treatments-xml/data/CD/40/D0/CD40D06FFFD7780C658A505BFAE6FAB8.xml

<document id="ADA5B7F39DC06302B7CF98E41C747E4B" ID-CLB-Dataset="31057" ID-DOI="10.11646/zootaxa.4369.4.7" ID-GBIF-Dataset="0ff3b724-fbb2-458f-a55b-d28910952a66" ID-ISSN="1175-5326" ID-Zenodo-Dep="1136494" ID-ZooBank="CE0D1E89-B2CC-4871-ACE3-23C41343AA5C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1515401419884" checkinUser="plazi" docAuthor="Komai, Tomoyuki &amp; Fujita, Yoshihisa" docDate="2018" docId="CD40D06FFFD7780C658A505BFAE6FAB8" docLanguage="en" docName="zootaxa.4369.4.7.pdf" docOrigin="Zootaxa 4369 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Caligoneus cavernicola Komai &amp; Fujita 2018, n. sp." docType="treatment" docVersion="10" lastPageNumber="584" masterDocId="3179A817FFD57805651D5607FF83FFEF" masterDocTitle="A new genus and new species of alpheid shrimp from a marine cave in the Ryukyu Islands, Japan, with additional record of Salmoneus antricola Komai, Yamada &amp; Yunokawa, 2015 (Crustacea: Decapoda: Caridea)" masterLastPageNumber="586" masterPageNumber="575" pageNumber="577" updateTime="1698512995346" updateUser="ExternalLinkService">
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<mods:title id="AD6E63567D08ADFB304C45F100E4C254">A new genus and new species of alpheid shrimp from a marine cave in the Ryukyu Islands, Japan, with additional record of Salmoneus antricola Komai, Yamada &amp; Yunokawa, 2015 (Crustacea: Decapoda: Caridea)</mods:title>
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<mods:namePart id="90686FF38EA9C6ECEA522AE4D8966E62">Komai, Tomoyuki</mods:namePart>
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<treatment id="CD40D06FFFD7780C658A505BFAE6FAB8" ID-DOI="http://doi.org/10.5281/zenodo.5982862" ID-GBIF-Taxon="139669237" ID-Zenodo-Dep="5982862" LSID="urn:lsid:plazi:treatment:CD40D06FFFD7780C658A505BFAE6FAB8" httpUri="http://treatment.plazi.org/id/CD40D06FFFD7780C658A505BFAE6FAB8" lastPageId="9" lastPageNumber="584" pageId="2" pageNumber="577">
<subSubSection id="0DF332F2FFD77807658A505BFF7AF952" pageId="2" pageNumber="577" type="nomenclature">
<paragraph id="45566179FFD77807658A505BFE7EF998" blockId="2.[151,652,1628,1725]" box="[151,509,1628,1655]" pageId="2" pageNumber="577">
<heading id="1E1ED615FFD77807658A505BFE7EF998" bold="true" box="[151,509,1628,1655]" fontSize="11" level="1" pageId="2" pageNumber="577" reason="1">
<emphasis id="779DBD6BFFD77807658A505BFE7EF998" bold="true" box="[151,509,1628,1655]" pageId="2" pageNumber="577">
<taxonomicName id="82E91AFAFFD77807658A505BFE37F999" ID-CoL="85NCH" authority="Komai &amp; Fujita, 2018" authorityName="Komai &amp; Fujita" authorityYear="2018" box="[151,436,1628,1654]" class="Malacostraca" family="Alpheidae" genus="Caligoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="577" phylum="Arthropoda" rank="species" species="cavernicola" status="sp. nov.">
<emphasis id="779DBD6BFFD77807658A505BFE37F999" bold="true" box="[151,436,1628,1654]" italics="true" pageId="2" pageNumber="577">Caligoneus cavernicola</emphasis>
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<taxonomicNameLabel id="6CAE0010FFD7780764A7505AFE7EF998" box="[442,509,1629,1655]" pageId="2" pageNumber="577" rank="species">n. sp.</taxonomicNameLabel>
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[New Japanese name: Kurayami-teppou-ebi] 
<figureCitation id="DDD27DFCFFD77807658A50A3FF7AF952" box="[151,249,1700,1725]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="4.[151,250,1973,1995]" captionStartId-1="5.[151,250,1824,1846]" captionStartId-2="6.[151,250,1234,1256]" captionStartId-3="7.[151,250,1877,1899]" captionStartId-4="8.[151,250,1959,1981]" captionTargetBox-0="[225,1325,207,1939]" captionTargetBox-1="[244,1334,231,1786]" captionTargetBox-2="[186,1401,209,1202]" captionTargetBox-3="[213,1345,218,1829]" captionTargetBox-4="[150,1435,217,1919]" captionTargetId-0="figure@4.[196,1392,183,1962]" captionTargetId-1="figure@5.[216,1376,193,1815]" captionTargetId-2="figure@6.[151,1436,193,1229]" captionTargetId-3="figure@7.[187,1374,195,1844]" captionTargetId-4="figure@8.[150,1436,217,1920]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 1. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm), habitus in lateral view (antennular and antennal flagella partially omitted)." captionText-1="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." captionText-2="FIGURE 3. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm), mouthparts dissected from left side. A, mandible, internal view; B, same, external view; C, same, incisor process, distal surface; D, maxillule (coxal endite missing); E, maxilla; F, maxilliped 1; G,same, lateral view of exopod and endopod; H, maxilliped 2." captionText-3="FIGURE 4. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm). A, left maxilliped3,lateral view; B, same,coxaand proximal part of antepenultimate article, dorsal view;C, leftpereopod 1 (= cheliped), lateral view; D, same, chela, extensor view; E, right pereopod 1 (= cheliped), lateral view; F, same, chela, extensor view; G, left pereopod 2, lateral view; H, left pereopod 3, lateral view; I, same, distal part of propodus and dactylus, lateral view; J, left pereopod 4, lateral view; K, left pereopod 5, lateral view; L, same, distal part of propodus and dactylus, lateral view." captionText-4="FIGURE 5. Caligoneus cavernicola n.gen., n. sp., general habitus and colour in life. A, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, RUMF-ZC-02733, non-ovigerous individual(cl 5.6mm)." httpUri-0="https://zenodo.org/record/1136496/files/figure.png" httpUri-1="https://zenodo.org/record/1136498/files/figure.png" httpUri-2="https://zenodo.org/record/1136500/files/figure.png" httpUri-3="https://zenodo.org/record/1136502/files/figure.png" httpUri-4="https://zenodo.org/record/1136504/files/figure.png" pageId="2" pageNumber="577">Figs 1–5</figureCitation>
</paragraph>
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<paragraph id="45566179FFD77807658A50EBFDCBF8A2" blockId="2.[151,1437,1772,2013]" pageId="2" pageNumber="577">
<materialsCitation id="F5816B24FFD77807658A50EBFDCBF8A2" ID-GBIF-Occurrence="1802522159" collectingDate="2015-07-18" collectionCode="SCUBA" collectorName="Ohoba No. &amp; K. Yunokawa &amp; Fujita." country="Japan" county="Ryukyu Islands" elevation="12" location="Material" municipality="Ie Island" pageId="2" pageNumber="577" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
<emphasis id="779DBD6BFFD77807658A50EBFEFEF8EA" bold="true" box="[151,381,1772,1797]" pageId="2" pageNumber="577">
<location id="403637A2FFD77807658A50EBFF7CF8EA" LSID="urn:lsid:plazi:treatment:CD40D06FFFD7780C658A505BFAE6FAB8:403637A2FFD77807658A50EBFF7CF8EA" box="[151,255,1772,1797]" country="Japan" county="Ryukyu Islands" municipality="Ie Island" name="Material" pageId="2" pageNumber="577" stateProvince="Okinawa">Material</location>
examined
</emphasis>
. 
<typeStatus id="9A52DFDBFFD77807649250EBFE7FF8EA" box="[399,508,1772,1797]" pageId="2" pageNumber="577" type="holotype">Holotype</typeStatus>
: CBM-ZC 14136, ovigerous individual (cl 
<quantity id="8211CC9CFFD7780766EB50EBFBD7F8EB" box="[1014,1108,1772,1797]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="577" unit="mm" value="5.7">5.7 mm</quantity>
), unnamed cave (sometimes called as “ 
<collectorName id="E81C04AFFFD7780764105108FE0FF8C7" box="[269,396,1807,1832]" pageId="2" pageNumber="577">Ohoba No.</collectorName>
2 
<location id="403637A2FFD7780764BF5117FE62F8C7" LSID="urn:lsid:plazi:treatment:CD40D06FFFD7780C658A505BFAE6FAB8:403637A2FFD7780764BF5117FE62F8C7" box="[418,481,1808,1832]" country="Japan" county="Ryukyu Islands" municipality="Ie Island" name="Cave" pageId="2" pageNumber="577" stateProvince="Okinawa">Cave</location>
”), 
<collectingMunicipality id="A532FB03FFD77807671F5117FDE4F8C7" box="[514,615,1807,1832]" pageId="2" pageNumber="577">Ie Island</collectingMunicipality>
, 
<collectingRegion id="872DAF9BFFD7780767695108FD5AF8C7" box="[628,729,1807,1832]" country="Japan" name="Okinawa" pageId="2" pageNumber="577">Okinawa</collectingRegion>
Islands, 
<collectingCounty id="AC3719F5FFD77807665F5108FC77F8C7" box="[834,1012,1807,1832]" pageId="2" pageNumber="577">Ryukyu Islands</collectingCounty>
, 
<collectingCountry id="3DFE21E9FFD77807611C5117FBC7F8C8" box="[1025,1092,1808,1831]" name="Japan" pageId="2" pageNumber="577">Japan</collectingCountry>
, 
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<quantity id="8211CC9CFFD77807614D5117FB0EF8C8" box="[1104,1165,1808,1832]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="577" unit="m" value="12.0">12 m</quantity>
</elevation>
, 
<collectingDate id="2113BE51FFD7780761865117FAB1F8C7" box="[1179,1330,1807,1832]" pageId="2" pageNumber="577" value="2015-07-18">
<date id="315747B9FFD7780761865117FAB1F8C7" box="[1179,1330,1807,1832]" pageId="2" pageNumber="577" value="2015-07-18">18 July 2015</date>
</collectingDate>
, 
<collectionCode id="23F8F9BCFFD7780760225117FA1FF8C7" box="[1343,1436,1808,1832]" pageId="2" pageNumber="577">SCUBA</collectionCode>
diving, coll. 
<collectorName id="E81C04AFFFD7780764385132FE3FF8A2" box="[293,444,1844,1869]" pageId="2" pageNumber="577">K. Yunokawa</collectorName>
&amp; Y. 
<collectorName id="E81C04AFFFD7780764E25133FDCBF8A2" box="[511,584,1844,1869]" pageId="2" pageNumber="577">Fujita.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="45566179FFD7780765DA515FFA1BF832" blockId="2.[151,1437,1772,2013]" pageId="2" pageNumber="577">
<materialsCitation id="F5816B24FFD7780765DA515FFE8CF857" ID-GBIF-Occurrence="1802522158" collectingDate="2015-08-03" collectionCode="SCUBA" collectorName="K. Yunokawa" country="Japan" county="Ryukyu Islands" elevation="12" location="Material" municipality="Ie Island" pageId="2" pageNumber="577" specimenCount="2" stateProvince="Okinawa" typeStatus="paratype">
<typeStatus id="9A52DFDBFFD7780765DA515FFEB5F89F" box="[199,310,1880,1904]" pageId="2" pageNumber="577" type="paratype">Paratypes</typeStatus>
: RUMF-ZC-02733, non-ovigerous individual (cl 
<quantity id="8211CC9CFFD77807667F5150FC3EF89F" box="[866,957,1879,1904]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="2" pageNumber="577" unit="mm" value="5.6">5.6 mm</quantity>
), same data as holotype; CBM-ZC 14137, ovigerous individual (cl 
<quantity id="8211CC9CFFD7780764AB517BFD96F87B" box="[438,533,1916,1941]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="2" pageNumber="577" unit="mm" value="5.8">5.8 mm</quantity>
), same locality as 
<typeStatus id="9A52DFDBFFD7780767E5517BFCD8F87A" box="[760,859,1916,1941]" pageId="2" pageNumber="577" type="holotype">holotype</typeStatus>
, 
<elevation id="CEC4864AFFD778076671517BFC2FF87B" box="[876,940,1916,1940]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="577" unit="m" value="12.0">
<quantity id="8211CC9CFFD778076671517BFC2FF87B" box="[876,940,1916,1940]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="577" unit="m" value="12.0">12 m</quantity>
</elevation>
, 
<collectingDate id="2113BE51FFD7780766A3517BFBECF87A" box="[958,1135,1916,1941]" pageId="2" pageNumber="577" value="2015-08-03">
<date id="315747B9FFD7780766A3517BFBECF87A" box="[958,1135,1916,1941]" pageId="2" pageNumber="577" value="2015-08-03">3 August 2015</date>
</collectingDate>
, 
<collectionCode id="23F8F9BCFFD77807619C517BFB5EF87B" box="[1153,1245,1916,1940]" pageId="2" pageNumber="577">SCUBA</collectionCode>
diving, coll. 
<collectorName id="E81C04AFFFD77807609F517AFE8CF857" pageId="2" pageNumber="577">K. Yunokawa</collectorName>
</materialsCitation>
; 
<materialsCitation id="F5816B24FFD7780764045198FE67F832" ID-GBIF-Occurrence="1802522161" collectingDate="2017-06-23" collectionCode="SCUBA" collectorName="Y. Fujita" country="Japan" county="Ryukyu Islands" elevation="12" location="Material" municipality="Ie Island" pageId="2" pageNumber="577" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
RUMF-ZC-04569, non-ovigerous individual (cl 
<quantity id="8211CC9CFFD7780766335198FC05F857" box="[814,902,1951,1976]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="577" unit="mm" value="5.7">5.7 mm</quantity>
), same locality as holotype, 
<elevation id="CEC4864AFFD7780761A051A7FB7BF858" box="[1213,1272,1952,1976]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="577" unit="m" value="12.0">
<quantity id="8211CC9CFFD7780761A051A7FB7BF858" box="[1213,1272,1952,1976]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="577" unit="m" value="12.0">12 m</quantity>
</elevation>
, 
<collectingDate id="2113BE51FFD77807601E51A7FA1BF857" box="[1283,1432,1952,1976]" pageId="2" pageNumber="577" value="2017-06-23">
<date id="315747B9FFD77807601E51A7FA1BF857" box="[1283,1432,1952,1976]" pageId="2" pageNumber="577" value="2017-06-23">23 June 2017</date>
</collectingDate>
, 
<collectionCode id="23F8F9BCFFD77807658A51C3FF70F833" box="[151,243,1988,2012]" pageId="2" pageNumber="577">SCUBA</collectionCode>
diving, coll. 
<collectorName id="E81C04AFFFD77807649E51C2FE67F832" box="[387,484,1988,2013]" pageId="2" pageNumber="577">Y. Fujita</collectorName>
</materialsCitation>
; 
<materialsCitation id="F5816B24FFD7780764F251C3FA1BF832" ID-GBIF-Occurrence="1802522160" box="[495,1432,1988,2013]" collectingDate="2017-06-23" collectionCode="RUMF-ZC" collectorName="Y. Fujita" country="Japan" county="Ryukyu Islands" elevation="12" location="Material" municipality="Ie Island" pageId="2" pageNumber="577" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
RUMF-ZC-04570, ovigerous individual (cl 
<quantity id="8211CC9CFFD7780766CD51C3FBABF833" box="[976,1064,1988,2013]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="577" unit="mm" value="5.7">5.7 mm</quantity>
), same data as 
<collectionCode id="23F8F9BCFFD7780761D651C3FAC5F833" box="[1227,1350,1988,2012]" pageId="2" pageNumber="577">RUMF-ZC</collectionCode>
0 4569.
</materialsCitation>
</paragraph>
<paragraph id="45566179FFD6780665DA569FFE05FF3A" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="578">
Non-type: RUMF-ZC-04568, non-ovigerous individual (cl 6.0 mm; damaged for 
<collectionCode id="23F8F9BCFFD678066142569FFB1FFF5F" box="[1119,1180,152,176]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="3" pageNumber="578">DNA</collectionCode>
extraction), same data as 
<collectionCode id="23F8F9BCFFD6780665A856BBFEB1FF3B" box="[181,306,188,212]" pageId="3" pageNumber="578">RUMF-ZC</collectionCode>
0 4569.
</paragraph>
</subSubSection>
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<paragraph id="45566179FFD6780665DA56D8FA91FE1A" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="578">
<emphasis id="779DBD6BFFD6780665DA56D8FEDAFF17" bold="true" box="[199,345,223,248]" pageId="3" pageNumber="578">Description.</emphasis>
<typeStatus id="9A52DFDBFFD67806647F56D8FE4BFF17" box="[354,456,223,248]" pageId="3" pageNumber="578" type="holotype">
<emphasis id="779DBD6BFFD67806647F56D8FE4BFF17" box="[354,456,223,248]" italics="true" pageId="3" pageNumber="578">Holotype</emphasis>
</typeStatus>
. Body (
<figureCitation id="DDD27DFCFFD67806673556D8FDE8FF17" box="[552,619,223,248]" captionStart="FIGURE 1" captionStartId="4.[151,250,1973,1995]" captionTargetBox="[225,1325,207,1939]" captionTargetId="figure@4.[196,1392,183,1962]" captionTargetPageId="4" captionText="FIGURE 1. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm), habitus in lateral view (antennular and antennal flagella partially omitted)." httpUri="https://zenodo.org/record/1136496/files/figure.png" pageId="3" pageNumber="578">Fig. 1</figureCitation>
) moderately stout. Rostrum (
<figureCitation id="DDD27DFCFFD6780666DC56D8FBBFFF17" box="[961,1084,223,248]" captionStart="FIGURE 2" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[244,1334,231,1786]" captionTargetId="figure@5.[216,1376,193,1815]" captionTargetPageId="5" captionText="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="3" pageNumber="578">Fig. 2A–C</figureCitation>
) not reaching distal margin of article 1 of antennular peduncle, trigonal in cross-section with carinate dorsal and lateral margins, longer than wide at base, directed forward; tip sharply pointed; dorsal mid-line somewhat elevated, mid-dorsal carina extending to gastric region beyond postrostral spine; lateral carina not reaching rostral apex, merging into orbital margin; ventral surface slightly convex but not medially carinate, unarmed or armed with 1 minute subdistal spine; lateral margins nearly straight in dorsal view; ventral surface unarmed. Carapace (
<figureCitation id="DDD27DFCFFD6780666BD5793FC72FE43" box="[928,1009,404,429]" captionStart="FIGURE 1" captionStartId="4.[151,250,1973,1995]" captionTargetBox="[225,1325,207,1939]" captionTargetId="figure@4.[196,1392,183,1962]" captionTargetPageId="4" captionText="FIGURE 1. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm), habitus in lateral view (antennular and antennal flagella partially omitted)." httpUri="https://zenodo.org/record/1136496/files/figure.png" pageId="3" pageNumber="578">Figs. 1</figureCitation>
, 
<figureCitation id="DDD27DFCFFD67806611F5793FBC5FE43" box="[1026,1094,404,428]" captionStart="FIGURE 2" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[244,1334,231,1786]" captionTargetId="figure@5.[216,1376,193,1815]" captionTargetPageId="5" captionText="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="3" pageNumber="578">2A–C</figureCitation>
) glabrous, with 1 small middorsal spine at postrostral position; anterolateral margin gently sinuous, with small extra-orbital spine lateral to eyestalk; cardiac notch shallow; orbital spine directed forward; pterygostomial angle broadly rounded.
</paragraph>
<paragraph id="45566179FFD6780665DA57F8FC79FD1F" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="578">
Pleon (
<figureCitation id="DDD27DFCFFD67806640457F8FEDFFDF7" box="[281,348,511,536]" captionStart="FIGURE 1" captionStartId="4.[151,250,1973,1995]" captionTargetBox="[225,1325,207,1939]" captionTargetId="figure@4.[196,1392,183,1962]" captionTargetPageId="4" captionText="FIGURE 1. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm), habitus in lateral view (antennular and antennal flagella partially omitted)." httpUri="https://zenodo.org/record/1136496/files/figure.png" pageId="3" pageNumber="578">Fig. 1</figureCitation>
) with anterior 3 pleura rounded ventrally and posteroventrally; pleuron 4 somewhat produced posteriorly but unarmed; pleuron 5 with acutely pointed posteroventral angle. Pleomere 6 without articulated flap at posteroventral angle; preanal plate rounded; posterolateral process truncate. Telson (
<figureCitation id="DDD27DFCFFD6780661615440FB4FFD8F" box="[1148,1228,583,608]" captionStart="FIGURE 1" captionStartId="4.[151,250,1973,1995]" captionTargetBox="[225,1325,207,1939]" captionTargetId="figure@4.[196,1392,183,1962]" captionTargetPageId="4" captionText="FIGURE 1. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm), habitus in lateral view (antennular and antennal flagella partially omitted)." httpUri="https://zenodo.org/record/1136496/files/figure.png" pageId="3" pageNumber="578">Figs. 1</figureCitation>
, 
<figureCitation id="DDD27DFCFFD6780661C0544FFA83FD8F" box="[1245,1280,584,608]" captionStart="FIGURE 2" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[244,1334,231,1786]" captionTargetId="figure@5.[216,1376,193,1815]" captionTargetPageId="5" captionText="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="3" pageNumber="578">2D</figureCitation>
) 1.1 times as long as pleomere 6, with 2 pairs of dorsolateral spiniform setae on posterior half situated laterally from blunt dorsolateral ridge; posterior margin with very deep, narrow, V-shaped cleft, remaining margin truncate, bearing 2 greatly unequal spiniform setae, lateral about twice as long as mesial (
<figureCitation id="DDD27DFCFFD6780666CF54B3FBAEFD23" box="[978,1069,692,717]" captionStart="FIGURE 2" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[244,1334,231,1786]" captionTargetId="figure@5.[216,1376,193,1815]" captionTargetPageId="5" captionText="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="3" pageNumber="578">Fig. 2E</figureCitation>
); dorsal surface with 2 shallow longitudinal sulci, one on each side of mid-line, reaching to posterior margin.
</paragraph>
<paragraph id="45566179FFD6780665DA54FBFC66FCD7" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="578">
Eyestalks (
<figureCitation id="DDD27DFCFFD67806645F54FBFE3FFCFB" box="[322,444,764,789]" captionStart="FIGURE 2" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[244,1334,231,1786]" captionTargetId="figure@5.[216,1376,193,1815]" captionTargetPageId="5" captionText="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="3" pageNumber="578">Fig. 2A, B</figureCitation>
) exposed both in dorsal and lateral views, each with small distal spinule at dorsomesial angle; cornea anterolateral, substantially reduced in size, darkly pigmented.
</paragraph>
<paragraph id="45566179FFD6780665DA5543FC32FC02" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="578">
Antennular peduncle (
<figureCitation id="DDD27DFCFFD6780664DE5543FDBDFCB3" box="[451,574,836,861]" captionStart="FIGURE 2" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[244,1334,231,1786]" captionTargetId="figure@5.[216,1376,193,1815]" captionTargetPageId="5" captionText="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="3" pageNumber="578">Fig. 2A, C</figureCitation>
) stout, overreaching distal margin of antennal scaphocerite by 0.3 length of article 3. Article 1 with small forwardly directed spine on ventromesial ridge at midlength; stylocerite terminating in slender, acute spine reaching to distal margin of article 1. Distomesial angle of article 2 somewhat produced distally. Flagella very long, more than twice body length; lateral flagellum (
<figureCitation id="DDD27DFCFFD6780666F355A8FBC4FC27" box="[1006,1095,943,968]" captionStart="FIGURE 2" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[244,1334,231,1786]" captionTargetId="figure@5.[216,1376,193,1815]" captionTargetPageId="5" captionText="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="3" pageNumber="578">Fig. 2A</figureCitation>
) biramous, fused portion very short; accessory ramus consisting of about 10 articles with aesthetascs.
</paragraph>
<paragraph id="45566179FFD6780665DA55FFFAAFFBB7" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="578">
Antenna (
<figureCitation id="DDD27DFCFFD67806642A55F0FE56FBFF" box="[311,469,1015,1040]" captionStart="FIGURE 2" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[244,1334,231,1786]" captionTargetId="figure@5.[216,1376,193,1815]" captionTargetPageId="5" captionText="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="3" pageNumber="578">Fig. 2A, C, F</figureCitation>
) with basicerite bearing small ventrolateral distal spine. Scaphocerite subovate with nearly straight lateral margin and gently convex mesial margin, 2.4 times as long as wide; distal blade broadly rounded, reaching well beyond small distolateral tooth. Carpocerite reaching mid-length of scaphocerite.
</paragraph>
<paragraph id="45566179FFD6780665DA5263FC4EFB92" blockId="3.[151,1437,151,2013]" box="[199,973,1124,1149]" pageId="3" pageNumber="578">Epistomial sclerites unarmed. Labrum with non-crested median lobe.</paragraph>
<paragraph id="45566179FFD6780665DA5280FEF9F9E7" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="578">
Mandible (
<figureCitation id="DDD27DFCFFD6780664595280FE3DFB4F" box="[324,446,1159,1184]" captionStart="FIGURE 3" captionStartId="6.[151,250,1234,1256]" captionTargetBox="[186,1401,209,1202]" captionTargetId="figure@6.[151,1436,193,1229]" captionTargetPageId="6" captionText="FIGURE 3. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm), mouthparts dissected from left side. A, mandible, internal view; B, same, external view; C, same, incisor process, distal surface; D, maxillule (coxal endite missing); E, maxilla; F, maxilliped 1; G,same, lateral view of exopod and endopod; H, maxilliped 2." httpUri="https://zenodo.org/record/1136500/files/figure.png" pageId="3" pageNumber="578">Fig. 3A–C</figureCitation>
) with broad incisor process, terminal margin with 6 unequal blunt teeth; inner surface excavate, accommodate 2-articulated palp; distal article of palp subovate, marginally setose; molar process relatively narrow. Maxillule (
<figureCitation id="DDD27DFCFFD6780664FD52C8FDBBFB07" box="[480,568,1231,1256]" captionStart="FIGURE 3" captionStartId="6.[151,250,1234,1256]" captionTargetBox="[186,1401,209,1202]" captionTargetId="figure@6.[151,1436,193,1229]" captionTargetPageId="6" captionText="FIGURE 3. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm), mouthparts dissected from left side. A, mandible, internal view; B, same, external view; C, same, incisor process, distal surface; D, maxillule (coxal endite missing); E, maxilla; F, maxilliped 1; G,same, lateral view of exopod and endopod; H, maxilliped 2." httpUri="https://zenodo.org/record/1136500/files/figure.png" pageId="3" pageNumber="578">Fig. 3D</figureCitation>
) with basial endite bearing both spiniform and stiff setae, arranged in double row, on mesial margin. Maxilla (
<figureCitation id="DDD27DFCFFD67806670852F3FDEFFAE3" box="[533,620,1268,1293]" captionStart="FIGURE 3" captionStartId="6.[151,250,1234,1256]" captionTargetBox="[186,1401,209,1202]" captionTargetId="figure@6.[151,1436,193,1229]" captionTargetPageId="6" captionText="FIGURE 3. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm), mouthparts dissected from left side. A, mandible, internal view; B, same, external view; C, same, incisor process, distal surface; D, maxillule (coxal endite missing); E, maxilla; F, maxilliped 1; G,same, lateral view of exopod and endopod; H, maxilliped 2." httpUri="https://zenodo.org/record/1136500/files/figure.png" pageId="3" pageNumber="578">Fig. 3E</figureCitation>
) with poorly developed, slightly bilobed, coxal endite; endopod short, curved mesially, not extending beyond distal margin of basial endite; posterior lobe of scaphognathite short, rounded. Maxilliped 1 (
<figureCitation id="DDD27DFCFFD6780664A8533BFDB6FABB" box="[437,565,1340,1365]" captionStart="FIGURE 3" captionStartId="6.[151,250,1234,1256]" captionTargetBox="[186,1401,209,1202]" captionTargetId="figure@6.[151,1436,193,1229]" captionTargetPageId="6" captionText="FIGURE 3. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm), mouthparts dissected from left side. A, mandible, internal view; B, same, external view; C, same, incisor process, distal surface; D, maxillule (coxal endite missing); E, maxilla; F, maxilliped 1; G,same, lateral view of exopod and endopod; H, maxilliped 2." httpUri="https://zenodo.org/record/1136500/files/figure.png" pageId="3" pageNumber="578">Fig. 3F, G</figureCitation>
) with unequally bilobed coxal endite; basial endite generally oval, with longitudinal row of setae on mid-line of outer surface; exopod with narrow caridean lobe, flagellum well developed; epipod large, bilobed. Maxilliped 2 (
<figureCitation id="DDD27DFCFFD6780667A45383FC92FA73" box="[697,785,1412,1437]" captionStart="FIGURE 3" captionStartId="6.[151,250,1234,1256]" captionTargetBox="[186,1401,209,1202]" captionTargetId="figure@6.[151,1436,193,1229]" captionTargetPageId="6" captionText="FIGURE 3. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm), mouthparts dissected from left side. A, mandible, internal view; B, same, external view; C, same, incisor process, distal surface; D, maxillule (coxal endite missing); E, maxilla; F, maxilliped 1; G,same, lateral view of exopod and endopod; H, maxilliped 2." httpUri="https://zenodo.org/record/1136500/files/figure.png" pageId="3" pageNumber="578">Fig. 3H</figureCitation>
) somewhat slender; basis and ischium fused; carpus with produced mesial angle; propodus and dactylus partially fused, with row of slender spiniform setae on mesial margin; epipod large, devoid of podobranch; exopodal flagellum well developed, with 1 long plumose seta arising at about mid-length.
</paragraph>
<paragraph id="45566179FFD6780665DA5013FD14F90F" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="578">
Maxilliped 3 (
<figureCitation id="DDD27DFCFFD6780664715013FE6AF9C3" box="[364,489,1556,1581]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[213,1345,218,1829]" captionTargetId="figure@7.[187,1374,195,1844]" captionTargetPageId="7" captionText="FIGURE 4. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm). A, left maxilliped3,lateral view; B, same,coxaand proximal part of antepenultimate article, dorsal view;C, leftpereopod 1 (= cheliped), lateral view; D, same, chela, extensor view; E, right pereopod 1 (= cheliped), lateral view; F, same, chela, extensor view; G, left pereopod 2, lateral view; H, left pereopod 3, lateral view; I, same, distal part of propodus and dactylus, lateral view; J, left pereopod 4, lateral view; K, left pereopod 5, lateral view; L, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/1136502/files/figure.png" pageId="3" pageNumber="578">Fig. 4A, B</figureCitation>
) with endopod slender, not reaching distal margin of antennal scaphocerite. Coxa with lateral plate rounded-subquadrate (
<figureCitation id="DDD27DFCFFD67806676B5030FD54F9BF" box="[630,727,1591,1616]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[213,1345,218,1829]" captionTargetId="figure@7.[187,1374,195,1844]" captionTargetPageId="7" captionText="FIGURE 4. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm). A, left maxilliped3,lateral view; B, same,coxaand proximal part of antepenultimate article, dorsal view;C, leftpereopod 1 (= cheliped), lateral view; D, same, chela, extensor view; E, right pereopod 1 (= cheliped), lateral view; F, same, chela, extensor view; G, left pereopod 2, lateral view; H, left pereopod 3, lateral view; I, same, distal part of propodus and dactylus, lateral view; J, left pereopod 4, lateral view; K, left pereopod 5, lateral view; L, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/1136502/files/figure.png" pageId="3" pageNumber="578">Fig. 4B</figureCitation>
). Antepenultimate article sinuously curved in dorsal view, proximal half flattened dorsoventrally. Penultimate article subcylindrical. Ultimate article 1.5 times as long as penultimate article, tapering distally from mid-length, terminating in acute tip, latter bearing 1 long subterminal seta but no stout spiniform setae; mesial face with several transverse rows of stiff serrulate setae. Exopod not reaching distal end of antepenultimate article.
</paragraph>
<paragraph id="45566179FFD6780665DA50EBFB13F832" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="578">
Pereopods 1 (= chelipeds) (
<figureCitation id="DDD27DFCFFD67806670E50EBFD17F8EB" box="[531,660,1772,1797]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[213,1345,218,1829]" captionTargetId="figure@7.[187,1374,195,1844]" captionTargetPageId="7" captionText="FIGURE 4. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm). A, left maxilliped3,lateral view; B, same,coxaand proximal part of antepenultimate article, dorsal view;C, leftpereopod 1 (= cheliped), lateral view; D, same, chela, extensor view; E, right pereopod 1 (= cheliped), lateral view; F, same, chela, extensor view; G, left pereopod 2, lateral view; H, left pereopod 3, lateral view; I, same, distal part of propodus and dactylus, lateral view; J, left pereopod 4, lateral view; K, left pereopod 5, lateral view; L, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/1136502/files/figure.png" pageId="3" pageNumber="578">Fig. 4C, E</figureCitation>
) subequal in size, symmetrical in shape, slender, simple, carried extended or loosely folded, in full extension only slightly overreaching distal margin of antennal scaphocerite. Ischium with 4 (left) or 6 (right) spiniform setae on ventrolateral surface. Merus slightly longer than ischium, subcylindrical. Carpus subequal in length to merus, slightly widening distally, 8 times as long as distal width. Chela (
<figureCitation id="DDD27DFCFFD678066582517BFE94F87B" box="[159,279,1916,1941]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[213,1345,218,1829]" captionTargetId="figure@7.[187,1374,195,1844]" captionTargetPageId="7" captionText="FIGURE 4. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm). A, left maxilliped3,lateral view; B, same,coxaand proximal part of antepenultimate article, dorsal view;C, leftpereopod 1 (= cheliped), lateral view; D, same, chela, extensor view; E, right pereopod 1 (= cheliped), lateral view; F, same, chela, extensor view; G, left pereopod 2, lateral view; H, left pereopod 3, lateral view; I, same, distal part of propodus and dactylus, lateral view; J, left pereopod 4, lateral view; K, left pereopod 5, lateral view; L, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/1136502/files/figure.png" pageId="3" pageNumber="578">Fig. 4D, F</figureCitation>
) tapering distally, 0.7 times as long as carpus; palm subcylindrical; fingers 1.4 times as long as palm, not crossing distally; occlusal margin of fixed finger with 3 very small teeth proximal to mid-length, some teeth directed slightly backwards; dactylus slightly curved distally, unarmed on occlusal margin.
</paragraph>
<caption id="119631F1FFD17801658A51B2FC9DF806" httpUri="https://zenodo.org/record/1136496/files/figure.png" pageId="4" pageNumber="579" targetBox="[225,1325,207,1939]" targetPageId="4">
<paragraph id="45566179FFD17801658A51B2FC9DF806" blockId="4.[151,1436,1972,2025]" pageId="4" pageNumber="579">
<emphasis id="779DBD6BFFD17801658A51B2FE96F825" bold="true" box="[151,277,1973,1995]" pageId="4" pageNumber="579">FIGURE 1.</emphasis>
<taxonomicName id="82E91AFAFFD17801640051B3FD92F825" authority="Komai &amp; Fujita, 2018" authorityName="Komai &amp; Fujita" authorityYear="2018" box="[285,529,1972,1994]" class="Malacostraca" family="Alpheidae" genus="Caligoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="579" phylum="Arthropoda" rank="species" species="cavernicola" status="gen. nov.">
<emphasis id="779DBD6BFFD17801640051B3FD92F825" box="[285,529,1972,1994]" italics="true" pageId="4" pageNumber="579">Caligoneus cavernicola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CAE0010FFD17801670451B2FD27F825" box="[537,676,1973,1994]" pageId="4" pageNumber="579" rank="species">
<emphasis id="779DBD6BFFD17801670451B2FD27F825" bold="true" box="[537,676,1973,1994]" pageId="4" pageNumber="579">n.gen., n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm), habitus in lateral view (antennular and antennal flagella partially omitted).
</paragraph>
</caption>
<caption id="119631F1FFD07800658A5127FE06F801" httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="5" pageNumber="580" targetBox="[244,1334,231,1786]" targetPageId="5">
<paragraph id="45566179FFD07800658A5127FE06F801" blockId="5.[151,1436,1824,2030]" pageId="5" pageNumber="580">
<emphasis id="779DBD6BFFD07800658A5127FE97F8DA" bold="true" box="[151,276,1824,1846]" pageId="5" pageNumber="580">FIGURE 2.</emphasis>
<taxonomicName id="82E91AFAFFD0780064005127FD93F8D9" authority="Komai &amp; Fujita, 2018" authorityName="Komai &amp; Fujita" authorityYear="2018" box="[285,528,1824,1846]" class="Malacostraca" family="Alpheidae" genus="Caligoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="580" phylum="Arthropoda" rank="species" species="cavernicola" status="gen. nov.">
<emphasis id="779DBD6BFFD0780064005127FD93F8D9" box="[285,528,1824,1846]" italics="true" pageId="5" pageNumber="580">Caligoneus cavernicola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CAE0010FFD0780067055127FD21F8DA" box="[536,674,1824,1845]" pageId="5" pageNumber="580" rank="species">
<emphasis id="779DBD6BFFD0780067055127FD21F8DA" bold="true" box="[536,674,1824,1845]" pageId="5" pageNumber="580">n.gen., n. sp.</emphasis>
</taxonomicNameLabel>
A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137, ovigerous individual (cl 5.8 mm). A, anterior part of carapace and cephalic appendages, lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view.
</paragraph>
</caption>
<caption id="119631F1FFD37803658A52D5FA03FAC9" httpUri="https://zenodo.org/record/1136500/files/figure.png" pageId="6" pageNumber="581" targetBox="[186,1401,209,1202]" targetPageId="6">
<paragraph id="45566179FFD37803658A52D5FA03FAC9" blockId="6.[151,1436,1234,1318]" pageId="6" pageNumber="581">
<emphasis id="779DBD6BFFD37803658A52D5FE97FB08" bold="true" box="[151,276,1234,1256]" pageId="6" pageNumber="581">FIGURE 3.</emphasis>
<taxonomicName id="82E91AFAFFD37803640152D5FD8DFB07" authority="Komai &amp; Fujita, 2018" authorityName="Komai &amp; Fujita" authorityYear="2018" box="[284,526,1234,1256]" class="Malacostraca" family="Alpheidae" genus="Caligoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="581" phylum="Arthropoda" rank="species" species="cavernicola" status="gen. nov.">
<emphasis id="779DBD6BFFD37803640152D5FD8DFB07" box="[284,526,1234,1256]" italics="true" pageId="6" pageNumber="581">Caligoneus cavernicola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CAE0010FFD37803670852D4FD1CFB07" box="[533,671,1235,1256]" pageId="6" pageNumber="581" rank="species">
<emphasis id="779DBD6BFFD37803670852D4FD1CFB07" bold="true" box="[533,671,1235,1256]" pageId="6" pageNumber="581">n.gen., n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm), mouthparts dissected from left side. A, mandible, internal view; B, same, external view; C, same, incisor process, distal surface; D, maxillule (coxal endite missing); E, maxilla; F, maxilliped 1; G, same, lateral view of exopod and endopod; H, maxilliped 2.
</paragraph>
</caption>
<paragraph id="45566179FFD3780365DA5355FEE5FA15" blockId="6.[151,1436,1362,2035]" pageId="6" pageNumber="581">
Pereopod 2 (
<figureCitation id="DDD27DFCFFD3780364455355FE32FA84" box="[344,433,1362,1387]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[213,1345,218,1829]" captionTargetId="figure@7.[187,1374,195,1844]" captionTargetPageId="7" captionText="FIGURE 4. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm). A, left maxilliped3,lateral view; B, same,coxaand proximal part of antepenultimate article, dorsal view;C, leftpereopod 1 (= cheliped), lateral view; D, same, chela, extensor view; E, right pereopod 1 (= cheliped), lateral view; F, same, chela, extensor view; G, left pereopod 2, lateral view; H, left pereopod 3, lateral view; I, same, distal part of propodus and dactylus, lateral view; J, left pereopod 4, lateral view; K, left pereopod 5, lateral view; L, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/1136502/files/figure.png" pageId="6" pageNumber="581">Fig. 4G</figureCitation>
) slender, overreaching distal margin of antennal scale by half-length of carpus. Ischium with 3 spiniform setae on ventrolateral surface. Merus subequal in length to ischium, flexor side slightly inflated distally. Carpus 0.8 times as long as ischium and merus combined, subdivided into 5 articles, ratio of carpal articles from proximal to distal equal to: 5: 0.8: 0.8: 1.3: 1. Chela slightly longer than distal-most carpal article, simple, with tufts of setae.
</paragraph>
<paragraph id="45566179FFD3780365DA5001FBE0F88C" blockId="6.[151,1436,1362,2035]" pageId="6" pageNumber="581">
Pereopods 3–5 very slender, elongate, pereopods 3 and 4 subequal in length, pereopod 5 distinctly longer than each of pereopods 3 and 4. Pereopod 3 (
<figureCitation id="DDD27DFCFFD378036790502DFD68F9AC" box="[653,747,1578,1603]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[213,1345,218,1829]" captionTargetId="figure@7.[187,1374,195,1844]" captionTargetPageId="7" captionText="FIGURE 4. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm). A, left maxilliped3,lateral view; B, same,coxaand proximal part of antepenultimate article, dorsal view;C, leftpereopod 1 (= cheliped), lateral view; D, same, chela, extensor view; E, right pereopod 1 (= cheliped), lateral view; F, same, chela, extensor view; G, left pereopod 2, lateral view; H, left pereopod 3, lateral view; I, same, distal part of propodus and dactylus, lateral view; J, left pereopod 4, lateral view; K, left pereopod 5, lateral view; L, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/1136502/files/figure.png" pageId="6" pageNumber="581">Fig. 4H</figureCitation>
) in full extension overreaching distal margin of antennal scaphocerite by 0.4 length of carpus; ischium without spiniform setae; merus unarmed, flexor side slightly inflated distally; carpus slenderer than merus, 1.3 times as long as merus, lacking distodorsal projection; propodus with 5 widely spaced, minute spiniform setae (including one adjacent to propodo-dactylar articulation) on flexor margin; dactylus 0.1 times as long as propodus, slender (9 times as long as wide), simple, almost straight (
<figureCitation id="DDD27DFCFFD37803602650BDFA0CF93C" box="[1339,1423,1722,1747]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[213,1345,218,1829]" captionTargetId="figure@7.[187,1374,195,1844]" captionTargetPageId="7" captionText="FIGURE 4. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm). A, left maxilliped3,lateral view; B, same,coxaand proximal part of antepenultimate article, dorsal view;C, leftpereopod 1 (= cheliped), lateral view; D, same, chela, extensor view; E, right pereopod 1 (= cheliped), lateral view; F, same, chela, extensor view; G, left pereopod 2, lateral view; H, left pereopod 3, lateral view; I, same, distal part of propodus and dactylus, lateral view; J, left pereopod 4, lateral view; K, left pereopod 5, lateral view; L, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/1136502/files/figure.png" pageId="6" pageNumber="581">
Fig. 
<date id="315747B9FFD37803606950BCFA0CF93C" box="[1396,1423,1723,1747]" pageId="6" pageNumber="581">4I</date>
</figureCitation>
). Pereopod 4 (
<figureCitation id="DDD27DFCFFD37803643550D9FEFBF918" box="[296,376,1758,1783]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[213,1345,218,1829]" captionTargetId="figure@7.[187,1374,195,1844]" captionTargetPageId="7" captionText="FIGURE 4. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm). A, left maxilliped3,lateral view; B, same,coxaand proximal part of antepenultimate article, dorsal view;C, leftpereopod 1 (= cheliped), lateral view; D, same, chela, extensor view; E, right pereopod 1 (= cheliped), lateral view; F, same, chela, extensor view; G, left pereopod 2, lateral view; H, left pereopod 3, lateral view; I, same, distal part of propodus and dactylus, lateral view; J, left pereopod 4, lateral view; K, left pereopod 5, lateral view; L, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/1136502/files/figure.png" pageId="6" pageNumber="581">Fig. 4J</figureCitation>
) similar to pereopod 3, overreaching distal margin of antennal scaphocerite by full length of propodus; ischium unarmed. Pereopod 5 (
<figureCitation id="DDD27DFCFFD3780367605105FD5BF8F4" box="[637,728,1794,1819]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[213,1345,218,1829]" captionTargetId="figure@7.[187,1374,195,1844]" captionTargetPageId="7" captionText="FIGURE 4. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm). A, left maxilliped3,lateral view; B, same,coxaand proximal part of antepenultimate article, dorsal view;C, leftpereopod 1 (= cheliped), lateral view; D, same, chela, extensor view; E, right pereopod 1 (= cheliped), lateral view; F, same, chela, extensor view; G, left pereopod 2, lateral view; H, left pereopod 3, lateral view; I, same, distal part of propodus and dactylus, lateral view; J, left pereopod 4, lateral view; K, left pereopod 5, lateral view; L, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/1136502/files/figure.png" pageId="6" pageNumber="581">Fig. 4K</figureCitation>
) overreaching distal margin of scaphocerite by 0.4 length of carpus; ischium unarmed; carpus and propodus elongate, with setal brush on distal 0.4 of flexor margin (
<figureCitation id="DDD27DFCFFD37803602A5121FA0CF8D0" box="[1335,1423,1830,1855]" captionStart="FIGURE 4" captionStartId="7.[151,250,1877,1899]" captionTargetBox="[213,1345,218,1829]" captionTargetId="figure@7.[187,1374,195,1844]" captionTargetPageId="7" captionText="FIGURE 4. Caligoneus cavernicola n.gen., n. sp., holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm). A, left maxilliped3,lateral view; B, same,coxaand proximal part of antepenultimate article, dorsal view;C, leftpereopod 1 (= cheliped), lateral view; D, same, chela, extensor view; E, right pereopod 1 (= cheliped), lateral view; F, same, chela, extensor view; G, left pereopod 2, lateral view; H, left pereopod 3, lateral view; I, same, distal part of propodus and dactylus, lateral view; J, left pereopod 4, lateral view; K, left pereopod 5, lateral view; L, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/1136502/files/figure.png" pageId="6" pageNumber="581">Fig. 4L</figureCitation>
); dactylus similar to those of pereopods 3 and 4, less than 0.2 times as long as propodus.
</paragraph>
<paragraph id="45566179FFD3780365DA5169FA95F81C" blockId="6.[151,1436,1362,2035]" pageId="6" pageNumber="581">
Pleopod 1 (
<figureCitation id="DDD27DFCFFD3780364565169FE27F868" box="[331,420,1902,1927]" captionStart="FIGURE 2" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[244,1334,231,1786]" captionTargetId="figure@5.[216,1376,193,1815]" captionTargetPageId="5" captionText="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="6" pageNumber="581">Fig. 2G</figureCitation>
) with endopod about 0.3 length of exopod, gently curved inward, tapering to blunt apex (
<figureCitation id="DDD27DFCFFD3780365825195FF79F844" box="[159,250,1938,1963]" captionStart="FIGURE 2" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[244,1334,231,1786]" captionTargetId="figure@5.[216,1376,193,1815]" captionTargetPageId="5" captionText="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="6" pageNumber="581">Fig. 2H</figureCitation>
). Pleopod 2 (
<figureCitation id="DDD27DFCFFD3780364925195FE5FF844" box="[399,476,1938,1963]" captionStart="FIGURE 2" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[244,1334,231,1786]" captionTargetId="figure@5.[216,1376,193,1815]" captionTargetPageId="5" captionText="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="6" pageNumber="581">
Fig. 
<date id="315747B9FFD3780364DE5194FE5FF844" box="[451,476,1939,1963]" pageId="6" pageNumber="581">2I</date>
</figureCitation>
) with appendices interna and masculina present in both non-ovigerous and ovigerous individuals; appendix masculina (
<figureCitation id="DDD27DFCFFD37803670F51B1FDE2F820" box="[530,609,1974,1999]" captionStart="FIGURE 2" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[244,1334,231,1786]" captionTargetId="figure@5.[216,1376,193,1815]" captionTargetPageId="5" captionText="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="6" pageNumber="581">Fig. 2J</figureCitation>
) about 0.8 length of appendix interna, slightly widened to distal 0.2, then tapering to subacute apex, with row of setae distolaterally; endopod about 0.8 times as long as exopod.
</paragraph>
<caption id="119631F1FFD27802658A5152FA90F80A" httpUri="https://zenodo.org/record/1136502/files/figure.png" pageId="7" pageNumber="582" targetBox="[213,1345,218,1829]" targetPageId="7">
<paragraph id="45566179FFD27802658A5152FA90F80A" blockId="7.[151,1436,1876,2021]" pageId="7" pageNumber="582">
<emphasis id="779DBD6BFFD27802658A5152FE9AF885" bold="true" box="[151,281,1877,1899]" pageId="7" pageNumber="582">FIGURE 4.</emphasis>
<taxonomicName id="82E91AFAFFD2780264385153FD94F885" authority="Komai &amp; Fujita, 2018" authorityName="Komai &amp; Fujita" authorityYear="2018" box="[293,535,1876,1898]" class="Malacostraca" family="Alpheidae" genus="Caligoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="582" phylum="Arthropoda" rank="species" species="cavernicola" status="gen. nov.">
<emphasis id="779DBD6BFFD2780264385153FD94F885" box="[293,535,1876,1898]" italics="true" pageId="7" pageNumber="582">Caligoneus cavernicola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CAE0010FFD27802673E5152FD31F885" box="[547,690,1877,1898]" pageId="7" pageNumber="582" rank="species">
<emphasis id="779DBD6BFFD27802673E5152FD31F885" bold="true" box="[547,690,1877,1898]" pageId="7" pageNumber="582">n.gen., n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm). A, left maxilliped 3, lateral view; B, same, coxa and proximal part of antepenultimate article, dorsal view; C, left pereopod 1 (= cheliped), lateral view; D, same, chela, extensor view; E, right pereopod 1 (= cheliped), lateral view; F, same, chela, extensor view; G, left pereopod 2, lateral view; H, left pereopod 3, lateral view; I, same, distal part of propodus and dactylus, lateral view; J, left pereopod 4, lateral view; K, left pereopod 5, lateral view; L, same, distal part of propodus and dactylus, lateral view.
</paragraph>
</caption>
<caption id="119631F1FFDD780D658A51A0FBD1F833" httpUri="https://zenodo.org/record/1136504/files/figure.png" pageId="8" pageNumber="583" targetBox="[150,1435,217,1919]" targetPageId="8">
<paragraph id="45566179FFDD780D658A51A0FBD1F833" blockId="8.[151,1436,1959,2012]" pageId="8" pageNumber="583">
<emphasis id="779DBD6BFFDD780D658A51A0FE91F852" bold="true" box="[151,274,1959,1981]" pageId="8" pageNumber="583">FIGURE 5.</emphasis>
<taxonomicName id="82E91AFAFFDD780D640451A0FD88F852" authority="Komai &amp; Fujita, 2018" authorityName="Komai &amp; Fujita" authorityYear="2018" box="[281,523,1959,1981]" class="Malacostraca" family="Alpheidae" genus="Caligoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="583" phylum="Arthropoda" rank="species" species="cavernicola" status="gen. nov.">
<emphasis id="779DBD6BFFDD780D640451A0FD88F852" box="[281,523,1959,1981]" italics="true" pageId="8" pageNumber="583">Caligoneus cavernicola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CAE0010FFDD780D670C51AFFD1BF852" box="[529,664,1960,1981]" pageId="8" pageNumber="583" rank="species">
<emphasis id="779DBD6BFFDD780D670C51AFFD1BF852" bold="true" box="[529,664,1960,1981]" pageId="8" pageNumber="583">
<taxonomicNameLabel id="6CAE0010FFDD780D670C51AFFDD0F852" box="[529,595,1960,1981]" pageId="8" pageNumber="583" rank="genus">n.gen.</taxonomicNameLabel>
, n. sp.
</emphasis>
</taxonomicNameLabel>
, general habitus and colour in life. A, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, RUMF-ZC-02733, non-ovigerous individual (cl 5.6 mm).
</paragraph>
</caption>
<paragraph id="45566179FFDC780C65DA5690FBE8FEF2" blockId="9.[151,1437,151,1368]" pageId="9" pageNumber="584">
Uropod (
<figureCitation id="DDD27DFCFFDC780C64325690FE09FF5F" box="[303,394,151,176]" captionStart="FIGURE 2" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[244,1334,231,1786]" captionTargetId="figure@5.[216,1376,193,1815]" captionTargetPageId="5" captionText="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="9" pageNumber="584">Fig. 2D</figureCitation>
) with lateral lobe of protopod unarmed. Endopod and exopod narrowly subovoid, former shorter than latter, reaching posterior margin of telson. Exopod with faintly concave, setose lateral margin terminating distally in minute spine covering base of minute distolateral spiniform seta; diaeresis sinuous, without spine mesial to distolateral spine; distal lamella extending far beyond distolateral spine.
</paragraph>
<paragraph id="45566179FFDC780C65DA572FFD13FED0" blockId="9.[151,1437,151,1368]" box="[199,656,295,320]" pageId="9" pageNumber="584">
Eggs at stage near hatching 0.9× 
<quantity id="8211CC9CFFDC780C672E5720FD0FFED0" box="[563,652,295,320]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="584" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
<paragraph id="45566179FFDC780C65DA574BFC76FD6B" blockId="9.[151,1437,151,1368]" pageId="9" pageNumber="584">
<emphasis id="779DBD6BFFDC780C65DA574BFEB6FE8A" bold="true" box="[199,309,332,357]" pageId="9" pageNumber="584">Variation</emphasis>
. The remaining five specimens (four 
<typeStatus id="9A52DFDBFFDC780C67FB574AFCD2FE8A" box="[742,849,333,357]" pageId="9" pageNumber="584" type="paratype">paratypes</typeStatus>
and one non-type) are generally very similar to the 
<typeStatus id="9A52DFDBFFDC780C658A5768FF7EFE67" box="[151,253,367,392]" pageId="9" pageNumber="584" type="holotype">holotype</typeStatus>
. One of them (
<typeStatus id="9A52DFDBFFDC780C64BE5777FD8BFE67" box="[419,520,368,392]" pageId="9" pageNumber="584" type="paratype">paratype</typeStatus>
, CBM-ZC 14137) has two postrostral spines (
<figureCitation id="DDD27DFCFFDC780C61135768FBE5FE67" box="[1038,1126,367,392]" captionStart="FIGURE 2" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[244,1334,231,1786]" captionTargetId="figure@5.[216,1376,193,1815]" captionTargetPageId="5" captionText="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="9" pageNumber="584">Fig. 2B</figureCitation>
), whilst the other four have one postrostral spine, as in the 
<typeStatus id="9A52DFDBFFDC780C671C5793FDE4FE42" box="[513,615,404,429]" pageId="9" pageNumber="584" type="holotype">holotype</typeStatus>
. In three specimens (two 
<typeStatus id="9A52DFDBFFDC780C668F5792FB83FE42" box="[914,1024,405,429]" pageId="9" pageNumber="584" type="paratype">paratypes</typeStatus>
, RUMF-ZC 0 4569, 0 4570, and one non-type, RUMF-ZC 04568), the rostrum has a minuscule subdistal tooth on the ventral margin, such a tooth being absent in the 
<typeStatus id="9A52DFDBFFDC780C642E57DBFE16FE1A" box="[307,405,476,501]" pageId="9" pageNumber="584" type="holotype">holotype</typeStatus>
and the remaining two 
<typeStatus id="9A52DFDBFFDC780C67B757DAFC96FE1A" box="[682,789,477,501]" pageId="9" pageNumber="584" type="paratype">paratypes</typeStatus>
(CBM-ZC 14137, RUMF-ZC-02733). The ischia of the pereopods 1 and 2 are quite variable in the armature, having four to six and three or four spiniform setae, respectively. The ischium of the pereopod 3 can be either unarmed [
<typeStatus id="9A52DFDBFFDC780C668E5423FC7BFDD2" box="[915,1016,548,573]" pageId="9" pageNumber="584" type="holotype">holotype</typeStatus>
and two 
<typeStatus id="9A52DFDBFFDC780C617D5422FB48FDD2" box="[1120,1227,549,573]" pageId="9" pageNumber="584" type="paratype">paratypes</typeStatus>
(CBM-ZC 14137, RUMF-ZC-02733)] or armed with one or two spiniform setae [two 
<typeStatus id="9A52DFDBFFDC780C668B544FFB82FD8F" box="[918,1025,584,608]" pageId="9" pageNumber="584" type="paratype">paratypes</typeStatus>
(RUMF-ZC 0 4569, 04570) and one non-type (RUMF-ZC 04568)]. Eggs in non-eyed stage measure 0.7× 
<quantity id="8211CC9CFFDC780C668B546BFC73FD6B" box="[918,1008,620,644]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="584" unit="mm" value="0.4">0.4 mm</quantity>
.
</paragraph>
<paragraph id="45566179FFDC780C65DA5488FB30FD1F" blockId="9.[151,1437,151,1368]" pageId="9" pageNumber="584">
<emphasis id="779DBD6BFFDC780C65DA5488FEF7FD47" bold="true" box="[199,372,655,680]" pageId="9" pageNumber="584">Colour in life</emphasis>
. Body and appendages (
<figureCitation id="DDD27DFCFFDC780C67BB5488FCAFFD47" box="[678,812,655,680]" captionStart="FIGURE 5" captionStartId="8.[151,250,1959,1981]" captionTargetBox="[150,1435,217,1919]" captionTargetId="figure@8.[150,1436,217,1920]" captionTargetPageId="8" captionText="FIGURE 5. Caligoneus cavernicola n.gen., n. sp., general habitus and colour in life. A, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, RUMF-ZC-02733, non-ovigerous individual(cl 5.6mm)." httpUri="https://zenodo.org/record/1136504/files/figure.png" pageId="9" pageNumber="584">Fig. 5A, B</figureCitation>
) generally semi-translucent with scattered red dots, sometimes more intense on carapace and pleomeres to telson, forming diffuse transverse bands (
<figureCitation id="DDD27DFCFFDC780C603254B3FA0DFD22" box="[1327,1422,692,717]" captionStart="FIGURE 5" captionStartId="8.[151,250,1959,1981]" captionTargetBox="[150,1435,217,1919]" captionTargetId="figure@8.[150,1436,217,1920]" captionTargetPageId="8" captionText="FIGURE 5. Caligoneus cavernicola n.gen., n. sp., general habitus and colour in life. A, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, RUMF-ZC-02733, non-ovigerous individual(cl 5.6mm)." httpUri="https://zenodo.org/record/1136504/files/figure.png" pageId="9" pageNumber="584">Fig. 5B</figureCitation>
); yellowish ovary and hepatopancreas visible throughout integument; cornea darkly pigmented.
</paragraph>
</subSubSection>
<subSubSection id="0DF332F2FFDC780C65DA54FBFC32FBB4" pageId="9" pageNumber="584" type="distribution">
<paragraph id="45566179FFDC780C65DA54FBFE82FCD8" blockId="9.[151,1437,151,1368]" pageId="9" pageNumber="584">
<emphasis id="779DBD6BFFDC780C65DA54FBFEDAFCFA" bold="true" box="[199,345,764,789]" pageId="9" pageNumber="584">Distribution</emphasis>
. At present known only from a marine cave off Ie 
<collectingCountry id="3DFE21E9FFDC780C66B854FBFC6FFCFA" box="[933,1004,764,789]" name="Iceland" pageId="9" pageNumber="584">Island</collectingCountry>
, 
<collectingRegion id="872DAF9BFFDC780C66E454FBFBDDFCFA" box="[1017,1118,764,789]" country="Japan" name="Okinawa" pageId="9" pageNumber="584">Okinawa</collectingRegion>
Islands, 
<collectingCountry id="3DFE21E9FFDC780C61DB54FAFA8AFCFB" box="[1222,1289,765,788]" name="Japan" pageId="9" pageNumber="584">Japan</collectingCountry>
, at depths of 
<quantity id="8211CC9CFFDC780C658A5527FF7EFCD8" box="[151,253,800,824]" metricMagnitude="1" metricUnit="m" metricValue="1.45" metricValueMax="1.7" metricValueMin="1.2" pageId="9" pageNumber="584" unit="m" value="14.5" valueMax="17.0" valueMin="12.0">
12– 
<specimenCount id="53EFAAF0FFDC780C65DF5527FF7EFCD8" box="[194,253,800,824]" pageId="9" pageNumber="584" type="generic">17 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="45566179FFDC780C65DA5543FC32FBB4" blockId="9.[151,1437,151,1368]" pageId="9" pageNumber="584">
<emphasis id="779DBD6BFFDC780C65DA5543FEA9FCB2" bold="true" box="[199,298,836,861]" pageId="9" pageNumber="584">Habitat.</emphasis>
The 
<typeStatus id="9A52DFDBFFDC780C64785542FE16FCB2" box="[357,405,837,861]" pageId="9" pageNumber="584">type</typeStatus>
locality of 
<taxonomicName id="82E91AFAFFDC780C67045543FCA6FCB2" authority="Komai &amp; Fujita, 2018" authorityName="Komai &amp; Fujita" authorityYear="2018" box="[537,805,836,861]" class="Malacostraca" family="Alpheidae" genus="Caligoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="584" phylum="Arthropoda" rank="species" species="cavernicola" status="sp. nov.">
<emphasis id="779DBD6BFFDC780C67045543FCA6FCB2" box="[537,805,836,861]" italics="true" pageId="9" pageNumber="584">Caligoneus cavernicola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CAE0010FFDC780C66305543FCEEFCB2" box="[813,877,836,861]" pageId="9" pageNumber="584" rank="species">
<emphasis id="779DBD6BFFDC780C66305543FCEEFCB2" bold="true" box="[813,877,836,861]" pageId="9" pageNumber="584">n. sp.</emphasis>
</taxonomicNameLabel>
is a deep marine cave located on a reef slope off northeastern coast of Ie 
<collectingCountry id="3DFE21E9FFDC780C64B35560FE75FC6F" box="[430,502,871,896]" name="Iceland" pageId="9" pageNumber="584">Island</collectingCountry>
. The entrance to the cave lies at a depth of about 
<quantity id="8211CC9CFFDC780C6123556FFBFAFC90" box="[1086,1145,872,896]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="584" unit="m" value="17.0">17 m</quantity>
below the sea level. The interior of the cave is 
<quantity id="8211CC9CFFDC780C6493558BFE44FC4B" box="[398,455,908,933]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="584" unit="m" value="45.0">45 m</quantity>
long and bifurcated into “main tube” and “large hall” at about 
<quantity id="8211CC9CFFDC780C6193558BFB4BFC4B" box="[1166,1224,908,932]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="584" unit="m" value="30.0">30 m</quantity>
from the entrance. The “large hall” is about 
<quantity id="8211CC9CFFDC780C64AD55B4FE75FC25" box="[432,502,947,971]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="584" unit="m" value="200.0">200 m</quantity>
2 and its height to the roof is about 
<quantity id="8211CC9CFFDC780C666355B4FC34FC25" box="[894,951,947,971]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="584" unit="m" value="12.0">12 m</quantity>
(the roof is about 
<quantity id="8211CC9CFFDC780C619955B4FB2CFC25" box="[1156,1199,947,971]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="584" unit="m" value="8.0">8 m</quantity>
below the sea level). Shrimps were mostly found in small holes inside the “large hall”. The syntopic decapod fauna includes 
<taxonomicName id="82E91AFAFFDC780C603E55D1FF7DFBFC" authorityName="Komai, Yamada &amp; Yunokawa" authorityYear="2015" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="584" phylum="Arthropoda" rank="species" species="antricola">
<emphasis id="779DBD6BFFDC780C603E55D1FF7DFBFC" italics="true" pageId="9" pageNumber="584">Salmoneus antricola</emphasis>
</taxonomicName>
(Caridea: 
<taxonomicName id="82E91AFAFFDC780C646255FDFE74FBFC" authorityName="Komai &amp; Fujita" authorityYear="2018" box="[383,503,1018,1043]" class="Malacostraca" family="Alpheidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="584" phylum="Arthropoda" rank="family">Alpheidae</taxonomicName>
), 
<taxonomicName id="82E91AFAFFDC780C670D55FDFB97FBFC" authority="Komai &amp; Yamada, 2011" authorityName="Komai &amp; Yamada" authorityYear="2011" box="[528,1044,1018,1043]" class="Malacostraca" family="Bresiliidae" genus="Bresilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="584" phylum="Arthropoda" rank="species" species="rufioculus">
<emphasis id="779DBD6BFFDC780C670D55FDFD65FBFC" box="[528,742,1018,1043]" italics="true" pageId="9" pageNumber="584">Bresilia rufioculus</emphasis>
<bibRefCitation id="21781C88FFDC780C67EE55FDFB97FBFC" author="Komai" box="[755,1044,1018,1043]" pageId="9" pageNumber="584" refString="Komai, T. &amp; Yamada, Y. (2011) A new species of the caridean genus Bresilia Calman (Decapoda: Bresiliidae) discovered from a shallow-water submarine cave in Okinawa Islands, Japan. Bulletin of the National Museum of Nature and Science, Series A (Zoology), Supplement, 5, 71 - 82." type="journal article" year="2011">Komai &amp; Yamada, 2011</bibRefCitation>
</taxonomicName>
(Caridea: 
<taxonomicName id="82E91AFAFFDC780C618B55FDFA97FBFC" box="[1174,1300,1018,1043]" class="Malacostraca" family="Bresiliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="584" phylum="Arthropoda" rank="family">Bresiliidae</taxonomicName>
), 
<taxonomicName id="82E91AFAFFDC780C603155FCFE77FBD8" authority="Fujita &amp; Naruse, 2011" authorityName="Fujita &amp; Naruse" authorityYear="2011" class="Malacostraca" family="Portunidae" genus="Catoptrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="584" phylum="Arthropoda" rank="species" species="iejima">
<emphasis id="779DBD6BFFDC780C603155FCFF5EFBD8" italics="true" pageId="9" pageNumber="584">Catoptrus iejima</emphasis>
<bibRefCitation id="21781C88FFDC780C65F45219FE77FBD8" author="Fujita" box="[233,500,1054,1079]" pageId="9" pageNumber="584" refString="Fujita, Y. &amp; Naruse, T. (2011) Catoptrus iejima, a new species of cavernicolous swimming crab (Crustacea: Brachyura: Portunidae) from a submarine cave at Ie Island, Ryukyu Islands, Japan. Zootaxa, 2918, 29 - 38." type="journal article" year="2011">Fujita &amp; Naruse, 2011</bibRefCitation>
</taxonomicName>
(Brachyura: 
<taxonomicName id="82E91AFAFFDC780C678F5219FC93FBD8" box="[658,784,1054,1079]" class="Malacostraca" family="Portunidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="584" phylum="Arthropoda" rank="family">Portunidae</taxonomicName>
), 
<taxonomicName id="82E91AFAFFDC780C66355218FA89FBD8" authority="Ng &amp; Takeda, 2003" authorityName="Ng &amp; Takeda" authorityYear="2003" box="[808,1290,1054,1080]" class="Malacostraca" family="Portunidae" genus="Atoportunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="584" phylum="Arthropoda" rank="species" species="gustavi">
<emphasis id="779DBD6BFFDC780C66355218FB93FBD8" box="[808,1040,1055,1079]" italics="true" pageId="9" pageNumber="584">Atoportunus gustavi</emphasis>
Ng &amp; 
<bibRefCitation id="21781C88FFDC780C61735219FA89FBD8" author="Takeda" box="[1134,1290,1054,1079]" pageId="9" pageNumber="584" refString="Ng, P. K. L. &amp; Takeda, M. (2003) Atoportunus, a remarkable new genus of cryptic swimming crab with descriptions of two new species from the Indo-West Pacific. Micronesica, 35 / 36, 417 - 430." type="journal article" year="2003">Takeda, 2003</bibRefCitation>
</taxonomicName>
(Brachyura: 
<taxonomicName id="82E91AFAFFDC780C658A5245FE96FBB4" box="[151,277,1090,1115]" class="Malacostraca" family="Portunidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="584" phylum="Arthropoda" rank="family">Portunidae</taxonomicName>
), and 
<taxonomicName id="82E91AFAFFDC780C644A5245FC2DFBB4" authority="Naruse &amp; Fujita, 2015" authorityName="Naruse &amp; Fujita" authorityYear="2015" box="[343,942,1090,1115]" class="Malacostraca" family="Xanthidae" genus="Lipkemera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="584" phylum="Arthropoda" rank="species" species="iejima">
<emphasis id="779DBD6BFFDC780C644A5245FD98FBB5" box="[343,539,1090,1115]" italics="true" pageId="9" pageNumber="584">Lipkemera iejima</emphasis>
<bibRefCitation id="21781C88FFDC780C673E5244FC9EFBB4" author="Naruse" box="[547,797,1090,1115]" pageId="9" pageNumber="584" refString="Naruse, T. &amp; Fujita, Y. (2015) Lipkemera iejima, a new cavernicolous crab (Brachyura: Xanthidae) from a submarine cave at Ie Island, central Ryukyu Islands, Japan. Crustacean Research, 44, 21 - 27. https: // doi. org / 10.18353 / crustacea. 44.0 _ 21" type="journal article" year="2015">Naruse &amp; Fujita, 2015</bibRefCitation>
(Xanthidae)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="0DF332F2FFDC780C65DA5261FD37FB4C" pageId="9" pageNumber="584" type="etymology">
<paragraph id="45566179FFDC780C65DA5261FD37FB4C" blockId="9.[151,1437,151,1368]" pageId="9" pageNumber="584">
<emphasis id="779DBD6BFFDC780C65DA5261FEC5FB90" bold="true" box="[199,326,1126,1151]" pageId="9" pageNumber="584">Etymology</emphasis>
. Named after the cave habitat of the new species, the Latin cavern (= cave) and –cola (suffix, meaning dweller). Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="0DF332F2FFDC780C65DA52A9FAE6FAB8" pageId="9" pageNumber="584" type="discussion">
<paragraph id="45566179FFDC780C65DA52A9FAE6FAB8" blockId="9.[151,1437,151,1368]" pageId="9" pageNumber="584">
<emphasis id="779DBD6BFFDC780C65DA52A9FEB7FB28" bold="true" box="[199,308,1198,1223]" pageId="9" pageNumber="584">Remarks</emphasis>
. Among the six specimens examined, three are ovigerous. Like species of 
<taxonomicName id="82E91AFAFFDC780C61F752A9FAE7FB28" authorityName="Holthuis" authorityYear="1955" box="[1258,1380,1198,1223]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="584" phylum="Arthropoda" rank="genus">
<emphasis id="779DBD6BFFDC780C61F752A9FAE7FB28" box="[1258,1380,1198,1223]" italics="true" pageId="9" pageNumber="584">Salmoneus</emphasis>
</taxonomicName>
and 
<taxonomicName id="82E91AFAFFDC780C658A52D5FEC6FB04" authorityName="Anker" authorityYear="2010" box="[151,325,1234,1259]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="584" phylum="Arthropoda" rank="genus">
<emphasis id="779DBD6BFFDC780C658A52D5FEC6FB04" box="[151,325,1234,1259]" italics="true" pageId="9" pageNumber="584">Triacanthoneus</emphasis>
</taxonomicName>
, all individuals including the ovigerous ones have a well-developed appendix masculina just next to the appendix interna (
<figureCitation id="DDD27DFCFFDC780C64AB52F1FD8AFAE0" box="[438,521,1270,1295]" captionStart="FIGURE 2" captionStartId="5.[151,250,1824,1846]" captionTargetBox="[244,1334,231,1786]" captionTargetId="figure@5.[216,1376,193,1815]" captionTargetPageId="5" captionText="FIGURE 2. Caligoneus cavernicola n.gen., n. sp. A, C–J, holotype, CBM-ZC 14136, ovigerous individual (cl 5.7 mm); B, paratype, CBM-ZC 14137,ovigerous individual (cl 5.8 mm).A, anterior part of carapace and cephalic appendages,lateral view (antennular and antennal flagella partially omitted); B, anterior part of carapace, lateral view; C, anterior part of carapace and cephalic appendages, dorsal view; D, telson and right uropod, dorsal view (marginal setae on uropodal rami partially omitted); E, posterior end of telson, dorsal view; F, left antennal peduncle, ventral view (marginal setae on scaphocerite omitted); G, left pleopod 1, dorsal view; H, same, endopod, dorsomesial view; I, left pleopod 2, dorsal view; J, same, appendices interna and masculina, dorsal view." httpUri="https://zenodo.org/record/1136498/files/figure.png" pageId="9" pageNumber="584">Fig. 2J</figureCitation>
), on the pleopod 2, indicating a possibility of some form of hermaphroditism. Several morphological features of the new species clearly represent adaptations to cave environments, such as the reduced cornea located at the distolateral portion of the eyestalk and the very slender elongate pereopods 3–5.
</paragraph>
</subSubSection>
</treatment>
</document>