295 lines
38 KiB
XML
295 lines
38 KiB
XML
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<mods:title id="BE276EBCD63C018B8391BBA86DDC8224">A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species</mods:title>
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<mods:namePart id="A348DF0ABA9F9D0A3EB3BE58C42113F5">Gloeer, Peter</mods:namePart>
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<mods:affiliation id="299E6A1A70EA2339D66ADF644289D3E7">Biodiversity Research Laboratory, Hetlingen, Germany</mods:affiliation>
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<mods:namePart id="F0F4D348F418C4244F6A7C9C1CDF63C0">Guerlek, Mustafa E</mods:namePart>
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<mods:affiliation id="B11641830151D62082A020B6FAD79892">Mehmet Akif Ersoy niversity, Burdur, Turkey</mods:affiliation>
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<mods:namePart id="478CC9AE9CE3F63916250406B176E998">Neubauer, Thomas A</mods:namePart>
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<mods:affiliation id="2C44BA13B27FB34465CE28A54C80A9FF">Justus Liebig University Giessen, Giessen, Germany & Naturalis Biodiversity Centre, Leiden, Netherlands</mods:affiliation>
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<treatment id="CE9DBC3ED0E85E748C24DE140FBF9B83" ID-GBIF-Taxon="161939041" LSID="urn:lsid:plazi:treatment:CE9DBC3ED0E85E748C24DE140FBF9B83" httpUri="http://treatment.plazi.org/id/CE9DBC3ED0E85E748C24DE140FBF9B83" lastPageNumber="25" pageId="0" pageNumber="25" scope_class="Gastropoda" scope_family="Neritidae" scope_subFamily="Neritininae" scope_superFamily="Neritoidea">
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<subSubSection id="D5CAE94F325491333E43F21FA601C7C8" pageId="0" pageNumber="25" type="nomenclature">
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<paragraph id="CF51A79A4FA58FE9B550166565C2572C" pageId="0" pageNumber="25">
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<taxonomicName id="F286F8BE6C67C4A3F57B3C5C5EBDB8B0" ID-CoL="7BVNJ" LSID="CE9DBC3E-D0E8-5E74-8C24-DE140FBF9B83" authority="(G. B. Sowerby II, 1849)" baseAuthorityName="G. B. Sowerby II" baseAuthorityYear="1849" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus macri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="macri">Theodoxus macri (G.B. Sowerby II, 1849)</taxonomicName>
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<figureCitation id="FF043D9FA65E2F503258AAB51472C027" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Theodoxus macri (G. B. Sowerby II, 1849). A - D. Specimen collected close to Harbiye Falls, Harbiye, Antakya, Turkey (UGSB 24179); E - H. Specimen from Lake Yenisehir, Reyhanli, Turkey (UGSB 24180); I, J. Specimen from Lake Balik, Adalar, Turkey (UGSB 24181). All specimens (A-J) were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure18" httpUri="https://binary.pensoft.net/fig/376467" pageId="0" pageNumber="25">Figure 18A-H</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="224C0F8D542B3498256B6DDC7F658277" pageId="0" pageNumber="25" type="reference_group">
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<paragraph id="2F6F84B94559C5A5BB23604FCEF6412A" pageId="0" pageNumber="25">
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<taxonomicName id="51465CC3F05B973A13E6AD6243B53575" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina Macri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="Macri">Neritina Macri</taxonomicName>
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G.B. Sowerby
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<date id="0A79ACFF7DB524F2084059183A8EBD8F" value="1849-02">II 1849</date>
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: 531, pl. 116, fig. 222.
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</paragraph>
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<paragraph id="9791EBE9975D3D4197772707FBC37C07" pageId="0" pageNumber="25">
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<taxonomicName id="8D32CA7B4F9EFD5D926589FA919E8920" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
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Macrii [sic]:
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<bibRefCitation id="A33BE398347780AE28A34E459BEDB4AC" author="Martens, Ev" journalOrPublisher="Bauer & Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
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||
: 88-90, pl. 13, fig. 13 (partim, non pl. 4, figs 11-13, pl. 13, figs 27-29); Blanckenhorn 1889: 81 (synonymy tentative);
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<bibRefCitation id="0BE8D6F697FE572EBC3C8EDCCA318D65" author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
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||
: 5, pl. 212, figs 1327, 1328 (partim, excluding synonyms).
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</paragraph>
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<paragraph id="612929CA68F3D7AC8FA176328B3FAD37" pageId="0" pageNumber="25">
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non
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<taxonomicName id="7B3DE3598B2C48DE6F7964A1591E1510" class="Gastropoda" family="Neritidae" genus="Theodoxia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxia" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Theodoxia</taxonomicName>
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Macrii [sic]:
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<bibRefCitation id="405FD59E46DBE0F543C656500D41292D" author="Germain, L" journalOrPublisher="Bailliere et fils, Paris" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/13187112" refId="B59" refString="Germain, L, 1921. Mollusques terrestres et fluviatiles de Syrie. Tome premier - Introduction et Gasteropodes. J.-B. Bailliere et fils, Paris, https://biodiversitylibrary.org/page/13187112" title="Mollusques terrestres et fluviatiles de Syrie. Tome premier - Introduction et Gasteropodes. J. - B." url="https://biodiversitylibrary.org/page/13187112" year="1921">Germain 1921</bibRefCitation>
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||
: 516-518.
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</paragraph>
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||
<paragraph id="9D67E24A8129FD41D3FFB24971F7CA1F" pageId="0" pageNumber="25">
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non
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<taxonomicName id="37FA62C3D401AD53D32ECD19E2750BC9" baseAuthorityName="Sowerby" baseAuthorityYear="1849" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus macrii" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="macrii">Theodoxus macrii</taxonomicName>
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[sic]:
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<bibRefCitation id="3EF1B04D7E3DC3D193CFE0DCA2537F4D" author="Amr, ZS" journalOrPublisher="Denisia" pageId="0" pageNumber="25" pagination="221 - 227" publicationUrl="http://www.zobodat.at/stable/pdf/DENISIA_0014_0221-0227.pdf" refId="B10" refString="Amr, ZS, Abu Baker, M, 2004. Freshwater snails of Jordan. Denisia 14: 221 - 227, http://www.zobodat.at/stable/pdf/DENISIA_0014_0221-0227.pdf" title="Freshwater snails of Jordan." url="http://www.zobodat.at/stable/pdf/DENISIA_0014_0221-0227.pdf" volume="14" year="2004">Amr and Abu Baker 2004</bibRefCitation>
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||
: 222, fig. 2;
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<bibRefCitation id="A729845DE993376594E7EE2DF3E09299" author="Handal, EN" journalOrPublisher="Jordan Journal of Natural History" pageId="0" pageNumber="25" pagination="23 - 29" refId="B70" refString="Handal, EN, Amr, Z, Qumsiyeh, MB, 2015. Some records of freshwater snail from the Occupied Palestinian territories. Jordan Journal of Natural History 1 (2): 23 - 29" title="Some records of freshwater snail from the Occupied Palestinian territories." volume="1" year="2015">Handal et al. 2015</bibRefCitation>
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: 25-26, fig. 1B.
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</paragraph>
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<paragraph id="79CA125DD85BF9612DE4B915AA2EB006" pageId="0" pageNumber="25">
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?
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<taxonomicName id="C50BD5C063B303615E307D431E3694A9" baseAuthorityName="Sowerby" baseAuthorityYear="1849" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus macrii" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="macrii">Theodoxus macrii</taxonomicName>
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[sic]:
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<bibRefCitation id="84071C97061E6A7C5B862D7BAF04FCDA" author="Gloeer, P" journalOrPublisher="Peter Gloeer, Hetlingen, Germany" pageId="0" pageNumber="25" refId="B62" refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany" title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails." year="2019">
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||
<normalizedToken id="9707383C1D96A57649F3F5518F2F9C78" originalValue="Glöer">Gloeer</normalizedToken>
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||
2019
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||
</bibRefCitation>
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||
: 46, fig. 32.
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="7126B0652E20DD12F72AB56388B4ECEC" pageId="0" pageNumber="25" type="type locality">
|
||
<paragraph id="FF0345824FEF5FC5D2AE6B3669EC0574" pageId="0" pageNumber="25">
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<typeStatus id="BC43D6C169DB7F3003E412E3271DBB62">Type</typeStatus>
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locality.
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</paragraph>
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<paragraph id="DAF05D7F854EA235EC84E126BCDAFAA3" pageId="0" pageNumber="25">Asia Minor (= Anatolia).</paragraph>
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</subSubSection>
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<subSubSection id="D2E5E289068939E3D6C9E210AC8A9153" pageId="0" pageNumber="25" type="type material">
|
||
<paragraph id="FD0F650FE431ACEB8141BFBB69CB7428" pageId="0" pageNumber="25">
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<typeStatus id="0BBBBC59DEF98F9FCED22579C4C3B214">Type</typeStatus>
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material.
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</paragraph>
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<paragraph id="10B63A120683BD35016F7CBFE16A1CE4" pageId="0" pageNumber="25">
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According to
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<bibRefCitation id="E62102D22C6BF2F5B79DDD67FBFAA46F" author="Dance, SP" journalOrPublisher="Faber and Faber, London" pageId="0" pageNumber="25" refId="B41" refString="Dance, SP, 1966. Shell collecting. An illustrated history. Faber and Faber, London" title="Shell collecting. An illustrated history." year="1966">Dance (1966)</bibRefCitation>
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||
, most material described by G.B. Sowerby II is deposited in
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<collectionCode id="8486A26CA0220182978B15D23BE4E893" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]">BMNH</collectionCode>
|
||
, and parts also in the Tomlin Collection of
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<collectionCode id="E6B44866DE0A22EE5978D2F8F10F38CB" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" type="">NMW</collectionCode>
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. Unfortunately, the
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<typeStatus id="08C612AF554539B0BD11A01828C593DE">types</typeStatus>
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could not been located in either of these two institutions (A. Salvador, H. Wood pers. comm. 07/2018).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="5768B5F7227CA68577E9AE48A1746148" pageId="0" pageNumber="25" type="remarks">
|
||
<paragraph id="486D79697047F30A6EA8FA64E55DB95F" pageId="0" pageNumber="25">Remarks.</paragraph>
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<paragraph id="706E7B7548E50C6D4FAC7E1DF2517180" pageId="0" pageNumber="25">
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The identity of this species is doubtful at the moment. G.B. Sowerby II (1849) based it on a black periostracum, oval shell, and a grey, more inclined columellar plate. The short description and sole figure, as well as the imprecise locality information ("Asia Minor") and the apparent lack of
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<typeStatus id="25E4F68713D902ED851E7A5F56471911">type</typeStatus>
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||
material, render an attribution of newly collected specimens to that species uncertain.
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<bibRefCitation id="9BD980FFA1B26C87004B5F4D861711E9" DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al.'s (2019a)</bibRefCitation>
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material from eastern Cilicia and south-east Anatolia fit well in terms of shell shape and periostracum colouration, and we tentatively consider them to belong to
|
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<taxonomicName id="F167E31D3860A97AA50E7E14DA8B6FAA" lsidName="T. macri" pageId="0" pageNumber="25" rank="species" species="macri">
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<emphasis id="815C0DC7F06F2C9C54231630119FACA2" italics="true" pageId="0" pageNumber="25">T. macri</emphasis>
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||
</taxonomicName>
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(Fig.
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<figureCitation id="B28A7E5587F2400EF2083D59AFB5C557" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Theodoxus macri (G. B. Sowerby II, 1849). A - D. Specimen collected close to Harbiye Falls, Harbiye, Antakya, Turkey (UGSB 24179); E - H. Specimen from Lake Yenisehir, Reyhanli, Turkey (UGSB 24180); I, J. Specimen from Lake Balik, Adalar, Turkey (UGSB 24181). All specimens (A-J) were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure18" httpUri="https://binary.pensoft.net/fig/376467" pageId="0" pageNumber="25">18</figureCitation>
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). The previous perception of
|
||
<taxonomicName id="4331EA4C4F6192B8A9A88ABD6F524CA8" lsidName="T. macri" pageId="0" pageNumber="25" rank="species" species="macri">
|
||
<emphasis id="3C0EFF647FDF2FCE2A33C44A5F8F3979" italics="true" pageId="0" pageNumber="25">T. macri</emphasis>
|
||
</taxonomicName>
|
||
was a different one though. In the literature of the late 19th century, it has been commonly considered a more widely distributed Middle Eastern species, with the junior synonyms
|
||
<taxonomicName id="B16CB0787D34EEACB362ED36A882EB2B" lsidName="N. karasuna" pageId="0" pageNumber="25" rank="species" species="karasuna">
|
||
<emphasis id="9AD51788D2480BB78174AA71D6A398C6" italics="true" pageId="0" pageNumber="25">N. karasuna</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6E743981354B6EDDA53167CA6C8B36BD" lsidName="N. michonii" pageId="0" pageNumber="25" rank="species" species="michonii">
|
||
<emphasis id="5803F276F708FEA13A6552111BE08454" italics="true" pageId="0" pageNumber="25">N. michonii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="E021F33FEE31F04C305A5DF54C2221E6" author="Martens, Ev" journalOrPublisher="Bauer & Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="279230BEEFA239ADE6BB8CF9CFA99EC2" author="Westerlund, CA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/10887359" refId="B160" refString="Westerlund, CA, 1886. Fauna der in der palaearctischen Region (Europa, Kaukasien, Sibirien, Turan, Persien, Kurdistan, Armenien, Mesopotamien, Kleinasien, Syrien, Arabien, Egypten, Tripolis, Tunesien, Algerien und Marocco) lebenden Binnenconchylien. VI. Fam. Ampullariidae, Paludinidae, Hydrobiidae, Melaniidae, Valvatidae & Neritidae. Hakan Ohlsson, Lund, 156 + 13 pp. https://biodiversitylibrary.org/page/10887359" title="Fauna der in der palaearctischen Region (Europa, Kaukasien, Sibirien, Turan, Persien, Kurdistan, Armenien, Mesopotamien, Kleinasien, Syrien, Arabien, Egypten, Tripolis, Tunesien, Algerien und Marocco) lebenden Binnenconchylien. VI. Fam. Ampullariidae, Paludinidae, Hydrobiidae, Melaniidae, Valvatidae & Neritidae. Hakan Ohlsson, Lund, 156 + 13 pp." url="https://biodiversitylibrary.org/page/10887359" year="1886">Westerlund 1886</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="0747EE236EFDC969F9DBD8110C64E715" author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
|
||
). More recently,
|
||
<bibRefCitation id="FF10E972FBE124B8018D101A1DA8FF52" author="Dagan, D" journalOrPublisher="Israel Journal of Zoology" pageId="0" pageNumber="25" pagination="223 - 229" refId="B39" refString="Dagan, D, 1971. Taxonomic discrimination between certain species of the genus Theodoxus (Gastropoda, Neritidae). Israel Journal of Zoology 20 (3): 223 - 229" title="Taxonomic discrimination between certain species of the genus Theodoxus (Gastropoda, Neritidae)." volume="20" year="1971">Dagan (1971)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="2D6E44AD76412E65FB000C57A4E9257C" author="Tchernov, E" journalOrPublisher="Malacologia" pageId="0" pageNumber="25" pagination="147 - 184" publicationUrl="https://biodiversitylibrary.org/page/13122806" refId="B154" refString="Tchernov, E, 1975. The molluscs of the Sea of Galilee. Malacologia 15 (1): 147 - 184, https://biodiversitylibrary.org/page/13122806" title="The molluscs of the Sea of Galilee." url="https://biodiversitylibrary.org/page/13122806" volume="15" year="1975">Tchernov (1975)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="D6CB692768735B9CF6C6EEC541A0E56E" author="Bandel, K" journalOrPublisher="Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg" pageId="0" pageNumber="25" pagination="65 - 164" refId="B20" refString="Bandel, K, 2001. The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda). Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg 85: 65 - 164" title="The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda)." volume="85" year="2001">Bandel (2001)</bibRefCitation>
|
||
considered
|
||
<taxonomicName id="21CE17854EBAABC674C74A0C6F5AF84C" lsidName="T. macri" pageId="0" pageNumber="25" rank="species" species="macri">
|
||
<emphasis id="E0AD69CF229AF363C84C2684B7EF3CA5" italics="true" pageId="0" pageNumber="25">T. macri</emphasis>
|
||
</taxonomicName>
|
||
a junior synonym of
|
||
<taxonomicName id="20D938D5820EA1F3ECAF57A49F0FEA20" lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
|
||
<emphasis id="0B18F0744C1F3C3F8B69CFD4B137C1C3" italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
|
||
</taxonomicName>
|
||
, but based only on material from Israel and Jordan. Molecular analyses suggest that
|
||
<taxonomicName id="E9459BE84B59068F560B4EA6E8BCF074" authorityName="Montfort" authorityYear="1810" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">
|
||
<emphasis id="0CB292E3F5517B8E5AC4529571E2E071" italics="true" pageId="0" pageNumber="25">Theodoxus</emphasis>
|
||
</taxonomicName>
|
||
from Jordan, Israel, and Palestine belong to a single species,
|
||
<taxonomicName id="6D4B4065C969E2CC6E27A8BD40B2E997" lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
|
||
<emphasis id="C9E1DFDC8C2A5AAF29FA7A25ABBA33C2" italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
|
||
</taxonomicName>
|
||
, which is distinct from all other species (
|
||
<bibRefCitation id="39AC9C865AB28E1F9B41DE0347A99033" DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. 2019a</bibRefCitation>
|
||
; Fig.
|
||
<figureCitation id="66491CC1B351AB789AA3B44E31E64ABD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
|
||
). Although the true identity of
|
||
<taxonomicName id="286C2165EFA165063CB6264AC926EA50" lsidName="T. macri" pageId="0" pageNumber="25" rank="species" species="macri">
|
||
<emphasis id="F46CB171400AE80954EEDE0223AFE989" italics="true" pageId="0" pageNumber="25">T. macri</emphasis>
|
||
</taxonomicName>
|
||
remains dubious at the moment, these results indicate that
|
||
<taxonomicName id="FA11C204D60212D94B712B1D11A87D74" lsidName="T. macri" pageId="0" pageNumber="25" rank="species" species="macri">
|
||
<emphasis id="70C6A5DB373EAE4561B334ACAFADB1D0" italics="true" pageId="0" pageNumber="25">T. macri</emphasis>
|
||
</taxonomicName>
|
||
does not occur outside of Cilicia and south-eastern Anatolia. Probably all previous records of
|
||
<taxonomicName id="3BFDA9909BF3EB6F0FF265D7BB488A1C" lsidName="T. macri" pageId="0" pageNumber="25" rank="species" species="macri">
|
||
<emphasis id="A522C181E00543649C085835CB55AE9A" italics="true" pageId="0" pageNumber="25">T. macri</emphasis>
|
||
</taxonomicName>
|
||
from the Middle East (e.g.
|
||
<bibRefCitation id="50381C8DDCBB5A8E775125A89635F8A3" DOI="https://doi.org/10.1093/oxfordjournals.mollus.a064815" author="Najim, AT" journalOrPublisher="Journal of Molluscan Studies" pageId="0" pageNumber="25" pagination="159 - 163" refId="B112" refString="Najim, AT, 1959. Notes on the distribution of some molluscs in Iraq. Journal of Molluscan Studies 33 (4): 159 - 163, DOI: https://doi.org/10.1093/oxfordjournals.mollus.a064815" title="Notes on the distribution of some molluscs in Iraq." url="https://doi.org/10.1093/oxfordjournals.mollus.a064815" volume="33" year="1959">Najim 1959</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EA2977D0FA88358DD124360A3C71F249" DOI="https://doi.org/10.3923/pjbs.2006.549.552" author="Elkarmi, AZ" journalOrPublisher="Pakistan Journal of Biological Sciences" pageId="0" pageNumber="25" pagination="549 - 552" refId="B50" refString="Elkarmi, AZ, Ismail, NS, 2006. Population structure and shell morphometrics of the gastropod Theodoxus macri (Neritidae: Prosobranchia) from Azraq Oasis, Jordan. Pakistan Journal of Biological Sciences 9 (3): 549 - 552, DOI: https://doi.org/10.3923/pjbs.2006.549.552" title="Population structure and shell morphometrics of the gastropod Theodoxus macri (Neritidae: Prosobranchia) from Azraq Oasis, Jordan." url="https://doi.org/10.3923/pjbs.2006.549.552" volume="9" year="2006">Elkarmi and Ismail 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="8A741050AC124E7EBF5873DC30E35337" author="Handal, EN" journalOrPublisher="Jordan Journal of Natural History" pageId="0" pageNumber="25" pagination="23 - 29" refId="B70" refString="Handal, EN, Amr, Z, Qumsiyeh, MB, 2015. Some records of freshwater snail from the Occupied Palestinian territories. Jordan Journal of Natural History 1 (2): 23 - 29" title="Some records of freshwater snail from the Occupied Palestinian territories." volume="1" year="2015">Handal et al. 2015</bibRefCitation>
|
||
) refer to
|
||
<taxonomicName id="D743C75DC172EC2A0B6E9E74FEE60F65" lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
|
||
<emphasis id="52E271FFD7422C81DD87A37645209C8D" italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
|
||
</taxonomicName>
|
||
. If this is corroborated,
|
||
<taxonomicName id="473538CB8571562DB70BB1578AA56513" lsidName="T. macri" pageId="0" pageNumber="25" rank="species" species="macri">
|
||
<emphasis id="C4BDEB8DDB75444D55282E2812E67171" italics="true" pageId="0" pageNumber="25">T. macri</emphasis>
|
||
</taxonomicName>
|
||
likely diverged from a common ancestor with
|
||
<taxonomicName id="D6E7A977DDB5B17E52F6C04B95FA6F94" lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
|
||
<emphasis id="F3C071523D142A73C513CE643B40FF5B" italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
|
||
</taxonomicName>
|
||
in the early Pleistocene (
|
||
<bibRefCitation id="06EB43AEE8233F5E11442513AF8F808D" DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. 2019a</bibRefCitation>
|
||
; Fig.
|
||
<figureCitation id="6A4BDC100076DC4A766F636C8E57BE7A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="4E8429DFB0E8798E1A4D486E7D686290" doi="10.3897/zse.96.48312.figure18" httpUri="https://binary.pensoft.net/fig/376467" pageId="0" pageNumber="25" start="Figure 18" startId="F18">
|
||
<paragraph id="F275A1E577F0BE88BE625C443BA33C97" pageId="0" pageNumber="25">
|
||
<emphasis id="FA695BD13E6E504E87903477B5C62DBB" bold="true" pageId="0" pageNumber="25">Figure 18.</emphasis>
|
||
<taxonomicName id="A153F468FB9E6C0BB812A8F27A0591E4" baseAuthorityName="G. B. Sowerby II" baseAuthorityYear="1849" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus macri" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="macri">
|
||
<emphasis id="96A0D2A251A5B95E4C78E8E4A4BE9600" italics="true" pageId="0" pageNumber="25">Theodoxus macri</emphasis>
|
||
</taxonomicName>
|
||
(G.B. Sowerby
|
||
<collectionCode id="63AD8AA1D55F9AC84FBC47761EACE687">II</collectionCode>
|
||
, 1849).
|
||
<emphasis id="A7A3F0DD787324857A2B11D5795F8EFF" bold="true" pageId="0" pageNumber="25">A</emphasis>
|
||
-
|
||
<emphasis id="BA01F0B2D8D9D104DCBD9757CB9BC660" bold="true" pageId="0" pageNumber="25">D</emphasis>
|
||
. Specimen collected close to Harbiye Falls, Harbiye, Antakya, Turkey (
|
||
<collectionCode id="6786E2481750B0D7235A6C567B6268F3" collectionName="UGSB">UGSB</collectionCode>
|
||
24179);
|
||
<emphasis id="2AC826CA3F63872BA772A8359F745FC3" bold="true" pageId="0" pageNumber="25">E</emphasis>
|
||
-
|
||
<emphasis id="936F6D74A535D2F77070343057B55C60" bold="true" pageId="0" pageNumber="25">H</emphasis>
|
||
. Specimen from Lake
|
||
<normalizedToken id="076A5D26225B2375DF338C2B5E68A118" originalValue="Yenişehir">Yenisehir</normalizedToken>
|
||
,
|
||
<normalizedToken id="3964065FF32135E4C45C2AA590823138" originalValue="Reyhanlı">Reyhanli</normalizedToken>
|
||
, Turkey (
|
||
<collectionCode id="17A74E13623A1D2ED990BFDD18DCFF32" collectionName="UGSB">UGSB</collectionCode>
|
||
24180);
|
||
<emphasis id="25BF0B5E5DD01BDF96098C42403CBBED" bold="true" pageId="0" pageNumber="25">I</emphasis>
|
||
,
|
||
<emphasis id="89DE65B69142CC46B992CCD1B172CEB3" bold="true" pageId="0" pageNumber="25">J</emphasis>
|
||
. Specimen from Lake
|
||
<normalizedToken id="02143E31553783DD87B27F5AD7817276" originalValue="Balık">Balik</normalizedToken>
|
||
, Adalar, Turkey (
|
||
<collectionCode id="C24737701970FB461A067AEDC06D396A" collectionName="UGSB">UGSB</collectionCode>
|
||
24181). All specimens (A-J) were used in the phylogeny (Fig.
|
||
<figureCitation id="2FE1E15501BA455E6D77E8E255CDF6CC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
|
||
). Scale bars:
|
||
<quantity id="24A913C170A5F9166369E2726F253DED" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2090D9BCDC652C39EB849FCC0F8218F8" pageId="0" pageNumber="25">
|
||
Note on authority: G.B. Sowerby II (1849) attributed the name to
|
||
<normalizedToken id="63CB05EDC01064FFA297444807A746C9" originalValue="Récluz">Recluz</normalizedToken>
|
||
based on a manuscript name, but Sowerby remains the sole author of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5D44E3FB93EB9E3AB04FA9FCDECCD2D4" pageId="0" pageNumber="25" type="distribution">
|
||
<paragraph id="2B1D8ABB8905C42FA93594CD135EC963" pageId="0" pageNumber="25">Distribution.</paragraph>
|
||
<paragraph id="3A04500193F00CB63A66188605F70828" pageId="0" pageNumber="25">
|
||
The distribution range of
|
||
<taxonomicName id="6691930DA6B9BE706C43D4878299DC4D" lsidName="T. macri" pageId="0" pageNumber="25" rank="species" species="macri">
|
||
<emphasis id="7418087057BB0FC5C2D82F0091237641" italics="true" pageId="0" pageNumber="25">T. macri</emphasis>
|
||
</taxonomicName>
|
||
cannot be fully elucidated at present (see Remarks). The material studied by
|
||
<bibRefCitation id="BE213864BEACD5801487B7CEB2077205" DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. (2019a)</bibRefCitation>
|
||
derives primarily from the drainage systems in eastern Cilicia and nearby drainages in south-east Anatolia (Fig.
|
||
<figureCitation id="2F505DBAD5746BBD2A8A3E0730E6E77B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S 1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A - D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2)." figureDoi="10.3897/zse.96.48312.figure3" httpUri="https://binary.pensoft.net/fig/376452" pageId="0" pageNumber="25">3A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |