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<document id="EC55697625D4E0ACA1C3B58321C0F7E5" ID-DOI="10.1080/00222930500393368" ID-ISSN="1464-5262" ID-Zenodo-Dep="5220802" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629364456045" checkinUser="felipe" docAuthor="Adams, Richard J., Price, Roger D. &amp; Clayton, Dale H." docDate="2005" docId="CB3C8797C3188701FE151CB2FDEAFC15" docLanguage="en" docName="JNatHist.39.41.3545-3618.pdf" docOrigin="Journal of Natural History 39 (41)" docSource="http://dx.doi.org/10.1080/00222930500393368" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Columbicola becheti Tendeiro 1965" docType="treatment" docVersion="3" lastPageNumber="3579" masterDocId="3705FFEFC33A8725FFF11922FFA3FF97" masterDocTitle="Taxonomic revision of Old World members of the feather louse genus Columbicola (Phthiraptera: Ischnocera), including descriptions of eight new species" masterLastPageNumber="3618" masterPageNumber="3545" pageNumber="3577" updateTime="1699069527252" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="761859CFDF486DEF579CC827DA151D44">Taxonomic revision of Old World members of the feather louse genus Columbicola (Phthiraptera: Ischnocera), including descriptions of eight new species</mods:title>
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<mods:namePart id="81CE85270BDC481AC192E14875DB44EB">Adams, Richard J.</mods:namePart>
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<mods:namePart id="D12D914E3F164A5DC767AF1F39172F2C">Price, Roger D.</mods:namePart>
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<mods:namePart id="F9F9D23D3D7B04EB870970AC743616AB">Clayton, Dale H.</mods:namePart>
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<mods:title id="D53A09F19EC9AF407B9B37961CDCCDA8">Journal of Natural History</mods:title>
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<mods:date id="4B5FAD83D846F1C69920289AAE003F72">2005</mods:date>
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<paragraph id="432A3681C3188707FE151CB2FCBBFA3F" blockId="34.[138,1185,1424,1675]" box="[484,792,1424,1448]" pageId="34" pageNumber="3577">
<emphasis id="71E1EA93C3188707FE151CB2FCBBFA3F" bold="true" box="[484,792,1424,1448]" pageId="34" pageNumber="3577">
8.
<taxonomicName id="84954D02C3188707FDF21CB2FDFCFA3F" ID-CoL="XC8Q" authorityName="Tendeiro" authorityYear="1965" box="[515,607,1424,1448]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="34" pageNumber="3577" phylum="Arthropoda" rank="species" species="becheti">
<emphasis id="71E1EA93C3188707FDF21CB2FDFCFA3F" bold="true" box="[515,607,1424,1448]" italics="true" pageId="34" pageNumber="3577">becheti</emphasis>
</taxonomicName>
species group
</emphasis>
</paragraph>
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<subSubSection id="0B8F650AC3188701FF7B1C93FDEAFC15" lastPageId="36" lastPageNumber="3579" pageId="34" pageNumber="3577" type="description">
<paragraph id="432A3681C3188707FF7B1C93FC8FF91C" blockId="34.[138,1185,1424,1675]" pageId="34" pageNumber="3577">
This group contains a single species from the host genus
<taxonomicName id="84954D02C3188707FCCB1C93FC29FA5E" authorityName="Hodgson" authorityYear="1836" box="[826,906,1457,1481]" class="Aves" family="Columbidae" genus="Ducula" kingdom="Animalia" order="Columbiformes" pageId="34" pageNumber="3577" phylum="Chordata" rank="genus">
<emphasis id="71E1EA93C3188707FCCB1C93FC29FA5E" box="[826,906,1457,1481]" italics="true" pageId="34" pageNumber="3577">Ducula</emphasis>
</taxonomicName>
. It has both sexes with thickened sagittal band posterior to dorsoanterior head plate (
<figureCitation id="DBAE2A04C3188707FC961CF3FB80FA7E" box="[871,1059,1489,1513]" captionStart="Figures 95111" captionStartId="39.[125,194,1529,1548]" captionTargetBox="[142,1153,133,1491]" captionTargetId="figure-0@39.[142,1155,133,1493]" captionTargetPageId="39" captionText="Figures 95111. Columbicola effeminatus: (95) female dorsal head; (96) male genitalia; (97) female ventral terminalia. C. becheti: (98) male dorsal head; (99) male genitalia; (100) female dorsal head; (101) female ventral terminalia. C. fortis: (102) male dorsal head; (103) female dorsal head; (104) male genitalia; (105) female ventral terminalia. C. tasmaniensis: (106) male genitalia; (107) female ventral terminalia. C. mjoebergi: (108) male dorsal head; (109) female dorsal head; (110) male genitalia; (111) female ventral terminalia." figureDoi="http://doi.org/10.5281/zenodo.5220814" httpUri="https://zenodo.org/record/5220814/files/figure.png" pageId="34" pageNumber="3577">Figures 98, 100</figureCitation>
); marginal head carina rounded, complete; body elongate (
<figureCitation id="DBAE2A04C3188707FD5A1CD0FCAFF99D" box="[683,780,1522,1546]" captionStart="Figures 118" captionStartId="13.[125,194,1411,1430]" captionTargetBox="[172,1126,156,1375]" captionTargetId="figure-0@13.[172,1126,154,1375]" captionTargetPageId="13" captionText="Figures 118. Columbicola columbae: (1) dorsal-ventral male. Columbicola male: (2) metanotal margin with two long, two short setal pattern; (3) metanotal margin with three long, one short setal pattern; (4) dorsoanterior head—a5anterior medial head setae (AMHS), b5posterior medial head setae (PMHS); (5) genitalia, width (GW); (6) body measurements—head length (HL), dorsoanterior head plate width (APW), head width (HW), scape length (SL), prothorax width (PW), metathorax width (MW), total length (TL). C. columbae: (7) male dorsal head (arrow5medioposterior head setae); (8) male genitalia; (9) female dorsal head; (10) female ventral terminalia. C. bacillus: (11) male genitalia; (12) female ventral terminalia. C. stresemanni: (13) male dorsal head; (14) female dorsal head. C. claviformis: (15) male dorsal head; (16) male genitalia; (17) female ventral terminalia. C. tschulyschman: (18) male dorsal head." figureDoi="http://doi.org/10.5281/zenodo.5220804" httpUri="https://zenodo.org/record/5220804/files/figure.png" pageId="34" pageNumber="3577">Figure 1</figureCitation>
); and each side of metanotum with two long, two short setae (
<figureCitation id="DBAE2A04C3188707FE471F30FDB9F9BD" box="[438,538,1554,1578]" captionStart="Figures 118" captionStartId="13.[125,194,1411,1430]" captionTargetBox="[172,1126,156,1375]" captionTargetId="figure-0@13.[172,1126,154,1375]" captionTargetPageId="13" captionText="Figures 118. Columbicola columbae: (1) dorsal-ventral male. Columbicola male: (2) metanotal margin with two long, two short setal pattern; (3) metanotal margin with three long, one short setal pattern; (4) dorsoanterior head—a5anterior medial head setae (AMHS), b5posterior medial head setae (PMHS); (5) genitalia, width (GW); (6) body measurements—head length (HL), dorsoanterior head plate width (APW), head width (HW), scape length (SL), prothorax width (PW), metathorax width (MW), total length (TL). C. columbae: (7) male dorsal head (arrow5medioposterior head setae); (8) male genitalia; (9) female dorsal head; (10) female ventral terminalia. C. bacillus: (11) male genitalia; (12) female ventral terminalia. C. stresemanni: (13) male dorsal head; (14) female dorsal head. C. claviformis: (15) male dorsal head; (16) male genitalia; (17) female ventral terminalia. C. tschulyschman: (18) male dorsal head." figureDoi="http://doi.org/10.5281/zenodo.5220804" httpUri="https://zenodo.org/record/5220804/files/figure.png" pageId="34" pageNumber="3577">Figure 2</figureCitation>
). Male antenna with scape barely enlarged, lacking third segment spur (
<figureCitation id="DBAE2A04C3188707FEC81F11FE0FF9DC" box="[313,428,1587,1611]" captionStart="Figures 95111" captionStartId="39.[125,194,1529,1548]" captionTargetBox="[142,1153,133,1491]" captionTargetId="figure-0@39.[142,1155,133,1493]" captionTargetPageId="39" captionText="Figures 95111. Columbicola effeminatus: (95) female dorsal head; (96) male genitalia; (97) female ventral terminalia. C. becheti: (98) male dorsal head; (99) male genitalia; (100) female dorsal head; (101) female ventral terminalia. C. fortis: (102) male dorsal head; (103) female dorsal head; (104) male genitalia; (105) female ventral terminalia. C. tasmaniensis: (106) male genitalia; (107) female ventral terminalia. C. mjoebergi: (108) male dorsal head; (109) female dorsal head; (110) male genitalia; (111) female ventral terminalia." figureDoi="http://doi.org/10.5281/zenodo.5220814" httpUri="https://zenodo.org/record/5220814/files/figure.png" pageId="34" pageNumber="3577">Figure 98</figureCitation>
); genitalic mesosome compact, triangular; anterior portion with four pores, each side with reduced lateral sclerites (
<figureCitation id="DBAE2A04C3188707FD211F71FCE7F9FC" box="[720,836,1619,1643]" captionStart="Figures 95111" captionStartId="39.[125,194,1529,1548]" captionTargetBox="[142,1153,133,1491]" captionTargetId="figure-0@39.[142,1155,133,1493]" captionTargetPageId="39" captionText="Figures 95111. Columbicola effeminatus: (95) female dorsal head; (96) male genitalia; (97) female ventral terminalia. C. becheti: (98) male dorsal head; (99) male genitalia; (100) female dorsal head; (101) female ventral terminalia. C. fortis: (102) male dorsal head; (103) female dorsal head; (104) male genitalia; (105) female ventral terminalia. C. tasmaniensis: (106) male genitalia; (107) female ventral terminalia. C. mjoebergi: (108) male dorsal head; (109) female dorsal head; (110) male genitalia; (111) female ventral terminalia." figureDoi="http://doi.org/10.5281/zenodo.5220814" httpUri="https://zenodo.org/record/5220814/files/figure.png" pageId="34" pageNumber="3577">Figure 99</figureCitation>
). Female with subgenital plate groove broad, rounded, lacking lateral setae (
<figureCitation id="DBAE2A04C3188707FD6A1F51FCBDF91C" box="[667,798,1651,1675]" captionStart="Figures 95111" captionStartId="39.[125,194,1529,1548]" captionTargetBox="[142,1153,133,1491]" captionTargetId="figure-0@39.[142,1155,133,1493]" captionTargetPageId="39" captionText="Figures 95111. Columbicola effeminatus: (95) female dorsal head; (96) male genitalia; (97) female ventral terminalia. C. becheti: (98) male dorsal head; (99) male genitalia; (100) female dorsal head; (101) female ventral terminalia. C. fortis: (102) male dorsal head; (103) female dorsal head; (104) male genitalia; (105) female ventral terminalia. C. tasmaniensis: (106) male genitalia; (107) female ventral terminalia. C. mjoebergi: (108) male dorsal head; (109) female dorsal head; (110) male genitalia; (111) female ventral terminalia." figureDoi="http://doi.org/10.5281/zenodo.5220814" httpUri="https://zenodo.org/record/5220814/files/figure.png" pageId="34" pageNumber="3577">Figure 101</figureCitation>
).
</paragraph>
<caption id="17EA6609C3198706FF8C1F20FD6BF9E8" ID-DOI="http://doi.org/10.5281/zenodo.5220812" ID-Zenodo-Dep="5220812" httpUri="https://zenodo.org/record/5220812/files/figure.png" pageId="35" pageNumber="3578" startId="35.[125,194,1538,1557]" targetBox="[140,1160,133,1502]" targetPageId="35">
<paragraph id="432A3681C3198706FF8C1F20FD6BF9E8" blockId="35.[125,1173,1538,1663]" pageId="35" pageNumber="3578">
Figures 7894.
<taxonomicName id="84954D02C3198706FEE71F20FE68F982" box="[278,459,1538,1557]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="35" pageNumber="3578" phylum="Arthropoda" rank="species" species="beccarii">
<emphasis id="71E1EA93C3198706FEE71F20FE68F982" box="[278,459,1538,1557]" italics="true" pageId="35" pageNumber="3578">Columbicola beccarii</emphasis>
</taxonomicName>
: (78) male dorsal head; (79) female dorsal head; (80) male genitalia.
<taxonomicName id="84954D02C3198706FB8F1F20FF46F9A7" authorityName="Eichler" authorityYear="1942" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="35" pageNumber="3578" phylum="Arthropoda" rank="species" species="taschenbergi">
<emphasis id="71E1EA93C3198706FB8F1F20FF46F9A7" italics="true" pageId="35" pageNumber="3578">C. taschenbergi</emphasis>
</taxonomicName>
: (81) male dorsal head; (82) male genitalia.
<taxonomicName id="84954D02C3198706FD621F3FFD5AF9A7" authorityName="Tendeiro" authorityYear="1973" box="[659,761,1565,1584]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="35" pageNumber="3578" phylum="Arthropoda" rank="species" species="mckeani">
<emphasis id="71E1EA93C3198706FD621F3FFD5AF9A7" box="[659,761,1565,1584]" italics="true" pageId="35" pageNumber="3578">C. mckeani</emphasis>
</taxonomicName>
: (83) male dorsal head; (84) female dorsal head; (85) male genitalia; (86) female ventral terminalia.
<taxonomicName id="84954D02C3198706FD651F15FCA7F9DD" baseAuthorityName="Piaget" baseAuthorityYear="1880" box="[660,772,1591,1610]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="35" pageNumber="3578" phylum="Arthropoda" rank="species" species="exilicornis">
<emphasis id="71E1EA93C3198706FD651F15FCA7F9DD" box="[660,772,1591,1610]" italics="true" pageId="35" pageNumber="3578">C. exilicornis</emphasis>
</taxonomicName>
: (87) male dorsal head; (88) female dorsal head; (89) male genitalia; (90) female ventral terminalia.
<taxonomicName id="84954D02C3198706FD6B1F70FD50F9F2" authorityName="Adams &amp; Price &amp; Clayton" authorityYear="2005" box="[666,755,1618,1637]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="35" pageNumber="3578" phylum="Arthropoda" rank="species" species="arnoldi">
<emphasis id="71E1EA93C3198706FD6B1F70FD50F9F2" box="[666,755,1618,1637]" italics="true" pageId="35" pageNumber="3578">C. arnoldi</emphasis>
</taxonomicName>
: (91) male dorsal head; (92) male genitalia; (93) female dorsal head.
<taxonomicName id="84954D02C3198706FE991F4EFE45F9E8" authorityName="Tendeiro" authorityYear="1984" box="[360,486,1644,1663]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="35" pageNumber="3578" phylum="Arthropoda" rank="species" species="effeminatus">
<emphasis id="71E1EA93C3198706FE991F4EFE45F9E8" box="[360,486,1644,1663]" italics="true" pageId="35" pageNumber="3578">C. effeminatus</emphasis>
</taxonomicName>
: (94) male dorsal head.
</paragraph>
</caption>
<paragraph id="432A3681C31E8701FE1519AAFCC2FF37" blockId="36.[165,1107,136,225]" box="[484,865,136,160]" pageId="36" pageNumber="3579">
<taxonomicName id="84954D02C31E8701FE1519AAFCC2FF37" authority="Tendeiro" authorityName="Tendeiro" authorityYear="1965" box="[484,865,136,160]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="36" pageNumber="3579" phylum="Arthropoda" rank="species" species="becheti">
<emphasis id="71E1EA93C31E8701FE1519AAFD4EFF37" bold="true" box="[484,749,136,160]" italics="true" pageId="36" pageNumber="3579">Columbicola becheti</emphasis>
Tendeiro
</taxonomicName>
</paragraph>
<paragraph id="432A3681C31E8701FDB0198BFBF0FF76" blockId="36.[165,1107,136,225]" pageId="36" pageNumber="3579">
(
<figureCitation id="DBAE2A04C31E8701FDB9198BFD5DFF56" box="[584,766,169,193]" captionStart="Figures 95111" captionStartId="39.[125,194,1529,1548]" captionTargetBox="[142,1153,133,1491]" captionTargetId="figure-0@39.[142,1155,133,1493]" captionTargetPageId="39" captionText="Figures 95111. Columbicola effeminatus: (95) female dorsal head; (96) male genitalia; (97) female ventral terminalia. C. becheti: (98) male dorsal head; (99) male genitalia; (100) female dorsal head; (101) female ventral terminalia. C. fortis: (102) male dorsal head; (103) female dorsal head; (104) male genitalia; (105) female ventral terminalia. C. tasmaniensis: (106) male genitalia; (107) female ventral terminalia. C. mjoebergi: (108) male dorsal head; (109) female dorsal head; (110) male genitalia; (111) female ventral terminalia." figureDoi="http://doi.org/10.5281/zenodo.5220814" httpUri="https://zenodo.org/record/5220814/files/figure.png" pageId="36" pageNumber="3579">Figures 98101</figureCitation>
)
<taxonomicName id="84954D02C31E8701FF5419EBFDD7FF76" authority="Tendeiro 1965: 279" authorityName="Tendeiro" authorityPageNumber="279" authorityYear="1965" box="[165,628,201,225]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="36" pageNumber="3579" phylum="Arthropoda" rank="species" species="becheti">
<emphasis id="71E1EA93C31E8701FF5419EBFEDFFF76" box="[165,380,201,225]" italics="true" pageId="36" pageNumber="3579">Columbicola becheti</emphasis>
<bibRefCitation id="27044B70C31E8701FE7419EBFDD7FF76" author="Tendeiro J" box="[389,628,201,225]" pageId="36" pageNumber="3579" pagination="1 - 460" refId="ref39197" refString="Tendeiro J. 1965. Estudos sobre Malofagos. Revisao monografica do genero Columbicola Ewing (Ischnocera, Philopteridae). Memorias da Junta de Investigacoes do Ultramar, No. 32 (1962). p 1 - 460." type="journal article" year="1965">Tendeiro 1965: 279</bibRefCitation>
</taxonomicName>
.
<typeStatus id="9C2E8823C31E8701FD7019EBFD1EFF76" box="[641,701,201,225]" pageId="36" pageNumber="3579">Type</typeStatus>
host:
<taxonomicName id="84954D02C31E8701FCF619EBFBEDFF76" authority="(G. R. Gray)" baseAuthorityName="G. R. Gray" baseAuthorityYear="1859" box="[775,1102,201,225]" class="Aves" family="Columbidae" genus="Ducula" kingdom="Animalia" order="Columbiformes" pageId="36" pageNumber="3579" phylum="Chordata" rank="species" species="goliath">
<emphasis id="71E1EA93C31E8701FCF619EBFC0AFF76" box="[775,937,201,225]" italics="true" pageId="36" pageNumber="3579">Ducula goliath</emphasis>
(G. R. Gray)
</taxonomicName>
.
</paragraph>
<paragraph id="432A3681C31E8701FF7B183BFEA0FEA6" blockId="36.[138,259,281,305]" box="[138,259,281,305]" pageId="36" pageNumber="3579">
<emphasis id="71E1EA93C31E8701FF7B183BFEA0FEA6" box="[138,259,281,305]" italics="true" pageId="36" pageNumber="3579">Description</emphasis>
</paragraph>
<paragraph id="432A3681C31E8701FF7B1868FBC0FD96" blockId="36.[138,1185,330,513]" pageId="36" pageNumber="3579">
<materialsCitation id="F3FD3CDCC31E8701FF7B1868FBFCFD96" collectionCode="APW, HW, PW, MW" latitude="0.098" longLatPrecision="78" longitude="-0.127" pageId="36" pageNumber="3579" specimenCount="1">
Male head as in
<figureCitation id="DBAE2A04C31E8701FEB51868FE17FEF5" box="[324,436,330,354]" captionStart="Figures 95111" captionStartId="39.[125,194,1529,1548]" captionTargetBox="[142,1153,133,1491]" captionTargetId="figure-0@39.[142,1155,133,1493]" captionTargetPageId="39" captionText="Figures 95111. Columbicola effeminatus: (95) female dorsal head; (96) male genitalia; (97) female ventral terminalia. C. becheti: (98) male dorsal head; (99) male genitalia; (100) female dorsal head; (101) female ventral terminalia. C. fortis: (102) male dorsal head; (103) female dorsal head; (104) male genitalia; (105) female ventral terminalia. C. tasmaniensis: (106) male genitalia; (107) female ventral terminalia. C. mjoebergi: (108) male dorsal head; (109) female dorsal head; (110) male genitalia; (111) female ventral terminalia." figureDoi="http://doi.org/10.5281/zenodo.5220814" httpUri="https://zenodo.org/record/5220814/files/figure.png" pageId="36" pageNumber="3579">Figure 98</figureCitation>
;
<collectionCode id="2584AE44C31E8701FE4E1868FDA2FEF5" box="[447,513,330,354]" pageId="36" pageNumber="3579">APW</collectionCode>
,
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<geoCoordinate id="26A15046C31E8701FDBF1868FD3DFEF5" box="[590,670,330,354]" degrees="0.171" direction="west" orientation="longitude" pageId="36" pageNumber="3579" precision="55" value="-0.171">0.171</geoCoordinate>
(0.158);
<collectionCode id="2584AE44C31E8701FCF81868FC9DFEF5" box="[777,830,330,354]" pageId="36" pageNumber="3579">HW</collectionCode>
, 0.350.37 (0.355); HL, 0.61 0.65 (0.632); HL/
<collectionCode id="2584AE44C31E8701FEAE1848FE31FE15" box="[351,402,362,386]" pageId="36" pageNumber="3579">HW</collectionCode>
, 1.741.83 (1.78); SL, 0.930.98 (0.955). Thorax with
<collectionCode id="2584AE44C31E8701FBD51848FBF0FE15" box="[1060,1107,362,386]" country="Thailand" name="Paleontological Collections" pageId="36" pageNumber="3579">PW</collectionCode>
, 0.27 0.29 (0.280);
<collectionCode id="2584AE44C31E8701FECD18ABFED6FE36" box="[316,373,393,417]" country="Netherlands" name="Museum Wasmann" pageId="36" pageNumber="3579">MW</collectionCode>
, 0.340.40 (0.367). Genitalia as in
<figureCitation id="DBAE2A04C31E8701FCB318ABFC11FE36" box="[834,946,393,417]" captionStart="Figures 95111" captionStartId="39.[125,194,1529,1548]" captionTargetBox="[142,1153,133,1491]" captionTargetId="figure-0@39.[142,1155,133,1493]" captionTargetPageId="39" captionText="Figures 95111. Columbicola effeminatus: (95) female dorsal head; (96) male genitalia; (97) female ventral terminalia. C. becheti: (98) male dorsal head; (99) male genitalia; (100) female dorsal head; (101) female ventral terminalia. C. fortis: (102) male dorsal head; (103) female dorsal head; (104) male genitalia; (105) female ventral terminalia. C. tasmaniensis: (106) male genitalia; (107) female ventral terminalia. C. mjoebergi: (108) male dorsal head; (109) female dorsal head; (110) male genitalia; (111) female ventral terminalia." figureDoi="http://doi.org/10.5281/zenodo.5220814" httpUri="https://zenodo.org/record/5220814/files/figure.png" pageId="36" pageNumber="3579">Figure 99</figureCitation>
; GW,
<geoCoordinate id="26A15046C31E8701FBFF18ABFBF3FE36" box="[1038,1104,393,417]" degrees="0.098" direction="north" orientation="latitude" pageId="36" pageNumber="3579" precision="55" value="0.098">0.098</geoCoordinate>
<geoCoordinate id="26A15046C31E8701FBA118ABFB02FE36" box="[1104,1185,393,417]" degrees="0.127" direction="west" orientation="longitude" pageId="36" pageNumber="3579" precision="55" value="-0.127">0.127</geoCoordinate>
(0.119). TL, 2.742.93 (2.83). Female head as in
<figureCitation id="DBAE2A04C31E8701FD1E188BFCCDFE56" box="[751,878,425,449]" captionStart="Figures 95111" captionStartId="39.[125,194,1529,1548]" captionTargetBox="[142,1153,133,1491]" captionTargetId="figure-0@39.[142,1155,133,1493]" captionTargetPageId="39" captionText="Figures 95111. Columbicola effeminatus: (95) female dorsal head; (96) male genitalia; (97) female ventral terminalia. C. becheti: (98) male dorsal head; (99) male genitalia; (100) female dorsal head; (101) female ventral terminalia. C. fortis: (102) male dorsal head; (103) female dorsal head; (104) male genitalia; (105) female ventral terminalia. C. tasmaniensis: (106) male genitalia; (107) female ventral terminalia. C. mjoebergi: (108) male dorsal head; (109) female dorsal head; (110) male genitalia; (111) female ventral terminalia." figureDoi="http://doi.org/10.5281/zenodo.5220814" httpUri="https://zenodo.org/record/5220814/files/figure.png" pageId="36" pageNumber="3579">Figure 100</figureCitation>
;
<collectionCode id="2584AE44C31E8701FC8F188BFC10FE56" box="[894,947,425,449]" pageId="36" pageNumber="3579">HW</collectionCode>
, 0.350.36 (0.355); HL, 0.630.65 (0.635); HL/
<collectionCode id="2584AE44C31E8701FE2B18EBFDAEFE76" box="[474,525,457,481]" pageId="36" pageNumber="3579">HW</collectionCode>
, 1.751.81 (1.79). Thorax with
<collectionCode id="2584AE44C31E8701FC7818EBFC1BFE76" box="[905,952,457,481]" country="Thailand" name="Paleontological Collections" pageId="36" pageNumber="3579">PW</collectionCode>
, 0.280.29 (0.282);
<collectionCode id="2584AE44C31E8701FF7B18CBFF60FD96" box="[138,195,489,513]" country="Netherlands" name="Museum Wasmann" pageId="36" pageNumber="3579">MW</collectionCode>
, 0.350.38 (0.365). Ventral terminalia as in
<figureCitation id="DBAE2A04C31E8701FD2018CBFCF2FD96" box="[721,849,489,513]" captionStart="Figures 95111" captionStartId="39.[125,194,1529,1548]" captionTargetBox="[142,1153,133,1491]" captionTargetId="figure-0@39.[142,1155,133,1493]" captionTargetPageId="39" captionText="Figures 95111. Columbicola effeminatus: (95) female dorsal head; (96) male genitalia; (97) female ventral terminalia. C. becheti: (98) male dorsal head; (99) male genitalia; (100) female dorsal head; (101) female ventral terminalia. C. fortis: (102) male dorsal head; (103) female dorsal head; (104) male genitalia; (105) female ventral terminalia. C. tasmaniensis: (106) male genitalia; (107) female ventral terminalia. C. mjoebergi: (108) male dorsal head; (109) female dorsal head; (110) male genitalia; (111) female ventral terminalia." figureDoi="http://doi.org/10.5281/zenodo.5220814" httpUri="https://zenodo.org/record/5220814/files/figure.png" pageId="36" pageNumber="3579">Figure 101</figureCitation>
. TL, 2.923.11 (2.99)
</materialsCitation>
.
</paragraph>
<paragraph id="432A3681C31E8701FF7B1B1BFF4BFDC6" blockId="36.[138,232,569,593]" box="[138,232,569,593]" pageId="36" pageNumber="3579">
<emphasis id="71E1EA93C31E8701FF7B1B1BFF4BFDC6" box="[138,232,569,593]" italics="true" pageId="36" pageNumber="3579">Material</emphasis>
</paragraph>
<paragraph id="432A3681C31E8701FF7B1B4BFD3FFD36" blockId="36.[138,1185,617,673]" pageId="36" pageNumber="3579">
<materialsCitation id="F3FD3CDCC31E8701FF7B1B4BFDCBFD36" country="New Caledonia" location="New Caledonia" pageId="36" pageNumber="3579" specimenCount="10" specimenCount-female="5" specimenCount-male="5" typeStatus="holotype">
<specimenCount id="5593FD08C31E8701FF7B1B4BFF46FD16" box="[138,229,617,641]" pageId="36" pageNumber="3579" type="male">4 males</specimenCount>
,
<specimenCount id="5593FD08C31E8701FF001B4BFEFEFD16" box="[241,349,617,641]" pageId="36" pageNumber="3579" type="female">4 females</specimenCount>
(including
<typeStatus id="9C2E8823C31E8701FE131B4BFDE5FD16" box="[482,582,617,641]" pageId="36" pageNumber="3579" type="holotype">holotype</typeStatus>
male,
<typeStatus id="9C2E8823C31E8701FD621B4BFD4EFD16" box="[659,749,617,641]" pageId="36" pageNumber="3579" type="allotype">allotype</typeStatus>
female,
<specimenCount id="5593FD08C31E8701FCA11B4BFC02FD16" box="[848,929,617,641]" pageId="36" pageNumber="3579" type="male">1 male</specimenCount>
,
<specimenCount id="5593FD08C31E8701FC5D1B4BFBADFD16" box="[940,1038,617,641]" pageId="36" pageNumber="3579" type="female">1 female</specimenCount>
<typeStatus id="9C2E8823C31E8701FBE41B4BFB20FD16" box="[1045,1155,617,641]" pageId="36" pageNumber="3579" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="84954D02C31E8701FF7B1BABFF55FD36" authorityName="Tendeiro" authorityYear="1965" box="[138,246,649,673]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="36" pageNumber="3579" phylum="Arthropoda" rank="species" species="becheti">
<emphasis id="71E1EA93C31E8701FF7B1BABFF55FD36" box="[138,246,649,673]" italics="true" pageId="36" pageNumber="3579">C. becheti</emphasis>
</taxonomicName>
), ex
<taxonomicName id="84954D02C31E8701FEC51BA8FE00FD36" baseAuthorityName="G. R. Gray" baseAuthorityYear="1859" box="[308,419,649,673]" class="Aves" family="Columbidae" genus="Ducula" kingdom="Animalia" order="Columbiformes" pageId="36" pageNumber="3579" phylum="Chordata" rank="species" species="goliath">
<emphasis id="71E1EA93C31E8701FEC51BA8FE00FD36" box="[308,419,649,673]" italics="true" pageId="36" pageNumber="3579">D. goliath</emphasis>
</taxonomicName>
,
<collectingCountry id="3B827611C31E8701FE421BABFDCBFD36" box="[435,616,649,673]" name="New Caledonia" pageId="36" pageNumber="3579">New Caledonia</collectingCountry>
</materialsCitation>
(2).
</paragraph>
<paragraph id="432A3681C31E8701FF7B1BF8FF4AFD65" blockId="36.[138,233,730,754]" box="[138,233,730,754]" pageId="36" pageNumber="3579">
<emphasis id="71E1EA93C31E8701FF7B1BF8FF4AFD65" box="[138,233,730,754]" italics="true" pageId="36" pageNumber="3579">Remarks</emphasis>
</paragraph>
<paragraph id="432A3681C31E8701FF7B1A28FDEAFC15" blockId="36.[138,1185,778,898]" pageId="36" pageNumber="3579">
The very large size of this louse, in conjunction with the structure of the genitalia and the male antenna, is distinctive. Despite having only two long metathoracic setae, the existence of small lateral sclerites around the mesosome hints at a distant relationship between this louse and those in the
<taxonomicName id="84954D02C31E8701FE671A48FE57FC15" baseAuthorityName="Rudow" baseAuthorityYear="1869" box="[406,500,874,898]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="36" pageNumber="3579" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="71E1EA93C31E8701FE671A48FE57FC15" box="[406,500,874,898]" italics="true" pageId="36" pageNumber="3579">longiceps</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
</treatment>
</document>