265 lines
31 KiB
XML
265 lines
31 KiB
XML
<document id="BB67064C2689D320DE057577D146DFC4" ID-CLB-Dataset="32505" ID-DOI="10.11646/zootaxa.4305.1.1" ID-GBIF-Dataset="24ae3650-4b27-4fcb-9f51-1787fd8aa2a4" ID-ISSN="1175-5326" ID-Zenodo-Dep="842587" ID-ZooBank="1192C3A0-5CCB-4A86-903C-A2B82906A5F9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1502688179640" checkinUser="plazi" docAuthor="Cáceres-Chamizo, Julia P., Sanner, Joann, Tilbrook, Kevin J. & Ostrovsky, Andrew N." docDate="2017" docId="CF0AB852FFD4E904FF03FC929130E66D" docLanguage="en" docName="zootaxa.4305.1.1.pdf" docOrigin="Zootaxa 4305 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Exechonella similis Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky 2017, n. sp." docType="treatment" docVersion="7" lastPageNumber="45" masterDocId="3333C02AFFFEE928FF94FFF59553E431" masterDocTitle="Revision of the Recent species of Exechonella Canu & Bassler in Duvergier, 1924 and Actisecos Canu & Bassler, 1927 (Bryozoa, Cheilostomata): systematics, biogeography and evolutionary trends in skeletal morphology" masterLastPageNumber="79" masterPageNumber="1" pageNumber="43" updateTime="1698466532424" updateUser="ExternalLinkService">
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<mods:titleInfo id="717737E323EC1342C18E27EF88266DB4">
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<mods:title id="6A7CA277AC9478AB5120106F4E319DB2">Revision of the Recent species of Exechonella Canu & Bassler in Duvergier, 1924 and Actisecos Canu & Bassler, 1927 (Bryozoa, Cheilostomata): systematics, biogeography and evolutionary trends in skeletal morphology</mods:title>
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</mods:titleInfo>
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<mods:name id="643EE5089C2B558DA0C8EE73BC4BA408" type="personal">
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<mods:role id="1FD4957FDAC286CD8386F2269321B5AA">
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<mods:roleTerm id="615CEB68E46D7B62811E7C18ABB63F1E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E4292C017D3840EF8156087DF684954F">Cáceres-Chamizo, Julia P.</mods:namePart>
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</mods:name>
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<mods:name id="B2990FD482682CD63AB4738C5BC79C36" type="personal">
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<mods:roleTerm id="2B00AFC861DB3B2C1745C1437426130C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3289B0E392B2C9241F6E0FA7D00F22CC">Sanner, Joann</mods:namePart>
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</mods:name>
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<mods:name id="E47533329824E0AF25CA0504E6102E08" type="personal">
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<mods:roleTerm id="D6D4B3688706DAD1FBDBCFFC41098E00">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8DC337610371AD761605DF4469000723">Tilbrook, Kevin J.</mods:namePart>
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</mods:name>
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<mods:name id="B1908C516F8A9BACDDDCBC37D5F08694" type="personal">
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<mods:role id="5671CDF8DB90351B91FE006690F99D41">
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<mods:roleTerm id="A0DDCCC2C6080FCE86EEA29A0978C015">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FC9A79BE2F04C38C26BD228C1DE87F7D">Ostrovsky, Andrew N.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E755BB16D851B55D099B744477056C09">text</mods:typeOfResource>
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<mods:titleInfo id="7528E2ED9703018B82547C6EC4EFD93C">
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<mods:title id="2D201602931E98675C0B9903FCE086B0">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E5D0612A92FB516694A70FB74B24746A">
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<mods:date id="2E15BB40FFDF343C8A1210F1EEC5D578">2017</mods:date>
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<mods:detail id="8D4063257FECE2F1EC3FEDEAED358277" type="volume">
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<mods:number id="8E616AC9499AE334CC0CB5E1E1A95BB8">4305</mods:number>
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</mods:detail>
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<mods:detail id="7DA20E03CD1C939B3AA6DB34C8BD2144" type="issue">
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<mods:number id="6A8FD2416CF6F9F60970641993BC6575">1</mods:number>
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<mods:end id="AEF1F7EBE697E91803538D20F238606C">79</mods:end>
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<mods:classification id="76E5EFF6EE0A0C30EC8622449A13C9FE">journal article</mods:classification>
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<mods:identifier id="8A0011B2E3E2667FF0997F4FB0F820D1" type="CLB-Dataset">32505</mods:identifier>
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<mods:identifier id="C57962DAB4EFEE73B6C959B267CAC9CB" type="DOI">10.11646/zootaxa.4305.1.1</mods:identifier>
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<mods:identifier id="C958020CFFE2E65E2F90624A465DF1B7" type="GBIF-Dataset">24ae3650-4b27-4fcb-9f51-1787fd8aa2a4</mods:identifier>
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<mods:identifier id="9A9BD8A7D2AC70DC497A6CADF3D48495" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="CF0AB852FFD4E904FF03FC929130E66D" ID-DOI="http://doi.org/10.5281/zenodo.6017338" ID-GBIF-Taxon="132632419" ID-Zenodo-Dep="6017338" LSID="urn:lsid:plazi:treatment:CF0AB852FFD4E904FF03FC929130E66D" httpUri="http://treatment.plazi.org/id/CF0AB852FFD4E904FF03FC929130E66D" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43">
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<subSubSection id="0FB95ACFFFD4E902FF03FC92943BE792" pageId="42" pageNumber="43" type="nomenclature">
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<paragraph id="471C0944FFD4E902FF03FC92949DE7B3" blockId="42.[151,462,871,931]" box="[151,462,871,898]" pageId="42" pageNumber="43">
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<heading id="1C54BE28FFD4E902FF03FC92949DE7B3" bold="true" box="[151,462,871,898]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
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<emphasis id="75D7D556FFD4E902FF03FC92949DE7B3" bold="true" box="[151,462,871,898]" pageId="42" pageNumber="43">
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<taxonomicName id="80A372C7FFD4E902FF03FC9294D0E7B0" authority="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky, 2017" authorityName="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky" authorityYear="2017" box="[151,387,871,897]" class="Gymnolaemata" family="Exechonellidae" genus="Exechonella" kingdom="Animalia" order="Cheilostomatida" pageId="42" pageNumber="43" phylum="Bryozoa" rank="species" species="similis" status="sp. nov.">
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<emphasis id="75D7D556FFD4E902FF03FC9294D0E7B0" bold="true" box="[151,387,871,897]" italics="true" pageId="42" pageNumber="43">Exechonella similis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6EE4682DFFD4E902FE1FFC9D949DE7B3" box="[395,462,872,898]" pageId="42" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="471C0944FFD4E902FF03FC7F943BE792" blockId="42.[151,462,871,931]" box="[151,360,906,931]" pageId="42" pageNumber="43">
|
||
(
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<figureCitation id="DF9815C1FFD4E902FF0BFC7F95A1E792" box="[159,242,906,931]" captionStart="FIGURE 17" captionStartId="43.[151,250,1791,1813]" captionTargetBox="[258,1312,195,1751]" captionTargetId="figure@43.[258,1312,195,1751]" captionTargetPageId="43" captionText="FIGURE 17. Exechonella similis n. sp. Great Barrier Reef, Lizard Island (A: paratype DPUV 2016 - 0001 - 0002; F, G: paratype 2016 - 0001 - 0003; B ‒ E, H: holotype MTQ G 100216). A, general view of non-cleaned colony from above. Peripheral zooidal buds with membranous frontal wall are well seen. Peristomes with processes are seen in some zooids in the central colony part. B, E, F, close-up of autozooids. Some lateralmost foramina with avicularia shown by arrows in B. C, peripheral part of the colony, showing peristome shape and multiporous mural septula. D, close-up of autozooid showing details of primary orifice and peristome. G, non-cleaned lateralmost foramen with avicularium. Arrowhead shows an edge of mandible. H, close-up of lateralmost foramen with avicularium associated with kenozooid in cleaned colony. Pores of kenozooid have centrally perforated cuticular plates. Scale bars: A = 1 mm; B ‒ E = 100 µm; F = 500 µm; G = 50 µm; H = 10 µm." httpUri="https://zenodo.org/record/842621/files/figure.png" pageId="42" pageNumber="43">Fig. 17</figureCitation>
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,
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<tableCitation id="0A213CFFFFD4E902FF69FC7F940CE792" box="[253,351,906,931]" captionStart="TABLE 16" captionStartId="44.[151,239,404,427]" captionText="TABLE 16. Measurements (in µm, except number of foramina) of holotype specimen of Exechonella similis n. sp. Abbreviations: ancestrula length (AncL), ancestrula width (AncW), autozooid length (AzL), autozooid width (AzW), diameter of a foramen including rim (FoD), number of frontal foramina (FoN), diameter of the opening of a foramen (OD), primary orifice length (OrL), primary orifice width (OrW). Mean (m), standard deviation (sd), range (r) and number of measurements (n)." pageId="42" pageNumber="43">Table 16</tableCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0FB95ACFFFD4E902FF03FC2797A9E7D9" box="[151,762,978,1000]" pageId="42" pageNumber="43" type="description">
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||
<paragraph id="471C0944FFD4E902FF03FC2797A9E7D9" blockId="42.[151,895,978,1031]" box="[151,762,978,1000]" pageId="42" pageNumber="43">
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||
<taxonomicName id="80A372C7FFD4E902FF03FC27973BE7D9" authority=": Tilbrook et al. 2001" authorityName="Tilbrook et al." authorityYear="2001" box="[151,616,978,1000]" class="Gymnolaemata" family="Exechonellidae" genus="Exechonella" kingdom="Animalia" order="Cheilostomatida" pageId="42" pageNumber="43" phylum="Bryozoa" rank="species" species="brasiliensis">
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<emphasis id="75D7D556FFD4E902FF03FC2794DCE7D9" box="[151,399,978,1000]" italics="true" pageId="42" pageNumber="43">Exechonella brasiliensis</emphasis>
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: Tilbrook
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<emphasis id="75D7D556FFD4E902FE6EFC269774E7D9" box="[506,551,978,1000]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
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||
. 2001
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||
</taxonomicName>
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||
, p. 65, fig.
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||
<quantity id="805BA4A1FFD4E902FD4EFC2797A4E7D9" box="[730,759,978,1000]" metricMagnitude="-3" metricUnit="kg" metricValue="8.0" pageId="42" pageNumber="43" unit="g" value="8.0">8g</quantity>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0FB95ACFFFD4E902FF03FC04962CE036" box="[151,895,1009,1031]" pageId="42" pageNumber="43" type="reference_group">
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<paragraph id="471C0944FFD4E902FF03FC04962CE036" blockId="42.[151,895,978,1031]" box="[151,895,1009,1031]" pageId="42" pageNumber="43">
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||
?
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||
<taxonomicName id="80A372C7FFD4E902FF3CFC049790E036" ID-CoL="3DKG6" authority=": Winston & Heimberg 1986" authorityName=": Winston & Heimberg" authorityYear="1986" box="[168,707,1009,1031]" class="Gymnolaemata" family="Exechonellidae" genus="Exechonella" kingdom="Animalia" order="Cheilostomatida" pageId="42" pageNumber="43" phylum="Bryozoa" rank="species" species="brasiliensis">
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||
<emphasis id="75D7D556FFD4E902FF3CFC0494F2E036" box="[168,417,1009,1031]" italics="true" pageId="42" pageNumber="43">Exechonella brasiliensis</emphasis>
|
||
:
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||
<bibRefCitation id="233274B5FFD4E902FE39FC049790E036" author="Winston" box="[429,707,1009,1031]" pageId="42" pageNumber="43" refString="Winston, J. E. & Heimberg, B. F. (1986) Bryozoans from Bali, Lombok and Komodo. American Museum Novitates, 2847, 1 - 49." type="journal article" year="1986">Winston & Heimberg 1986</bibRefCitation>
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||
</taxonomicName>
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||
, p. 15, figs 26–27.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0FB95ACFFFD4E902FF03FBC19178E089" pageId="42" pageNumber="43" type="materials_examined">
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<paragraph id="471C0944FFD4E902FF03FBC19178E089" blockId="42.[151,1437,1076,2037]" pageId="42" pageNumber="43">
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<materialsCitation id="F7CB0319FFD4E902FF03FBC1912BE041" ID-GBIF-Occurrence="1657176750" collectingDate="16-10-02" collectingDateMax="2012-10-08" collectingDateMin="16-10-02" county="Cobey Hole" elevation="16" location="Coral Sea" municipality="Lizard Island" pageId="42" pageNumber="43" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="75D7D556FFD4E902FF03FBC194D0E07C" bold="true" box="[151,387,1076,1101]" pageId="42" pageNumber="43">Material examined.</emphasis>
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||
<emphasis id="75D7D556FFD4E902FE19FBC194A7E07C" box="[397,500,1076,1101]" italics="true" pageId="42" pageNumber="43">
|
||
<typeStatus id="9818B7E6FFD4E902FE19FBC194A7E07C" box="[397,500,1076,1101]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
: MTQ G
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||
<date id="331D2F84FFD4E902FDF6FBC197EAE07C" box="[610,697,1076,1101]" pageId="42" pageNumber="43" value="16-10-02">
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||
<collectingDate id="2359D66CFFD4E902FDF6FBC197EAE07C" box="[610,697,1076,1101]" pageId="42" pageNumber="43" value="16-10-02">100216</collectingDate>
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||
</date>
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||
, on coral rubble (mounted on SEM stub and coated with gold).
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||
<location id="427C5F9FFFD4E902FF03FBA2945BE041" LSID="urn:lsid:plazi:treatment:CF0AB852FFD4E904FF03FC929130E66D:427C5F9FFFD4E902FF03FBA2945BE041" box="[151,264,1111,1136]" county="Cobey Hole" municipality="Lizard Island" name="Coral Sea" pageId="42" pageNumber="43">Coral Sea</location>
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||
,
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||
<location id="427C5F9FFFD4E902FE81FBAD94B8E041" LSID="urn:lsid:plazi:treatment:CF0AB852FFD4E904FF03FC929130E66D:427C5F9FFFD4E902FE81FBAD94B8E041" box="[277,491,1111,1136]" county="Cobey Hole" municipality="Lizard Island" name="Great Barrier Reef" pageId="42" pageNumber="43">Great Barrier Reef</location>
|
||
,
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||
<collectingMunicipality id="A778933EFFD4E902FE6CFBA297C3E041" box="[504,656,1111,1136]" pageId="42" pageNumber="43">Lizard Island</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="AE7D71C8FFD4E902FD09FBA29674E041" box="[669,807,1111,1136]" pageId="42" pageNumber="43">Cobey Hole</collectingCounty>
|
||
, depth
|
||
<quantity id="805BA4A1FFD4E902FCEFFBA296E4E05E" box="[891,951,1111,1136]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="42" pageNumber="43" unit="m" value="16.0">
|
||
<elevation id="CC8EEE77FFD4E902FCEFFBA296E4E05E" box="[891,951,1111,1136]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="42" pageNumber="43" unit="m" value="16.0">16 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="331D2F84FFD4E902FC50FBAD912BE041" box="[964,1144,1111,1136]" pageId="42" pageNumber="43" value="2012-10-08">
|
||
<collectingDate id="2359D66CFFD4E902FC50FBAD912BE041" box="[964,1144,1111,1136]" pageId="42" pageNumber="43" value="2012-10-08">8 October 2012</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="F7CB0319FFD4E902FB10FBAC9174E089" ID-GBIF-Occurrence="1657176784" collectingDate="2012-10-04" collectionCode="DPUV" county="Watson Bay" elevation="7" location="Coral Sea" municipality="Lizard Island" pageId="42" pageNumber="43" specimenCode="DPUV 2016- 0001-0002, DPUV 2016-0001-0003" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="75D7D556FFD4E902FB10FBAC91A6E041" box="[1156,1269,1113,1136]" italics="true" pageId="42" pageNumber="43">
|
||
<typeStatus id="9818B7E6FFD4E902FB10FBAC91A6E041" box="[1156,1269,1113,1136]" pageId="42" pageNumber="43" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<specimenCode id="1705A13FFFD4E902FA92FBAD9440E0A5" pageId="42" pageNumber="43">
|
||
<collectionCode id="21B29181FFD4E902FA92FBAD9001E05E" box="[1286,1362,1112,1135]" pageId="42" pageNumber="43">DPUV</collectionCode>
|
||
2016- 0001-0002
|
||
</specimenCode>
|
||
,
|
||
<specimenCode id="1705A13FFFD4E902FEB6FB889767E0A4" box="[290,564,1148,1173]" pageId="42" pageNumber="43">
|
||
<collectionCode id="21B29181FFD4E902FEB6FB88943DE0A5" box="[290,366,1149,1172]" pageId="42" pageNumber="43">DPUV</collectionCode>
|
||
2016-0001-0003
|
||
</specimenCode>
|
||
(on bivalve shells, non-cleaned, mounted on SEM-stubs, uncoated).
|
||
<location id="427C5F9FFFD4E902FACAFB899591E089" LSID="urn:lsid:plazi:treatment:CF0AB852FFD4E904FF03FC929130E66D:427C5F9FFFD4E902FACAFB899591E089" county="Watson Bay" municipality="Lizard Island" name="Coral Sea" pageId="42" pageNumber="43">Coral Sea</location>
|
||
,
|
||
<location id="427C5F9FFFD4E902FF59FB5594F2E089" LSID="urn:lsid:plazi:treatment:CF0AB852FFD4E904FF03FC929130E66D:427C5F9FFFD4E902FF59FB5594F2E089" box="[205,417,1183,1208]" county="Watson Bay" municipality="Lizard Island" name="Great Barrier Reef" pageId="42" pageNumber="43">Great Barrier Reef</location>
|
||
,
|
||
<collectingMunicipality id="A778933EFFD4E902FE38FB6A9710E089" box="[428,579,1183,1208]" pageId="42" pageNumber="43">Lizard Island</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="AE7D71C8FFD4E902FDDAFB559785E089" box="[590,726,1184,1208]" pageId="42" pageNumber="43">Watson Bay</collectingCounty>
|
||
, depth
|
||
<quantity id="805BA4A1FFD4E902FCB3FB6A963AE086" box="[807,873,1183,1208]" metricMagnitude="0" metricUnit="m" metricValue="6.5" pageId="42" pageNumber="43" unit="m" value="6.5">
|
||
<elevation id="CC8EEE77FFD4E902FCB3FB6A963AE086" box="[807,873,1183,1208]" metricMagnitude="0" metricUnit="m" metricValue="6.5" pageId="42" pageNumber="43" unit="m" value="6.5">6.5 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="331D2F84FFD4E902FCE1FB559174E089" box="[885,1063,1183,1208]" pageId="42" pageNumber="43" value="2012-10-04">
|
||
<collectingDate id="2359D66CFFD4E902FCE1FB559174E089" box="[885,1063,1183,1208]" pageId="42" pageNumber="43" value="2012-10-04">4 October 2012</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FB95ACFFFD4E902FF53FB319746E131" pageId="42" pageNumber="43" type="etymology">
|
||
<paragraph id="471C0944FFD4E902FF53FB319746E131" blockId="42.[151,1437,1076,2037]" pageId="42" pageNumber="43">
|
||
<emphasis id="75D7D556FFD4E902FF53FB31941DE0EC" bold="true" box="[199,334,1220,1245]" pageId="42" pageNumber="43">Etymology.</emphasis>
|
||
The name given because of very close similarity of this species to
|
||
<taxonomicName id="80A372C7FFD4E902FBD3FB3091E0E0ED" authority="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky, 2017" authorityName="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky" authorityYear="2017" box="[1095,1203,1221,1244]" class="Gymnolaemata" family="Exechonellidae" genus="Exechonella" kingdom="Animalia" order="Cheilostomatida" pageId="42" pageNumber="43" phylum="Bryozoa" rank="species" species="azeezii" status="sp. nov.">
|
||
<emphasis id="75D7D556FFD4E902FBD3FB3091E0E0ED" box="[1095,1203,1221,1244]" italics="true" pageId="42" pageNumber="43">E. azeezii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75D7D556FFD4E902FB28FB3191A8E0EC" bold="true" box="[1212,1275,1220,1245]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="6EE4682DFFD4E902FB28FB3191A8E0EC" box="[1212,1275,1220,1245]" pageId="42" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Derived from the Latin word “
|
||
<taxonomicName id="80A372C7FFD4E902FED9FB1294CFE131" authorityName="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky" authorityYear="2017" box="[333,412,1255,1280]" class="Gymnolaemata" family="Exechonellidae" genus="Exechonella" kingdom="Animalia" order="Cheilostomatida" pageId="42" pageNumber="43" phylum="Bryozoa" rank="species" species="similis">similis</taxonomicName>
|
||
” (similar).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FB95ACFFFD4E902FF53FAF99673E354" pageId="42" pageNumber="43" type="description">
|
||
<paragraph id="471C0944FFD4E902FF53FAF99673E354" blockId="42.[151,1437,1076,2037]" pageId="42" pageNumber="43">
|
||
<emphasis id="75D7D556FFD4E902FF53FAF9940AE114" bold="true" box="[199,345,1292,1317]" pageId="42" pageNumber="43">Description.</emphasis>
|
||
Colonies encrusting, unilaminar, multiserial. Zooids pentagonal or hexagonal separated by a narrow deep groove. Primary orifice almost subcircular, wider than long, poster as wide as the anter but slightly smaller in size, both predominantly of rounded outline. Anter wall underlain by an inner lamina that ends in the mid-lateral side, to form triangular or oval elongate condyles, with pointed or rounded tips extending medially and beyond the edge of the step-like curve below. Peristome is low, collar-like, slightly flared, with a pustulose external surface. The proximal edge of the peristome normally with a prominent or soft central fold-like projection. In the ancestrular area peristome could have up to four such projections, distal, proximal and two lateral. Frontal shield convex, pustulose, evenly covered with 21–36 circular or oval, well-separated foramina, with a relatively narrow raised rim with a peripheral inner wall surface and smooth or slightly wrinkled gymnocystal sloping walls, surrounding a central lumen. Fusions between foraminal rims are not common in most zooids. Small round or oval marginal pores are obvious. Vertical zooidal walls with multiporous mural septula with one row of communication pores. Lateral avicularia are frequent, developing in either or both lateralmost foramina, which are larger than of the other foramina; each avicularium is situated on the outer (often raised) edge of a foramen obvious as a shallower or deeper depression (sometimes surrounded by a low rim) with a central button-like elevation with thick walls (sometimes cylindrical with thin walls) and a central pore. Oval or triangular kenozooids are frequent, often associated with avicularia, with a frontal surface having 4–7 small pores each with centrally perforated cuticular plate. Ancestrula autozooid-like, smaller or the same size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FB95ACFFFD4E904FF53F89A94C7E52D" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43" type="discussion">
|
||
<paragraph id="471C0944FFD4E904FF53F89A94C7E52D" blockId="42.[151,1437,1076,2037]" lastBlockId="44.[151,1437,151,357]" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43">
|
||
<emphasis id="75D7D556FFD4E902FF53F89A9468E3B9" bold="true" box="[199,315,1903,1928]" pageId="42" pageNumber="43">Remarks.</emphasis>
|
||
<taxonomicName id="80A372C7FFD4E902FED6F89A974EE3B9" authority="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky, 2017" authorityName="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky" authorityYear="2017" box="[322,541,1903,1928]" class="Gymnolaemata" family="Exechonellidae" genus="Exechonella" kingdom="Animalia" order="Cheilostomatida" pageId="42" pageNumber="43" phylum="Bryozoa" rank="species" species="similis" status="sp. nov.">
|
||
<emphasis id="75D7D556FFD4E902FED6F89A974EE3B9" box="[322,541,1903,1928]" italics="true" pageId="42" pageNumber="43">Exechonella similis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6EE4682DFFD4E902FDB0F89A9730E3B6" box="[548,611,1903,1928]" pageId="42" pageNumber="43" rank="species">
|
||
<emphasis id="75D7D556FFD4E902FDB0F89A970EE3B9" bold="true" box="[548,605,1903,1928]" pageId="42" pageNumber="43">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is reminiscent of
|
||
<taxonomicName id="80A372C7FFD4E902FCB9F88496C3E3B9" authority="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky, 2017" authorityName="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky" authorityYear="2017" box="[813,912,1905,1928]" class="Gymnolaemata" family="Exechonellidae" genus="Exechonella" kingdom="Animalia" order="Cheilostomatida" pageId="42" pageNumber="43" phylum="Bryozoa" rank="species" species="azeezi" status="sp. nov.">
|
||
<emphasis id="75D7D556FFD4E902FCB9F88496C3E3B9" box="[813,912,1905,1928]" italics="true" pageId="42" pageNumber="43">E. azeezi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75D7D556FFD4E902FC0CF89A9685E3B9" bold="true" box="[920,982,1903,1928]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="6EE4682DFFD4E902FC0CF89A9685E3B9" box="[920,982,1903,1928]" pageId="42" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in most characters. The main difference between these two species is the shape of the primary orifice typically subcircular with the anter and poster having the same width in the former species, and the poster typically more angular and narrow in the latter species. Both
|
||
<taxonomicName id="80A372C7FFD4E902FA10F84C95B3E3C4" authorityName="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky" authorityYear="2017" class="Gymnolaemata" family="Exechonellidae" genus="Exechonella" kingdom="Animalia" order="Cheilostomatida" pageId="42" pageNumber="43" phylum="Bryozoa" rank="species" species="similis">
|
||
<emphasis id="75D7D556FFD4E902FA10F84C95B3E3C4" italics="true" pageId="42" pageNumber="43">E. similis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75D7D556FFD4E902FF73F829944DE3C4" bold="true" box="[231,286,2012,2037]" pageId="42" pageNumber="43">n. sp</emphasis>
|
||
and
|
||
<taxonomicName id="80A372C7FFD4E902FEC1F82894EBE3C5" authority="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky, 2017" authorityName="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky" authorityYear="2017" box="[341,440,2013,2036]" class="Gymnolaemata" family="Exechonellidae" genus="Exechonella" kingdom="Animalia" order="Cheilostomatida" pageId="42" pageNumber="43" phylum="Bryozoa" rank="species" species="azeezi" status="sp. nov.">
|
||
<emphasis id="75D7D556FFD4E902FEC1F82894EBE3C5" box="[341,440,2013,2036]" italics="true" pageId="42" pageNumber="43">E. azeezi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75D7D556FFD4E902FE2BF82994ADE3C4" bold="true" box="[447,510,2012,2037]" pageId="42" pageNumber="43">
|
||
<taxonomicNameLabel id="6EE4682DFFD4E902FE2BF82994ADE3C4" box="[447,510,2012,2037]" pageId="42" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differ from E.
|
||
<taxonomicName id="80A372C7FFD4E902FD32F8299674E3C4" authorityName=": Winston & Heimberg" authorityYear="1986" box="[678,807,2012,2037]" class="Gymnolaemata" family="Exechonellidae" genus="Exechonella" kingdom="Animalia" order="Cheilostomatida" pageId="42" pageNumber="43" phylum="Bryozoa" rank="species" species="brasiliensis">
|
||
<emphasis id="75D7D556FFD4E902FD32F8299674E3C4" box="[678,807,2012,2037]" italics="true" pageId="42" pageNumber="43">brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
by the primary orifice that is wider than long, pustulose surface of the peristome and the smaller foramina, button-like shape of the central element of avicularium, and the larger size of the lateralmost foramina (with avicularium) in comparison with the rest of the frontal foramina. The specimen from Komodo described by
|
||
<bibRefCitation id="233274B5FFD2E904FDE2FF2A96B9E4C9" author="Winston" box="[630,1002,223,248]" pageId="44" pageNumber="45" refString="Winston, J. E. & Heimberg, B. F. (1986) Bryozoans from Bali, Lombok and Komodo. American Museum Novitates, 2847, 1 - 49." type="journal article" year="1986">Winston and Heimberg (1986)</bibRefCitation>
|
||
should be additionally studied for comparative purposes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="13DC59CCFFD5E903FF03F90A91FEE3DD" httpUri="https://zenodo.org/record/842621/files/figure.png" pageId="43" pageNumber="44" targetBox="[258,1312,195,1751]" targetPageId="43">
|
||
<paragraph id="471C0944FFD5E903FF03F90A91FEE3DD" blockId="43.[151,1437,1791,2028]" pageId="43" pageNumber="44">
|
||
<emphasis id="75D7D556FFD5E903FF03F90A944CE324" bold="true" box="[151,287,1791,1813]" pageId="43" pageNumber="44">FIGURE 17.</emphasis>
|
||
<taxonomicName id="80A372C7FFD5E903FEB1F90A94B8E324" authority="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky, 2017" authorityName="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky" authorityYear="2017" box="[293,491,1791,1813]" class="Gymnolaemata" family="Exechonellidae" genus="Exechonella" kingdom="Animalia" order="Cheilostomatida" pageId="43" pageNumber="44" phylum="Bryozoa" rank="species" species="similis" status="sp. nov.">
|
||
<emphasis id="75D7D556FFD5E903FEB1F90A94B8E324" box="[293,491,1791,1813]" italics="true" pageId="43" pageNumber="44">Exechonella similis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75D7D556FFD5E903FE65F90A9779E324" bold="true" box="[497,554,1791,1813]" pageId="43" pageNumber="44">
|
||
<taxonomicNameLabel id="6EE4682DFFD5E903FE65F90A9779E324" box="[497,554,1791,1813]" pageId="43" pageNumber="44" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Great Barrier Reef, Lizard Island (A: paratype DPUV 2016-0001-0002; F, G: paratype 2016-0001-0003; B‒E, H: holotype MTQ G100216). A, general view of non-cleaned colony from above. Peripheral zooidal buds with membranous frontal wall are well seen. Peristomes with processes are seen in some zooids in the central colony part. B, E, F, close-up of autozooids. Some lateralmost foramina with avicularia shown by arrows in B. C, peripheral part of the colony, showing peristome shape and multiporous mural septula. D, close-up of autozooid showing details of primary orifice and peristome. G, non-cleaned lateralmost foramen with avicularium. Arrowhead shows an edge of mandible. H, close-up of lateralmost foramen with avicularium associated with kenozooid in cleaned colony. Pores of kenozooid have centrally perforated cuticular plates. Scale bars: A = 1 mm; B‒E = 100 µm; F = 500 µm; G = 50 µm; H = 10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0FB95ACFFFD2E904FF53FED29130E66D" pageId="44" pageNumber="45" type="distribution">
|
||
<paragraph id="471C0944FFD2E904FF53FED29742E554" blockId="44.[151,1437,151,357]" pageId="44" pageNumber="45">
|
||
<emphasis id="75D7D556FFD2E904FF53FED2940AE571" bold="true" box="[199,345,295,320]" pageId="44" pageNumber="45">Distribution</emphasis>
|
||
.
|
||
<taxonomicName id="80A372C7FFD2E904FEFEFED2971BE571" authority="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky, 2017" authorityName="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky" authorityYear="2017" box="[362,584,295,320]" class="Gymnolaemata" family="Exechonellidae" genus="Exechonella" kingdom="Animalia" order="Cheilostomatida" pageId="44" pageNumber="45" phylum="Bryozoa" rank="species" species="similis" status="sp. nov.">
|
||
<emphasis id="75D7D556FFD2E904FEFEFED2971BE571" box="[362,584,295,320]" italics="true" pageId="44" pageNumber="45">Exechonella similis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75D7D556FFD2E904FDC7FED297C6E571" bold="true" box="[595,661,295,320]" pageId="44" pageNumber="45">
|
||
<taxonomicNameLabel id="6EE4682DFFD2E904FDC7FED297C6E571" box="[595,661,295,320]" pageId="44" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a west Pacific distribution including the areas of
|
||
<collectingCountry id="3FB449D4FFD2E904FA99FEDD903AE50E" box="[1293,1385,296,319]" name="Vanuatu" pageId="44" pageNumber="45">Vanuatu</collectingCountry>
|
||
and Lizard Island, Great Barrier Reef.
|
||
</paragraph>
|
||
<caption id="13DC59CCFFD2E904FF03FE619482E61A" pageId="44" pageNumber="45">
|
||
<paragraph id="471C0944FFD2E904FF03FE619482E61A" blockId="44.[151,1436,403,555]" pageId="44" pageNumber="45">
|
||
<emphasis id="75D7D556FFD2E904FF03FE619448E59A" bold="true" box="[151,283,404,427]" pageId="44" pageNumber="45">TABLE 16.</emphasis>
|
||
Measurements (in µm, except number of foramina) of holotype specimen of
|
||
<taxonomicName id="80A372C7FFD2E904FBEAFE669002E59A" authority="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky, 2017" authorityName="Cáceres-Chamizo & Sanner & Tilbrook & Ostrovsky" authorityYear="2017" box="[1150,1361,403,427]" class="Gymnolaemata" family="Exechonellidae" genus="Exechonella" kingdom="Animalia" order="Cheilostomatida" pageId="44" pageNumber="45" phylum="Bryozoa" rank="species" species="similis" status="sp. nov.">
|
||
<emphasis id="75D7D556FFD2E904FBEAFE669002E59A" box="[1150,1361,403,427]" italics="true" pageId="44" pageNumber="45">Exechonella similis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="75D7D556FFD2E904FAC8FE6190CFE59A" bold="true" box="[1372,1436,404,427]" pageId="44" pageNumber="45">
|
||
<taxonomicNameLabel id="6EE4682DFFD2E904FAC8FE6190CFE59A" box="[1372,1436,404,427]" pageId="44" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Abbreviations: ancestrula length (AncL), ancestrula width (AncW), autozooid length (AzL), autozooid width (AzW), diameter of a foramen including rim (FoD), number of frontal foramina (FoN), diameter of the opening of a foramen (OD), primary orifice length (OrL), primary orifice width (OrW). Mean (m), standard deviation (sd), range (r) and number of measurements (n).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="471C0944FFD2E904FC80FDB39130E66D" blockId="44.[788,1123,582,604]" box="[788,1123,582,604]" pageId="44" pageNumber="45">Lizard Island, Great Barrier Reef</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |