. presence) of a broad and conspicuous dark margin around the caudal fin, a rounded (vs. triangular) anal fin, broader membranes of the dorsal fin which give it a rounded (vs. triangular) appearance, and filamentous extensions on the dorsal fin not reaching beyond middle of adipose-fin base (vs. very long filamentous extensions of the first two dorsalfin rays that reach beyond the posterior base of the adipose fin). It differs from
<taxonomicName id="80B1C3F7FFC8FF96FC12F669FBF98297" box="[1020,1145,383,406]" class="Actinopterygii" family="Bagridae" genus="Hemibagrus" kingdom="Animalia" order="Siluriformes" pageId="33" pageNumber="238" phylum="Chordata" rank="species" species="nemurus">
<emphasis id="75C56466FFC8FF96FC12F669FBF98297" box="[1020,1145,383,406]" italics="true" pageId="33" pageNumber="238">H. nemurus</emphasis>
</taxonomicName>
in having a relatively longer and narrower head (head width 5560% HL vs. 6069) and a smaller dorsal to adipose distance (7.710.9% SL vs. 10.216.2), and from
<taxonomicName id="80B1C3F7FFC8FF96FBC8F6C9FB4582F7" authorityName="Ng &amp; Rainboth" authorityYear="1999" box="[1062,1221,479,502]" class="Actinopterygii" family="Bagridae" genus="Hemibagrus" kingdom="Animalia" order="Siluriformes" pageId="33" pageNumber="238" phylum="Chordata" rank="species" species="spilopterus">
<emphasis id="75C56466FFC8FF96FBC8F6C9FB4582F7" box="[1062,1221,479,502]" italics="true" pageId="33" pageNumber="238">H. spilopterus</emphasis>
</taxonomicName>
in having a smaller dorsal to adipose distance (7.710.9% SL vs. 10.817.6), a flat (vs. gently convex) interorbital space, and dorsal-fin rays (excluding filamentous extensions) reaching beyond (vs. not reaching or just reaching) the adipose-fin origin.
</paragraph>
</subSubSection>
<subSubSection id="0FABEBFFFFC8FF96FCC3F58BFACE8597" pageId="33" pageNumber="238" type="description">
<paragraph id="470EB874FFC8FF96FCC3F58BFC3486D7" blockId="33.[811,1452,669,1494]" pageId="33" pageNumber="238">
<emphasis id="75C56466FFC8FF96FCC3F58BFC3781B7" bold="true" box="[813,951,669,694]" pageId="33" pageNumber="238">
<emphasis id="75C56466FFC8FF96FCC3F58BFC3081B4" bold="true" box="[813,944,669,693]" italics="true" pageId="33" pageNumber="238">Description</emphasis>
.
</emphasis>
— Biometric and meristic data as in Table 13. General description as for genus. Head depressed and broad, body moderately compressed; head width 5560% HL. Interorbital space flat. Eye diameter 1015% HL. Premaxillary tooth band not exposed when mouth is closed. Dorsal profile rising evenly but not steeply from tip of snout to origin of dorsal fin, then sloping gently ventrally from there to end of caudal peduncle. Ventral profile horizontal to origin of anal fin, then sloping dorsally to end of caudal peduncle. Adipose-fin base short, approximately as long as anal-fin base; maximum height 5.07.2% SL (2.33.2 times in length of adipose-fin base). Adipose-fin origin separated from base of last dorsal-fin ray by distance of 7.710.9% SL. Dorsal spine stout, with 610 serrations along posterior edge. Anteriormost branch of each dorsal-fin ray longer than other branches, imparting jagged distal margin to fin; membranes broad, imparting rounded appearance. Adpressed dorsal fin (excluding filamentous extensions of rays) reaching beyond adipose-fin origin; filamentous extensions not reaching beyond middle of adipose-fin base. Pectoral spine stout, with 1318 large serrations along posterior edge. Anal fin rounded, origin slightly posterior to adipose origin. Caudal fin forked; upper and lower lobes rounded. Maxillary barbels reaching barbels reaching at least vertical through base of last anal-fin ray (221265% HL). Maximum recorded size
<quantity id="80491591FFC8FF96FCC3F2A8FC0686D4" box="[813,902,1470,1494]" metricMagnitude="-1" metricUnit="m" metricValue="4.2" pageId="33" pageNumber="238" unit="mm" value="420.0">420 mm</quantity>
SL.
</paragraph>
<paragraph id="470EB874FFC8FF96FCC3F2EBFACE8597" blockId="33.[813,1452,1533,1686]" pageId="33" pageNumber="238">
<emphasis id="75C56466FFC8FF96FCC3F2EBFC078517" bold="true" box="[813,903,1533,1558]" pageId="33" pageNumber="238">
<emphasis id="75C56466FFC8FF96FCC3F2EBFCFF8517" bold="true" box="[813,895,1533,1558]" italics="true" pageId="33" pageNumber="238">Colour</emphasis>
.
</emphasis>
— Dorsal surface of head and body uniform brownish-grey; ventral surface of head and body dirty white; adipose fin brownish-grey, without black spot; fin-rays and inter-radial membranes of all fins grey; dorsal surface of barbels brownish-grey, ventral surface dirty white.
</paragraph>
</subSubSection>
<caption id="13CEE8FCFFC8FF96FCC2FF18FC0D8B41" ID-DOI="http://doi.org/10.5281/zenodo.5351836" ID-Zenodo-Dep="5351836" httpUri="https://zenodo.org/record/5351836/files/figure.png" pageId="33" pageNumber="238" startId="33.[812,847,2062,2084]" targetBox="[813,1453,1794,2037]" targetPageId="33">
<paragraph id="470EB874FFC8FF96FCC2FF18FC0D8B41" blockId="33.[812,1453,2062,2112]" pageId="33" pageNumber="238">
Fig. 24.
<taxonomicName id="80B1C3F7FFC8FF96FC95FF19FB1D8B22" authority=", CMK" authorityName="CMK" box="[891,1181,2062,2083]" class="Actinopterygii" family="Bagridae" genus="Hemibagrus" kingdom="Animalia" order="Siluriformes" pageId="33" pageNumber="238" phylum="Chordata" rank="species" species="filamentus">
<emphasis id="75C56466FFC8FF96FC95FF19FBDB8B22" box="[891,1115,2063,2083]" italics="true" pageId="33" pageNumber="238">Hemibagrus filamentus</emphasis>
, CMK
</taxonomicName>
13393, 110.2 mm SL, Laos: Nam San.
</paragraph>
</caption>
<subSubSection id="0FABEBFFFFCBFF95FF66F2ECFE3A85F0" pageId="34" pageNumber="239" type="distribution">
<paragraph id="470EB874FFCBFF95FF66F2ECFF6E8552" blockId="34.[136,775,1530,1619]" pageId="34" pageNumber="239">
<emphasis id="75C56466FFCBFF95FF66F2ECFE948512" bold="true" box="[136,276,1530,1555]" pageId="34" pageNumber="239">
<emphasis id="75C56466FFCBFF95FF66F2ECFE8D8512" bold="true" box="[136,269,1530,1555]" italics="true" pageId="34" pageNumber="239">Distribution</emphasis>
.
</emphasis>
<taxonomicName id="80B1C3F7FFCBFF95FED1F2EAFDB88512" box="[319,568,1532,1555]" class="Actinopterygii" family="Bagridae" genus="Hemibagrus" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="239" phylum="Chordata" rank="species" species="filamentus">
<emphasis id="75C56466FFCBFF95FED1F2EAFDB88512" box="[319,568,1532,1555]" italics="true" pageId="34" pageNumber="239">Hemibagrus filamentus</emphasis>
</taxonomicName>
is known from the middle and lower Mekong in
<collectingCountry id="3FA6F8E4FFCBFF95FE2BF10DFE7B8533" box="[453,507,1563,1586]" name="Laos" pageId="34" pageNumber="239">Laos</collectingCountry>
,
<collectingCountry id="3FA6F8E4FFCBFF95FDE8F10DFDE58532" box="[518,613,1563,1587]" name="Thailand" pageId="34" pageNumber="239">Thailand</collectingCountry>
and
<collectingCountry id="3FA6F8E4FFCBFF95FD77F10DFC878532" box="[665,775,1563,1587]" name="Cambodia" pageId="34" pageNumber="239">Cambodia</collectingCountry>
(
<figureCitation id="DF8AA4F1FFCBFF95FF7EF12DFF618552" box="[144,225,1595,1619]" captionStart="Fig" captionStartId="27.[812,847,2034,2056]" captionTargetBox="[812,1454,1230,2011]" captionTargetId="figure-8@27.[811,1454,1230,2011]" captionTargetPageId="27" captionText="Fig.22. Map showing collection localities of Hemibagrus nemurus, H. capitulum and H. filamentus specimens examined in this study." figureDoi="http://doi.org/10.5281/zenodo.5351830" httpUri="https://zenodo.org/record/5351830/files/figure.png" pageId="34" pageNumber="239">Fig. 22</figureCitation>
).
</paragraph>
<paragraph id="470EB874FFCBFF95FF66F16FFE3A85F0" blockId="34.[136,775,1657,1777]" pageId="34" pageNumber="239">
<emphasis id="75C56466FFCBFF95FF66F16FFEEA8593" bold="true" box="[136,362,1657,1682]" pageId="34" pageNumber="239">
<emphasis id="75C56466FFCBFF95FF66F16FFEE38593" bold="true" box="[136,355,1657,1682]" italics="true" pageId="34" pageNumber="239">Habitat and biology</emphasis>
.
</emphasis>
<taxonomicName id="80B1C3F7FFCBFF95FE7DF16AFD0C8592" box="[403,652,1660,1683]" class="Actinopterygii" family="Bagridae" genus="Hemibagrus" kingdom="Animalia" order="Siluriformes" pageId="34" pageNumber="239" phylum="Chordata" rank="species" species="filamentus">
<emphasis id="75C56466FFCBFF95FE7DF16AFD0C8592" box="[403,652,1660,1683]" italics="true" pageId="34" pageNumber="239">Hemibagrus filamentus</emphasis>
</taxonomicName>
is found in slow-flowing or standing waters, moving into flooded forests during periods of high water levels. It feeds on crustaceans and fishes (
<bibRefCitation id="2320C585FFCBFF95FEECF1CFFE2F85F0" author="Rainboth, W" box="[258,431,1753,1777]" pageId="34" pageNumber="239" refId="ref75901" refString="Rainboth, W. J., 1996. Fishes of the Cambodian Mekong. FAO Species Identification Field Guide for Fishery Purposes. FAO, Rome. Xi + 265, 27 pls." type="book" year="1996">Rainboth, 1996</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>