605 lines
101 KiB
XML
605 lines
101 KiB
XML
<document id="9CDF712402233C660C40EC4233896ABA" ID-CLB-Dataset="30781" ID-DOI="10.11646/zootaxa.4378.3.7" ID-GBIF-Dataset="9f591632-961c-4e3d-83ac-74cfb01de68b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1170041" ID-ZooBank="110496AC-8F8B-473F-A633-4EC0E4081838" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1518183044212" checkinUser="plazi" docAuthor="Komai, Tomoyuki" docDate="2018" docId="CC201F316307FFDFFF7EFC5DFAB90BB4" docLanguage="en" docName="zootaxa.4378.3.7.pdf" docOrigin="Zootaxa 4378 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thuylamea shiranui Komai 2018, n. sp." docType="treatment" docUuid="D6ACA18B-A941-4A23-AA18-123E67306F8A" docUuidSource="ZooBank" docVersion="8" lastPageNumber="395" masterDocId="301967496306FFD7FFE9FF93FFFD0C20" masterDocTitle="A new species of the alpheid shrimp genus Thuylamea Nguyên, 2001 (Crustacea: Decapoda: Caridea) from Yatsushiro Sea, Kyushu, Japan" masterLastPageNumber="396" masterPageNumber="387" pageNumber="388" updateTime="1698520882903" updateUser="ExternalLinkService">
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<mods:titleInfo id="28BE33154FC6D69253C76C467D000FC1">
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<mods:title id="102DF59084A8294CDD0AA1C57D1811F2">A new species of the alpheid shrimp genus Thuylamea Nguyên, 2001 (Crustacea: Decapoda: Caridea) from Yatsushiro Sea, Kyushu, Japan</mods:title>
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</mods:titleInfo>
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<mods:name id="E9281728C4AE7114F26E303523A48545" type="personal">
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<mods:role id="4530EB0DBD5DDA4D89E2D98499662844">
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<mods:roleTerm id="E2F495948638725DF698BD3CB40A3B64">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DC85E05F311C0425F59B79338D1CED2D">Komai, Tomoyuki</mods:namePart>
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<mods:typeOfResource id="95ACFE544AC62DD13614C30B77FF8E66">text</mods:typeOfResource>
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<mods:title id="E8BF471D62328991D8822F4B08BDEDE1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7DDBE87808CB783E7D0ACC53B092F348">
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<mods:date id="1DCEDC90FBAC4BD1B004591CCB4FD344">2018</mods:date>
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<mods:number id="335CBBBA3163725783F726C4029A5295">2018-02-09</mods:number>
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<mods:number id="5894CA27B6E565FECB31BB00C85A3F35">4378</mods:number>
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<mods:detail id="FDF4A79BFCB55804DD86B488A4DB7CE1" type="issue">
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<mods:number id="339B256B21CCB8D9211002663E43F882">3</mods:number>
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<mods:classification id="2420471263E58B3D0F05EF93B596FEDF">journal article</mods:classification>
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<mods:identifier id="BC39D24DE5C4C47CBA323A086B24627B" type="DOI">10.11646/zootaxa.4378.3.7</mods:identifier>
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<treatment id="CC201F316307FFDFFF7EFC5DFAB90BB4" ID-DOI="http://doi.org/10.5281/zenodo.5975385" ID-GBIF-Taxon="140569376" ID-Zenodo-Dep="5975385" LSID="urn:lsid:zoobank.org:act:D6ACA18B-A941-4A23-AA18-123E67306F8A" httpUri="http://treatment.plazi.org/id/CC201F316307FFDFFF7EFC5DFAB90BB4" lastPageId="8" lastPageNumber="395" pageId="1" pageNumber="388">
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<subSubSection id="0C93FDAC6307FFD6FF7EFC5DFF05080E" pageId="1" pageNumber="388" type="nomenclature">
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<paragraph id="4436AE276307FFD6FF7EFC5DFE2A0FC8" blockId="1.[151,635,974,1070]" box="[151,471,974,1000]" pageId="1" pageNumber="388">
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<heading id="1F7E194B6307FFD6FF7EFC5DFE2A0FC8" bold="true" box="[151,471,974,1000]" fontSize="11" level="1" pageId="1" pageNumber="388" reason="1">
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<emphasis id="76FD72356307FFD6FF7EFC5DFE2A0FC8" bold="true" box="[151,471,974,1000]" pageId="1" pageNumber="388">
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<taxonomicName id="8389D5A46307FFD6FF7EFC5DFE730FC8" ID-CoL="85Z5W" authority="Komai, 2018" authorityName="Komai" authorityYear="2018" box="[151,398,974,1000]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="388" phylum="Arthropoda" rank="species" species="shiranui" status="sp. nov.">
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<emphasis id="76FD72356307FFD6FF7EFC5DFE730FC8" bold="true" box="[151,398,974,1000]" italics="true" pageId="1" pageNumber="388">Thuylamea shiranui</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6DCECF4E6307FFD6FE7CFC5DFE2A0FC8" box="[405,471,974,1000]" pageId="1" pageNumber="388" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4436AE276307FFD6FF7EFC61FF05080E" blockId="1.[151,635,974,1070]" pageId="1" pageNumber="388">
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[New Japanese name: Shiranui-teppou-ebi]
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<figureCitation id="DCB2B2A26307FFD6FF7EFB86FF05080E" box="[151,248,1045,1070]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,250,1255,1277]" captionStartId-1="3.[151,250,1519,1541]" captionStartId-2="4.[151,250,1944,1966]" captionStartId-3="5.[151,250,1405,1427]" captionStartId-4="6.[151,250,1719,1741]" captionStartId-5="7.[151,250,1277,1299]" captionTargetBox-0="[175,1415,212,1210]" captionTargetBox-1="[172,1413,226,1477]" captionTargetBox-2="[175,1395,634,1896]" captionTargetBox-3="[175,1403,208,1357]" captionTargetBox-4="[178,1411,249,1669]" captionTargetBox-5="[151,1435,517,1255]" captionTargetId-0="figure@2.[151,1435,193,1234]" captionTargetId-1="figure@3.[151,1435,193,1498]" captionTargetId-2="figure@4.[151,1435,589,1923]" captionTargetId-3="figure@5.[151,1435,185,1394]" captionTargetId-4="figure@6.[151,1435,193,1698]" captionTargetId-5="figure@7.[151,1435,517,1255]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="FIGURE 1. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, carapace and cephalic appendages, lateral (right) view; B,pleon, telson and pleonal appendages,lateral (right) view." captionText-1="FIGURE 2. Thuylamea shiranui n. sp.. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146. A, carapace, dorsal view (left side damaged); B, anterior part of carapace and cephalic appendages, dorsal view (perpendicular for antennae); C, telson, dorsal view; D, right antennal peduncle, ventral view (setae omitted); E, distal par of outer flagellum of right antennule, dorsomesial view; F, epistome, ventral view; G, left uropod, dorsal view (perpendicular; setae partially omitted); eyes, ventro-frontal view (antennae broken off)." captionText-2="FIGURE3. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, left cheliped, lateral view; B, same, mesial view; C, same, chela, dorsal view; D, same, chela, ventral view (setae omitted); E, same, fingers, dorsal view; F, same, fingers, ventral view (setae omitted)." captionText-3="FIGURE 4. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, right cheliped, lateral view; B, same, mesial view; C, same, chela, dorsal view; D, same, chela, ventral view (setae omitted)." captionText-4="FIGURE 5. Thuylamea shiranui n. sp. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146; I, J, paratype, male (cl 4.3 mm), CBM-ZC 14149. A, left maxilliped 3, lateral view; B, same, basal part with arthrobranch, ventral view;C, left pereopod 3, lateral view; D, same, dactylus, lateral view; E, right pereopod 5, lateral view; F, left pleopod 1, ventral view (setae omitted); G, left pleopod 2, ventral view (setae omitted); H, left pereopod 2, lateral view; I, right pleopod 1, ventral view (setae on exopod omitted); J, left pleopod 2, dorsal view (marginal setae on rami omitted)." captionText-5="FIGURE 6. Thuylamea shiranui n. sp., paratype, female (cl 6.7 mm), CBM-ZC 14148, cephalothorax in dorsolateral view, showing left side of ovary extending to carapacedorsum." httpUri-0="https://zenodo.org/record/1170043/files/figure.png" httpUri-1="https://zenodo.org/record/1170045/files/figure.png" httpUri-2="https://zenodo.org/record/1170047/files/figure.png" httpUri-3="https://zenodo.org/record/1170049/files/figure.png" httpUri-4="https://zenodo.org/record/1170051/files/figure.png" httpUri-5="https://zenodo.org/record/1170053/files/figure.png" pageId="1" pageNumber="388">Figs 1–6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="0C93FDAC6307FFD6FF7EFBCEFD2809B6" pageId="1" pageNumber="388" type="materials_examined">
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<paragraph id="4436AE276307FFD6FF7EFBCEFB7008BA" blockId="1.[151,1436,1117,1970]" pageId="1" pageNumber="388">
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<materialsCitation id="F4E1A47A6307FFD6FF7EFBCEFB7008BA" ID-GBIF-Occurrence="1806525924" collectingDate="2017-05-30" elevation="29" latitude="32.35763" location="Gosho-ura Island" longLatPrecision="1" longitude="130.38176" pageId="1" pageNumber="388" specimenCount="1" typeStatus="holotype">
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<emphasis id="76FD72356307FFD6FF7EFBCEFE7D0856" bold="true" box="[151,384,1117,1142]" pageId="1" pageNumber="388">Material examined.</emphasis>
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<typeStatus id="9B3210856307FFD6FE6FFBCEFE0E0856" box="[390,499,1117,1142]" pageId="1" pageNumber="388" type="holotype">Holotype</typeStatus>
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: CBM-ZC 14145, female (cl
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<quantity id="837103C26307FFD6FCD0FBCEFC6F0855" box="[825,914,1117,1142]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="1" pageNumber="388" unit="mm" value="5.1">5.1 mm</quantity>
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; pereopods 2 and 4 missing), NE of
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<location id="4156F8FC6307FFD6FACFFBCEFF2308BA" LSID="urn:lsid:plazi:treatment:CC201F316307FFDFFF7EFC5DFAB90BB4:4156F8FC6307FFD6FACFFBCEFF2308BA" latitude="32.35763" longLatPrecision="1" longitude="130.38176" name="Gosho-ura Island" pageId="1" pageNumber="388">Gosho-ura Island</location>
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,
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<geoCoordinate id="21BDC8E06307FFD6FF00FB12FE8408BA" box="[233,377,1153,1178]" direction="north" orientation="latitude" pageId="1" pageNumber="388" precision="1" value="32.35763">32°21.458'N</geoCoordinate>
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,
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||
<geoCoordinate id="21BDC8E06307FFD6FE6CFB12FDDC08BA" box="[389,545,1153,1178]" direction="east" orientation="longitude" pageId="1" pageNumber="388" precision="1" value="130.38176">130°22.906′E</geoCoordinate>
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,
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<elevation id="CFA449146307FFD6FDC5FB11FD9A08B9" box="[556,615,1154,1178]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="388" unit="m" value="29.0">
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<quantity id="837103C26307FFD6FDC5FB11FD9A08B9" box="[556,615,1154,1178]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="388" unit="m" value="29.0">29 m</quantity>
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</elevation>
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, silt, stn 17, Smith-McIntyre grab,
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<collectingDate id="2073710F6307FFD6FC19FB11FB7408BA" box="[1008,1161,1154,1178]" pageId="1" pageNumber="388" value="2017-05-30">
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<date id="303788E76307FFD6FC19FB11FB7408BA" box="[1008,1161,1154,1178]" pageId="1" pageNumber="388" value="2017-05-30">30 May 2017</date>
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</collectingDate>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="4436AE276307FFD6FF2EFB35FD2809B6" blockId="1.[151,1436,1117,1970]" pageId="1" pageNumber="388">
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Paratype: CBM-ZC 14146, 1 specimen, sex not determinable (cl
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<quantity id="837103C26307FFD6FC48FB35FC06089D" box="[929,1019,1190,1214]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="388" unit="mm" value="3.7">3.7 mm</quantity>
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; heavily damaged, antennae missing, only left maxilliped 3, left pereopod 1, 2 and 5 preserved),
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<materialsCitation id="F4E1A47A6307FFD6FCCFFB59FC550926" ID-GBIF-Occurrence="1806525930" collectingDate="2017-05-30" elevation="29" latitude="32.35763" location="Gosho-ura Island" longLatPrecision="1" longitude="130.38176" pageId="1" pageNumber="388" specimenCount="1" specimenCount-female="1">
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same data as holotype; CBM-ZC 14147,
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<specimenCount id="528F65AE6307FFD6FB07FB59FAB208C2" box="[1262,1359,1225,1250]" pageId="1" pageNumber="388" type="female">1 female</specimenCount>
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(cl
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<quantity id="837103C26307FFD6FA90FB59FF3B0925" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="388" unit="mm" value="4.7">4.7 mm</quantity>
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; antennae and maxillipeds 3 missing, only pereopods 1 present)
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</materialsCitation>
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,
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<materialsCitation id="F4E1A47A6307FFD6FC5CFB7DFADA096E" ID-GBIF-Occurrence="1806525935" collectingDate="2017-05-30" elevation="29" latitude="32.38895" location="Gosho-ura Island" longLatPrecision="1" longitude="130.22745" pageId="1" pageNumber="388" specimenCount="1" specimenCount-female="1">
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same data as holotype; CBM-ZC 14 148,
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<specimenCount id="528F65AE6307FFD6FA67FB7DFF1E090A" pageId="1" pageNumber="388" type="female">1 female</specimenCount>
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(cl
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<quantity id="837103C26307FFD6FEF2FA82FE870909" box="[283,378,1297,1322]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="1" pageNumber="388" unit="mm" value="6.7">6.7 mm</quantity>
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; maxilliped 3 damaged; all pereopods except for right pereopod 1 missing), off Hondo, Amakusa,
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<geoCoordinate id="21BDC8E06307FFD6FEE7FAA6FE62096E" box="[270,415,1333,1358]" direction="north" orientation="latitude" pageId="1" pageNumber="388" precision="1" value="32.38895">32°23.337′N</geoCoordinate>
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,
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||
<geoCoordinate id="21BDC8E06307FFD6FE43FAA6FDBB096E" box="[426,582,1333,1358]" direction="east" orientation="longitude" pageId="1" pageNumber="388" precision="1" value="130.22745">130°13.647′E</geoCoordinate>
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,
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<elevation id="CFA449146307FFD6FDB8FAA5FD5D096D" box="[593,672,1334,1358]" metricMagnitude="1" metricUnit="m" metricValue="2.87" pageId="1" pageNumber="388" unit="m" value="28.7">
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||
<quantity id="837103C26307FFD6FDB8FAA5FD5D096D" box="[593,672,1334,1358]" metricMagnitude="1" metricUnit="m" metricValue="2.87" pageId="1" pageNumber="388" unit="m" value="28.7">28.7 m</quantity>
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</elevation>
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, stn 14, Smith-McIntyre grab, silt and sand,
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<collectingDate id="2073710F6307FFD6FB67FAA5FADA096E" box="[1166,1319,1334,1358]" pageId="1" pageNumber="388" value="2017-05-30">
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||
<date id="303788E76307FFD6FB67FAA5FADA096E" box="[1166,1319,1334,1358]" pageId="1" pageNumber="388" value="2017-05-30">30 May 2017</date>
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</collectingDate>
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</materialsCitation>
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; CBM-ZC 14149, 1 male (cl
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<quantity id="837103C26307FFD6FE8AFAC9FE400951" box="[355,445,1370,1394]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="1" pageNumber="388" unit="mm" value="4.3">4.3 mm</quantity>
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; all pereopods missing), E of Himedo, Kami-Amakusa, 32°25.704’N, 130°27.712’E,
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<quantity id="837103C26307FFD6FA69FAC9FF5209B5" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="388" unit="m" value="27.0">27 m</quantity>
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, stn 13, Smith-McIntyre grab, silt,
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<date id="303788E76307FFD6FDD1FAEDFD2C09B6" box="[568,721,1406,1430]" pageId="1" pageNumber="388" value="2017-05-31">31 May 2017</date>
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||
.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="0C93FDAC6307FFDFFF2EFA32FE170D3D" lastPageId="8" lastPageNumber="395" pageId="1" pageNumber="388" type="description">
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||
<paragraph id="4436AE276307FFD6FF2EFA32FAE0099A" blockId="1.[151,1436,1117,1970]" box="[199,1309,1441,1466]" pageId="1" pageNumber="388">
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||
<emphasis id="76FD72356307FFD6FF2EFA32FDC6099A" bold="true" box="[199,571,1441,1466]" pageId="1" pageNumber="388">
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Description of female
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<typeStatus id="9B3210856307FFD6FE24FA32FDCB099A" box="[461,566,1441,1466]" pageId="1" pageNumber="388" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Body (
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<figureCitation id="DCB2B2A26307FFD6FD64FA32FCF8099A" box="[653,773,1441,1466]" captionStart="FIGURE 1" captionStartId="2.[151,250,1255,1277]" captionTargetBox="[175,1415,212,1210]" captionTargetId="figure@2.[151,1435,193,1234]" captionTargetPageId="2" captionText="FIGURE 1. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, carapace and cephalic appendages, lateral (right) view; B,pleon, telson and pleonal appendages,lateral (right) view." httpUri="https://zenodo.org/record/1170043/files/figure.png" pageId="1" pageNumber="388">Fig. 1A, B</figureCitation>
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) elongate, subcylindrical; integument glabrous.
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</paragraph>
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<paragraph id="4436AE276307FFD6FF2EFA55FF2E0AB2" blockId="1.[151,1436,1117,1970]" pageId="1" pageNumber="388">
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Rostrum (
|
||
<figureCitation id="DCB2B2A26307FFD6FED1FA56FE4E09FE" box="[312,435,1477,1502]" captionStart="FIGURE 2" captionStartId="3.[151,250,1519,1541]" captionTargetBox="[172,1413,226,1477]" captionTargetId="figure@3.[151,1435,193,1498]" captionTargetPageId="3" captionText="FIGURE 2. Thuylamea shiranui n. sp.. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146. A, carapace, dorsal view (left side damaged); B, anterior part of carapace and cephalic appendages, dorsal view (perpendicular for antennae); C, telson, dorsal view; D, right antennal peduncle, ventral view (setae omitted); E, distal par of outer flagellum of right antennule, dorsomesial view; F, epistome, ventral view; G, left uropod, dorsal view (perpendicular; setae partially omitted); eyes, ventro-frontal view (antennae broken off)." httpUri="https://zenodo.org/record/1170045/files/figure.png" pageId="1" pageNumber="388">Fig. 2A, B</figureCitation>
|
||
) very short, broadly triangular, flattened dorsoventrally, directed slightly downward; tip sharply pointed; lateral margin merging into evenly concave orbital margin; ventral surface unarmed. Carapace (
|
||
<figureCitation id="DCB2B2A26307FFD6FF76F99EFF000A06" box="[159,253,1549,1574]" captionStart="FIGURE 1" captionStartId="2.[151,250,1255,1277]" captionTargetBox="[175,1415,212,1210]" captionTargetId="figure@2.[151,1435,193,1234]" captionTargetPageId="2" captionText="FIGURE 1. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, carapace and cephalic appendages, lateral (right) view; B,pleon, telson and pleonal appendages,lateral (right) view." httpUri="https://zenodo.org/record/1170043/files/figure.png" pageId="1" pageNumber="388">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="DCB2B2A26307FFD6FEE3F99DFED30A06" box="[266,302,1550,1574]" captionStart="FIGURE 2" captionStartId="3.[151,250,1519,1541]" captionTargetBox="[172,1413,226,1477]" captionTargetId="figure@3.[151,1435,193,1498]" captionTargetPageId="3" captionText="FIGURE 2. Thuylamea shiranui n. sp.. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146. A, carapace, dorsal view (left side damaged); B, anterior part of carapace and cephalic appendages, dorsal view (perpendicular for antennae); C, telson, dorsal view; D, right antennal peduncle, ventral view (setae omitted); E, distal par of outer flagellum of right antennule, dorsomesial view; F, epistome, ventral view; G, left uropod, dorsal view (perpendicular; setae partially omitted); eyes, ventro-frontal view (antennae broken off)." httpUri="https://zenodo.org/record/1170045/files/figure.png" pageId="1" pageNumber="388">2A</figureCitation>
|
||
) with broadly rounded anterolateral margin, without armature; length about twice of height; dorsum generally rounded in anterior half, dorsal profile in lateral view nearly straight, having shallow median sulcus between gastric and cardiac regions; posterodorsal margin slightly elevated in broad transverse ridge; cardiac notch deep.
|
||
</paragraph>
|
||
<paragraph id="4436AE276307FFD6FF2EF90EFE940B92" blockId="1.[151,1436,1117,1970]" pageId="1" pageNumber="388">
|
||
Pleon (
|
||
<figureCitation id="DCB2B2A26307FFD6FEF4F90EFE840A96" box="[285,377,1693,1718]" captionStart="FIGURE 1" captionStartId="2.[151,250,1255,1277]" captionTargetBox="[175,1415,212,1210]" captionTargetId="figure@2.[151,1435,193,1234]" captionTargetPageId="2" captionText="FIGURE 1. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, carapace and cephalic appendages, lateral (right) view; B,pleon, telson and pleonal appendages,lateral (right) view." httpUri="https://zenodo.org/record/1170043/files/figure.png" pageId="1" pageNumber="388">Fig. 1B</figureCitation>
|
||
) with anterior 3 pleura rounded ventrally or posteroventrally; pleura 4 and 5 produced posteriorly but rounded, unarmed. Pleomere 6 1.3 times as long as pleomere 5, 1.3 times as long as high, without articulated plate at posteroventral angle; dorsal surface with short transverse groove anteriorly; posteroventral angle blunt; preanal plate unarmed; posterolateral process rounded. Telson (
|
||
<figureCitation id="DCB2B2A26307FFD6FC03F89AFBBD0B02" box="[1002,1088,1801,1826]" captionStart="FIGURE 2" captionStartId="3.[151,250,1519,1541]" captionTargetBox="[172,1413,226,1477]" captionTargetId="figure@3.[151,1435,193,1498]" captionTargetPageId="3" captionText="FIGURE 2. Thuylamea shiranui n. sp.. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146. A, carapace, dorsal view (left side damaged); B, anterior part of carapace and cephalic appendages, dorsal view (perpendicular for antennae); C, telson, dorsal view; D, right antennal peduncle, ventral view (setae omitted); E, distal par of outer flagellum of right antennule, dorsomesial view; F, epistome, ventral view; G, left uropod, dorsal view (perpendicular; setae partially omitted); eyes, ventro-frontal view (antennae broken off)." httpUri="https://zenodo.org/record/1170045/files/figure.png" pageId="1" pageNumber="388">Fig. 2C</figureCitation>
|
||
) 1.1 times as long as pleomere 6 and 2.3 times as long as wide, slightly narrowing posterior to mid-length, with 2 pairs of small dorsolateral spiniform setae (anterior pair located at about mid-length of telson, posterior pair at 0.6 length of telson); posterior margin produced, rounded, with row of long setae flanked by 2 lateral pairs of minute spiniform setae; dorsal surface nearly flat.
|
||
</paragraph>
|
||
<caption id="10F6FEAF6304FFD5FF7EFB74FC34093C" httpUri="https://zenodo.org/record/1170043/files/figure.png" pageId="2" pageNumber="389" startId="2.[151,250,1255,1277]" targetBox="[175,1415,212,1210]" targetPageId="2">
|
||
<paragraph id="4436AE276304FFD5FF7EFB74FC34093C" blockId="2.[151,1436,1255,1308]" pageId="2" pageNumber="389">
|
||
<emphasis id="76FD72356304FFD5FF7EFB74FEEE08DD" bold="true" box="[151,275,1255,1277]" pageId="2" pageNumber="389">FIGURE 1.</emphasis>
|
||
<taxonomicName id="8389D5A46304FFD5FEF0FB74FE1908DD" authority="Komai, 2018" authorityName="Komai" authorityYear="2018" box="[281,484,1255,1277]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="shiranui" status="sp. nov.">
|
||
<emphasis id="76FD72356304FFD5FEF0FB74FE1908DD" box="[281,484,1255,1277]" italics="true" pageId="2" pageNumber="389">Thuylamea shiranui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6DCECF4E6304FFD5FE05FB74FDD808DD" box="[492,549,1255,1277]" pageId="2" pageNumber="389" rank="species">
|
||
<emphasis id="76FD72356304FFD5FE05FB74FDD808DD" bold="true" box="[492,549,1255,1277]" pageId="2" pageNumber="389">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype, female (cl 5.1 mm), CBM-ZC 14145. A, carapace and cephalic appendages, lateral (right) view; B, pleon, telson and pleonal appendages, lateral (right) view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4436AE276304FFD5FF2EFAD0FA740984" blockId="2.[151,1437,1347,2020]" pageId="2" pageNumber="389">
|
||
Eyestalks (
|
||
<figureCitation id="DCB2B2A26304FFD5FEAFFAD0FE5A097C" box="[326,423,1347,1372]" captionStart="FIGURE 1" captionStartId="2.[151,250,1255,1277]" captionTargetBox="[175,1415,212,1210]" captionTargetId="figure@2.[151,1435,193,1234]" captionTargetPageId="2" captionText="FIGURE 1. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, carapace and cephalic appendages, lateral (right) view; B,pleon, telson and pleonal appendages,lateral (right) view." httpUri="https://zenodo.org/record/1170043/files/figure.png" pageId="2" pageNumber="389">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="DCB2B2A26304FFD5FE51FAD7FE21097C" box="[440,476,1348,1372]" captionStart="FIGURE 2" captionStartId="3.[151,250,1519,1541]" captionTargetBox="[172,1413,226,1477]" captionTargetId="figure@3.[151,1435,193,1498]" captionTargetPageId="3" captionText="FIGURE 2. Thuylamea shiranui n. sp.. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146. A, carapace, dorsal view (left side damaged); B, anterior part of carapace and cephalic appendages, dorsal view (perpendicular for antennae); C, telson, dorsal view; D, right antennal peduncle, ventral view (setae omitted); E, distal par of outer flagellum of right antennule, dorsomesial view; F, epistome, ventral view; G, left uropod, dorsal view (perpendicular; setae partially omitted); eyes, ventro-frontal view (antennae broken off)." httpUri="https://zenodo.org/record/1170045/files/figure.png" pageId="2" pageNumber="389">2B</figureCitation>
|
||
); supplemented by
|
||
<figureCitation id="DCB2B2A26304FFD5FD2CFAD0FCE2097C" box="[709,799,1347,1372]" captionStart="FIGURE 2" captionStartId="3.[151,250,1519,1541]" captionTargetBox="[172,1413,226,1477]" captionTargetId="figure@3.[151,1435,193,1498]" captionTargetPageId="3" captionText="FIGURE 2. Thuylamea shiranui n. sp.. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146. A, carapace, dorsal view (left side damaged); B, anterior part of carapace and cephalic appendages, dorsal view (perpendicular for antennae); C, telson, dorsal view; D, right antennal peduncle, ventral view (setae omitted); E, distal par of outer flagellum of right antennule, dorsomesial view; F, epistome, ventral view; G, left uropod, dorsal view (perpendicular; setae partially omitted); eyes, ventro-frontal view (antennae broken off)." httpUri="https://zenodo.org/record/1170045/files/figure.png" pageId="2" pageNumber="389">Fig. 2H</figureCitation>
|
||
of
|
||
<typeStatus id="9B3210856304FFD5FCA7FAD7FC4D097C" box="[846,944,1348,1372]" pageId="2" pageNumber="389" type="paratype">paratype</typeStatus>
|
||
) short, stout, subtriangular with produced anteromesial angles, contiguous, covered with frontal margin of carapace in dorsal view, cornea partially visible in lateral view, small, rounded, darkly pigmented, with scattered whitish spots possibly representing reduced facets.
|
||
</paragraph>
|
||
<paragraph id="4436AE276304FFD5FF2EFA3CFE1E0AC8" blockId="2.[151,1437,1347,2020]" pageId="2" pageNumber="389">
|
||
Antennular peduncle (
|
||
<figureCitation id="DCB2B2A26304FFD5FE38FA3CFDCB09E8" box="[465,566,1455,1480]" captionStart="FIGURE 1" captionStartId="2.[151,250,1255,1277]" captionTargetBox="[175,1415,212,1210]" captionTargetId="figure@2.[151,1435,193,1234]" captionTargetPageId="2" captionText="FIGURE 1. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, carapace and cephalic appendages, lateral (right) view; B,pleon, telson and pleonal appendages,lateral (right) view." httpUri="https://zenodo.org/record/1170043/files/figure.png" pageId="2" pageNumber="389">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="DCB2B2A26304FFD5FDA2FA23FD9009E8" box="[587,621,1456,1480]" captionStart="FIGURE 2" captionStartId="3.[151,250,1519,1541]" captionTargetBox="[172,1413,226,1477]" captionTargetId="figure@3.[151,1435,193,1498]" captionTargetPageId="3" captionText="FIGURE 2. Thuylamea shiranui n. sp.. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146. A, carapace, dorsal view (left side damaged); B, anterior part of carapace and cephalic appendages, dorsal view (perpendicular for antennae); C, telson, dorsal view; D, right antennal peduncle, ventral view (setae omitted); E, distal par of outer flagellum of right antennule, dorsomesial view; F, epistome, ventral view; G, left uropod, dorsal view (perpendicular; setae partially omitted); eyes, ventro-frontal view (antennae broken off)." httpUri="https://zenodo.org/record/1170045/files/figure.png" pageId="2" pageNumber="389">2B</figureCitation>
|
||
) about 0.4 times as long as carapace, exceeding distal margin of scaphocerite by half-length of article 2, but not reaching end of antennal carpocerite. Article 1 about twice as long as wide, gently inflated proximally with well-developed statocyst lobe; ventromesial ridge with strong subdistal spine directed forward; stylocerite closely adjoining lateral side of article 1, gradually tapering to spine falling slightly short of distal margin of article 1. Article 2 somewhat depressed, subequal in length to article 1, 3.0 times as long as wide. Article 3 short, also somewhat depressed, less than half length of article 2. Outer flagellum subequal in length to peduncle, bifurcate at eighth or ninth article; shorter ramus consisting of 2 or 3 articles, bearing some aesthetascs; longer ramus slender, multiarticulate; mesial flagellum much slenderer and slightly shorter than lateral flagellum.
|
||
</paragraph>
|
||
<paragraph id="4436AE276304FFD5FF2EF967FD010BBC" blockId="2.[151,1437,1347,2020]" pageId="2" pageNumber="389">
|
||
Antenna (
|
||
<figureCitation id="DCB2B2A26304FFD5FEDCF960FE6F0B2C" box="[309,402,1779,1804]" captionStart="FIGURE 1" captionStartId="2.[151,250,1255,1277]" captionTargetBox="[175,1415,212,1210]" captionTargetId="figure@2.[151,1435,193,1234]" captionTargetPageId="2" captionText="FIGURE 1. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, carapace and cephalic appendages, lateral (right) view; B,pleon, telson and pleonal appendages,lateral (right) view." httpUri="https://zenodo.org/record/1170043/files/figure.png" pageId="2" pageNumber="389">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="DCB2B2A26304FFD5FE49F967FE180B2C" box="[416,485,1780,1804]" captionStart="FIGURE 2" captionStartId="3.[151,250,1519,1541]" captionTargetBox="[172,1413,226,1477]" captionTargetId="figure@3.[151,1435,193,1498]" captionTargetPageId="3" captionText="FIGURE 2. Thuylamea shiranui n. sp.. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146. A, carapace, dorsal view (left side damaged); B, anterior part of carapace and cephalic appendages, dorsal view (perpendicular for antennae); C, telson, dorsal view; D, right antennal peduncle, ventral view (setae omitted); E, distal par of outer flagellum of right antennule, dorsomesial view; F, epistome, ventral view; G, left uropod, dorsal view (perpendicular; setae partially omitted); eyes, ventro-frontal view (antennae broken off)." httpUri="https://zenodo.org/record/1170045/files/figure.png" pageId="2" pageNumber="389">2B, D</figureCitation>
|
||
) with basicerite stout, with small, acute ventrolateral distal tooth. Distomesial angle of article 3 strongly produced distally. Carpocerite elongate (about 5 times as long as wide), subcylindrical, exceeding far beyond distal margin of scaphocerite by 0.3 of its length. Scaphocerite short, about 0.3 times as long as carapace, 2.5 times as long as wide, elongate subovate; lateral margin nearly straight, terminating in small spine falling slightly short of broadly rounded distal lamella.
|
||
</paragraph>
|
||
<paragraph id="4436AE276304FFD5FF2EF834FD2F0BE0" blockId="2.[151,1437,1347,2020]" box="[199,722,1959,1984]" pageId="2" pageNumber="389">
|
||
Epistome (
|
||
<figureCitation id="DCB2B2A26304FFD5FEA9F834FE6E0BE0" box="[320,403,1959,1984]" captionStart="FIGURE 2" captionStartId="3.[151,250,1519,1541]" captionTargetBox="[172,1413,226,1477]" captionTargetId="figure@3.[151,1435,193,1498]" captionTargetPageId="3" captionText="FIGURE 2. Thuylamea shiranui n. sp.. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146. A, carapace, dorsal view (left side damaged); B, anterior part of carapace and cephalic appendages, dorsal view (perpendicular for antennae); C, telson, dorsal view; D, right antennal peduncle, ventral view (setae omitted); E, distal par of outer flagellum of right antennule, dorsomesial view; F, epistome, ventral view; G, left uropod, dorsal view (perpendicular; setae partially omitted); eyes, ventro-frontal view (antennae broken off)." httpUri="https://zenodo.org/record/1170045/files/figure.png" pageId="2" pageNumber="389">Fig. 2F</figureCitation>
|
||
) with laminar median tooth.
|
||
</paragraph>
|
||
<paragraph id="4436AE276304FFD5FF2EF858FA6F0BC4" blockId="2.[151,1437,1347,2020]" box="[199,1426,1995,2020]" pageId="2" pageNumber="389">
|
||
Mouthparts not dissected, similar to those of
|
||
<taxonomicName id="8389D5A46304FFD5FD29F85EFCC50BC4" authorityName="Nguyen" authorityYear="2001" box="[704,824,1995,2020]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="389" phylum="Arthropoda" rank="species" species="camelus">
|
||
<emphasis id="76FD72356304FFD5FD29F85EFCC50BC4" box="[704,824,1995,2020]" italics="true" pageId="2" pageNumber="389">T. camelus</emphasis>
|
||
</taxonomicName>
|
||
in external observation (cf.
|
||
<bibRefCitation id="2018D3D66304FFD5FB9AF85FFAF20BC4" box="[1139,1295,1996,2020]" pageId="2" pageNumber="389" refId="ref4172" refString="Nguyen, V. X. (2001) A new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from South Vietnam. Zoologische Mededelingen, Leiden, 75, 217 - 228." type="book chapter">Nguyen 2001</bibRefCitation>
|
||
: figs 7–11).
|
||
</paragraph>
|
||
<caption id="10F6FEAF6305FFD4FF7EFA7CFBB10A5F" httpUri="https://zenodo.org/record/1170045/files/figure.png" pageId="3" pageNumber="390" startId="3.[151,250,1519,1541]" targetBox="[172,1413,226,1477]" targetPageId="3">
|
||
<paragraph id="4436AE276305FFD4FF7EFA7CFBB10A5F" blockId="3.[151,1436,1519,1663]" pageId="3" pageNumber="390">
|
||
<emphasis id="76FD72356305FFD4FF7EFA7CFEEE0A25" bold="true" box="[151,275,1519,1541]" pageId="3" pageNumber="390">FIGURE 2.</emphasis>
|
||
<taxonomicName id="8389D5A46305FFD4FEF3FA7CFE180A25" authority="Komai, 2018" authorityName="Komai" authorityYear="2018" box="[282,485,1519,1541]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="390" phylum="Arthropoda" rank="species" species="shiranui" status="sp. nov.">
|
||
<emphasis id="76FD72356305FFD4FEF3FA7CFE180A25" box="[282,485,1519,1541]" italics="true" pageId="3" pageNumber="390">Thuylamea shiranui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6DCECF4E6305FFD4FE05FA7CFDD80A25" box="[492,549,1519,1541]" pageId="3" pageNumber="390" rank="species">
|
||
<emphasis id="76FD72356305FFD4FE05FA7CFDD80A25" bold="true" box="[492,549,1519,1541]" pageId="3" pageNumber="390">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146. A, carapace, dorsal view (left side damaged); B, anterior part of carapace and cephalic appendages, dorsal view (perpendicular for antennae); C, telson, dorsal view; D, right antennal peduncle, ventral view (setae omitted); E, distal par of outer flagellum of right antennule, dorsomesial view; F, epistome, ventral view; G, left uropod, dorsal view (perpendicular; setae partially omitted); eyes, ventro-frontal view (antennae broken off).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4436AE276305FFD4FF2EF93FFE620BBD" blockId="3.[151,1436,1708,2021]" pageId="3" pageNumber="390">
|
||
Maxilliped 3 endopod (
|
||
<figureCitation id="DCB2B2A26305FFD4FE33F93FFDC10AE5" box="[474,572,1708,1733]" captionStart="FIGURE 1" captionStartId="2.[151,250,1255,1277]" captionTargetBox="[175,1415,212,1210]" captionTargetId="figure@2.[151,1435,193,1234]" captionTargetPageId="2" captionText="FIGURE 1. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, carapace and cephalic appendages, lateral (right) view; B,pleon, telson and pleonal appendages,lateral (right) view." httpUri="https://zenodo.org/record/1170043/files/figure.png" pageId="3" pageNumber="390">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="DCB2B2A26305FFD4FDA4F93FFD8D0AE5" box="[589,624,1708,1733]" captionStart="FIGURE 5" captionStartId="6.[151,250,1719,1741]" captionTargetBox="[178,1411,249,1669]" captionTargetId="figure@6.[151,1435,193,1698]" captionTargetPageId="6" captionText="FIGURE 5. Thuylamea shiranui n. sp. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146; I, J, paratype, male (cl 4.3 mm), CBM-ZC 14149. A, left maxilliped 3, lateral view; B, same, basal part with arthrobranch, ventral view;C, left pereopod 3, lateral view; D, same, dactylus, lateral view; E, right pereopod 5, lateral view; F, left pleopod 1, ventral view (setae omitted); G, left pleopod 2, ventral view (setae omitted); H, left pereopod 2, lateral view; I, right pleopod 1, ventral view (setae on exopod omitted); J, left pleopod 2, dorsal view (marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1170051/files/figure.png" pageId="3" pageNumber="390">5A</figureCitation>
|
||
) moderately slender, reaching as far as distolateral spine of antennal scaphocerite. Coxa with lateral plate triangular with forwardly directed apex, bearing strap-like epipod arising from ventral surface (
|
||
<figureCitation id="DCB2B2A26305FFD4FEB2F967FE460B2D" box="[347,443,1780,1805]" captionStart="FIGURE 5" captionStartId="6.[151,250,1719,1741]" captionTargetBox="[178,1411,249,1669]" captionTargetId="figure@6.[151,1435,193,1698]" captionTargetPageId="6" captionText="FIGURE 5. Thuylamea shiranui n. sp. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146; I, J, paratype, male (cl 4.3 mm), CBM-ZC 14149. A, left maxilliped 3, lateral view; B, same, basal part with arthrobranch, ventral view;C, left pereopod 3, lateral view; D, same, dactylus, lateral view; E, right pereopod 5, lateral view; F, left pleopod 1, ventral view (setae omitted); G, left pleopod 2, ventral view (setae omitted); H, left pereopod 2, lateral view; I, right pleopod 1, ventral view (setae on exopod omitted); J, left pleopod 2, dorsal view (marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1170051/files/figure.png" pageId="3" pageNumber="390">Fig. 5B</figureCitation>
|
||
). Antepenultimate article sinuously curved in dorsal view, proximal half flattened dorsoventrally. Penultimate article (= carpus) subcylindrical, slightly widened distally. Ultimate article 1.4 times as long as penultimate article, gradually tapering distally, terminating in blunt tip, bearing some short terminal setae; mesial face with several transverse tracts of stiff serrulate setae. Exopod flagellum-like, overreaching distal end of antepenultimate article.
|
||
</paragraph>
|
||
<paragraph id="4436AE276305FFD3FF2EF83BFC310E38" blockId="3.[151,1436,1708,2021]" lastBlockId="4.[151,1437,151,536]" lastPageId="4" lastPageNumber="391" pageId="3" pageNumber="390">
|
||
Pereopods 1 (chelipeds) (
|
||
<figureCitation id="DCB2B2A26305FFD4FE0AF83BFDD60BE1" box="[483,555,1960,1985]" captionStart="FIGURE3" captionStartId="4.[151,250,1944,1966]" captionTargetBox="[175,1395,634,1896]" captionTargetId="figure@4.[151,1435,589,1923]" captionTargetPageId="4" captionText="FIGURE3. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, left cheliped, lateral view; B, same, mesial view; C, same, chela, dorsal view; D, same, chela, ventral view (setae omitted); E, same, fingers, dorsal view; F, same, fingers, ventral view (setae omitted)." httpUri="https://zenodo.org/record/1170047/files/figure.png" pageId="3" pageNumber="390">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="DCB2B2A26305FFD4FDD0F83AFDB50BE1" box="[569,584,1961,1985]" captionStart="FIGURE 4" captionStartId="5.[151,250,1405,1427]" captionTargetBox="[175,1403,208,1357]" captionTargetId="figure@5.[151,1435,185,1394]" captionTargetPageId="5" captionText="FIGURE 4. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, right cheliped, lateral view; B, same, mesial view; C, same, chela, dorsal view; D, same, chela, ventral view (setae omitted)." httpUri="https://zenodo.org/record/1170049/files/figure.png" pageId="3" pageNumber="390">4</figureCitation>
|
||
) enlarged, subequal in length, symmetrical between right and left, extended forward, with dactylus articulated laterally to palm. Ischium short, strongly compressed, unarmed. Merus more strongly compressed proximally; dorsal margin sinuous; ventral margin nearly straight. Carpus short, cup-shaped, about half-length of merus. Chela subequal in length to carapace. Palm subcylindrical, slightly compressed dorsoventrally, 4.1 times (left) or 4.3 times (right) as long as wide, generally smooth on surfaces; ventral surface with shallow, saddle-like transverse depression proximal to base of fingers; dactylar articulation armed with triangular tooth on either side (left chela;
|
||
<figureCitation id="DCB2B2A26302FFD3FD84FEB4FD220D60" box="[621,735,295,320]" captionStart="FIGURE3" captionStartId="4.[151,250,1944,1966]" captionTargetBox="[175,1395,634,1896]" captionTargetId="figure@4.[151,1435,589,1923]" captionTargetPageId="4" captionText="FIGURE3. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, left cheliped, lateral view; B, same, mesial view; C, same, chela, dorsal view; D, same, chela, ventral view (setae omitted); E, same, fingers, dorsal view; F, same, fingers, ventral view (setae omitted)." httpUri="https://zenodo.org/record/1170047/files/figure.png" pageId="4" pageNumber="391">Fig. 3E, F</figureCitation>
|
||
) or unarmed (right chela;
|
||
<figureCitation id="DCB2B2A26302FFD3FBEEFEB4FB7C0D60" box="[1031,1153,295,320]" captionStart="FIGURE 4" captionStartId="5.[151,250,1405,1427]" captionTargetBox="[175,1403,208,1357]" captionTargetId="figure@5.[151,1435,185,1394]" captionTargetPageId="5" captionText="FIGURE 4. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, right cheliped, lateral view; B, same, mesial view; C, same, chela, dorsal view; D, same, chela, ventral view (setae omitted)." httpUri="https://zenodo.org/record/1170049/files/figure.png" pageId="4" pageNumber="391">Fig. 4C, D</figureCitation>
|
||
); adhesive plaque poorly developed. Fixed finger slender, tapering to slightly curved unguis; ventral surface with shallow concavity proximally; occlusal margin proximally with low, rounded tooth, otherwise without conspicuous armature. Dactylus broad, strongly flattened, laminar, not tapering distally, slightly twisted, about 0.7 times as long as palm and about 3.7 times as long as wide; occlusal margin slightly serrulate with 1 low, broadly rounded proximal tooth, ending in a small spiniform claw exceeded by broadly rounded dactylar terminal margin; outer and trminal margins sharply edged, with row of short setae becoming more numerous distally.
|
||
</paragraph>
|
||
<caption id="10F6FEAF6302FFD3FF7EF80BFDF10BCC" httpUri="https://zenodo.org/record/1170047/files/figure.png" pageId="4" pageNumber="391" startId="4.[151,250,1944,1966]" targetBox="[175,1395,634,1896]" targetPageId="4">
|
||
<paragraph id="4436AE276302FFD3FF7EF80BFDF10BCC" blockId="4.[151,1436,1944,2028]" pageId="4" pageNumber="391">
|
||
<emphasis id="76FD72356302FFD3FF7EF80BFEEF0B8E" bold="true" box="[151,274,1944,1966]" pageId="4" pageNumber="391">FIGURE 3.</emphasis>
|
||
<taxonomicName id="8389D5A46302FFD3FEF0F80BFE190B8E" authority="Komai, 2018" authorityName="Komai" authorityYear="2018" box="[281,484,1944,1966]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="391" phylum="Arthropoda" rank="species" species="shiranui" status="sp. nov.">
|
||
<emphasis id="76FD72356302FFD3FEF0F80BFE190B8E" box="[281,484,1944,1966]" italics="true" pageId="4" pageNumber="391">Thuylamea shiranui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6DCECF4E6302FFD3FE03F80BFDDE0B8E" box="[490,547,1944,1966]" pageId="4" pageNumber="391" rank="species">
|
||
<emphasis id="76FD72356302FFD3FE03F80BFDDE0B8E" bold="true" box="[490,547,1944,1966]" pageId="4" pageNumber="391">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype, female (cl 5.1 mm), CBM-ZC 14145. A, left cheliped, lateral view; B, same, mesial view; C, same, chela, dorsal view; D, same, chela, ventral view (setae omitted); E, same, fingers, dorsal view; F, same, fingers, ventral view (setae omitted).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="10F6FEAF6303FFD2FF7EFAEEFBB80992" httpUri="https://zenodo.org/record/1170049/files/figure.png" pageId="5" pageNumber="392" startId="5.[151,250,1405,1427]" targetBox="[175,1403,208,1357]" targetPageId="5">
|
||
<paragraph id="4436AE276303FFD2FF7EFAEEFBB80992" blockId="5.[151,1435,1405,1458]" pageId="5" pageNumber="392">
|
||
<emphasis id="76FD72356303FFD2FF7EFAEEFEE809B3" bold="true" box="[151,277,1405,1427]" pageId="5" pageNumber="392">FIGURE 4.</emphasis>
|
||
<taxonomicName id="8389D5A46303FFD2FEF7FAEEFE1009B3" authority="Komai, 2018" authorityName="Komai" authorityYear="2018" box="[286,493,1405,1427]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="392" phylum="Arthropoda" rank="species" species="shiranui" status="sp. nov.">
|
||
<emphasis id="76FD72356303FFD2FEF7FAEEFE1009B3" box="[286,493,1405,1427]" italics="true" pageId="5" pageNumber="392">Thuylamea shiranui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6DCECF4E6303FFD2FE1CFAEEFDCC09B3" box="[501,561,1405,1427]" pageId="5" pageNumber="392" rank="species">
|
||
<emphasis id="76FD72356303FFD2FE1CFAEEFDCC09B3" bold="true" box="[501,561,1405,1427]" pageId="5" pageNumber="392">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype, female (cl 5.1 mm), CBM-ZC 14145. A, right cheliped, lateral view; B, same, mesial view; C, same, chela, dorsal view; D, same, chela, ventral view (setae omitted).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4436AE276303FFD2FF2EFA4DFB710AEF" blockId="5.[151,1437,1502,2031]" pageId="5" pageNumber="392">
|
||
Only left pereopod 3 and right pereopod 5 preserved. Pereopod 3 (
|
||
<figureCitation id="DCB2B2A26303FFD2FC47FA4DFBF909D7" box="[942,1028,1502,1527]" captionStart="FIGURE 5" captionStartId="6.[151,250,1719,1741]" captionTargetBox="[178,1411,249,1669]" captionTargetId="figure@6.[151,1435,193,1698]" captionTargetPageId="6" captionText="FIGURE 5. Thuylamea shiranui n. sp. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146; I, J, paratype, male (cl 4.3 mm), CBM-ZC 14149. A, left maxilliped 3, lateral view; B, same, basal part with arthrobranch, ventral view;C, left pereopod 3, lateral view; D, same, dactylus, lateral view; E, right pereopod 5, lateral view; F, left pleopod 1, ventral view (setae omitted); G, left pleopod 2, ventral view (setae omitted); H, left pereopod 2, lateral view; I, right pleopod 1, ventral view (setae on exopod omitted); J, left pleopod 2, dorsal view (marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1170051/files/figure.png" pageId="5" pageNumber="392">Fig. 5C</figureCitation>
|
||
) moderately stout; ischium widened distally, unarmed; merus strongly compressed laterally, narrowing distally; carpus slightly widened distally, distinctly shorter than merus, bearing extensor distal process; propodus subequal in length to carpus; dactylus (
|
||
<figureCitation id="DCB2B2A26303FFD2FA86F9B5FF470A43" captionStart="FIGURE 5" captionStartId="6.[151,250,1719,1741]" captionTargetBox="[178,1411,249,1669]" captionTargetId="figure@6.[151,1435,193,1698]" captionTargetPageId="6" captionText="FIGURE 5. Thuylamea shiranui n. sp. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146; I, J, paratype, male (cl 4.3 mm), CBM-ZC 14149. A, left maxilliped 3, lateral view; B, same, basal part with arthrobranch, ventral view;C, left pereopod 3, lateral view; D, same, dactylus, lateral view; E, right pereopod 5, lateral view; F, left pleopod 1, ventral view (setae omitted); G, left pleopod 2, ventral view (setae omitted); H, left pereopod 2, lateral view; I, right pleopod 1, ventral view (setae on exopod omitted); J, left pleopod 2, dorsal view (marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1170051/files/figure.png" pageId="5" pageNumber="392">Fig. 5D</figureCitation>
|
||
) slender, 0.8 times as long as propodus, slightly compressed dorsoventrally, very slightly curved. Pereopod 5 (
|
||
<figureCitation id="DCB2B2A26303FFD2FF76F9FDFF0A0AA7" box="[159,247,1646,1671]" captionStart="FIGURE 5" captionStartId="6.[151,250,1719,1741]" captionTargetBox="[178,1411,249,1669]" captionTargetId="figure@6.[151,1435,193,1698]" captionTargetPageId="6" captionText="FIGURE 5. Thuylamea shiranui n. sp. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146; I, J, paratype, male (cl 4.3 mm), CBM-ZC 14149. A, left maxilliped 3, lateral view; B, same, basal part with arthrobranch, ventral view;C, left pereopod 3, lateral view; D, same, dactylus, lateral view; E, right pereopod 5, lateral view; F, left pleopod 1, ventral view (setae omitted); G, left pleopod 2, ventral view (setae omitted); H, left pereopod 2, lateral view; I, right pleopod 1, ventral view (setae on exopod omitted); J, left pleopod 2, dorsal view (marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1170051/files/figure.png" pageId="5" pageNumber="392">Fig. 5E</figureCitation>
|
||
) generally similar to pereopod 3, but distinctly shorter; carpus subequal in length to merus; propodus subequal in length to carpus, bearing grooming apparatus consisting of obliquely transverse rows of stiff serrulate setae increasing in length distally; dactylus slightly twisted, 0.8 times as long as propodus.
|
||
</paragraph>
|
||
<paragraph id="4436AE276303FFD2FF2EF949FAB30B7F" blockId="5.[151,1437,1502,2031]" pageId="5" pageNumber="392">
|
||
Pleopod 1 (
|
||
<figureCitation id="DCB2B2A26303FFD2FEA0F949FE610AD3" box="[329,412,1754,1779]" captionStart="FIGURE 5" captionStartId="6.[151,250,1719,1741]" captionTargetBox="[178,1411,249,1669]" captionTargetId="figure@6.[151,1435,193,1698]" captionTargetPageId="6" captionText="FIGURE 5. Thuylamea shiranui n. sp. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146; I, J, paratype, male (cl 4.3 mm), CBM-ZC 14149. A, left maxilliped 3, lateral view; B, same, basal part with arthrobranch, ventral view;C, left pereopod 3, lateral view; D, same, dactylus, lateral view; E, right pereopod 5, lateral view; F, left pleopod 1, ventral view (setae omitted); G, left pleopod 2, ventral view (setae omitted); H, left pereopod 2, lateral view; I, right pleopod 1, ventral view (setae on exopod omitted); J, left pleopod 2, dorsal view (marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1170051/files/figure.png" pageId="5" pageNumber="392">Fig. 5F</figureCitation>
|
||
) with small endopod about 0.3 length of exopod, not tapering, with slight notch on mesial margin, terminal margin rounded; exopod subequal in length to protopod. Pleopod 2 (
|
||
<figureCitation id="DCB2B2A26303FFD2FB63F96DFB1B0B37" box="[1162,1254,1790,1815]" captionStart="FIGURE 5" captionStartId="6.[151,250,1719,1741]" captionTargetBox="[178,1411,249,1669]" captionTargetId="figure@6.[151,1435,193,1698]" captionTargetPageId="6" captionText="FIGURE 5. Thuylamea shiranui n. sp. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146; I, J, paratype, male (cl 4.3 mm), CBM-ZC 14149. A, left maxilliped 3, lateral view; B, same, basal part with arthrobranch, ventral view;C, left pereopod 3, lateral view; D, same, dactylus, lateral view; E, right pereopod 5, lateral view; F, left pleopod 1, ventral view (setae omitted); G, left pleopod 2, ventral view (setae omitted); H, left pereopod 2, lateral view; I, right pleopod 1, ventral view (setae on exopod omitted); J, left pleopod 2, dorsal view (marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1170051/files/figure.png" pageId="5" pageNumber="392">Fig. 5G</figureCitation>
|
||
) with protopod slightly longer than rami; endopod slightly longer than exopod, with appendix interna arising at mid-length of mesial margin and distally curved mesially. Appendices internae of pleopods 3–5 similarly curved mesially.
|
||
</paragraph>
|
||
<paragraph id="4436AE276303FFD2FF2EF8F9FE7D0BCF" blockId="5.[151,1437,1502,2031]" pageId="5" pageNumber="392">
|
||
Uropod (
|
||
<figureCitation id="DCB2B2A26303FFD2FEC4F8F9FE7B0BA3" box="[301,390,1898,1923]" captionStart="FIGURE 2" captionStartId="3.[151,250,1519,1541]" captionTargetBox="[172,1413,226,1477]" captionTargetId="figure@3.[151,1435,193,1498]" captionTargetPageId="3" captionText="FIGURE 2. Thuylamea shiranui n. sp.. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146. A, carapace, dorsal view (left side damaged); B, anterior part of carapace and cephalic appendages, dorsal view (perpendicular for antennae); C, telson, dorsal view; D, right antennal peduncle, ventral view (setae omitted); E, distal par of outer flagellum of right antennule, dorsomesial view; F, epistome, ventral view; G, left uropod, dorsal view (perpendicular; setae partially omitted); eyes, ventro-frontal view (antennae broken off)." httpUri="https://zenodo.org/record/1170045/files/figure.png" pageId="5" pageNumber="392">Fig. 2G</figureCitation>
|
||
) with protopod divided into 2 lobes by deep, narrow incision, each lobe terminating in tiny spine. Endopod and exopod suboval, subequal in length, overreaching posterior tip of telson. Exopod slightly thick, setose lateral margin terminating distally in light angle, forming base of tiny distolateral spiniform seta; diaeresis transverse, unarmed.
|
||
</paragraph>
|
||
<caption id="10F6FEAF6300FFD1FF7EF924FF090B47" httpUri="https://zenodo.org/record/1170051/files/figure.png" pageId="6" pageNumber="393" startId="6.[151,250,1719,1741]" targetBox="[178,1411,249,1669]" targetPageId="6">
|
||
<paragraph id="4436AE276300FFD1FF7EF924FF090B47" blockId="6.[151,1436,1719,1895]" pageId="6" pageNumber="393">
|
||
<emphasis id="76FD72356300FFD1FF7EF924FEEE0AED" bold="true" box="[151,275,1719,1741]" pageId="6" pageNumber="393">FIGURE 5.</emphasis>
|
||
<taxonomicName id="8389D5A46300FFD1FEF3F924FE1A0AED" authority="Komai, 2018" authorityName="Komai" authorityYear="2018" box="[282,487,1719,1741]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="393" phylum="Arthropoda" rank="species" species="shiranui" status="sp. nov.">
|
||
<emphasis id="76FD72356300FFD1FEF3F924FE1A0AED" box="[282,487,1719,1741]" italics="true" pageId="6" pageNumber="393">Thuylamea shiranui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6DCECF4E6300FFD1FE04F924FDDA0AED" box="[493,551,1719,1741]" pageId="6" pageNumber="393" rank="species">
|
||
<emphasis id="76FD72356300FFD1FE04F924FDDA0AED" bold="true" box="[493,551,1719,1741]" pageId="6" pageNumber="393">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146; I, J, paratype, male (cl 4.3 mm), CBM-ZC 14149. A, left maxilliped 3, lateral view; B, same, basal part with arthrobranch, ventral view; C, left pereopod 3, lateral view; D, same, dactylus, lateral view; E, right pereopod 5, lateral view; F, left pleopod 1, ventral view (setae omitted); G, left pleopod 2, ventral view (setae omitted); H, left pereopod 2, lateral view; I, right pleopod 1, ventral view (setae on exopod omitted); J, left pleopod 2, dorsal view (marginal setae on rami omitted).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4436AE276300FFD1FF2EF806FAF20BF0" blockId="6.[151,1436,1940,2037]" pageId="6" pageNumber="393">
|
||
Notes on paratypes. All paratypes are more or less damaged; pereopod 2 (left) preserved only in the smallest specimen (cl
|
||
<quantity id="837103C26300FFD1FEC5F82BFE790BEF" box="[300,388,1976,2000]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="393" unit="mm" value="3.7">3.7 mm</quantity>
|
||
, sex not determinable; CBM-ZC14146); none of the specimens with pereopod 4.
|
||
</paragraph>
|
||
<paragraph id="4436AE276300FFD0FF2EF84EFC110DF0" blockId="6.[151,1436,1940,2037]" lastBlockId="7.[151,1437,151,464]" lastPageId="7" lastPageNumber="394" pageId="6" pageNumber="393">
|
||
<typeStatus id="9B3210856300FFD1FF2EF84EFEC90BD5" box="[199,308,2013,2037]" pageId="6" pageNumber="393" type="paratype">Paratypes</typeStatus>
|
||
generally similar to
|
||
<typeStatus id="9B3210856300FFD1FDCDF84FFD770BD5" box="[548,650,2012,2037]" pageId="6" pageNumber="393" type="holotype">holotype</typeStatus>
|
||
. Carapace dorsum flattened in anterior half and rounded in posterior half in male (CBM-ZC 14149), whereas rounded entirely in smallest specimen (CBM-ZC 14146); in other two female
|
||
<typeStatus id="9B3210856301FFD0FF03FF2EFEA80CF5" box="[234,341,189,213]" pageId="7" pageNumber="394" type="paratype">paratypes</typeStatus>
|
||
(CBM-ZC 14147, 14148), carapace dorsum similar to that of
|
||
<typeStatus id="9B3210856301FFD0FBE7FF2FFB890CF5" box="[1038,1140,188,213]" pageId="7" pageNumber="394" type="holotype">holotype</typeStatus>
|
||
. Eyestalks as figured (
|
||
<figureCitation id="DCB2B2A26301FFD0FA86FF2FFF400CD8" captionStart="FIGURE 2" captionStartId="3.[151,250,1519,1541]" captionTargetBox="[172,1413,226,1477]" captionTargetId="figure@3.[151,1435,193,1498]" captionTargetPageId="3" captionText="FIGURE 2. Thuylamea shiranui n. sp.. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146. A, carapace, dorsal view (left side damaged); B, anterior part of carapace and cephalic appendages, dorsal view (perpendicular for antennae); C, telson, dorsal view; D, right antennal peduncle, ventral view (setae omitted); E, distal par of outer flagellum of right antennule, dorsomesial view; F, epistome, ventral view; G, left uropod, dorsal view (perpendicular; setae partially omitted); eyes, ventro-frontal view (antennae broken off)." httpUri="https://zenodo.org/record/1170045/files/figure.png" pageId="7" pageNumber="394">Fig. 2H</figureCitation>
|
||
). Antennular peduncle overreaching distal margin of antennal scale by 0.2–0.4 length of article 2; lateral antennular flagellum bifurcate at 8th article in male
|
||
<typeStatus id="9B3210856301FFD0FD35FE96FCC00D3D" box="[732,829,261,285]" pageId="7" pageNumber="394" type="paratype">paratype</typeStatus>
|
||
(CBM-ZC 14149) and at 11th article in largest female
|
||
<typeStatus id="9B3210856301FFD0FF7EFEBBFF0A0D60" box="[151,247,296,320]" pageId="7" pageNumber="394" type="paratype">paratype</typeStatus>
|
||
(CBM-ZC 14148) (in other two specimens, the antennule and antennae are missing). Cheliped palm 4.3– 4.4 times as long as wide; dactylar articulation of palm without tooth in all preserved chela (
|
||
<figureCitation id="DCB2B2A26301FFD0FB2BFEDFFAC20D44" box="[1218,1343,332,357]" captionStart="FIGURE 4" captionStartId="5.[151,250,1405,1427]" captionTargetBox="[175,1403,208,1357]" captionTargetId="figure@5.[151,1435,185,1394]" captionTargetPageId="5" captionText="FIGURE 4. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, right cheliped, lateral view; B, same, mesial view; C, same, chela, dorsal view; D, same, chela, ventral view (setae omitted)." httpUri="https://zenodo.org/record/1170049/files/figure.png" pageId="7" pageNumber="394">Fig. 4C, D</figureCitation>
|
||
) (1 pair from CBM-ZC 14147; one side from CBM-ZC 14146 and 14148). Pereopod 2 (
|
||
<figureCitation id="DCB2B2A26301FFD0FBFAFEFCFB970DA8" box="[1043,1130,367,392]" captionStart="FIGURE 5" captionStartId="6.[151,250,1719,1741]" captionTargetBox="[178,1411,249,1669]" captionTargetId="figure@6.[151,1435,193,1698]" captionTargetPageId="6" captionText="FIGURE 5. Thuylamea shiranui n. sp. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146; I, J, paratype, male (cl 4.3 mm), CBM-ZC 14149. A, left maxilliped 3, lateral view; B, same, basal part with arthrobranch, ventral view;C, left pereopod 3, lateral view; D, same, dactylus, lateral view; E, right pereopod 5, lateral view; F, left pleopod 1, ventral view (setae omitted); G, left pleopod 2, ventral view (setae omitted); H, left pereopod 2, lateral view; I, right pleopod 1, ventral view (setae on exopod omitted); J, left pleopod 2, dorsal view (marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1170051/files/figure.png" pageId="7" pageNumber="394">Fig. 5H</figureCitation>
|
||
) of smallest specimen with merus subequal in length to ischium; carpus divided into 5 articles, length ratio (proximal to distal) 2.9: 0.8: 0:8: 0.6: 1; chela about half-length of carpus; dactylus 1.7 times as long as palm.
|
||
</paragraph>
|
||
<caption id="10F6FEAF6301FFD0FF7EFB6EFD210912" httpUri="https://zenodo.org/record/1170053/files/figure.png" pageId="7" pageNumber="394" startId="7.[151,250,1277,1299]" targetBox="[151,1435,517,1255]" targetPageId="7">
|
||
<paragraph id="4436AE276301FFD0FF7EFB6EFD210912" blockId="7.[151,1436,1277,1330]" pageId="7" pageNumber="394">
|
||
<emphasis id="76FD72356301FFD0FF7EFB6EFEE80933" bold="true" box="[151,277,1277,1299]" pageId="7" pageNumber="394">FIGURE 6.</emphasis>
|
||
<taxonomicName id="8389D5A46301FFD0FEF4FB6EFE160933" authority="Komai, 2018" authorityName="Komai" authorityYear="2018" box="[285,491,1277,1299]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="394" phylum="Arthropoda" rank="species" species="shiranui" status="sp. nov.">
|
||
<emphasis id="76FD72356301FFD0FEF4FB6EFE160933" box="[285,491,1277,1299]" italics="true" pageId="7" pageNumber="394">Thuylamea shiranui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6DCECF4E6301FFD0FE1AFB6EFDD30933" box="[499,558,1277,1299]" pageId="7" pageNumber="394" rank="species">
|
||
<emphasis id="76FD72356301FFD0FE1AFB6EFDD30933" bold="true" box="[499,558,1277,1299]" pageId="7" pageNumber="394">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype, female (cl 6.7 mm), CBM-ZC 14148, cephalothorax in dorsolateral view, showing left side of ovary extending to carapace dorsum.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="10F6FEAF6301FFD0FF7EF8BAFC9A0B7E" httpUri="https://zenodo.org/record/1170055/files/figure.png" pageId="7" pageNumber="394" startId="7.[151,250,1833,1855]" targetBox="[166,1423,1394,1796]" targetPageId="7">
|
||
<paragraph id="4436AE276301FFD0FF7EF8BAFC9A0B7E" blockId="7.[151,1436,1833,1886]" pageId="7" pageNumber="394">
|
||
<emphasis id="76FD72356301FFD0FF7EF8BAFEE80B1F" bold="true" box="[151,277,1833,1855]" pageId="7" pageNumber="394">FIGURE 7.</emphasis>
|
||
<taxonomicName id="8389D5A46301FFD0FEF7F8BAFD750B1F" authority="Nguyen, 2001" authorityName="Nguyen" authorityYear="2001" box="[286,648,1833,1855]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="394" phylum="Arthropoda" rank="species" species="camelus">
|
||
<emphasis id="76FD72356301FFD0FEF7F8BAFE160B1F" box="[286,491,1833,1855]" italics="true" pageId="7" pageNumber="394">Thuylamea camelus</emphasis>
|
||
Nguyên, 2001
|
||
</taxonomicName>
|
||
, male (cl 4.2 mm), ZRC 1994.4387. A, carapace and cephalic appendages, lateral view; B, close up of posterodorsal part of carapace, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4436AE276301FFDFFF2EF814FDBD0CD8" blockId="7.[151,1436,1927,2024]" lastBlockId="8.[151,1437,151,1941]" lastPageId="8" lastPageNumber="395" pageId="7" pageNumber="394">
|
||
Male pleopod 1 (
|
||
<figureCitation id="DCB2B2A26301FFD0FE62F814FE240B80" box="[395,473,1927,1952]" captionStart="FIGURE 5" captionStartId="6.[151,250,1719,1741]" captionTargetBox="[178,1411,249,1669]" captionTargetId="figure@6.[151,1435,193,1698]" captionTargetPageId="6" captionText="FIGURE 5. Thuylamea shiranui n. sp. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146; I, J, paratype, male (cl 4.3 mm), CBM-ZC 14149. A, left maxilliped 3, lateral view; B, same, basal part with arthrobranch, ventral view;C, left pereopod 3, lateral view; D, same, dactylus, lateral view; E, right pereopod 5, lateral view; F, left pleopod 1, ventral view (setae omitted); G, left pleopod 2, ventral view (setae omitted); H, left pereopod 2, lateral view; I, right pleopod 1, ventral view (setae on exopod omitted); J, left pleopod 2, dorsal view (marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1170051/files/figure.png" pageId="7" pageNumber="394">
|
||
Fig.
|
||
<date id="303788E76301FFD0FE28F814FE240B80" box="[449,473,1927,1952]" pageId="7" pageNumber="394">5I</date>
|
||
</figureCitation>
|
||
) with endopod less than half-length of exopod, proportionally narrower than female endopod, with 3 plumose setae terminally and shorter scattered simple setae on ventral surface and mesial margin; exopod slightly longer than protopod. Male pleopod 2 (
|
||
<figureCitation id="DCB2B2A26301FFD0FCE4F85CFCA10BC8" box="[781,860,1999,2024]" captionStart="FIGURE 5" captionStartId="6.[151,250,1719,1741]" captionTargetBox="[178,1411,249,1669]" captionTargetId="figure@6.[151,1435,193,1698]" captionTargetPageId="6" captionText="FIGURE 5. Thuylamea shiranui n. sp. A–G, holotype, female (cl 5.1 mm), CBM-ZC 14145; H, paratype, sex indeterminable (cl 3.7 mm), CBM-ZC 14146; I, J, paratype, male (cl 4.3 mm), CBM-ZC 14149. A, left maxilliped 3, lateral view; B, same, basal part with arthrobranch, ventral view;C, left pereopod 3, lateral view; D, same, dactylus, lateral view; E, right pereopod 5, lateral view; F, left pleopod 1, ventral view (setae omitted); G, left pleopod 2, ventral view (setae omitted); H, left pereopod 2, lateral view; I, right pleopod 1, ventral view (setae on exopod omitted); J, left pleopod 2, dorsal view (marginal setae on rami omitted)." httpUri="https://zenodo.org/record/1170051/files/figure.png" pageId="7" pageNumber="394">Fig. 5J</figureCitation>
|
||
) with protopod slightly longer than rami; endopod with appendices interna and masculina arising at mid-length of mesial margin, appendix interna straight, rod-like, appendix masculina also rod-like, distinctly longer than appendix interna, bearing 5 simple setae in distal half; exopod slightly shorter than endopod.
|
||
</paragraph>
|
||
<paragraph id="4436AE27630EFFDFFF2EFE97FE170D3D" blockId="8.[151,1437,151,1941]" box="[199,490,260,285]" pageId="8" pageNumber="395">
|
||
<emphasis id="76FD7235630EFFDFFF2EFE97FE900D3D" bold="true" box="[199,365,260,285]" pageId="8" pageNumber="395">Colour in life.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0C93FDAC630EFFDFFF2EFEB4FB140D45" pageId="8" pageNumber="395" type="etymology">
|
||
<paragraph id="4436AE27630EFFDFFF2EFEB4FB140D45" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="395">
|
||
<emphasis id="76FD7235630EFFDFFF2EFEB4FEB30D60" bold="true" box="[199,334,295,320]" pageId="8" pageNumber="395">Etymology.</emphasis>
|
||
Derived from another Japanese name of the
|
||
<typeStatus id="9B321085630EFFDFFCB8FEBBFC7C0D60" box="[849,897,296,320]" pageId="8" pageNumber="395">type</typeStatus>
|
||
locality. “Shiranui” Sea is another name of the Yatsushiro Sea, referring to a kind of mysterious mirage occurring in the area, called as “
|
||
<taxonomicName id="8389D5A4630EFFDFFB90FEDFFB270D45" authorityName="Komai" authorityYear="2018" box="[1145,1242,332,357]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="shiranui">shiranui</taxonomicName>
|
||
”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0C93FDAC630EFFDFFF2EFEFCFAA80E40" pageId="8" pageNumber="395" type="distribution">
|
||
<paragraph id="4436AE27630EFFDFFF2EFEFCFAA80E40" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="395">
|
||
<emphasis id="76FD7235630EFFDFFF2EFEFCFE0D0DA8" bold="true" box="[199,496,367,392]" pageId="8" pageNumber="395">Distribution and habitat.</emphasis>
|
||
So far known only from Yatsushiro Sea, western Kyushu,
|
||
<collectingCountry id="3C9EEEB7630EFFDFFB96FEE3FB3F0DA7" box="[1151,1218,368,391]" name="Japan" pageId="8" pageNumber="395">Japan</collectingCountry>
|
||
; at depths of
|
||
<quantity id="837103C2630EFFDFFABFFEE3FF4D0D8C" metricMagnitude="1" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="8" pageNumber="395" unit="m" value="28.0" valueMax="29.0" valueMin="27.0">
|
||
27–
|
||
<specimenCount id="528F65AE630EFFDFFA69FEE3FF4D0D8C" pageId="8" pageNumber="395" type="generic">29 m</specimenCount>
|
||
</quantity>
|
||
. All specimens were collected from silt sediments using a Smith-McIntyre grab; other information on ecology is presently unavailable. Other benthic animals found in these samples were:
|
||
<taxonomicName id="8389D5A4630EFFDFFBFEFE24FB2B0DF0" authorityName="Pettibone" authorityYear="1992" box="[1047,1238,439,464]" class="Polychaeta" family="Sigalionidae" genus="Labiosthenolepis" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="395" phylum="Annelida" rank="genus">
|
||
<emphasis id="76FD7235630EFFDFFBFEFE24FB2B0DF0" box="[1047,1238,439,464]" italics="true" pageId="8" pageNumber="395">Labiosthenolepis</emphasis>
|
||
</taxonomicName>
|
||
sp. (Polychaeta:
|
||
<taxonomicName id="8389D5A4630EFFDFFF7EFE4FFEE40DD5" authorityName="Kinberg" authorityYear="1856" box="[151,281,476,501]" class="Polychaeta" family="Polynoidae" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="395" phylum="Annelida" rank="family">Polynoidae</taxonomicName>
|
||
; stn 13, 14, 17),
|
||
<taxonomicName id="8389D5A4630EFFDFFE01FE4FFCB10DD5" authority="(Kitamori)" authorityName="Kitamori" baseAuthorityName="Kitamori" baseAuthorityYear="1960" box="[488,844,476,501]" class="Polychaeta" family="Pilargidae" genus="Sigambra" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="395" phylum="Annelida" rank="species" species="hanaokai">
|
||
<emphasis id="76FD7235630EFFDFFE01FE4FFD350DD5" box="[488,712,476,501]" italics="true" pageId="8" pageNumber="395">Sigambra hanaokai</emphasis>
|
||
(Kitamori)
|
||
</taxonomicName>
|
||
(Polychaeta: Plargidae: stn 13, 14, 17),
|
||
<taxonomicName id="8389D5A4630EFFDFFAC3FE4FFE940E38" authority="Okunoda" authorityName="Okunoda" authorityYear="1937" class="Polychaeta" family="Magelonidae" genus="Magelona" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="395" phylum="Annelida" rank="species" species="japonica">
|
||
<emphasis id="76FD7235630EFFDFFAC3FE4FFF040E38" italics="true" pageId="8" pageNumber="395">Magelona japonica</emphasis>
|
||
Okunoda
|
||
</taxonomicName>
|
||
(Polychaeta:
|
||
<taxonomicName id="8389D5A4630EFFDFFDE1FE6CFD630E38" authorityName="Cunningham & Ramage" authorityYear="1888" box="[520,670,511,536]" class="Polychaeta" family="Magelonidae" kingdom="Animalia" order="Spionida" pageId="8" pageNumber="395" phylum="Annelida" rank="family">Magelonidae</taxonomicName>
|
||
; stn 13, 14, 17),
|
||
<taxonomicName id="8389D5A4630EFFDFFC8DFE6CFAF10E38" authority="Stimpson" authorityName="Stimpson" authorityYear="1858" box="[868,1292,511,536]" class="Malacostraca" family="Varunidae" genus="Asthenognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="inaequipes">
|
||
<emphasis id="76FD7235630EFFDFFC8DFE6CFB670E38" box="[868,1178,511,536]" italics="true" pageId="8" pageNumber="395">Asthenognathus inaequipes</emphasis>
|
||
Stimpson
|
||
</taxonomicName>
|
||
(Brachyura:
|
||
<taxonomicName id="8389D5A4630EFFDFFF7EFDB7FEF10E1D" box="[151,268,548,573]" class="Malacostraca" family="Varunidae" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="family">Varunidae</taxonomicName>
|
||
; stn 13, 17),
|
||
<taxonomicName id="8389D5A4630EFFDFFE76FDB7FDCE0E1D" box="[415,563,548,573]" class="Ophiuroidea" family="Amphiuridae" kingdom="Animalia" order="Ophiurida" pageId="8" pageNumber="395" phylum="Echinodermata" rank="family">Amphiuridae</taxonomicName>
|
||
sp. (Ophiuroidea; stn 13, 14, 17),
|
||
<taxonomicName id="8389D5A4630EFFDFFC53FDB7FB1E0E1D" authority="Marion" authorityName="Marion" box="[954,1251,548,573]" class="Enteropneusta" family="Spengelidae" genus="Glandiceps" kingdom="Animalia" pageId="8" pageNumber="395" phylum="Hemichordata" rank="species" species="hacksi">
|
||
<emphasis id="76FD7235630EFFDFFC53FDB7FB750E1D" box="[954,1160,548,573]" italics="true" pageId="8" pageNumber="395">Glandiceps hacksi</emphasis>
|
||
Marion
|
||
</taxonomicName>
|
||
(Enteropneusta:
|
||
<taxonomicName id="8389D5A4630EFFDFFF7EFDD4FED90E40" authorityName="Willey" authorityYear="1899" box="[151,292,583,608]" class="Enteropneusta" family="Spengelidae" kingdom="Animalia" pageId="8" pageNumber="395" phylum="Hemichordata" rank="family">Spengelidae</taxonomicName>
|
||
; stn 13, 17), and
|
||
<taxonomicName id="8389D5A4630EFFDFFE0AFDD4FC480E40" authority="(Bleeker)" authorityName="Bleeker" baseAuthorityName="Bleeker" baseAuthorityYear="1860" box="[483,949,583,608]" class="Actinopterygii" family="Gobiidae" genus="Paratrypauchen" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="395" phylum="Chordata" rank="species" species="microcephalus">
|
||
<emphasis id="76FD7235630EFFDFFE0AFDD4FCB90E40" box="[483,836,583,608]" italics="true" pageId="8" pageNumber="395">Paratrypauchen microcephalus</emphasis>
|
||
(Bleeker)
|
||
</taxonomicName>
|
||
(Teleostei:
|
||
<taxonomicName id="8389D5A4630EFFDFFBD1FDD4FB5F0E40" box="[1080,1186,583,608]" class="Actinopterygii" family="Gobiidae" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="395" phylum="Chordata" rank="family">Gobiidae</taxonomicName>
|
||
; stn 13, 14, 17).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0C93FDAC630EFFDFFF2EFDFFFAB90BB4" pageId="8" pageNumber="395" type="discussion">
|
||
<paragraph id="4436AE27630EFFDFFF2EFDFFFCF30FA0" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="395">
|
||
<emphasis id="76FD7235630EFFDFFF2EFDFFFEC60EA5" bold="true" box="[199,315,620,645]" pageId="8" pageNumber="395">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="9B321085630EFFDFFE9FFDFEFE5B0EA5" box="[374,422,621,645]" pageId="8" pageNumber="395">type</typeStatus>
|
||
series of the new species consists of one small specimen of unidentified sex (cl
|
||
<quantity id="837103C2630EFFDFFADCFDFFFA6D0EA4" box="[1333,1424,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="8" pageNumber="395" unit="mm" value="3.7">3.7 mm</quantity>
|
||
), three female (none ovigerous) specimens (cl
|
||
<quantity id="837103C2630EFFDFFD72FD1CFCD80E88" box="[667,805,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" metricValueMax="6.7" metricValueMin="4.7" pageId="8" pageNumber="395" unit="mm" value="5.7" valueMax="6.7" valueMin="4.7">4.7–6.7 mm</quantity>
|
||
) and one male specimen (cl
|
||
<quantity id="837103C2630EFFDFFB9BFD03FB330E88" box="[1138,1230,656,680]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="8" pageNumber="395" unit="mm" value="4.3">4.3 mm</quantity>
|
||
). As noted above, none of them is intact. The largest female (CBM-ZC 14148,
|
||
<typeStatus id="9B321085630EFFDFFCD7FD26FC5C0EED" box="[830,929,693,717]" pageId="8" pageNumber="395" type="paratype">paratype</typeStatus>
|
||
, cl
|
||
<quantity id="837103C2630EFFDFFC21FD27FBDD0EEC" box="[968,1056,692,717]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="8" pageNumber="395" unit="mm" value="6.7">6.7 mm</quantity>
|
||
, total length about
|
||
<quantity id="837103C2630EFFDFFB1FFD27FABA0EEC" box="[1270,1351,692,716]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="395" unit="mm" value="20.0">20 mm</quantity>
|
||
) has an ovary with partially formed eggs, which is visible inside the cephalothorax (
|
||
<figureCitation id="DCB2B2A2630EFFDFFC00FD44FBD00ED0" box="[1001,1069,727,752]" captionStart="FIGURE 6" captionStartId="7.[151,250,1277,1299]" captionTargetBox="[151,1435,517,1255]" captionTargetId="figure@7.[151,1435,517,1255]" captionTargetPageId="7" captionText="FIGURE 6. Thuylamea shiranui n. sp., paratype, female (cl 6.7 mm), CBM-ZC 14148, cephalothorax in dorsolateral view, showing left side of ovary extending to carapacedorsum." httpUri="https://zenodo.org/record/1170053/files/figure.png" pageId="8" pageNumber="395">Fig. 6</figureCitation>
|
||
), suggesting the sexual maturity. The
|
||
<typeStatus id="9B321085630EFFDFFF20FD6EFF040F35" box="[201,249,765,789]" pageId="8" pageNumber="395">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="8389D5A4630EFFDFFE8AFD6EFE270F35" authorityName="Nguyen" authorityYear="2001" box="[355,474,764,789]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="camelus">
|
||
<emphasis id="76FD7235630EFFDFFE8AFD6EFE270F35" box="[355,474,764,789]" italics="true" pageId="8" pageNumber="395">T. camelus</emphasis>
|
||
</taxonomicName>
|
||
consists of five male specimens (total length
|
||
<quantity id="837103C2630EFFDFFC3CFD6FFBB20F34" box="[981,1103,764,788]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="3.9" metricValueMin="3.1" pageId="8" pageNumber="395" unit="mm" value="35.0" valueMax="39.0" valueMin="31.0">31–39 mm</quantity>
|
||
) (
|
||
<bibRefCitation id="2018D3D6630EFFDFFB8DFD6FFAFD0F34" box="[1124,1280,764,788]" pageId="8" pageNumber="395" refId="ref4172" refString="Nguyen, V. X. (2001) A new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from South Vietnam. Zoologische Mededelingen, Leiden, 75, 217 - 228." type="book chapter">Nguyen 2001</bibRefCitation>
|
||
). Considering the possible variation related to size and sex, a small male specimen of
|
||
<taxonomicName id="8389D5A4630EFFDFFC24FCB2FBBA0F18" authorityName="Nguyen" authorityYear="2001" box="[973,1095,799,824]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="camelus">
|
||
<emphasis id="76FD7235630EFFDFFC24FCB2FBBA0F18" box="[973,1095,799,824]" italics="true" pageId="8" pageNumber="395">T. camelus</emphasis>
|
||
</taxonomicName>
|
||
(ZRC 1994.4387, cl
|
||
<quantity id="837103C2630EFFDFFAD2FCB3FA690F18" box="[1339,1428,800,824]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="8" pageNumber="395" unit="mm" value="4.2">4.2 mm</quantity>
|
||
) was examined for comparison. The description of a female specimen from
|
||
<collectingRegion id="864D60C5630EFFDFFC01FCD7FB900F7D" box="[1000,1133,836,861]" country="China" name="Guangdong" pageId="8" pageNumber="395">Guangdong</collectingRegion>
|
||
, South
|
||
<collectingCountry id="3C9EEEB7630EFFDFFB2AFCD7FAF50F7D" box="[1219,1288,836,861]" name="China" pageId="8" pageNumber="395">China</collectingCountry>
|
||
, provided by
|
||
<bibRefCitation id="2018D3D6630EFFDFFF7EFCFBFE840FA0" box="[151,377,871,896]" pageId="8" pageNumber="395" refId="ref4207" refString="Wang, Y. & Sha, Z. (2016) Redescription of the type species of the genus Thuylamea Nguyen, 2001 (Decapoda, Caridea, Alpheidae), based on a female specimen. Crustaceana, 89, 1185 - 1191. https: // doi. org / 10.1163 / 15685403 - 00003572" type="journal article">Wang & Sha (2016)</bibRefCitation>
|
||
, is also referred for the comparison.
|
||
</paragraph>
|
||
<paragraph id="4436AE27630EFFDFFF2EFC1FFE8B0A55" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="395">
|
||
The present new species share many features with
|
||
<taxonomicName id="8389D5A4630EFFDFFD14FC1FFC200F85" authorityName="Nguyen" authorityYear="2001" box="[765,989,908,933]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="camelus">
|
||
<emphasis id="76FD7235630EFFDFFD14FC1FFC200F85" box="[765,989,908,933]" italics="true" pageId="8" pageNumber="395">Thuylamea camelus</emphasis>
|
||
</taxonomicName>
|
||
, e.g., the elongate, subcylindrical body; the greatly reduced rostrum; the unarmed frontal or anterolateral margins of the carapace; the absence of an articulated flap at the posteroventral angle of the pleomere 6; the partially exposed eyes, the elongate carpocerite of the antennal peduncle; and the enlarged, anteriorly extended, symmetrical chelipeds, with thin lamella-like dactyli positioned laterally. As has been described by
|
||
<bibRefCitation id="2018D3D6630EFFDFFD2CFB8EFC880814" box="[709,885,1052,1077]" pageId="8" pageNumber="395" refId="ref4172" refString="Nguyen, V. X. (2001) A new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from South Vietnam. Zoologische Mededelingen, Leiden, 75, 217 - 228." type="book chapter">Nguyen (2001)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="2018D3D6630EFFDFFC5EFB8EFB570815" box="[951,1194,1052,1077]" pageId="8" pageNumber="395" refId="ref4207" refString="Wang, Y. & Sha, Z. (2016) Redescription of the type species of the genus Thuylamea Nguyen, 2001 (Decapoda, Caridea, Alpheidae), based on a female specimen. Crustaceana, 89, 1185 - 1191. https: // doi. org / 10.1163 / 15685403 - 00003572" type="journal article">Wang & Sha (2016)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="8389D5A4630EFFDFFB52FB8EFAC50815" authorityName="Nguyen" authorityYear="2001" box="[1211,1336,1052,1077]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="camelus">
|
||
<emphasis id="76FD7235630EFFDFFB52FB8EFAC50815" box="[1211,1336,1052,1077]" italics="true" pageId="8" pageNumber="395">T. camelus</emphasis>
|
||
</taxonomicName>
|
||
is quite distinctive in having peculiar submedian crests near the posterodorsal margin of the carapace.
|
||
<bibRefCitation id="2018D3D6630EFFDFFB2AFBACFF1C085D" pageId="8" pageNumber="395" refId="ref4035" refString="Anker, A. & De Grave, S. (2016) An updated and annotated checklist of marine and brackish caridean shrimps of Singapore. The Raffles Bulletin of Zoology, 34 (Supplement), 343 - 454." type="journal article">Anker & De Grave (2016)</bibRefCitation>
|
||
remarked that the submedian crests on the posterior part of the carapace is poorly developed in their young male specimen from
|
||
<collectingCountry id="3C9EEEB7630EFFDFFE6DFB14FE040880" box="[388,505,1159,1184]" name="Singapore" pageId="8" pageNumber="395">Singapore</collectingCountry>
|
||
, suggesting the ontogenetic variation in the development of this peculiar structure. Reexamination of this specimen (ZRC 1994.4387) has confirmed that there is a pair of low tubercles (
|
||
<figureCitation id="DCB2B2A2630EFFDFFAFDFB3FFA7308E4" box="[1300,1422,1196,1221]" captionStart="FIGURE 7" captionStartId="7.[151,250,1833,1855]" captionTargetBox="[166,1423,1394,1796]" captionTargetId="figure@7.[151,1436,1379,1811]" captionTargetPageId="7" captionText="FIGURE 7. Thuylamea camelus Nguyên, 2001, male (cl 4.2 mm), ZRC 1994.4387. A, carapace and cephalic appendages, lateral view; B,close up of posterodorsal part of carapace, lateral view." httpUri="https://zenodo.org/record/1170055/files/figure.png" pageId="8" pageNumber="395">Fig. 7A, B</figureCitation>
|
||
), corresponding to the prominent crests in the adults, near the posterodorsal margin of the carapace. As described above, the posterior part of the carapace dorsum is smooth without any ornamentation in
|
||
<taxonomicName id="8389D5A4630EFFDFFB48FB66FAE1092D" authority="Komai, 2018" authorityName="Komai" authorityYear="2018" box="[1185,1308,1268,1293]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="shiranui" status="sp. nov.">
|
||
<emphasis id="76FD7235630EFFDFFB48FB66FAE1092D" box="[1185,1308,1268,1293]" italics="true" pageId="8" pageNumber="395">T. shiranui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6DCECF4E630EFFDFFACCFB67FA9A092D" box="[1317,1383,1268,1293]" pageId="8" pageNumber="395" rank="species">
|
||
<emphasis id="76FD7235630EFFDFFACCFB67FA9A092D" bold="true" box="[1317,1383,1268,1293]" pageId="8" pageNumber="395">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
The antennular peduncle is relatively longer in
|
||
<taxonomicName id="8389D5A4630EFFDFFD95FA8AFD080910" authority="Komai, 2018" authorityName="Komai" authorityYear="2018" box="[636,757,1303,1328]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="shiranui" status="sp. nov.">
|
||
<emphasis id="76FD7235630EFFDFFD95FA8AFD080910" box="[636,757,1303,1328]" italics="true" pageId="8" pageNumber="395">T. shiranui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6DCECF4E630EFFDFFD14FA84FCC00910" box="[765,829,1303,1328]" pageId="8" pageNumber="395" rank="species">
|
||
<emphasis id="76FD7235630EFFDFFD14FA84FCC00910" bold="true" box="[765,829,1303,1328]" pageId="8" pageNumber="395">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
than in
|
||
<taxonomicName id="8389D5A4630EFFDFFC72FA8AFBE90910" authorityName="Nguyen" authorityYear="2001" box="[923,1044,1303,1328]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="camelus">
|
||
<emphasis id="76FD7235630EFFDFFC72FA8AFBE90910" box="[923,1044,1303,1328]" italics="true" pageId="8" pageNumber="395">T. camelus</emphasis>
|
||
</taxonomicName>
|
||
[overreaching the distal margin of the antennal scaphocerite by the 0.2–0.5 length of the article
|
||
<quantity id="837103C2630EFFDFFCD4FAAFFC950975" box="[829,872,1340,1365]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="395" unit="in" value="2.0">2 in</quantity>
|
||
<taxonomicName id="8389D5A4630EFFDFFC87FAAEFC1B0975" authority="Komai, 2018" authorityName="Komai" authorityYear="2018" box="[878,998,1340,1365]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="shiranui" status="sp. nov.">
|
||
<emphasis id="76FD7235630EFFDFFC87FAAEFC1B0975" box="[878,998,1340,1365]" italics="true" pageId="8" pageNumber="395">T. shiranui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6DCECF4E630EFFDFFC04FAAFFBD60975" box="[1005,1067,1340,1365]" pageId="8" pageNumber="395" rank="species">
|
||
<emphasis id="76FD7235630EFFDFFC04FAAFFBD60975" bold="true" box="[1005,1067,1340,1365]" pageId="8" pageNumber="395">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="DCB2B2A2630EFFDFFBD2FAAFFB6D0974" box="[1083,1168,1340,1365]" captionStart="FIGURE 1" captionStartId="2.[151,250,1255,1277]" captionTargetBox="[175,1415,212,1210]" captionTargetId="figure@2.[151,1435,193,1234]" captionTargetPageId="2" captionText="FIGURE 1. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, carapace and cephalic appendages, lateral (right) view; B,pleon, telson and pleonal appendages,lateral (right) view." httpUri="https://zenodo.org/record/1170043/files/figure.png" pageId="8" pageNumber="395">Fig. 1A</figureCitation>
|
||
) versus just reaching or overreaching it by the half-length of the article
|
||
<quantity id="837103C2630EFFDFFD20FAF3FD040958" box="[713,761,1375,1400]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="395" unit="in" value="3.0">3 in</quantity>
|
||
<taxonomicName id="8389D5A4630EFFDFFCEAFAF2FC820958" authorityName="Nguyen" authorityYear="2001" box="[771,895,1375,1400]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="camelus">
|
||
<emphasis id="76FD7235630EFFDFFCEAFAF2FC820958" box="[771,895,1375,1400]" italics="true" pageId="8" pageNumber="395">T. camelus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="DCB2B2A2630EFFDFFC7BFACCFC0C0958" box="[914,1009,1375,1400]" captionStart="FIGURE 7" captionStartId="7.[151,250,1833,1855]" captionTargetBox="[166,1423,1394,1796]" captionTargetId="figure@7.[151,1436,1379,1811]" captionTargetPageId="7" captionText="FIGURE 7. Thuylamea camelus Nguyên, 2001, male (cl 4.2 mm), ZRC 1994.4387. A, carapace and cephalic appendages, lateral view; B,close up of posterodorsal part of carapace, lateral view." httpUri="https://zenodo.org/record/1170055/files/figure.png" pageId="8" pageNumber="395">Fig. 7A</figureCitation>
|
||
)]. Furthermore, comparison with the description of a female specimen (cl
|
||
<quantity id="837103C2630EFFDFFDD3FA17FD6309BC" box="[570,670,1412,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="8" pageNumber="395" unit="mm" value="11.5">11.5 mm</quantity>
|
||
) given by
|
||
<bibRefCitation id="2018D3D6630EFFDFFCF1FA16FC0609BD" box="[792,1019,1412,1437]" pageId="8" pageNumber="395" refId="ref4207" refString="Wang, Y. & Sha, Z. (2016) Redescription of the type species of the genus Thuylamea Nguyen, 2001 (Decapoda, Caridea, Alpheidae), based on a female specimen. Crustaceana, 89, 1185 - 1191. https: // doi. org / 10.1163 / 15685403 - 00003572" type="journal article">Wang & Sha (2016)</bibRefCitation>
|
||
shows that the shape of the cheliped dactylus is different between the two species. In
|
||
<taxonomicName id="8389D5A4630EFFDFFD2DFA3AFCC309E0" authority="Komai, 2018" authorityName="Komai" authorityYear="2018" box="[708,830,1447,1472]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="shiranui" status="sp. nov.">
|
||
<emphasis id="76FD7235630EFFDFFD2DFA3AFCC309E0" box="[708,830,1447,1472]" italics="true" pageId="8" pageNumber="395">T. shiranui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6DCECF4E630EFFDFFCAEFA34FC7A09E0" box="[839,903,1447,1472]" pageId="8" pageNumber="395" rank="species">
|
||
<emphasis id="76FD7235630EFFDFFCAEFA34FC7A09E0" bold="true" box="[839,903,1447,1472]" pageId="8" pageNumber="395">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, the claw is exceeded by the rounded terminal margin of the dactylus (
|
||
<figureCitation id="DCB2B2A2630EFFDFFE49FA5FFE0709C4" box="[416,506,1484,1509]" captionStart="FIGURE3" captionStartId="4.[151,250,1944,1966]" captionTargetBox="[175,1395,634,1896]" captionTargetId="figure@4.[151,1435,589,1923]" captionTargetPageId="4" captionText="FIGURE3. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, left cheliped, lateral view; B, same, mesial view; C, same, chela, dorsal view; D, same, chela, ventral view (setae omitted); E, same, fingers, dorsal view; F, same, fingers, ventral view (setae omitted)." httpUri="https://zenodo.org/record/1170047/files/figure.png" pageId="8" pageNumber="395">Figs 3E</figureCitation>
|
||
,
|
||
<figureCitation id="DCB2B2A2630EFFDFFDEFFA5FFDB109C4" box="[518,588,1484,1508]" captionStart="FIGURE 4" captionStartId="5.[151,250,1405,1427]" captionTargetBox="[175,1403,208,1357]" captionTargetId="figure@5.[151,1435,185,1394]" captionTargetPageId="5" captionText="FIGURE 4. Thuylamea shiranui n. sp., holotype, female (cl 5.1 mm), CBM-ZC 14145. A, right cheliped, lateral view; B, same, mesial view; C, same, chela, dorsal view; D, same, chela, ventral view (setae omitted)." httpUri="https://zenodo.org/record/1170049/files/figure.png" pageId="8" pageNumber="395">4C, D</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="8389D5A4630EFFDFFD36FA5EFCAA09C5" authorityName="Nguyen" authorityYear="2001" box="[735,855,1484,1509]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="camelus">
|
||
<emphasis id="76FD7235630EFFDFFD36FA5EFCAA09C5" box="[735,855,1484,1509]" italics="true" pageId="8" pageNumber="395">T. camelus</emphasis>
|
||
</taxonomicName>
|
||
, the unguis is terminal in the position (
|
||
<bibRefCitation id="2018D3D6630EFFDFFAE5FA5FFECF0A28" pageId="8" pageNumber="395" refId="ref4207" refString="Wang, Y. & Sha, Z. (2016) Redescription of the type species of the genus Thuylamea Nguyen, 2001 (Decapoda, Caridea, Alpheidae), based on a female specimen. Crustaceana, 89, 1185 - 1191. https: // doi. org / 10.1163 / 15685403 - 00003572" type="journal article">Wang & Sha 2016: fig. 1F</bibRefCitation>
|
||
). In the small male specimen from
|
||
<collectingCountry id="3C9EEEB7630EFFDFFD3CFA7CFCB70A28" box="[725,842,1519,1544]" name="Singapore" pageId="8" pageNumber="395">Singapore</collectingCountry>
|
||
, the claw of the cheliped dactylus is also terminal, although in the adult males, the claw is exceeded by the terminal margin of the dactylus (
|
||
<bibRefCitation id="2018D3D6630EFFDFFB9BF987FAA10A0C" box="[1138,1372,1556,1581]" pageId="8" pageNumber="395" refId="ref4172" refString="Nguyen, V. X. (2001) A new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from South Vietnam. Zoologische Mededelingen, Leiden, 75, 217 - 228." type="book chapter">Nguyên 2001: fig 13</bibRefCitation>
|
||
). The observation suggests that the shape of the dactylus is ontogenetically variable in males of
|
||
<taxonomicName id="8389D5A4630EFFDFFB25F9AAFAB40A70" authorityName="Nguyen" authorityYear="2001" box="[1228,1353,1591,1616]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="camelus">
|
||
<emphasis id="76FD7235630EFFDFFB25F9AAFAB40A70" box="[1228,1353,1591,1616]" italics="true" pageId="8" pageNumber="395">T. camelus</emphasis>
|
||
</taxonomicName>
|
||
and is sexually dimorphic.
|
||
</paragraph>
|
||
<paragraph id="4436AE27630EFFDFFF2EF9ECFCEF0A9D" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="395">
|
||
With the addition of the new species and the observed intraspecific variation in
|
||
<taxonomicName id="8389D5A4630EFFDFFB62F912FAF50AB8" authorityName="Nguyen" authorityYear="2001" box="[1163,1288,1663,1688]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="camelus">
|
||
<emphasis id="76FD7235630EFFDFFB62F912FAF50AB8" box="[1163,1288,1663,1688]" italics="true" pageId="8" pageNumber="395">T. camelus</emphasis>
|
||
</taxonomicName>
|
||
, the generic diagnosis of
|
||
<taxonomicName id="8389D5A4630EFFDFFECFF937FE5E0A9D" authorityName="Nguyen" authorityYear="2001" box="[294,419,1700,1725]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="76FD7235630EFFDFFECFF937FE5E0A9D" box="[294,419,1700,1725]" italics="true" pageId="8" pageNumber="395">Thuylamea</emphasis>
|
||
</taxonomicName>
|
||
(see above) is slightly emended.
|
||
</paragraph>
|
||
<paragraph id="4436AE27630EFFDFFF2EF95BFAB90BB4" blockId="8.[151,1437,151,1941]" pageId="8" pageNumber="395">
|
||
<bibRefCitation id="2018D3D6630EFFDFFF2EF95BFE890AC0" box="[199,372,1736,1760]" pageId="8" pageNumber="395" refId="ref4172" refString="Nguyen, V. X. (2001) A new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from South Vietnam. Zoologische Mededelingen, Leiden, 75, 217 - 228." type="book chapter">Nguyên (2001)</bibRefCitation>
|
||
emphasized the uniqueness of
|
||
<taxonomicName id="8389D5A4630EFFDFFD0DF954FC9D0AC0" authorityName="Nguyen" authorityYear="2001" box="[740,864,1735,1760]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="76FD7235630EFFDFFD0DF954FC9D0AC0" box="[740,864,1735,1760]" italics="true" pageId="8" pageNumber="395">Thuylamea</emphasis>
|
||
</taxonomicName>
|
||
, not specifying closely related taxa.
|
||
<bibRefCitation id="2018D3D6630EFFDFFAF9F954FF190B25" pageId="8" pageNumber="395" refId="ref3959" refString="Anker, A., Ahyong, S. T., Noel, P. Y. & Palmer, A. R. (2006) Morphological phylogeny of alpheid shrimps: parallel preadaptaion and the origin of a key morphological innovation, the snapping claw. Evolution, 60, 2507 - 2528. https: // doi. org / 10.1111 / j. 0014 - 3820.2006. tb 01886. x" type="journal article">
|
||
Anker
|
||
<emphasis id="76FD7235630EFFDFFA89F95AFA680AC0" box="[1376,1429,1735,1760]" italics="true" pageId="8" pageNumber="395">et al</emphasis>
|
||
. (2006)
|
||
</bibRefCitation>
|
||
, who performed phylogenetic analysis of
|
||
<taxonomicName id="8389D5A4630EFFDFFD3DF97FFCBA0B25" authorityName="Rafinesque" authorityYear="1815" box="[724,839,1772,1797]" class="Malacostraca" family="Alpheidae" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="family">Alpheidae</taxonomicName>
|
||
using morphological characters, estimated that the sister taxon of
|
||
<taxonomicName id="8389D5A4630EFFDFFED2F89CFDE40B08" authorityName="Nguyen" authorityYear="2001" box="[315,537,1807,1832]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="camelus">
|
||
<emphasis id="76FD7235630EFFDFFED2F89CFDE40B08" box="[315,537,1807,1832]" italics="true" pageId="8" pageNumber="395">Thuylamea camelus</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<taxonomicName id="8389D5A4630EFFDFFDD0F89CFBBF0B08" authority="Mendez & Wicksten, 1982" authorityName="Mendez & Wicksten" authorityYear="1982" box="[569,1090,1807,1832]" class="Malacostraca" family="Alpheidae" genus="Notalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="imarpe">
|
||
<emphasis id="76FD7235630EFFDFFDD0F89CFCF30B08" box="[569,782,1807,1832]" italics="true" pageId="8" pageNumber="395">Notalpheus imarpe</emphasis>
|
||
<bibRefCitation id="2018D3D6630EFFDFFCFDF89CFBBF0B08" box="[788,1090,1807,1832]" pageId="8" pageNumber="395" refId="ref4121" refString="Mendez, G. M. & Wicksten, M. K. (1982) Notalpheus imarpe: a new genus and species of snapping shrimp from western South America (Decapoda: Alpheidae). Proceedings of the Biological Society of Washington, 95, 709 - 713." type="journal article">Méndez & Wicksten, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
, a unique member of the genus
|
||
<taxonomicName id="8389D5A4630EFFDFFF7EF8A7FDB50B6C" authority="Mendez & Wicksten, 1982" authorityName="Mendez & Wicksten" authorityYear="1982" box="[151,584,1844,1869]" class="Malacostraca" family="Alpheidae" genus="Notalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="76FD7235630EFFDFFF7EF8A7FEEA0B6D" box="[151,279,1844,1869]" italics="true" pageId="8" pageNumber="395">Notalpheus</emphasis>
|
||
<bibRefCitation id="2018D3D6630EFFDFFEF4F8A7FDB50B6C" box="[285,584,1844,1869]" pageId="8" pageNumber="395" refId="ref4121" refString="Mendez, G. M. & Wicksten, M. K. (1982) Notalpheus imarpe: a new genus and species of snapping shrimp from western South America (Decapoda: Alpheidae). Proceedings of the Biological Society of Washington, 95, 709 - 713." type="journal article">Méndez & Wicksten, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
and that the next closely related taxon is
|
||
<taxonomicName id="8389D5A4630EFFDFFBFCF8A7FECD0B50" authority="De Grave & Anker, 2000" authorityName="De Grave & Anker" authorityYear="2000" class="Malacostraca" family="Alpheidae" genus="Orygmalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="species" species="politus">
|
||
<emphasis id="76FD7235630EFFDFFBFCF8A7FAF00B6D" box="[1045,1293,1844,1869]" italics="true" pageId="8" pageNumber="395">Orygmalpheus politus</emphasis>
|
||
<bibRefCitation id="2018D3D6630EFFDFFAFDF8A6FECD0B50" pageId="8" pageNumber="395" refId="ref4077" refString="De Grave, S. & Anker, A. (2000) A new genus and new species of alpheid shrimp (Crustacea: Decapoda) from Papua New Guinea. The Raffles Bulletin of Zoology, 48, 249 - 256." type="journal article">De Grave & Anker, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="9B321085630EFFDFFE92F8CBFE560B50" box="[379,427,1880,1904]" pageId="8" pageNumber="395">type</typeStatus>
|
||
species of the monotypic
|
||
<taxonomicName id="8389D5A4630EFFDFFCE1F8C4FC500B50" authorityName="De Grave & Anker" authorityYear="2000" box="[776,941,1879,1904]" class="Malacostraca" family="Alpheidae" genus="Orygmalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="76FD7235630EFFDFFCE1F8C4FC500B50" box="[776,941,1879,1904]" italics="true" pageId="8" pageNumber="395">Orygmalpheus</emphasis>
|
||
</taxonomicName>
|
||
. However, these two allied genera are morphologically quite different from
|
||
<taxonomicName id="8389D5A4630EFFDFFDD3F8EFFD4A0BB5" authorityName="Nguyen" authorityYear="2001" box="[570,695,1916,1941]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="395" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="76FD7235630EFFDFFDD3F8EFFD4A0BB5" box="[570,695,1916,1941]" italics="true" pageId="8" pageNumber="395">Thuylamea</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="2018D3D6630EFFDFFD03F8EFFBEE0BB4" box="[746,1043,1916,1941]" pageId="8" pageNumber="395" refId="ref4121" refString="Mendez, G. M. & Wicksten, M. K. (1982) Notalpheus imarpe: a new genus and species of snapping shrimp from western South America (Decapoda: Alpheidae). Proceedings of the Biological Society of Washington, 95, 709 - 713." type="journal article">Méndez & Wicksten 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="2018D3D6630EFFDFFBF6F8EEFAC40BB4" box="[1055,1337,1916,1941]" pageId="8" pageNumber="395" refId="ref4077" refString="De Grave, S. & Anker, A. (2000) A new genus and new species of alpheid shrimp (Crustacea: Decapoda) from Papua New Guinea. The Raffles Bulletin of Zoology, 48, 249 - 256." type="journal article">De Grave & Anker 2000</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |