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<document id="50581ADEAB6F5E995F042E8411948F76" ID-DOI="10.1080/00222930500393368" ID-ISSN="1464-5262" ID-Zenodo-Dep="5220802" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629364456045" checkinUser="felipe" docAuthor="Adams, Richard J., Price, Roger D. &amp; Clayton, Dale H." docDate="2005" docId="CB3C8797C31E8700FE381D9FFC68FE23" docLanguage="en" docName="JNatHist.39.41.3545-3618.pdf" docOrigin="Journal of Natural History 39 (41)" docSource="http://dx.doi.org/10.1080/00222930500393368" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Columbicola fortis" docType="treatment" docVersion="3" lastPageNumber="3580" masterDocId="3705FFEFC33A8725FFF11922FFA3FF97" masterDocTitle="Taxonomic revision of Old World members of the feather louse genus Columbicola (Phthiraptera: Ischnocera), including descriptions of eight new species" masterLastPageNumber="3618" masterPageNumber="3545" pageNumber="3579" updateTime="1699069527252" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="856A571EAD23B269FF8759FD84032088">Taxonomic revision of Old World members of the feather louse genus Columbicola (Phthiraptera: Ischnocera), including descriptions of eight new species</mods:title>
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<mods:namePart id="58A20B066230B2B14E9517B54D189F7B">Adams, Richard J.</mods:namePart>
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<mods:namePart id="49923AF2A2F09B4727E4582E5CC40BAE">Price, Roger D.</mods:namePart>
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<mods:namePart id="6971B7C646D3B4EA7037C3FABEC91E4E">Clayton, Dale H.</mods:namePart>
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<mods:title id="B02703BE282E03E8C91626B71039E4B6">Journal of Natural History</mods:title>
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<mods:date id="B3004CE6585020F27B8D2F806051951E">2005</mods:date>
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<mods:number id="81968EF3B39A2D75E0B7B1AE6A6968E2">2005-12-23</mods:number>
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<mods:number id="FF2485DB80CFC2451996C8B79795EE6A">39</mods:number>
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<treatment id="CB3C8797C31E8700FE381D9FFC68FE23" LSID="urn:lsid:plazi:treatment:CB3C8797C31E8700FE381D9FFC68FE23" httpUri="http://treatment.plazi.org/id/CB3C8797C31E8700FE381D9FFC68FE23" lastPageId="37" lastPageNumber="3580" pageId="36" pageNumber="3579">
<subSubSection id="0B8F650AC31E8701FE381D9FFCCDFB42" box="[457,878,1213,1237]" pageId="36" pageNumber="3579" type="nomenclature">
<paragraph id="432A3681C31E8701FE381D9FFCCDFB42" blockId="36.[165,1046,1213,1301]" box="[457,878,1213,1237]" pageId="36" pageNumber="3579">
<heading id="186281EDC31E8701FE381D9FFCCDFB42" box="[457,878,1213,1237]" centered="true" fontSize="10" level="2" pageId="36" pageNumber="3579" reason="2">
<taxonomicName id="84954D02C31E8701FE381D9FFCCDFB42" ID-CoL="5ZPZL" authority="(Taschenberg)" baseAuthorityName="Taschenberg" baseAuthorityYear="1882" box="[457,878,1213,1237]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="36" pageNumber="3579" phylum="Arthropoda" rank="species" species="fortis">
<emphasis id="71E1EA93C31E8701FE381D9FFD19FB42" bold="true" box="[457,698,1213,1237]" italics="true" pageId="36" pageNumber="3579">Columbicola fortis</emphasis>
(Taschenberg)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0B8F650AC31E8700FDC21DFFFC68FE23" lastPageId="37" lastPageNumber="3580" pageId="36" pageNumber="3579" type="description">
<paragraph id="432A3681C31E8701FDC21DFFFBB5FA82" blockId="36.[165,1046,1213,1301]" pageId="36" pageNumber="3579">
(
<figureCitation id="DBAE2A04C31E8701FDCA1DFFFCA3FB62" box="[571,768,1245,1269]" captionStart="Figures 95111" captionStartId="39.[125,194,1529,1548]" captionTargetBox="[142,1153,133,1491]" captionTargetId="figure-0@39.[142,1155,133,1493]" captionTargetPageId="39" captionText="Figures 95111. Columbicola effeminatus: (95) female dorsal head; (96) male genitalia; (97) female ventral terminalia. C. becheti: (98) male dorsal head; (99) male genitalia; (100) female dorsal head; (101) female ventral terminalia. C. fortis: (102) male dorsal head; (103) female dorsal head; (104) male genitalia; (105) female ventral terminalia. C. tasmaniensis: (106) male genitalia; (107) female ventral terminalia. C. mjoebergi: (108) male dorsal head; (109) female dorsal head; (110) male genitalia; (111) female ventral terminalia." figureDoi="http://doi.org/10.5281/zenodo.5220814" httpUri="https://zenodo.org/record/5220814/files/figure.png" pageId="36" pageNumber="3579">Figures 102105</figureCitation>
)
<taxonomicName id="84954D02C31E8701FF541DDFFDC0FA82" authority="Taschenberg 1882: 126" authorityName="Taschenberg" authorityPageNumber="126" authorityYear="1882" box="[165,611,1277,1301]" class="Insecta" family="Philopteridae" genus="Lipeurus" kingdom="Animalia" order="Psocodea" pageId="36" pageNumber="3579" phylum="Arthropoda" rank="species" species="fortis">
<emphasis id="71E1EA93C31E8701FF541DDFFEE3FA82" box="[165,320,1277,1301]" italics="true" pageId="36" pageNumber="3579">Lipeurus fortis</emphasis>
<bibRefCitation id="27044B70C31E8701FEB81DDFFDC0FA82" author="Taschenberg O" box="[329,611,1277,1301]" pageId="36" pageNumber="3579" pagination="1 - 244" refId="ref39091" refString="Taschenberg O. 1882. Die Mallophagen mit besonderer Berucksichtigung der von Dr. Meyer gesammelten Arten. Nova Acta der Kaiserlich Leopoldinisch-Carolinisch-Deutschen Akademie der Naturforscher 44: 1 - 244." type="journal article" year="1882">Taschenberg 1882: 126</bibRefCitation>
</taxonomicName>
.
<typeStatus id="9C2E8823C31E8701FD9E1DDFFD08FA82" box="[623,683,1277,1301]" pageId="36" pageNumber="3579">Type</typeStatus>
host:
<taxonomicName id="84954D02C31E8701FD071DDFFBB5FA82" authority="Gould." authorityName="Gould." box="[758,1046,1277,1301]" class="Aves" family="Columbidae" genus="Otidiphaps" kingdom="Animalia" order="Columbiformes" pageId="36" pageNumber="3579" phylum="Chordata" rank="species" species="nobilis">
<emphasis id="71E1EA93C31E8701FD071DDFFC19FA82" box="[758,954,1277,1301]" italics="true" pageId="36" pageNumber="3579">Otidiphaps nobilis</emphasis>
Gould.
</taxonomicName>
</paragraph>
<paragraph id="432A3681C31E8701FF7B1C6FFEA0FAF2" blockId="36.[138,259,1357,1381]" box="[138,259,1357,1381]" pageId="36" pageNumber="3579">
<emphasis id="71E1EA93C31E8701FF7B1C6FFEA0FAF2" box="[138,259,1357,1381]" italics="true" pageId="36" pageNumber="3579">Description</emphasis>
</paragraph>
<paragraph id="432A3681C31E8701FF7B1C5CFEB4F902" blockId="36.[138,1185,1406,1685]" pageId="36" pageNumber="3579">
<materialsCitation id="F3FD3CDCC31E8701FF7B1C5CFEB0F902" collectionCode="APW, HW, PW, MW" latitude="0.22" longLatPrecision="78" longitude="-0.245" pageId="36" pageNumber="3579" specimenCount="1">
Male head broadly bellshaped (
<figureCitation id="DBAE2A04C31E8701FDFB1C5CFD2FFA01" box="[522,652,1406,1430]" captionStart="Figures 95111" captionStartId="39.[125,194,1529,1548]" captionTargetBox="[142,1153,133,1491]" captionTargetId="figure-0@39.[142,1155,133,1493]" captionTargetPageId="39" captionText="Figures 95111. Columbicola effeminatus: (95) female dorsal head; (96) male genitalia; (97) female ventral terminalia. C. becheti: (98) male dorsal head; (99) male genitalia; (100) female dorsal head; (101) female ventral terminalia. C. fortis: (102) male dorsal head; (103) female dorsal head; (104) male genitalia; (105) female ventral terminalia. C. tasmaniensis: (106) male genitalia; (107) female ventral terminalia. C. mjoebergi: (108) male dorsal head; (109) female dorsal head; (110) male genitalia; (111) female ventral terminalia." figureDoi="http://doi.org/10.5281/zenodo.5220814" httpUri="https://zenodo.org/record/5220814/files/figure.png" pageId="36" pageNumber="3579">Figure 102</figureCitation>
);
<collectionCode id="2584AE44C31E8701FD551C5CFD45FA01" box="[676,742,1406,1430]" pageId="36" pageNumber="3579">APW</collectionCode>
, 0.216;
<collectionCode id="2584AE44C31E8701FCB61C5CFCDFFA01" box="[839,892,1406,1430]" pageId="36" pageNumber="3579">HW</collectionCode>
, 0.400.42 (0.410); HL, 0.560.58 (0.570); HL/
<collectionCode id="2584AE44C31E8701FE6D1CBCFE6CFA21" box="[412,463,1438,1462]" pageId="36" pageNumber="3579">HW</collectionCode>
, 1.381.40 (1.39); SL,
<geoCoordinate id="26A15046C31E8701FD2A1CBCFCBEFA21" box="[731,797,1438,1462]" degrees="0.170" direction="north" orientation="latitude" pageId="36" pageNumber="3579" precision="55" value="0.17">0.170</geoCoordinate>
<geoCoordinate id="26A15046C31E8701FCEC1CBCFCCDFA21" box="[797,878,1438,1462]" degrees="0.176" direction="west" orientation="longitude" pageId="36" pageNumber="3579" precision="55" value="-0.176">0.176</geoCoordinate>
(0.173). Thorax with
<collectionCode id="2584AE44C31E8701FB9F1CBCFB3EFA21" box="[1134,1181,1438,1462]" country="Thailand" name="Paleontological Collections" pageId="36" pageNumber="3579">PW</collectionCode>
, 0.350.36 (0.357);
<collectionCode id="2584AE44C31E8701FE701C9CFE19FA41" box="[385,442,1470,1494]" country="Netherlands" name="Museum Wasmann" pageId="36" pageNumber="3579">MW</collectionCode>
, 0.420.43 (0.427). Genitalia (
<figureCitation id="DBAE2A04C31E8701FCBA1C9CFC6FFA41" box="[843,972,1470,1494]" captionStart="Figures 95111" captionStartId="39.[125,194,1529,1548]" captionTargetBox="[142,1153,133,1491]" captionTargetId="figure-0@39.[142,1155,133,1493]" captionTargetPageId="39" captionText="Figures 95111. Columbicola effeminatus: (95) female dorsal head; (96) male genitalia; (97) female ventral terminalia. C. becheti: (98) male dorsal head; (99) male genitalia; (100) female dorsal head; (101) female ventral terminalia. C. fortis: (102) male dorsal head; (103) female dorsal head; (104) male genitalia; (105) female ventral terminalia. C. tasmaniensis: (106) male genitalia; (107) female ventral terminalia. C. mjoebergi: (108) male dorsal head; (109) female dorsal head; (110) male genitalia; (111) female ventral terminalia." figureDoi="http://doi.org/10.5281/zenodo.5220814" httpUri="https://zenodo.org/record/5220814/files/figure.png" pageId="36" pageNumber="3579">Figure 104</figureCitation>
) with parameres ribbon-like, wrapping behind mesosome; GW,
<geoCoordinate id="26A15046C31E8701FD2B1CFCFCB8FA61" box="[730,795,1502,1526]" degrees="0.123" direction="north" orientation="latitude" pageId="36" pageNumber="3579" precision="55" value="0.123">0.123</geoCoordinate>
<geoCoordinate id="26A15046C31E8701FCED1CFCFCCEFA61" box="[796,877,1502,1526]" degrees="0.137" direction="west" orientation="longitude" pageId="36" pageNumber="3579" precision="55" value="-0.137">0.137</geoCoordinate>
(0.131). TL, 2.202.25 (2.23). Female similar to male except as follows. Head as in
<figureCitation id="DBAE2A04C31E8701FCBF1CDFFC6EF982" box="[846,973,1533,1557]" captionStart="Figures 95111" captionStartId="39.[125,194,1529,1548]" captionTargetBox="[142,1153,133,1491]" captionTargetId="figure-0@39.[142,1155,133,1493]" captionTargetPageId="39" captionText="Figures 95111. Columbicola effeminatus: (95) female dorsal head; (96) male genitalia; (97) female ventral terminalia. C. becheti: (98) male dorsal head; (99) male genitalia; (100) female dorsal head; (101) female ventral terminalia. C. fortis: (102) male dorsal head; (103) female dorsal head; (104) male genitalia; (105) female ventral terminalia. C. tasmaniensis: (106) male genitalia; (107) female ventral terminalia. C. mjoebergi: (108) male dorsal head; (109) female dorsal head; (110) male genitalia; (111) female ventral terminalia." figureDoi="http://doi.org/10.5281/zenodo.5220814" httpUri="https://zenodo.org/record/5220814/files/figure.png" pageId="36" pageNumber="3579">Figure 103</figureCitation>
; dorsal head setae shorter;
<collectionCode id="2584AE44C31E8701FF1B1F3FFE8FF9A2" box="[234,300,1565,1589]" pageId="36" pageNumber="3579">APW</collectionCode>
,
<geoCoordinate id="26A15046C31E8701FEC61F3FFEDAF9A2" box="[311,377,1565,1589]" degrees="0.220" direction="north" orientation="latitude" pageId="36" pageNumber="3579" precision="55" value="0.22">0.220</geoCoordinate>
<geoCoordinate id="26A15046C31E8701FE8B1F3FFE69F9A2" box="[378,458,1565,1589]" degrees="0.245" direction="west" orientation="longitude" pageId="36" pageNumber="3579" precision="55" value="-0.245">0.245</geoCoordinate>
(0.230);
<collectionCode id="2584AE44C31E8701FDC71F3FFDC8F9A2" box="[566,619,1565,1589]" pageId="36" pageNumber="3579">HW</collectionCode>
, 0.420.45 (0.431); HL, 0.610.65 (0.626); HL/
<collectionCode id="2584AE44C31E8701FF7B1F1FFF1CF9C2" box="[138,191,1597,1621]" pageId="36" pageNumber="3579">HW</collectionCode>
, 1.401.48 (1.45). Thorax with
<collectionCode id="2584AE44C31E8701FDBC1F1FFDDFF9C2" box="[589,636,1597,1621]" country="Thailand" name="Paleontological Collections" pageId="36" pageNumber="3579">PW</collectionCode>
, 0.350.37 (0.360);
<collectionCode id="2584AE44C31E8701FC891F1FFC12F9C2" box="[888,945,1597,1621]" country="Netherlands" name="Museum Wasmann" pageId="36" pageNumber="3579">MW</collectionCode>
, 0.440.47 (0.448). Ventral terminalia as in
<figureCitation id="DBAE2A04C31E8701FE6A1F7FFDB9F9E2" box="[411,538,1629,1653]" captionStart="Figures 95111" captionStartId="39.[125,194,1529,1548]" captionTargetBox="[142,1153,133,1491]" captionTargetId="figure-0@39.[142,1155,133,1493]" captionTargetPageId="39" captionText="Figures 95111. Columbicola effeminatus: (95) female dorsal head; (96) male genitalia; (97) female ventral terminalia. C. becheti: (98) male dorsal head; (99) male genitalia; (100) female dorsal head; (101) female ventral terminalia. C. fortis: (102) male dorsal head; (103) female dorsal head; (104) male genitalia; (105) female ventral terminalia. C. tasmaniensis: (106) male genitalia; (107) female ventral terminalia. C. mjoebergi: (108) male dorsal head; (109) female dorsal head; (110) male genitalia; (111) female ventral terminalia." figureDoi="http://doi.org/10.5281/zenodo.5220814" httpUri="https://zenodo.org/record/5220814/files/figure.png" pageId="36" pageNumber="3579">Figure 105</figureCitation>
; subgenital plate slightly expanded anteriorly. TL, 2.40 2.55 (2.48)
</materialsCitation>
.
</paragraph>
<paragraph id="432A3681C31F8700FF8C19ABFF78FF36" blockId="37.[125,219,137,161]" box="[125,219,137,161]" pageId="37" pageNumber="3580">
<emphasis id="71E1EA93C31F8700FF8C19ABFF78FF36" box="[125,219,137,161]" italics="true" pageId="37" pageNumber="3580">Material</emphasis>
</paragraph>
<paragraph id="432A3681C31F8700FF8C199BFCBEFF46" blockId="37.[125,797,185,209]" box="[125,797,185,209]" pageId="37" pageNumber="3580">
<materialsCitation id="F3FD3CDCC31F8700FF8C199BFD49FF46" box="[125,746,185,209]" country="Papua New Guinea" location="Papua New Guinea" pageId="37" pageNumber="3580" specimenCount="11" specimenCount-female="8" specimenCount-male="3">
<specimenCount id="5593FD08C31F8700FF8C199BFF79FF46" box="[125,218,185,209]" pageId="37" pageNumber="3580" type="male">3 males</specimenCount>
,
<specimenCount id="5593FD08C31F8700FF19199BFEFAFF46" box="[232,345,185,209]" pageId="37" pageNumber="3580" type="female">8 females</specimenCount>
, ex
<taxonomicName id="84954D02C31F8700FE78199BFE50FF46" box="[393,499,185,209]" class="Aves" family="Columbidae" genus="Otidiphaps" kingdom="Animalia" order="Columbiformes" pageId="37" pageNumber="3580" phylum="Chordata" rank="species" species="nobilis">
<emphasis id="71E1EA93C31F8700FE78199BFE50FF46" box="[393,499,185,209]" italics="true" pageId="37" pageNumber="3580">O. nobilis</emphasis>
</taxonomicName>
,
<collectingCountry id="3B827611C31F8700FDF5199BFD49FF46" box="[516,746,185,209]" name="Papua New Guinea" pageId="37" pageNumber="3580">Papua New Guinea</collectingCountry>
</materialsCitation>
(4).
</paragraph>
<paragraph id="432A3681C31F8700FF8C1828FF7FFEB5" blockId="37.[125,220,266,290]" box="[125,220,266,290]" pageId="37" pageNumber="3580">
<emphasis id="71E1EA93C31F8700FF8C1828FF7FFEB5" box="[125,220,266,290]" italics="true" pageId="37" pageNumber="3580">Remarks</emphasis>
</paragraph>
<paragraph id="432A3681C31F8700FF8C1819FC68FE23" blockId="37.[125,1172,315,436]" pageId="37" pageNumber="3580">
<bibRefCitation id="27044B70C31F8700FF8C1819FEE3FEC4" author="Tendeiro J" box="[125,320,315,339]" pageId="37" pageNumber="3580" pagination="1 - 460" refId="ref39197" refString="Tendeiro J. 1965. Estudos sobre Malofagos. Revisao monografica do genero Columbicola Ewing (Ischnocera, Philopteridae). Memorias da Junta de Investigacoes do Ultramar, No. 32 (1962). p 1 - 460." type="journal article" year="1965">Tendeiro (1965)</bibRefCitation>
was unable to study this species directly because the
<typeStatus id="9C2E8823C31F8700FC561819FC7BFEC4" box="[935,984,315,339]" pageId="37" pageNumber="3580">type</typeStatus>
specimens were deposited in the collection of the University of Halle and were almost certainly destroyed in World War II. The species was later redescribed and illustrated by
<bibRefCitation id="27044B70C31F8700FC5E185EFF73FE23" author="Emerson KC &amp; Price RD" pageId="37" pageNumber="3580" pagination="59 - 61" refId="ref38387" refString="Emerson KC, Price RD. 1979. Columbicola fortis, Cavifera abdita, and Quateia iranensis (Mallophaga) from the magnificent ground pigeon (Otidiphaps nobilis) from New Guinea. Pacific Insects 20: 59 - 61." type="journal article" year="1979">Emerson and Price (1979)</bibRefCitation>
; it is easily recognized by the structure of the head and genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>