270 lines
30 KiB
XML
270 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1182.97346" ID-Pensoft-Pub="1313-2970-1182-165" ID-Pensoft-UUID="DCECB4AAD9E95ADB80188D91F386DD1C" ID-ZooBank="86F5AC45D35E4DDFAB042BCBDEEE8407" ModsDocID="1313-2970-1182-165" checkinTime="1697273694970" checkinUser="pensoft" docAuthor="Choi, Yeon-Jae, Park, Sun-Jae, Lee, Seung-Gyu & Park, Jong-Seok" docDate="2023" docId="CD7668F65DCD54EB8AAD2B5830003613" docLanguage="en" docName="ZooKeys 1182: 165-181" docOrigin="ZooKeys 1182" docPubDate="2023-10-13" docSource="http://dx.doi.org/10.3897/zookeys.1182.97346" docTitle="Bryaxis uljinensis Choi, Park, Lee & Park 2023, sp. nov." docType="treatment" docUuid="CF039CE1-2159-4BF5-8E6A-5B9508CBEAC5" docUuidSource="ZooBank" docVersion="1" id="DCECB4AAD9E95ADB80188D91F386DD1C" lastPageNumber="165" masterDocId="DCECB4AAD9E95ADB80188D91F386DD1C" masterDocTitle="Five new species of Bryaxis Kugelann (Coleoptera, Staphylinidae, Pselaphinae) from Korea and a nomenclatural note on Bryaxis mahunkai Loebl" masterLastPageNumber="181" masterPageNumber="165" pageNumber="165" updateTime="1697273694970" updateUser="pensoft">
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<mods:mods id="58FF89CE90A311960848E859C6B15723" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D60C1F9A5ED3B71594DDAC3A472D4ACD">
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<mods:title id="1EC80F30A650EDE2874242E7AF452A49">Five new species of Bryaxis Kugelann (Coleoptera, Staphylinidae, Pselaphinae) from Korea and a nomenclatural note on Bryaxis mahunkai Loebl</mods:title>
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<mods:name id="B8EC15EF4CA9725542172987410A5F07" type="personal">
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<mods:namePart id="481ED3D6629393DC9A03220A366BB7E7">Choi, Yeon-Jae</mods:namePart>
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<mods:nameIdentifier id="F3D7A9919C27A6AD1B25DD2E0B788F62" type="ORCID">https://orcid.org/0000-0003-0172-1128</mods:nameIdentifier>
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<mods:affiliation id="A9F7B22CD3EB9CEE0C408E09EE63CD03">Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju 28644, Republic of Korea</mods:affiliation>
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</mods:name>
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<mods:name id="071AC451AA1C3DDF0D540099CAA55577" type="personal">
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<mods:roleTerm id="3E0BD816E5257882D04D5E230C47039C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="10A3B3CD55494E88A1CB9155204584C7">Park, Sun-Jae</mods:namePart>
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<mods:nameIdentifier id="9587A0EFDC8FFA5AE6C93EF0B63E7F7A" type="ORCID">https://orcid.org/0000-0003-3759-6517</mods:nameIdentifier>
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<mods:affiliation id="4CD08E4671523C4FCBEF8C7953C1DFEF">Climate Change and Environmental Biology Research Division, National Institute of Biological Resources, Incheon 22689, Republic of Korea</mods:affiliation>
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</mods:name>
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<mods:name id="FB868545AF042B1DC66AE7A79902C857" type="personal">
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<mods:role id="73713B76E5526FAFB9AFA19077894260">
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<mods:roleTerm id="2F109751AF5757AF5B098318644AA769">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="71C418BCAAF5DDCD398FCD0144EF7D92">Lee, Seung-Gyu</mods:namePart>
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<mods:nameIdentifier id="D3701BB311A6D7213C79404155455C31" type="ORCID">https://orcid.org/0000-0001-5576-2799</mods:nameIdentifier>
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<mods:affiliation id="FE2579E4056F4D32FCECFD7C27048642">Climate Change and Environmental Biology Research Division, National Institute of Biological Resources, Incheon 22689, Republic of Korea</mods:affiliation>
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</mods:name>
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<mods:name id="EB94262D5DC4AC741BC044C12B704DD5" type="personal">
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<mods:namePart id="83DFFBFAEE14145811A42FA7E62B5162">Park, Jong-Seok</mods:namePart>
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<mods:nameIdentifier id="06C83787B16BFC56915E5099639F1DBC" type="ORCID">https://orcid.org/0000-0003-2063-2534</mods:nameIdentifier>
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<mods:affiliation id="FD3C6F898B4F0A26C10F3599324BF06B">Department of Biological Sciences and Biotechnology, Chungbuk National University, Cheongju 28644, Republic of Korea</mods:affiliation>
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<mods:nameIdentifier id="DB303E30BFE58F220B04EB8D0B7486F1" type="email">jpark16@cbnu.ac.kr</mods:nameIdentifier>
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<mods:typeOfResource id="B5E67AC9EB9A37B4347F9C8C018DC530">text</mods:typeOfResource>
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<mods:title id="00F185409A3DFC4CDFA0F2A87BEC4A52">ZooKeys</mods:title>
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<mods:part id="876ED48C3018A4F6CF1AC9B3304EA484">
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<mods:date id="EC7940EC7FCB7CFC17CB9DB4548C4429">2023</mods:date>
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<mods:number id="730CFA14B100015D3FBEE4733E773033">2023-10-13</mods:number>
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<mods:detail id="3E66AEAD9A3E4BBC8B6DDDD7AE9F3591" type="volume">
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<mods:number id="18E792E78D0A21279E37D1A85012977A">1182</mods:number>
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<mods:start id="528258A50DA17B4CA27CC71BC662D59A">165</mods:start>
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<mods:end id="B13F40C6F12D75676438DA041D3704D7">181</mods:end>
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<mods:url id="B5135AEF14EA4C1D9CA9883F91D5863D">http://dx.doi.org/10.3897/zookeys.1182.97346</mods:url>
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<mods:classification id="D5B64F44DCB2B0D493164463756638BC">journal article</mods:classification>
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<mods:identifier id="31B9C8A515566DE9326A0BBBA5D30916" type="DOI">http://dx.doi.org/10.3897/zookeys.1182.97346</mods:identifier>
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<mods:identifier id="4E2C8302AE78D9D3569BA0DFB06D297B" type="Pensoft-Pub">1313-2970-1182-165</mods:identifier>
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<mods:identifier id="62B40DFCC2C7A74C4026A8CCDD3D1047" type="ZooBank">86F5AC45D35E4DDFAB042BCBDEEE8407</mods:identifier>
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<treatment id="CD7668F65DCD54EB8AAD2B5830003613" LSID="urn:lsid:zoobank.org:act:CF039CE1-2159-4BF5-8E6A-5B9508CBEAC5" httpUri="http://treatment.plazi.org/id/CD7668F65DCD54EB8AAD2B5830003613" lastPageNumber="165" pageId="0" pageNumber="165">
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<subSubSection id="3B3BABCA38C99949B714B966E0DF370A" pageId="0" pageNumber="165" type="nomenclature">
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<paragraph id="F7572E4F41D693324F6D6A6AEB9D705D" pageId="0" pageNumber="165">
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<taxonomicName id="07BB593510659CC1A0004A9891E0B550" LSID="https://zoobank.org/CF039CE1-2159-4BF5-8E6A-5B9508CBEAC5" authority="Choi, Park, Lee & Park" authorityName="Choi, Park, Lee & Park" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryaxis uljinensis" order="Coleoptera" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="uljinensis" status="sp. nov.">Bryaxis uljinensis Choi, Park, Lee & Park</taxonomicName>
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<taxonomicNameLabel id="E003E3799A4958A7C050BEB30717C5BA" pageId="0" pageNumber="165">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="8465C3A0373B3AE82A8FB65E20DAE2D1" pageId="0" pageNumber="165" type="description">
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<paragraph id="70D705D83730B87222E5846A1EE71DFA" pageId="0" pageNumber="165">
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<figureCitation id="182AD2EB2F553CBFD400DF9DC8C99C38" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bryaxis uljinensis Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure3" httpUri="https://binary.pensoft.net/fig/920946" pageId="0" pageNumber="165">Figs 3</figureCitation>
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<figureCitation id="3F267B39EE07867323A76824CC96817B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Bryaxis uljinensis sp. nov. (A, C, E) and B. mahunkai Loebl (B, D, F). A, B maxillary palpi C, D antennal scapes and pedicels E, F protibiae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure4" httpUri="https://binary.pensoft.net/fig/920947" pageId="0" pageNumber="165">, 4A, C, E</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="85C65F71E9692CB25144D469E4C484C0" pageId="0" pageNumber="165" type="type material">
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<paragraph id="13F4D0C433A14C66E450027A33CA2214" pageId="0" pageNumber="165">Type materials</paragraph>
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<paragraph id="123A4AD8D56A9E152335B31789D66085" pageId="0" pageNumber="165">
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<emphasis id="0765F9A8038F8FD9A29CD4E8919E6A1F" bold="true" pageId="0" pageNumber="165">
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(
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<emphasis id="42232FF6F059132185E4B5DE6DE23A34" italics="true" pageId="0" pageNumber="165">N</emphasis>
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= 7, 4
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</emphasis>
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♂♂,
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<emphasis id="974BE1895146F1C54B25EF0DDA95AB90" bold="true" pageId="0" pageNumber="165">3</emphasis>
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♀♀)
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<emphasis id="5ADDA8ADB085AC2F2E4F579A97C6027A" bold="true" pageId="0" pageNumber="165">
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.
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<emphasis id="7EDF9AA2491298A0484B8B86BFCAE8A3" italics="true" pageId="0" pageNumber="165">Holotype male</emphasis>
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.
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</emphasis>
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"Korea: Gyeongbuk Prov. Onjeong-myeon, Uljin-gun, 8 Jun 2019,
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<geoCoordinate id="55022E1196EF34F341A6086A0158A1C5" degrees="36" direction="north" minutes="43" orientation="latitude" precision="1" seconds="23.0" value="36.723057">36°43'23.0"N</geoCoordinate>
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,
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<geoCoordinate id="EDD01F1F4F6957F40CBFC8476A6A0477" degrees="129" direction="east" minutes="20" orientation="longitude" precision="1" seconds="16.0" value="129.33777">129°20'16.0"E</geoCoordinate>
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, 180 m, sifting leaf litter near stream, J.-S. Park" (NIBR).
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<emphasis id="2205E9B5E039892FCDC34A9D40A6B712" bold="true" pageId="0" pageNumber="165">
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<emphasis id="EA179DBA9D5A405A5E6CD8CAA9825BCA" italics="true" pageId="0" pageNumber="165">Paratypes</emphasis>
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.
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</emphasis>
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2♂♂ (CBNUIC, 1♂ slide mounted, 1♂ dried). "Korea: Gangwon Prov. Gujeol-ri, Yeoryang-myeon, Jeongseon-gun, 24 Apr 2020,
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<geoCoordinate id="D0B1728675B1252BC99941B3AC495FFD" degrees="37" direction="north" minutes="31" orientation="latitude" precision="1" seconds="08.7" value="37.519085">37°31'08.7"N</geoCoordinate>
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,
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<geoCoordinate id="4FCE9B6807B4A040BCAFA51BE08D5B6C" degrees="128" direction="east" minutes="46" orientation="longitude" precision="1" seconds="42.8" value="128.77855">128°46'42.8"E</geoCoordinate>
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, 591 m, sifting leaf & soil litter, U.-J. Byeon, T.-Y. Jang". 1♂, 3♀♀ (CBNUIC, dried). "Korea: Gangwon Prov. Gujeol-ri, Yeoryang-myeon, Jeongseon-gun, 24 Apr 2020,
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<geoCoordinate id="AA5168A32D2EDD4B8A53717931439DCB" degrees="37" direction="north" minutes="30" orientation="latitude" precision="1" seconds="57.6" value="37.516">37°30'57.6"N</geoCoordinate>
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,
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<geoCoordinate id="FF8D58B1E30D7EBA80337FE7BB2BA02A" degrees="128" direction="east" minutes="45" orientation="longitude" precision="1" seconds="18.8" value="128.75522">128°45'18.8"E</geoCoordinate>
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, 510 m, sifting ant colony, leaf & soil litter, Y.-J. Choi, U.-J. Byeon".
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</paragraph>
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</subSubSection>
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<subSubSection id="32FF69492198A689BABCC4A851D3F77F" pageId="0" pageNumber="165" type="diagnosis">
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<paragraph id="34B0B4CCDAD4C7B50FEDB0527FAC1862" pageId="0" pageNumber="165">Diagnosis.</paragraph>
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<paragraph id="E053001AED12974AE0E35D707618FEE8" pageId="0" pageNumber="165">
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Antennal pedicel strongly swollen, subglobose with subcylindrical glandular nodule on basal 1/3 of inner margin (Figs
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<figureCitation id="C2E84E054115C17F568EB9B2CAE999B6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bryaxis uljinensis Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure3" httpUri="https://binary.pensoft.net/fig/920946" pageId="0" pageNumber="165">3F</figureCitation>
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,
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<figureCitation id="804BE1EDB514B33692110838368134B2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Bryaxis uljinensis sp. nov. (A, C, E) and B. mahunkai Loebl (B, D, F). A, B maxillary palpi C, D antennal scapes and pedicels E, F protibiae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure4" httpUri="https://binary.pensoft.net/fig/920947" pageId="0" pageNumber="165">4C</figureCitation>
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, arrows); protibiae with spine on internal side of widest (Figs
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<figureCitation id="BD7043E0DEE05938BD59E0B1FC0CEA84" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bryaxis uljinensis Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure3" httpUri="https://binary.pensoft.net/fig/920946" pageId="0" pageNumber="165">3G</figureCitation>
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,
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<figureCitation id="B0B4FB881F97A82323754FB37B52F11A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Bryaxis uljinensis sp. nov. (A, C, E) and B. mahunkai Loebl (B, D, F). A, B maxillary palpi C, D antennal scapes and pedicels E, F protibiae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure4" httpUri="https://binary.pensoft.net/fig/920947" pageId="0" pageNumber="165">4E</figureCitation>
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, arrows); parameres of male genitalia robust and fan-shaped, bearing three setae on each (Fig.
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<figureCitation id="7FD66CCCFDF53E4135AD0054E7F4A49B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bryaxis uljinensis Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure3" httpUri="https://binary.pensoft.net/fig/920946" pageId="0" pageNumber="165">3I</figureCitation>
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).
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</paragraph>
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<caption id="CB561ED25835F4B29F2EE11135F877EF" doi="10.3897/zookeys.1182.97346.figure3" httpUri="https://binary.pensoft.net/fig/920946" pageId="0" pageNumber="165" start="Figure 3" startId="F3">
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<paragraph id="4832ED68938396D92BCF87702CD47E63" pageId="0" pageNumber="165">
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<emphasis id="3F12D443BF3D5FCE194AF0DF3633B3D2" bold="true" pageId="0" pageNumber="165">Figure 3.</emphasis>
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<taxonomicName id="A728750D1663E91733AA78243ED47803" authorityName="Choi, Park, Lee & Park" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryaxis uljinensis" order="Coleoptera" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="uljinensis">
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<emphasis id="AD9150B2E724D09910255EA2015BA925" italics="true" pageId="0" pageNumber="165">Bryaxis uljinensis</emphasis>
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</taxonomicName>
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Choi, Park, Lee & Park, sp. nov., male (
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<emphasis id="6F0EB2A29D56406F7614911CA581669C" bold="true" pageId="0" pageNumber="165">A, B, E-I</emphasis>
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), female (
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<emphasis id="72C68A318C692C587997816A559B3DA3" bold="true" pageId="0" pageNumber="165">C, D</emphasis>
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).
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<emphasis id="B017F453C7516E0D8A0DB3ADB114BAC8" bold="true" pageId="0" pageNumber="165">A, C</emphasis>
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dorsal habitus
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<emphasis id="47AAC313412CF79AF8EC552601E1E99B" bold="true" pageId="0" pageNumber="165">B, D</emphasis>
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ventral habitus
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<emphasis id="DCE95C85AB63B768ED1650D8664291F5" bold="true" pageId="0" pageNumber="165">E</emphasis>
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head
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<emphasis id="19C95F9BDB08918C97639A27297B61D4" bold="true" pageId="0" pageNumber="165">F</emphasis>
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antenna
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<emphasis id="581CE241BC9C4CD9BEA573ECBA74F149" bold="true" pageId="0" pageNumber="165">G</emphasis>
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fore leg
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<emphasis id="DE9953524FFCCAC87AFC77F24C9E2586" bold="true" pageId="0" pageNumber="165">H</emphasis>
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hind leg
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<emphasis id="ED25EF74D592FF67599B0AD692DA3A77" bold="true" pageId="0" pageNumber="165">I</emphasis>
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aedeagus. Scale bars: 1 mm (
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<emphasis id="0907E3F5ACFBC4E86B5098A78F2D8489" bold="true" pageId="0" pageNumber="165">A-D</emphasis>
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); 0.1 mm (
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<emphasis id="9711AF2F64CBD474F1726ED12595CE7F" bold="true" pageId="0" pageNumber="165">E, I</emphasis>
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); 0.5 mm (
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<emphasis id="D6E50A162214E4677608482FFB3A116F" bold="true" pageId="0" pageNumber="165">F-H</emphasis>
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).
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</paragraph>
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</caption>
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<caption id="156B7D41ADD243ADE7AEF32BA1F702F8" doi="10.3897/zookeys.1182.97346.figure4" httpUri="https://binary.pensoft.net/fig/920947" pageId="0" pageNumber="165" start="Figure 4" startId="F4">
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<paragraph id="BFE3AAEBD5E22A141D43A84D2ED0275C" pageId="0" pageNumber="165">
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<emphasis id="457B7FCDCC64DD066D9FC721DECDE989" bold="true" pageId="0" pageNumber="165">Figure 4.</emphasis>
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Diagnostic characters of
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<taxonomicName id="D11BB4F8055621566A39E801A5E4AE75" authorityName="Choi, Park, Lee & Park" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryaxis uljinensis" order="Coleoptera" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="uljinensis">
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<emphasis id="49D054A79CCF383655D17EA2202025F4" italics="true" pageId="0" pageNumber="165">Bryaxis uljinensis</emphasis>
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</taxonomicName>
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sp. nov. (
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<emphasis id="990AC6AC100CC480C8EC28C009EB07AB" bold="true" pageId="0" pageNumber="165">A, C, E</emphasis>
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) and
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<taxonomicName id="8097270FAEC2EF3137E38F84A4AA0884" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="B. mahunkai" order="Coleoptera" pageId="0" pageNumber="165" rank="species" species="mahunkai">
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<emphasis id="8F596370723BA0DD4B8AC895BCF383CA" italics="true" pageId="0" pageNumber="165">B. mahunkai</emphasis>
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</taxonomicName>
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<normalizedToken id="82C1D8948C63D752D65F9DAAFA882E57" originalValue="Löbl">Loebl</normalizedToken>
|
|
(
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|
<emphasis id="D774061EE377189EE28083DD3FE3023D" bold="true" pageId="0" pageNumber="165">B, D, F</emphasis>
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|
).
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<emphasis id="9167BD43D7F1AD75470950CFBD608955" bold="true" pageId="0" pageNumber="165">A, B</emphasis>
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|
maxillary palpi
|
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<emphasis id="F5CDBF80EAAB097B3B73E6E3B3757BE3" bold="true" pageId="0" pageNumber="165">C, D</emphasis>
|
|
antennal scapes and pedicels
|
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<emphasis id="E91F8447C94726FBFF3A28B3C0733021" bold="true" pageId="0" pageNumber="165">E, F</emphasis>
|
|
protibiae. Scale bars: 0.1 mm.
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|
</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="D54BB1F948CDF85958FE9B2BEC926D2B" pageId="0" pageNumber="165" type="description">
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<paragraph id="1A63D1B5544544AE6776452E341B680E" pageId="0" pageNumber="165">Description.</paragraph>
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<paragraph id="8354F21905A67D651F498C7A1A6FC311" pageId="0" pageNumber="165">
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Body reddish brown, antennae, maxillary palpi, legs slightly lighter than body, length 1.25-1.31 mm, maximum width 0.53-0.60 mm (Fig.
|
|
<figureCitation id="8ABE4997F0E759D58BAB398DC041EA4F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bryaxis uljinensis Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure3" httpUri="https://binary.pensoft.net/fig/920946" pageId="0" pageNumber="165">3A-D</figureCitation>
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|
). Setae on body yellowish, and short. Head 0.98 times as long as wide (Fig.
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|
<figureCitation id="C97A8D53182EE41808D874A81AB6AB0F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bryaxis uljinensis Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure3" httpUri="https://binary.pensoft.net/fig/920946" pageId="0" pageNumber="165">3E</figureCitation>
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|
). Frons with small U-shaped impression between antennal tubercles; frontal foveae absent; frontal rostrum slightly distinct anteriorly. Vertex convex; longitudinal carina present; vertexal foveae small. Eyes large, longer than twice that of tempora, composed of 32-34 facets. Maxillary palpi moderately developed; palpomeres II-III smooth; palpomere IV 0.22 mm long and about 3.16 times as long as wide, subcylindrical pseudosegment at apex. Antennae about 0.53 mm long; scapes short without modification, 0.92 times as long as pedicels; pedicels long as wide; antennomere III-VIII subequal in length; IX 0.63 times as long as wide; X transverse, 0.61 times as long as wide; XI largest, pointed at apex, 1.61 times as long as wide (Fig.
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|
<figureCitation id="DD839DC3178208ACB8C6FE42345D6D79" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bryaxis uljinensis Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure3" httpUri="https://binary.pensoft.net/fig/920946" pageId="0" pageNumber="165">3F</figureCitation>
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|
). Pronotum 0.88 times as long as wide and widest at basal 3/5, lateral antebasal foveae connected by antebasal sulcus. Elytra convex, 0.85 times as long as wide and widest at basal 1/3, each elytron with two basal foveae and subhumeral fovea. Legs robust; internal spine on widest of protibiae (Fig.
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|
<figureCitation id="75335E72D569A89463AA1D75D4160A87" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bryaxis uljinensis Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure3" httpUri="https://binary.pensoft.net/fig/920946" pageId="0" pageNumber="165">3G</figureCitation>
|
|
, arrow); metatibiae with spine on apical (Fig.
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|
<figureCitation id="F0260AB16D5CA3FECF22BF3E8166D252" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bryaxis uljinensis Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure3" httpUri="https://binary.pensoft.net/fig/920946" pageId="0" pageNumber="165">3H</figureCitation>
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|
, arrow). Aedeagus small, 0.27 mm long and 1.61 times as long as wide; penis small fusiform and dorsal diaphragm bulbous; parameres symmetrical; endophallus composed with two convergent, slender struts (Fig.
|
|
<figureCitation id="341C1D0B17FE51B3582E09A4A0D1F924" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Bryaxis uljinensis Choi, Park, Lee & Park, sp. nov., male (A, B, E-I), female (C, D). A, C dorsal habitus B, D ventral habitus E head F antenna G fore leg H hind leg I aedeagus. Scale bars: 1 mm (A-D); 0.1 mm (E, I); 0.5 mm (F-H)." figureDoi="10.3897/zookeys.1182.97346.figure3" httpUri="https://binary.pensoft.net/fig/920946" pageId="0" pageNumber="165">3I</figureCitation>
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|
).
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|
</paragraph>
|
|
<paragraph id="E9B2EBCD8C7A5DD64541360E4E334BF0" pageId="0" pageNumber="165">
|
|
<emphasis id="51D770F4B57D18A7D70DDE62AE36BF0C" bold="true" pageId="0" pageNumber="165">
|
|
<emphasis id="4A4950529BA4D46097E408C0AB552450" italics="true" pageId="0" pageNumber="165">Sexual dimorphism</emphasis>
|
|
.
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|
</emphasis>
|
|
Female eyes slightly longer than tempora, composed of 15 facets; antennal pedicels simple; protibial spines and metatibial spines absent.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E47D12C1165F0F52CAC525E576151806" pageId="0" pageNumber="165" type="remarks">
|
|
<paragraph id="27C4B9A82B99CDE82470E689AB5FFE3F" pageId="0" pageNumber="165">Remarks.</paragraph>
|
|
<paragraph id="349C424005B1B62E2A1B4454B722BC79" pageId="0" pageNumber="165">
|
|
Adults of this species are very similar to
|
|
<taxonomicName id="6DD71C040D7C32EDC861B7FA91507154" authorityName="Loebl" authorityYear="1975" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bryaxis mahunkai" order="Coleoptera" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="mahunkai">
|
|
<emphasis id="045E6C440E7C34FDAAA75B2DE4A9B5B3" italics="true" pageId="0" pageNumber="165">Bryaxis mahunkai</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken id="F1A64C8481425384512C835D88600412" originalValue="Löbl">Loebl</normalizedToken>
|
|
, 1975 in having strongly swollen antennal pedicels (Fig.
|
|
<figureCitation id="C860FD5128671D3254AE82CBC3D0298A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Bryaxis uljinensis sp. nov. (A, C, E) and B. mahunkai Loebl (B, D, F). A, B maxillary palpi C, D antennal scapes and pedicels E, F protibiae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure4" httpUri="https://binary.pensoft.net/fig/920947" pageId="0" pageNumber="165">4C, D</figureCitation>
|
|
). However, they can be distinguished by smooth maxillary palpomere II-III (Fig.
|
|
<figureCitation id="810F328B34D2F2D29822B29184D5B656" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Bryaxis uljinensis sp. nov. (A, C, E) and B. mahunkai Loebl (B, D, F). A, B maxillary palpi C, D antennal scapes and pedicels E, F protibiae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure4" httpUri="https://binary.pensoft.net/fig/920947" pageId="0" pageNumber="165">4A</figureCitation>
|
|
), apically symmetrical antennal scapes, pedicels less swollen apically and bearing smaller glandular nodules (Fig.
|
|
<figureCitation id="0F5B031870359BFF1E063D9BA13D9C19" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Bryaxis uljinensis sp. nov. (A, C, E) and B. mahunkai Loebl (B, D, F). A, B maxillary palpi C, D antennal scapes and pedicels E, F protibiae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure4" httpUri="https://binary.pensoft.net/fig/920947" pageId="0" pageNumber="165">4C</figureCitation>
|
|
, arrow), fore legs with a tibial spine (Fig.
|
|
<figureCitation id="130EA00878B7039676FCCBFC5EC6D29C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Diagnostic characters of Bryaxis uljinensis sp. nov. (A, C, E) and B. mahunkai Loebl (B, D, F). A, B maxillary palpi C, D antennal scapes and pedicels E, F protibiae. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure4" httpUri="https://binary.pensoft.net/fig/920947" pageId="0" pageNumber="165">4E</figureCitation>
|
|
, arrow), and parameres wider than the penis (aedeagus in
|
|
<taxonomicName id="AB6AED607F7CE72110D9FEC557687797" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="B. mahunkai" order="Coleoptera" pageId="0" pageNumber="165" rank="species" species="mahunkai">
|
|
<emphasis id="4884543282BBCCF912DE55245CB5FDEC" italics="true" pageId="0" pageNumber="165">B. mahunkai</emphasis>
|
|
</taxonomicName>
|
|
as wide as penis; Fig.
|
|
<figureCitation id="4DB63986A1AA112CCDDEA415EB6CF257" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Diagnostic characters of Bryaxis mahunkai Loebl. A, B antennal scapes and pedicels C, D aedeagi. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1182.97346.figure10" httpUri="https://binary.pensoft.net/fig/920953" pageId="0" pageNumber="165">10C, D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="FA4A5315077934537E66FF6F1A26358F" pageId="0" pageNumber="165" type="comments">
|
|
<paragraph id="896429BDC420F78BF90DE0FA2D5182E0" pageId="0" pageNumber="165">Comments.</paragraph>
|
|
<paragraph id="9DE77EB40128695D68EADF363ADA1B42" pageId="0" pageNumber="165">
|
|
The localities of
|
|
<taxonomicName id="03E9ACF6E5FB1EBC3C0F257CBB706A3F" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="B. uljinensis" order="Coleoptera" pageId="0" pageNumber="165" rank="species" species="uljinensis">
|
|
<emphasis id="8B1194BF754B30C42B2D21E11BD84716" italics="true" pageId="0" pageNumber="165">B. uljinensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. probably overlap with those of
|
|
<taxonomicName id="D50EDB3BF8AC870883DE153E96D30BD9" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="B. mahunkai" order="Coleoptera" pageId="0" pageNumber="165" rank="species" species="mahunkai">
|
|
<emphasis id="B65B741827A6A31E733BB6416E5855A0" italics="true" pageId="0" pageNumber="165">B. mahunkai</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken id="DB8984BE1D53EF0EB437A09ACE944C49" originalValue="Löbl">Loebl</normalizedToken>
|
|
given that the latter are nationally distributed (Fig.
|
|
<figureCitation id="657EF585B7BD0F3C8C88C479397101FC" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Collection localities. Bryaxis grandinodus sp. nov. (blue pentagon); B. uljinensis sp. nov. (purple cross); B. fabaiformis sp. nov. (sky-blue star); B. girinensis sp. nov. (green diamond); B. nemorosus sp. nov. (navy hexagon); B. mahunkai Loebl (yellow square); B. koltzei (Reitter) (red circle); B. kimjongkuki Nomura & Lee (orange triangle)." figureDoi="10.3897/zookeys.1182.97346.figure11" httpUri="https://binary.pensoft.net/fig/920954" pageId="0" pageNumber="165">11</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="43CEAA3A13FCCFC3F2F7DCBA41F7085C" pageId="0" pageNumber="165" type="etymology">
|
|
<paragraph id="EEE41A906592964523BECE3FFBF13A69" pageId="0" pageNumber="165">Etymology.</paragraph>
|
|
<paragraph id="ECACF66B97C37F93FAEB66ECC0D7FC00" pageId="0" pageNumber="165">This species is named after the type locality, Uljin-gun.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="30FF90A3E19514068BBBE268C8758522" pageId="0" pageNumber="165" type="habitat">
|
|
<paragraph id="273EA8982269ABD742F3BB0016823A39" pageId="0" pageNumber="165">Habitat.</paragraph>
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|
<paragraph id="D261365FEAA3578DB2805FCB9F926CFA" pageId="0" pageNumber="165">The holotype was collected by sifting leaf litter in mixed forest. Paratypes were collected by sifting leaf litter, soil, and an ant colony.</paragraph>
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</subSubSection>
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|
<subSubSection id="6167D8E17FD4263809F1B49E2390F9E3" pageId="0" pageNumber="165" type="distribution">
|
|
<paragraph id="9B7D41AFD342598D58B9809A1733105D" pageId="0" pageNumber="165">Distribution.</paragraph>
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<paragraph id="F436FE7DB0CA5611B83DF877F90A55F5" pageId="0" pageNumber="165">Korea (Uljin-gun, Gyeongsangbuk-do; Jeongseon-gun, Gangwon-do).</paragraph>
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</subSubSection>
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</treatment>
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</document> |