205 lines
30 KiB
XML
205 lines
30 KiB
XML
<document ID-CLB-Dataset="21464" ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="CEE3427E42F352E19E8335E0A879E507" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole joffreville Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="2290CAFF-30DF-4911-8EEC-7AF36C7A79CE" docUuidSource="ZooBank" docVersion="6" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1732651982216" updateUser="ExternalLinkService">
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<mods:titleInfo id="2194C27C418CC63E4AA5A531A24E4CBA">
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<mods:title id="610A2B95E921FA8091A11B79FE30F243">Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
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<mods:name id="F78A8D38EA1AB10B49AD3E73749850C7" type="personal">
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<mods:roleTerm id="799C327CD5ECB2B04C587C296413BDDF">Author</mods:roleTerm>
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<mods:namePart id="E35BDA83B169B2BC587137ED474F2C9C">Salata, Sebastian</mods:namePart>
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<mods:affiliation id="4E5352164B06DC9C991E0114179A37A2">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="9F9C4E1AF01A06E82AC53C1379AB3407" type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
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<mods:nameIdentifier id="2556BD5570F22301DC673B50E8EA48FF" type="email">sdsalata@gmail.com</mods:nameIdentifier>
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<mods:namePart id="9C48E6EB1883CAEAE2857DF87439419D">Fisher, Brian L.</mods:namePart>
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<mods:affiliation id="FFE558C4B07BD89F36D65F5E2F6ED257">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="03E865157E902BA53345EA75F8EB15AB" type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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<mods:typeOfResource id="D67606F5008B51BB79291FCBED183A32">text</mods:typeOfResource>
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<mods:title id="C6233F24F92E7B49B3CF4979098CA401">ZooKeys</mods:title>
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<mods:date id="27C96E904D1D07A89B5117E3525E3B70">2020</mods:date>
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<mods:detail id="D22EF0F1452A75AF9A6941EF4AC84FE1" type="volume">
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<mods:number id="9148BCB08734D5ED7A90CFC35A8F959C">949</mods:number>
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<mods:start id="BD6FA8B2C11C97B95208B191954BF4AD">1</mods:start>
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<mods:location id="79E62B763A8F8763113BA680F903432F">
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<mods:url id="5250C1CAAC5B70D1CCC14E7244AE603A">http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
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<mods:classification id="AEEF191C0521DDDC08954537F9B20FBA">journal article</mods:classification>
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<mods:identifier id="BF20AE3C3D8FE6B1FCA73167100F365A" type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
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<mods:identifier id="16B43C917BC4BC2DF02FACC4D6111286" type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
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<treatment id="CEE3427E42F352E19E8335E0A879E507" ID-GBIF-Taxon="165090160" LSID="urn:lsid:zoobank.org:act:2290CAFF-30DF-4911-8EEC-7AF36C7A79CE" httpUri="http://treatment.plazi.org/id/CEE3427E42F352E19E8335E0A879E507" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="EC6B7BCB94D086860B2C3308E896300A" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="8D86A18B6CF3A727D25E73F016189D7F" pageId="0" pageNumber="1">
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<taxonomicName id="34F699657AE20A66C7F4C12FD65DCF60" ID-CoL="CMLC4" LSID="CEE3427E-42F3-52E1-9E83-35E0A879E507" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole joffreville" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="joffreville" status="sp. nov.">Pheidole joffreville</taxonomicName>
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<taxonomicNameLabel id="636293C5678050283B53DEDA27225392" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="280C7F85A69D9B5DF0775F3B6592546B" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Pheidole joffreville sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0406352) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0406355)." figureDoi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1">Figs 35A-F</figureCitation>
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<figureCitation id="3A99B6FCD9563793E35FFFB047DA3EA0" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">, 63Q</figureCitation>
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<figureCitation id="12ADC7F898150C8AA02F87954CD786DD" captionStart="Figure 65" captionStartId="F65" captionText="Figure 65. Distribution. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T)." figureDoi="10.3897/zookeys.949.51269.figure65" httpUri="https://binary.pensoft.net/fig/432234" pageId="0" pageNumber="1">, 65Q</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="A0BB169C419515FDA9FD5AFAC53CF21B" pageId="0" pageNumber="1" type="type material">
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<paragraph id="12151A972D595EAC5BCF912A10B644ED" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="357867ACA3A4FB1A296B454B9621607F" pageId="0" pageNumber="1">
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<emphasis id="FB5A4890BD13B1B3ED63FFEE67F66B7E" bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Madagascar. • 1 major worker; Antsiranana; Parc National Montagne
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<normalizedToken id="4ED89672F97B60EA48F62027A9AC5580" originalValue="d’Ambre">d'Ambre</normalizedToken>
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, 3.6 km 235°SW Joffreville;
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<geoCoordinate id="C628EE6F58496FC3397394EB63BC4839" degrees="12.53444" direction="south" orientation="latitude" precision="1" value="-12.53444">-12.53444</geoCoordinate>
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,
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<geoCoordinate id="C780615D6571413F0EE8B7C1CBAF33B3" degrees="49.1795" direction="east" orientation="longitude" precision="5" value="49.1795">49.1795</geoCoordinate>
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; alt. 925 m; 20 Jan 2001; Fisher et al. leg.; montane rainforest, ex rotten log; BLF02568; CASENT0406355 (CASC).
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<emphasis id="A8986B7909D197D967533D60AD0CD942" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes.</emphasis>
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• 4w., 2s.; same data as for holotype, CASENT0406358, CASENT0406356, CASENT0406352, CASENT0406354, CASENT0406350 (CASC, MHNG).
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</paragraph>
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<caption id="C17017DC0A90D7CF9AD6C40FC53C3FE2" doi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1" start="Figure 35" startId="F35">
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<paragraph id="DAD4C692021E503B231AC44E79E31FDC" pageId="0" pageNumber="1">
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<emphasis id="ABF25FA6EA998A0A5C01CFDF80D0E818" bold="true" pageId="0" pageNumber="1">Figure 35.</emphasis>
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<taxonomicName id="8BD7D780100FEC025438E88D2EA65083" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole joffreville" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="joffreville">
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<emphasis id="BEFEA8B650CFB4F1BA7057086859DFDA" italics="true" pageId="0" pageNumber="1">Pheidole joffreville</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis id="322224FCE4D76946BAFCBACB57C7ED58" bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis id="5F5EFAD94CA45AAF4F2935900B47B514" bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis id="A8842EA01EFCE0A0822BD3C734D808DC" bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0406352) and full-face view (
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<emphasis id="68158D1E66B3C9C3D52657D8BDBF69CB" bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis id="40DC996C852FFA0E50B570624A229732" bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis id="48BB81CDD246D681BF977D365A95A61E" bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0406355).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="0E99FC778BC94007C842080B7E2BEDCA" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="65747B2E237818D62D6A33607EF4D0F8" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="5CE565BF427992FC9FF37747FF25A38D" pageId="0" pageNumber="1">
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Moderately large species.
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<emphasis id="D58134BDE9A5F20BB6E3811ED14ECAF1" bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Head in full-face view sub-oval and widening slightly posteriorly, with anterior and posterior sides slightly convex, in lateral view sub-oval; ventral and dorsal faces convex; sides of the head with sparse, moderately long, suberect pilosity; anteromedial part of frons with thick, longitudinal, interrupted, and sparse rugae, posteromedial frons with rugae longitudinally irregular, interspaces distinctly rugofoveolate, rugae on posteromedial part directed outward; lateral sides with thick, sparse, and irregular rugae with distinctly rugofoveolate interspaces; occipital lobes, and area posterolateral from eyes without smooth notches; scape, when laid back, exceeding the midlength of head by one-fifth of its length; inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, narrower, and approximately as high as inner teeth; inner and outer hypostomal teeth closely spaced and not connected by concavity; mesosoma foveolate; promesonotum with additional sparse, thick, and transverse rugulae on dorsum; gaster smooth; body brown.
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<emphasis id="9BBDC65BB2E418C2F7CEF7E1707AC06B" bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Head foveolate; vertex with a few arcing, sparse, and moderately thick rugae; scape, when laid back, exceeding the posterior head margin by one-third of its length; promesonotum low and moderately long, arched; promesonotal groove present; propodeal spines moderate, triangular; mesosoma foveolate; katepisternum with big smooth notch; body brown.
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</paragraph>
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</subSubSection>
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<subSubSection id="363E59D63552515B3F0A88E0A0D0FB67" pageId="0" pageNumber="1" type="description">
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<paragraph id="EF7FB98A01A516AE394779FD194694E0" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="96188FC2971B57D684812AD640520A02" pageId="0" pageNumber="1">
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<emphasis id="F41A692A4FA91463A6B5C6A741CC6055" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis id="D04510AB0E49402606829D49CACCCEB2" italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 3): HL: 1.04-1.1 (1.06); HW: 1.03-1.16 (1.06); SL: 0.73-0.75 (0.74); EL: 0.14-0.15 (0.14); WL: 0.9-0.93 (0.92); PSL: 0.17-0.19 (0.17); MTL: 0.62-0.67 (0.64); PNW: 0.44-0.47 (0.45); PTW: 0.15-0.15 (0.15); PPW: 0.31-0.34 (0.32); CI: 98.6-101.0 (100.1); SI: 67.3-70.9 (69.3); PSLI: 15.4-17.9 (16.3); PPI: 44.4-47.3 (46.3); PNI: 42.2-43.0 (42.5); MTI: 59.7-60.1 (59.9).
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</paragraph>
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<paragraph id="69C4EF78BD73E70B0CF97F1F10F5090F" pageId="0" pageNumber="1">
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<emphasis id="18B207945158D1F82F7DD66BF7084F33" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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In full-face view sub-oval, widening posteriorly, with anterior and posterior sides convex (Fig.
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<figureCitation id="D86D82D19189A4EFA523E89035F75723" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Pheidole joffreville sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0406352) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0406355)." figureDoi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1">35B</figureCitation>
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). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with sparse, moderately long, suberect pilosity; whole head with dense, long, decumbent to erect pilosity. Anteromedial part of frons with thick, longitudinal, interrupted and sparse rugae, posteromedial frons with rugae longitudinally irregular, interspaces distinctly rugofoveolate, rugae on posteromedial part directed outward; lateral sides with thick, sparse, and irregular rugae with distinctly rugofoveolate interspaces. Occipital lobes with thinner and denser irregular rugae and rugofoveolate interspaces. Area posterolateral from eyes with dense rugofoveolae and additional sparse rugae. Gena with relatively sparse, thick, longitudinal rugae and distinctly rugofoveolate interspaces. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina present; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by one-fifth of its length; pilosity subdecumbent to erect (Fig.
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<figureCitation id="51E6FEF1C336911D26603421C6F2E12C" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Pheidole joffreville sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0406352) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0406355)." figureDoi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1">35B, D</figureCitation>
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). Inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, narrower, and approximately as high as inner teeth; inner and outer hypostomal teeth closely spaced and not connected by concavity (Fig.
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<figureCitation id="EFFBF82EF7FF0254A1EC6C349CE9435A" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">63Q</figureCitation>
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).
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<emphasis id="970301E269CB800E5129FA5593CFE92C" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
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In lateral view, promesonotum short, angular, and moderately low, posterior mesonotum moderately steep, mesonotal process indistinct, tubercle-like; promesonotal groove absent; metanotal groove absent; propodeal spines moderate, with wide base and acute apex; humeral area produced (Fig.
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<figureCitation id="95A7074E1D58EC81962BDA23120B06FF" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Pheidole joffreville sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0406352) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0406355)." figureDoi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1">35D</figureCitation>
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). Surface shiny and foveolate; promesonotum with additional sparse, thick, and transverse rugulae on dorsum. Pilosity moderately sparse, long, and erect (Fig.
|
||
<figureCitation id="88ECE6343DB112076C8A8CEC602EDB03" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Pheidole joffreville sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0406352) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0406355)." figureDoi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1">35D, F</figureCitation>
|
||
).
|
||
<emphasis id="3100D9F5DFC65DE1442A0E6793C19DF7" bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
|
||
Shiny with dense foveolae; node finely foveolate, triangular, with rounded and thick apex, in rear view node dorsoventrally slightly convex; pilosity moderately sparse and erect (Fig.
|
||
<figureCitation id="1E68950DCEF36DC5D7CB367EAD494D81" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Pheidole joffreville sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0406352) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0406355)." figureDoi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1">35D, F</figureCitation>
|
||
).
|
||
<emphasis id="D24545AB695CA3C47B04352ABC34E82F" bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
|
||
Shiny and foveolate; dorsum with reduced sculpture and smooth notch; in dorsal view oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse, and erect (Fig.
|
||
<figureCitation id="6CA9BB852FE0C91A8961A0053B9370E3" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Pheidole joffreville sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0406352) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0406355)." figureDoi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1">35D, F</figureCitation>
|
||
).
|
||
<emphasis id="BD2FEAA6B4A58BDC3B994F3D13E382E0" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
Shiny and smooth; pilosity moderately sparse, long, and erect (Fig.
|
||
<figureCitation id="1741AB302F5B86C1BCACD6195CD4C363" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Pheidole joffreville sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0406352) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0406355)." figureDoi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1">35D, F</figureCitation>
|
||
).
|
||
<emphasis id="9484BFBC6127B6109140497B9F4BA820" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Brown with yellowish legs (Fig.
|
||
<figureCitation id="00ABA784DB4049307C51A94E9FCAA333" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Pheidole joffreville sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0406352) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0406355)." figureDoi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1">35D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="CC5603AF278F372027F0F1C54C2625E9" pageId="0" pageNumber="1">
|
||
<emphasis id="F9F0C8338AA9588E54E03843CB5E218B" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis id="2E18ED594A3E51A8DF595F68BFA9624D" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 4): HL: 0.6-0.63 (0.62); HW: 0.52-0.54 (0.53); SL: 0.7-0.74 (0.72); EL: 0.13-0.13 (0.13); WL: 0.74-0.8 (0.76); PSL: 0.11-0.12 (0.11); MTL: 0.53-0.55 (0.54); PNW: 0.35-0.37 (0.36); PTW: 0.09-0.1 (0.09); PPW: 0.14-0.15 (0.15); CI: 116.0-120.5 (117.6); SI: 133.0-138.5 (136.2); PSLI: 17.6-18.9 (18.3); PPI: 58.6-67.8 (63.7); PNI: 66.9-70.2 (68.3); MTI: 99.3-105.6 (102.4).
|
||
</paragraph>
|
||
<paragraph id="357742C07B7CC13FF9B252450E7DBD05" pageId="0" pageNumber="1">
|
||
<emphasis id="C8A6EEBD86C9B10A2FF8A8F997DB748A" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
|
||
Cephalic margin slightly convex (Fig.
|
||
<figureCitation id="B282A2FD485359DDC1A30CAEC5B4692B" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Pheidole joffreville sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0406352) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0406355)." figureDoi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1">35A</figureCitation>
|
||
). Pilosity relatively sparse, moderately long, subdecumbent to erect. Sculpture shiny and foveolate; vertex with a few arcing, sparse, and moderately thick rugae; antennal sockets with few thick, curved outward rugae and foveolate interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, exceeding the posterior head margin by one-third of its length; pilosity dense, subdecumbent to erect (Fig.
|
||
<figureCitation id="7BB07E563CD66671A5D0595A399D57E3" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Pheidole joffreville sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0406352) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0406355)." figureDoi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1">35A, C</figureCitation>
|
||
).
|
||
<emphasis id="4D6D89ACA1D8E61693F7039F0B533976" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum low and moderately long, arched; promesonotal groove present; metanotal groove distinct; propodeal spines moderate, triangular (Fig.
|
||
<figureCitation id="288376460F7BF5B96E0B3DFD77BF579D" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Pheidole joffreville sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0406352) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0406355)." figureDoi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1">35C</figureCitation>
|
||
). Sculpture shiny and foveolate; katepisternum with big smooth notch. Pilosity very sparse, moderately long, and erect (Fig.
|
||
<figureCitation id="78DF76B6C7CDA170A0D7F4638D1CA87E" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Pheidole joffreville sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0406352) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0406355)." figureDoi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1">35C, E</figureCitation>
|
||
).
|
||
<emphasis id="DCF50F806D1D6577CEFC5858A83EEC26" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
With sparse, erect pilosity (Fig.
|
||
<figureCitation id="D66CFB63FB31B047605EB700381D7981" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Pheidole joffreville sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0406352) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0406355)." figureDoi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1">35C, E</figureCitation>
|
||
).
|
||
<emphasis id="98E52F4EEE9CFB78E7310A70C23C5BC0" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Brown, legs, antenna and gaster yellowish (Fig.
|
||
<figureCitation id="E3EB028ACDE2C118B40B56FF7D194DD5" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Pheidole joffreville sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0406352) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0406355)." figureDoi="10.3897/zookeys.949.51269.figure35" httpUri="https://binary.pensoft.net/fig/432204" pageId="0" pageNumber="1">35C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4F384A9A2A60B86F13198FEF45FBD329" pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph id="EE395AA33C453508B3EA23840DBD2E0E" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph id="44A8FB79EAA1EE9D507DC8F45E6AA140" pageId="0" pageNumber="1">From the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="068A44A93BA843944B592F2E746C8F37" pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph id="FD95FC249732B15FF88BD50940564E63" pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph id="5B2A376A74ABF987BC13BD544F12EF96" pageId="0" pageNumber="1">The species was collected at 925 m in elevation, in montane rainforest. Nest was located in rotten log.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5D309446796B4819C6205C759C9EABCD" pageId="0" pageNumber="1" type="comments">
|
||
<paragraph id="F23C487EDCDADB5C1FC35AAAE9D44DFC" pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph id="57E319087FE4FFF6ACF76B1109E83488" pageId="0" pageNumber="1">
|
||
<taxonomicName id="8932FEFF6323C62DF55C423836114873" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole joffreville" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="joffreville">
|
||
<emphasis id="842402A6859FAFEDC31F84F9C0927598" italics="true" pageId="0" pageNumber="1">Pheidole joffreville</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., described from Parc National Montagne
|
||
<normalizedToken id="763464B5A5D68D4828460979134E6952" originalValue="d’Ambre">d'Ambre</normalizedToken>
|
||
in Antsiranana, has major workers with dense and thick rugae, anteromedially longitudinal and posteromedially irregular, with distinctly rugofoveolate interspaces, and brown body colouration. Morphologically is most similar to allopatric
|
||
<taxonomicName id="1ECDB86464DED9D71F14D61705B0C419" lsidName="P. mivory" pageId="0" pageNumber="1" rank="species" species="mivory">
|
||
<emphasis id="0AEE7EBD68F4E871084AF5B2A8A8F8B4" italics="true" pageId="0" pageNumber="1">P. mivory</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., known from Parc National de Marojejy in Antsiranana. Majors of both taxa are extremely similar and species separation should be supported or based exclusively on minors. Majors of
|
||
<taxonomicName id="4D39E005B0304231CCD5FD7DD4681526" lsidName="P. joffreville" pageId="0" pageNumber="1" rank="species" species="joffreville">
|
||
<emphasis id="92EF0F927791E7330CF9DC978C08602C" italics="true" pageId="0" pageNumber="1">P. joffreville</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differ from
|
||
<taxonomicName id="1044B298DF7DBF573A75AD630092930C" lsidName="P. mivory" pageId="0" pageNumber="1" rank="species" species="mivory">
|
||
<emphasis id="0E6DEBA5AD81CBE1341B415960E0FA34" italics="true" pageId="0" pageNumber="1">P. mivory</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in distinctly rugofoveolate interspaces on frons and lack of longitudinal rugae on lateral sides of frons and lack of shagreened sculpture on first gastral tergite; minor workers differ in distinctly foveolate mesosoma with only katepisternum with smooth notch.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |