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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.745.23685" ID-PMC="PMC5904534" ID-Pensoft-Pub="1313-2970-745-101" ID-Pensoft-UUID="047DFFCAFFA5F32EA97C873F4708943F" ID-PubMed="29670449" ID-Zenodo-Dep="1222435" ID-ZooBank="8785DC6BC2A244FD94B6243EB07C717F" ModsDocID="1313-2970-745-101" checkinTime="1521799180220" checkinUser="pensoft" docAuthor="Moore, Matthew R., Cave, Ronald D. &amp; Branham, Marc A." docDate="2018" docId="CC1B92F520A915BF1613549A1EE42F59" docLanguage="en" docName="ZooKeys 745: 101-378" docOrigin="ZooKeys 745" docPubDate="2018-03-22" docSource="http://dx.doi.org/10.3897/zookeys.745.23685" docTitle="Cyclocephala melane Bates 1888" docType="treatment" docVersion="7" id="047DFFCAFFA5F32EA97C873F4708943F" lastPageNumber="190" masterDocId="047DFFCAFFA5F32EA97C873F4708943F" masterDocTitle="Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini)" masterLastPageNumber="378" masterPageNumber="101" pageNumber="190" updateTime="1740595812083" updateUser="felipe">
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<mods:title id="A6BB03D7598E25562741600444C359ED">Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini)</mods:title>
</mods:titleInfo>
<mods:name id="3061600C6BBBFDDA2AF01F6F0BC5C22E" type="personal">
<mods:role id="289793828FA6712634A3F63350C13593">
<mods:roleTerm id="DCA2ACC4CCA6DC8B6D0C10156A2846B7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="014837B72CFD842E5A317E1762246DF5">Moore, Matthew R.</mods:namePart>
<mods:affiliation id="859494E8C7B5963AC368438E70DB3D22">Department of Entomology and Nematology, University of Florida, Building 1881 Natural Area Drive, Steinmetz Hall, Gainesville, FL 32611, USA</mods:affiliation>
<mods:nameIdentifier id="FE34080F78767F8349E3EC52DAF8173C" type="email">cyclocephala@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="9325B72DC032EEBB500DD99260C82A69" type="personal">
<mods:role id="247FDDE9267D97BF0941E10CDB8622DA">
<mods:roleTerm id="4EAB8B04B8E4C49E72ADD0ADFFB88D56">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="11545812F2B20651855CE6D87CA9288D">Cave, Ronald D.</mods:namePart>
<mods:affiliation id="1FC9ADD26CDFE36128CC55D7DA6D7483">Department of Entomology and Nematology, University of Florida, Indian River Research and Education Center, 2199 South Rock Road, Fort Pierce, FL 34945, USA</mods:affiliation>
</mods:name>
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<mods:roleTerm id="139B6E32F21C396C425675E311B85957">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E609872BF6FC6C1BAA51AFD708283F74">Branham, Marc A.</mods:namePart>
<mods:affiliation id="5EC299205B8DC326646A136A7CC66DEF">Department of Entomology and Nematology, University of Florida, Building 1881 Natural Area Drive, Steinmetz Hall, Gainesville, FL 32611, USA</mods:affiliation>
</mods:name>
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<mods:title id="464437FD071DE51191245CC00076AF1A">ZooKeys</mods:title>
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<mods:part id="A1233BC332D1A95596FD27ED0A3BAE92">
<mods:date id="2CDB35E577DF5B783B1622EDA75A34FA">2018</mods:date>
<mods:detail id="2A73B66AC934BDD2F8B64363B28FBDE7" type="pubDate">
<mods:number id="B0AD9DA26432084715D9E7D6067DC1BC">2018-03-22</mods:number>
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<mods:number id="7ACBE5720444BF387F75389CEC9744C8">745</mods:number>
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<mods:classification id="7B6CCE7A43B6052D95A46DCF8E8E02A5">journal article</mods:classification>
<mods:identifier id="9CDD2D08E2563D732F13DDB1B5592932" type="DOI">http://dx.doi.org/10.3897/zookeys.745.23685</mods:identifier>
<mods:identifier id="952EA4A841E72DC677182F5AE8232789" type="Pensoft-Pub">1313-2970-745-101</mods:identifier>
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<treatment id="CC1B92F520A915BF1613549A1EE42F59" ID-GBIF-Taxon="142480031" LSID="urn:lsid:plazi:treatment:CC1B92F520A915BF1613549A1EE42F59" httpUri="http://treatment.plazi.org/id/CC1B92F520A915BF1613549A1EE42F59" lastPageNumber="190" pageId="89" pageNumber="190">
<subSubSection id="1A229F1F24DEEB44FF0FF4A942671A7B" pageId="89" pageNumber="190" type="nomenclature">
<paragraph id="41C8D1DFF92EEB0454C904BE46F92F4D" pageId="89" pageNumber="190">
<taxonomicName id="3BA6DF71B96A6067429F843F29D2CAC1" ID-CoL="32TXG" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="melane">Cyclocephala melane Bates, 1888</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="A659C5A0D010C6F8193AD4A8C207695F" pageId="89" pageNumber="190" type="reference_group">
<paragraph id="4AB4078B7DB51BDFAF32170D73035679" pageId="89" pageNumber="190">
<taxonomicName id="24E35FE9279AD256FFEAB3DD7AA65BC3" ID-CoL="32TXG" authority="Bates, 1888: 310" authorityName="Bates" authorityPageNumber="310" authorityYear="1888" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="melane">Cyclocephala melane Bates, 1888: 310</taxonomicName>
[original combination].
</paragraph>
</subSubSection>
<subSubSection id="2B9E2CBB824BFCD1834E1B464B72768B" pageId="89" pageNumber="190" type="types">
<paragraph id="E04DB43FF7FD6B63F6E66AE105374AD1" pageId="89" pageNumber="190">Types.</paragraph>
<paragraph id="0C1983177A71C1AD231C110AD391DB77" pageId="89" pageNumber="190">
Type at BMNH (
<bibRefCitation id="5982A12B62AA9BF36238C6B1609B2090" author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
<normalizedToken id="F620F52BC548462C3F1CDB4A60E46EB1" originalValue="Endrődi">Endrodi</normalizedToken>
1966
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7352CF186E93859C41D61A54187A89C2" pageId="89" pageNumber="190" type="distribution">
<paragraph id="7CAAF08B2190041ACDC92AA352FCD2BD" pageId="89" pageNumber="190">Distribution.</paragraph>
<paragraph id="B23B2193D58AA3976EF24D1827806343" pageId="89" pageNumber="190">
COSTA RICA: Cartago, Heredia, Puntarenas, San
<normalizedToken id="B5CB57E654F03A259D16E00BC90386B8" originalValue="José">Jose</normalizedToken>
. PANAMA:
<normalizedToken id="EB050B46F53A7D721867B468E2AFCD39" originalValue="Chiriquí">Chiriqui</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="465FEBBFB289D05A98FA57FC3F2B9943" pageId="89" pageNumber="190" type="references">
<paragraph id="4E061F11C11508E349E10A729BF4F0BD" pageId="89" pageNumber="190">References.</paragraph>
<paragraph id="A0A789FFD1E49E37EEC4F87DF503FA6A" pageId="89" pageNumber="190">
<bibRefCitation id="E1A22B23038CC8D599737426FB5E7766" author="Bates, HW" pageId="164" pageNumber="265" refId="B65" refString="Bates, HW, 1888. Pectinicornia and Lamellicornia, Family Dynastidae. In: Salvin O, Godman FD (Eds) Biologia Centrali-Americana. Insecta, Coleoptera, volume 2, part 2. RH Porter, London, 296-342." title="Pectinicornia and Lamellicornia, Family Dynastidae. In: Salvin O, Godman FD (Eds) Biologia Centrali-Americana. Insecta, Coleoptera, volume 2, part 2. RH Porter, London, 296 - 342." year="1888">Bates 1888</bibRefCitation>
,
<bibRefCitation id="85D27E59454D524B9294283393FCBD4A" author="Arrow, GJ" pageId="164" pageNumber="265" refId="B51" refString="Arrow, GJ, 1937b. Pars 156: Scarabaeidae: Dynastinae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Volumen XXI. Scarabaeidae III. W Junk, Berlin, 1-124." title="Pars 156: Scarabaeidae: Dynastinae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Volumen XXI. Scarabaeidae III. W Junk, Berlin, 1 - 124." year="1937 b">Arrow 1937b</bibRefCitation>
,
<bibRefCitation id="8C3136535427746F3C7B7C0ED69DBB82" author="Blackwelder, RE" pageId="167" pageNumber="268" refId="B101" refString="Blackwelder, RE, 1944. Checklist of coleopterous insects of Mexico, Central America, the West Indies, and South America, part 2. Bulletin of the United States National Museum 185: 189-341." title="Checklist of coleopterous insects of Mexico, Central America, the West Indies, and South America, part 2. Bulletin of the United States National Museum 185: 189 - 341." year="1944">Blackwelder 1944</bibRefCitation>
,
<bibRefCitation id="CCD00AF055C4D8A9EAF763CB6B1797C8" author="Pike, KS" pageId="213" pageNumber="314" refId="B899" refString="Pike, KS, River, RL, Ratcliffe, BC, Oseto, CY, Mayo, ZB, 1976. A world bibliography of the genus Cyclocephala (Coleoptera: Scarabaeidae). Miscellaneous Publication of the University of Nebraska Agricultural Experiment Station 32: 1-32." title="A world bibliography of the genus Cyclocephala (Coleoptera: Scarabaeidae). Miscellaneous Publication of the University of Nebraska Agricultural Experiment Station 32: 1 - 32." year="1976">Pike et al. 1976</bibRefCitation>
,
<bibRefCitation id="358943DF62F5D2D4CCB6F282B0E6DD0C" author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
<normalizedToken id="D21CBFA13B29DDF6270CF943596ADF51" originalValue="Endrődi">Endrodi</normalizedToken>
1966
</bibRefCitation>
,
<bibRefCitation id="081C4E5F09C70C9F4303E04CF736A534" author="Endrodi, S" pageId="178" pageNumber="279" refId="B321" refString="Endrodi, S, 1985a. The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" title="The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" year="1985 a">1985a</bibRefCitation>
,
<bibRefCitation id="31BAFC150D71D7880CFB38B0EDB2FB7E" DOI="https://doi.org/10.2307/2992022" author="Croat, TB" pageId="171" pageNumber="272" refId="B208" refString="Croat, TB, 1997. A revision of Philodendron subgenus Philodendron (Araceae) for Mexico and Central America. Annals of the Missouri Botanical Garden 84: 311-704. https://doi.org/10.2307/2992022" title="A revision of Philodendron subgenus Philodendron (Araceae) for Mexico and Central America. Annals of the Missouri Botanical Garden 84: 311 - 704." url="https://doi.org/10.2307/2992022" year="1997">Croat 1997</bibRefCitation>
,
<bibRefCitation id="20180059709744BC7B86914960D6F272" author="Ratcliffe, BC" pageId="217" pageNumber="318" refId="B968" refString="Ratcliffe, BC, 1992a. Nine new species and 11 country records of Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae) from Panama and Costa Rica. The Coleopterists Bulletin 46: 216-235." title="Nine new species and 11 country records of Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae) from Panama and Costa Rica. The Coleopterists Bulletin 46: 216 - 235." year="1992 a">Ratcliffe 1992a</bibRefCitation>
,
<bibRefCitation id="0C92E06E24B8F38D7BA87BEA46848DA2" DOI="https://doi.org/10.11646/zootaxa.32.1.1" author="Ratcliffe, BC" pageId="218" pageNumber="319" refId="B972" refString="Ratcliffe, BC, 2002a. A checklist of the Scarabaeoidea (Coleoptera) of Panama. Zootaxa 32: 1-48. https://doi.org/10.11646/zootaxa.32.1.1" title="A checklist of the Scarabaeoidea (Coleoptera) of Panama. Zootaxa 32: 1 - 48." url="https://doi.org/10.11646/zootaxa.32.1.1" year="2002 a">2002a</bibRefCitation>
,
<bibRefCitation id="91FED27F8FA5403BA4F8628F1A27ACB9" author="Ratcliffe, BC" pageId="218" pageNumber="319" refId="B974" refString="Ratcliffe, BC, 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1-506." title="The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." year="2003">2003</bibRefCitation>
,
<bibRefCitation id="862DF65E766D1E4EA2579EE7899A9A7E" author="Krajcik, M" pageId="200" pageNumber="301" refId="B658" refString="Krajcik, M, 2005. Dynastinae of the World Checklist (Coleoptera: Scarabaeidae: Dynastinae). Animma.X, Supplement No. 2, Pilsen, 122 pp." title="Dynastinae of the World Checklist (Coleoptera: Scarabaeidae: Dynastinae). Animma. X, Supplement No. 2, Pilsen, 122 pp." year="2005">Krajcik 2005</bibRefCitation>
,
<bibRefCitation id="7C49EA60C32072B43B306A397E95BFE8" author="Krajcik, M" pageId="200" pageNumber="301" refId="B659" refString="Krajcik, M, 2012. Checklist of the world Scarabaeoidea. Animma.X Supplement 5, Pilsen, 278 pp." title="Checklist of the world Scarabaeoidea. Animma. X Supplement 5, Pilsen, 278 pp." year="2012">2012</bibRefCitation>
,
<bibRefCitation id="097EFDCD09393360FBD835F7A27D4051" DOI="https://doi.org/10.1673/031.013.10001" author="Moore, MR" pageId="208" pageNumber="309" refId="B796" refString="Moore, MR, Jameson, ML, 2013. Floral associations of cyclocephaline scarab beetles. Journal of Insect Science 13 (100): 1-43. https://doi.org/10.1673/031.013.10001" title="Floral associations of cyclocephaline scarab beetles. Journal of Insect Science 13 (100): 1 - 43." url="https://doi.org/10.1673/031.013.10001" year="2013">Moore and Jameson 2013</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E1A829BAA2AEEB29FBE1184494A6F16F" pageId="89" pageNumber="190" type="remarks">
<paragraph id="25EB8853BDED8217896E3B312D7F36A6" pageId="89" pageNumber="190">Remarks.</paragraph>
<paragraph id="CB7739E3BB3F536690DAA8905344C4CA" pageId="89" pageNumber="190">
The original spelling of this specific epithet is
<taxonomicName id="282F663F5CA705988BC9A2BA14505DBB" authority="(Bates 1888)" authorityName="Bates" authorityYear="1888" baseAuthorityName="Bates" baseAuthorityYear="1888" class="Insecta" family="Scarabaeidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="melane">
&quot;
<emphasis id="D4DD861BC654ABEE704E5F89170D3878" italics="true" pageId="89" pageNumber="190">melane</emphasis>
</taxonomicName>
&quot; (
<bibRefCitation id="34B18F6DAF0BC4B187045C580B6EB68D" author="Bates, HW" pageId="164" pageNumber="265" refId="B65" refString="Bates, HW, 1888. Pectinicornia and Lamellicornia, Family Dynastidae. In: Salvin O, Godman FD (Eds) Biologia Centrali-Americana. Insecta, Coleoptera, volume 2, part 2. RH Porter, London, 296-342." title="Pectinicornia and Lamellicornia, Family Dynastidae. In: Salvin O, Godman FD (Eds) Biologia Centrali-Americana. Insecta, Coleoptera, volume 2, part 2. RH Porter, London, 296 - 342." year="1888">Bates 1888</bibRefCitation>
). Subsequently, some authors have spelled the name as &quot;
<emphasis id="945F1D80A2992EC538631DE707F7836D" italics="true" pageId="89" pageNumber="190">melanae</emphasis>
&quot; without explanation (
<bibRefCitation id="E674DB008CB36ED44B2B401ED7489B10" author="Endrodi, S" pageId="177" pageNumber="278" refId="B301" refString="Endrodi, S, 1966. Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1-460." title="Monographie der Dynastinae (Coleoptera, Lamellicornia). I. Teil. Entomologische Abhandlungen des Staatlichen Museums fuer Tierkunde Dresden 33: 1 - 460." year="1966">
<normalizedToken id="861C543231476CD204A48AF2EED082DA" originalValue="Endrődi">Endrodi</normalizedToken>
1966
</bibRefCitation>
,
<bibRefCitation id="9FBEE261965F37F1855AF52A9F8CD924" author="Endrodi, S" pageId="178" pageNumber="279" refId="B321" refString="Endrodi, S, 1985a. The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" title="The Dynastinae of the World. Dr. W. Junk Publisher, Dordrecht, 800 pp. [+ 46 pls]" year="1985 a">1985a</bibRefCitation>
,
<bibRefCitation id="AF019C5C6E844E76258A830A2F8B2D9F" author="Ratcliffe, BC" pageId="217" pageNumber="318" refId="B968" refString="Ratcliffe, BC, 1992a. Nine new species and 11 country records of Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae) from Panama and Costa Rica. The Coleopterists Bulletin 46: 216-235." title="Nine new species and 11 country records of Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae) from Panama and Costa Rica. The Coleopterists Bulletin 46: 216 - 235." year="1992 a">Ratcliffe 1992a</bibRefCitation>
,
<bibRefCitation id="CF2BC221C6729D2262EA6E23BCA5AF97" DOI="https://doi.org/10.11646/zootaxa.32.1.1" author="Ratcliffe, BC" pageId="218" pageNumber="319" refId="B972" refString="Ratcliffe, BC, 2002a. A checklist of the Scarabaeoidea (Coleoptera) of Panama. Zootaxa 32: 1-48. https://doi.org/10.11646/zootaxa.32.1.1" title="A checklist of the Scarabaeoidea (Coleoptera) of Panama. Zootaxa 32: 1 - 48." url="https://doi.org/10.11646/zootaxa.32.1.1" year="2002 a">2002a</bibRefCitation>
,
<bibRefCitation id="9C40CCA0B2E06FC76D15A4BB08661D49" author="Ratcliffe, BC" pageId="218" pageNumber="319" refId="B974" refString="Ratcliffe, BC, 2003. The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1-506." title="The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16: 1 - 506." year="2003">2003</bibRefCitation>
,
<bibRefCitation id="E76439607028181A7D67D281568E7D8F" DOI="https://doi.org/10.2307/2992022" author="Croat, TB" pageId="171" pageNumber="272" refId="B208" refString="Croat, TB, 1997. A revision of Philodendron subgenus Philodendron (Araceae) for Mexico and Central America. Annals of the Missouri Botanical Garden 84: 311-704. https://doi.org/10.2307/2992022" title="A revision of Philodendron subgenus Philodendron (Araceae) for Mexico and Central America. Annals of the Missouri Botanical Garden 84: 311 - 704." url="https://doi.org/10.2307/2992022" year="1997">Croat 1997</bibRefCitation>
,
<bibRefCitation id="22CAD4B75EC67A8B6B93FE9416B3243F" DOI="https://doi.org/10.1673/031.013.10001" author="Moore, MR" pageId="208" pageNumber="309" refId="B796" refString="Moore, MR, Jameson, ML, 2013. Floral associations of cyclocephaline scarab beetles. Journal of Insect Science 13 (100): 1-43. https://doi.org/10.1673/031.013.10001" title="Floral associations of cyclocephaline scarab beetles. Journal of Insect Science 13 (100): 1 - 43." url="https://doi.org/10.1673/031.013.10001" year="2013">Moore and Jameson 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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