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<document id="AD3874D01A1C0733351287F761257B58" ID-DOI="10.1080/00222930500393368" ID-ISSN="1464-5262" ID-Zenodo-Dep="5220802" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629364456045" checkinUser="felipe" docAuthor="Adams, Richard J., Price, Roger D. &amp; Clayton, Dale H." docDate="2005" docId="CB3C8797C3088711FE151D3EFDD4FCDC" docLanguage="en" docName="JNatHist.39.41.3545-3618.pdf" docOrigin="Journal of Natural History 39 (41)" docSource="http://dx.doi.org/10.1080/00222930500393368" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Columbicola reedi Adams &amp; Price &amp; Clayton 2005, n. sp." docType="treatment" docVersion="3" lastPageNumber="3595" masterDocId="3705FFEFC33A8725FFF11922FFA3FF97" masterDocTitle="Taxonomic revision of Old World members of the feather louse genus Columbicola (Phthiraptera: Ischnocera), including descriptions of eight new species" masterLastPageNumber="3618" masterPageNumber="3545" pageNumber="3593" updateTime="1699069527252" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="3DF40D89E58DB4AE967BC40C8585AD33">Taxonomic revision of Old World members of the feather louse genus Columbicola (Phthiraptera: Ischnocera), including descriptions of eight new species</mods:title>
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<mods:namePart id="5FF36BBA7D371AC0282B8FF8CBAC41E0">Adams, Richard J.</mods:namePart>
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<mods:namePart id="ADB29D9EE16508623572A7E0DB09FAE9">Price, Roger D.</mods:namePart>
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<mods:namePart id="9E3F5D38CDF667C769B745E74B94C96A">Clayton, Dale H.</mods:namePart>
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<mods:title id="8B8E1679856BC42D97A89FE1291C16B5">Journal of Natural History</mods:title>
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<mods:date id="3757E8980E23C2D26DFA3982BE2D68DD">2005</mods:date>
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<mods:number id="0D0C66E297043B180070EE90A33BC69C">39</mods:number>
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<paragraph id="432A3681C3088717FE151D3EFCBFFBA3" blockId="50.[484,796,1052,1109]" box="[484,796,1052,1076]" pageId="50" pageNumber="3593">
<heading id="186281EDC3088717FE151D3EFCBFFBA3" box="[484,796,1052,1076]" centered="true" fontSize="10" level="2" pageId="50" pageNumber="3593" reason="4">
<taxonomicName id="84954D02C3088717FE151D3EFD71FBA3" authority="Adams &amp; Price &amp; Clayton, 2005" authorityName="Adams &amp; Price &amp; Clayton" authorityYear="2005" box="[484,722,1052,1076]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="50" pageNumber="3593" phylum="Arthropoda" rank="species" species="reedi" status="sp. nov.">
<emphasis id="71E1EA93C3088717FE151D3EFD71FBA3" bold="true" box="[484,722,1052,1076]" italics="true" pageId="50" pageNumber="3593">Columbicola reedi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6AD257E8C3088717FD2A1D3EFCBFFBA3" box="[731,796,1052,1076]" pageId="50" pageNumber="3593" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="0B8F650AC3088711FDE81D1FFDD4FCDC" lastPageId="52" lastPageNumber="3595" pageId="50" pageNumber="3593" type="description">
<paragraph id="432A3681C3088717FDE81D1FFD4EFBC2" blockId="50.[484,796,1052,1109]" box="[537,749,1085,1109]" pageId="50" pageNumber="3593">
(
<figureCitation id="DBAE2A04C3088717FDD01D1FFD47FBC2" box="[545,740,1085,1109]" captionStart="Figures 147164" captionStartId="56.[138,207,1530,1549]" captionTargetBox="[154,1168,133,1492]" captionTargetId="figure-1@56.[154,1170,133,1495]" captionTargetPageId="56" captionText="Figures 147164. Columbicola harrisoni: (147) female ventral terminalia. C. reedi: (148) male dorsal head; (149) female dorsal head; (150) male genitalia; (151) female ventral terminalia. C. gourae: (152) male dorsal head; (153) male genitalia; (154) female dorsal head; (155) female ventral terminalia. C. paradoxus: (156) male dorsal head; (157) male genitalia. C. clayae: (158) male dorsal head; (159) female dorsal head; (160) male genitalia; (161) female ventral terminalia. C. davisae: (162) male dorsal head; (163) female dorsal head; (164) male genitalia." figureDoi="http://doi.org/10.5281/zenodo.5220822" httpUri="https://zenodo.org/record/5220822/files/figure.png" pageId="50" pageNumber="3593">Figures 148151</figureCitation>
)
</paragraph>
<paragraph id="432A3681C3088717FF7B1DBEFF50FB24" blockId="50.[138,243,1179,1203]" box="[138,243,1179,1203]" pageId="50" pageNumber="3593">
<emphasis id="71E1EA93C3088717FF7B1DBEFF50FB24" box="[138,243,1179,1203]" italics="true" pageId="50" pageNumber="3593">Type host</emphasis>
</paragraph>
<paragraph id="432A3681C3088717FF7B1DEEFDB5FB73" blockId="50.[138,534,1228,1252]" box="[138,534,1228,1252]" pageId="50" pageNumber="3593">
<taxonomicName id="84954D02C3088717FF7B1DEEFDB1FB73" authority="(Temminck)" baseAuthorityName="Temminck" baseAuthorityYear="1821" box="[138,530,1228,1252]" class="Aves" family="Columbidae" genus="Ptilinopus" kingdom="Animalia" order="Columbiformes" pageId="50" pageNumber="3593" phylum="Chordata" rank="species" species="magnificus">
<emphasis id="71E1EA93C3088717FF7B1DEEFED2FB73" box="[138,369,1228,1252]" italics="true" pageId="50" pageNumber="3593">Ptilinopus magnificus</emphasis>
(Temminck)
</taxonomicName>
.
</paragraph>
<paragraph id="432A3681C3088717FF7B1C08FEA0FAD5" blockId="50.[138,259,1322,1346]" box="[138,259,1322,1346]" pageId="50" pageNumber="3593">
<emphasis id="71E1EA93C3088717FF7B1C08FEA0FAD5" box="[138,259,1322,1346]" italics="true" pageId="50" pageNumber="3593">Description</emphasis>
</paragraph>
<paragraph id="432A3681C3088717FF7B1C79FE01F902" blockId="50.[138,1185,1371,1685]" pageId="50" pageNumber="3593">
<materialsCitation id="F3FD3CDCC3088717FF7B1C79FE3CF902" collectionCode="APW, HW, PW, MW" latitude="0.132" longLatPrecision="78" longitude="-0.167" pageId="50" pageNumber="3593" specimenCount="1">
Male head as in
<figureCitation id="DBAE2A04C3088717FEAE1C79FE7DFAE4" box="[351,478,1371,1395]" captionStart="Figures 147164" captionStartId="56.[138,207,1530,1549]" captionTargetBox="[154,1168,133,1492]" captionTargetId="figure-1@56.[154,1170,133,1495]" captionTargetPageId="56" captionText="Figures 147164. Columbicola harrisoni: (147) female ventral terminalia. C. reedi: (148) male dorsal head; (149) female dorsal head; (150) male genitalia; (151) female ventral terminalia. C. gourae: (152) male dorsal head; (153) male genitalia; (154) female dorsal head; (155) female ventral terminalia. C. paradoxus: (156) male dorsal head; (157) male genitalia. C. clayae: (158) male dorsal head; (159) female dorsal head; (160) male genitalia; (161) female ventral terminalia. C. davisae: (162) male dorsal head; (163) female dorsal head; (164) male genitalia." figureDoi="http://doi.org/10.5281/zenodo.5220822" httpUri="https://zenodo.org/record/5220822/files/figure.png" pageId="50" pageNumber="3593">Figure 148</figureCitation>
; anterior margin rounded or squared, occasionally weakly indented;
<collectionCode id="2584AE44C3088717FEF21C59FEE6FA04" box="[259,325,1403,1427]" pageId="50" pageNumber="3593">APW</collectionCode>
,
<geoCoordinate id="26A15046C3088717FEA51C59FE35FA04" box="[340,406,1403,1427]" degrees="0.137" direction="north" orientation="latitude" pageId="50" pageNumber="3593" precision="55" value="0.137">0.137</geoCoordinate>
<geoCoordinate id="26A15046C3088717FE641C59FE45FA04" box="[405,486,1403,1427]" degrees="0.147" direction="west" orientation="longitude" pageId="50" pageNumber="3593" precision="55" value="-0.147">0.147</geoCoordinate>
(0.143);
<collectionCode id="2584AE44C3088717FDAB1C59FD2CFA04" box="[602,655,1403,1427]" pageId="50" pageNumber="3593">HW</collectionCode>
, 0.280.30 (0.292); HL, 0.560.65 (0.605); HL/
<collectionCode id="2584AE44C3088717FF4D1CB9FF4CFA24" box="[188,239,1435,1459]" pageId="50" pageNumber="3593">HW</collectionCode>
, 2.002.17 (2.07); SL,
<geoCoordinate id="26A15046C3088717FDF41CB9FDE4FA24" box="[517,583,1435,1459]" degrees="0.118" direction="north" orientation="latitude" pageId="50" pageNumber="3593" precision="55" value="0.118">0.118</geoCoordinate>
<geoCoordinate id="26A15046C3088717FDB71CB9FD34FA24" box="[582,663,1435,1459]" degrees="0.147" direction="west" orientation="longitude" pageId="50" pageNumber="3593" precision="55" value="-0.147">0.147</geoCoordinate>
(0.130); medioposterior head setae medium length, not reaching posterior head margin. Thorax with
<collectionCode id="2584AE44C3088717FCCA1C9EFCC9FA43" box="[827,874,1468,1492]" country="Thailand" name="Paleontological Collections" pageId="50" pageNumber="3593">PW</collectionCode>
, 0.220.26 (0.240);
<collectionCode id="2584AE44C3088717FB951C9EFB3EFA43" box="[1124,1181,1468,1492]" country="Netherlands" name="Museum Wasmann" pageId="50" pageNumber="3593">MW</collectionCode>
, 0.270.30 (0.292). Genitalia as in
<figureCitation id="DBAE2A04C3088717FDC21CFEFD10FA63" box="[563,691,1500,1524]" captionStart="Figures 147164" captionStartId="56.[138,207,1530,1549]" captionTargetBox="[154,1168,133,1492]" captionTargetId="figure-1@56.[154,1170,133,1495]" captionTargetPageId="56" captionText="Figures 147164. Columbicola harrisoni: (147) female ventral terminalia. C. reedi: (148) male dorsal head; (149) female dorsal head; (150) male genitalia; (151) female ventral terminalia. C. gourae: (152) male dorsal head; (153) male genitalia; (154) female dorsal head; (155) female ventral terminalia. C. paradoxus: (156) male dorsal head; (157) male genitalia. C. clayae: (158) male dorsal head; (159) female dorsal head; (160) male genitalia; (161) female ventral terminalia. C. davisae: (162) male dorsal head; (163) female dorsal head; (164) male genitalia." figureDoi="http://doi.org/10.5281/zenodo.5220822" httpUri="https://zenodo.org/record/5220822/files/figure.png" pageId="50" pageNumber="3593">Figure 150</figureCitation>
; mesosome narrow, lateral sclerites thin, angled anteriorly; GW,
<geoCoordinate id="26A15046C3088717FE561CDEFE4BF983" box="[423,488,1532,1556]" degrees="0.113" direction="north" orientation="latitude" pageId="50" pageNumber="3593" precision="55" value="0.113">0.113</geoCoordinate>
<geoCoordinate id="26A15046C3088717FE181CDEFD99F983" box="[489,570,1532,1556]" degrees="0.127" direction="west" orientation="longitude" pageId="50" pageNumber="3593" precision="55" value="-0.127">0.127</geoCoordinate>
(0.118). TL, 2.352.68 (2.52). Female head as in
<figureCitation id="DBAE2A04C3088717FF7B1F3EFEAAF9A3" box="[138,265,1564,1588]" captionStart="Figures 147164" captionStartId="56.[138,207,1530,1549]" captionTargetBox="[154,1168,133,1492]" captionTargetId="figure-1@56.[154,1170,133,1495]" captionTargetPageId="56" captionText="Figures 147164. Columbicola harrisoni: (147) female ventral terminalia. C. reedi: (148) male dorsal head; (149) female dorsal head; (150) male genitalia; (151) female ventral terminalia. C. gourae: (152) male dorsal head; (153) male genitalia; (154) female dorsal head; (155) female ventral terminalia. C. paradoxus: (156) male dorsal head; (157) male genitalia. C. clayae: (158) male dorsal head; (159) female dorsal head; (160) male genitalia; (161) female ventral terminalia. C. davisae: (162) male dorsal head; (163) female dorsal head; (164) male genitalia." figureDoi="http://doi.org/10.5281/zenodo.5220822" httpUri="https://zenodo.org/record/5220822/files/figure.png" pageId="50" pageNumber="3593">Figure 149</figureCitation>
, with anterior margin as for male;
<collectionCode id="2584AE44C3088717FD5E1F3EFD52F9A3" box="[687,753,1564,1588]" pageId="50" pageNumber="3593">APW</collectionCode>
,
<geoCoordinate id="26A15046C3088717FCF11F3EFCE7F9A3" box="[768,836,1564,1588]" degrees="0.132" direction="north" orientation="latitude" pageId="50" pageNumber="3593" precision="55" value="0.132">0.132</geoCoordinate>
<geoCoordinate id="26A15046C3088717FCB51F3EFC35F9A3" box="[836,918,1564,1588]" degrees="0.167" direction="west" orientation="longitude" pageId="50" pageNumber="3593" precision="55" value="-0.167">0.167</geoCoordinate>
;
<collectionCode id="2584AE44C3088717FC541F3EFC79F9A3" box="[933,986,1564,1588]" pageId="50" pageNumber="3593">HW</collectionCode>
, 0.280.32; HL, 0.580.65; HL/
<collectionCode id="2584AE44C3088717FEB41F1FFEDBF9C2" box="[325,376,1597,1621]" pageId="50" pageNumber="3593">HW</collectionCode>
, 2.032.07. Thorax with
<collectionCode id="2584AE44C3088717FD401F1FFD43F9C2" box="[689,736,1597,1621]" country="Thailand" name="Paleontological Collections" pageId="50" pageNumber="3593">PW</collectionCode>
, 0.230.27;
<collectionCode id="2584AE44C3088717FC881F1FFC11F9C2" box="[889,946,1597,1621]" country="Netherlands" name="Museum Wasmann" pageId="50" pageNumber="3593">MW</collectionCode>
, 0.280.34. Ventral terminalia as in
<figureCitation id="DBAE2A04C3088717FEA61F7FFE75F9E2" box="[343,470,1629,1653]" captionStart="Figures 147164" captionStartId="56.[138,207,1530,1549]" captionTargetBox="[154,1168,133,1492]" captionTargetId="figure-1@56.[154,1170,133,1495]" captionTargetPageId="56" captionText="Figures 147164. Columbicola harrisoni: (147) female ventral terminalia. C. reedi: (148) male dorsal head; (149) female dorsal head; (150) male genitalia; (151) female ventral terminalia. C. gourae: (152) male dorsal head; (153) male genitalia; (154) female dorsal head; (155) female ventral terminalia. C. paradoxus: (156) male dorsal head; (157) male genitalia. C. clayae: (158) male dorsal head; (159) female dorsal head; (160) male genitalia; (161) female ventral terminalia. C. davisae: (162) male dorsal head; (163) female dorsal head; (164) male genitalia." figureDoi="http://doi.org/10.5281/zenodo.5220822" httpUri="https://zenodo.org/record/5220822/files/figure.png" pageId="50" pageNumber="3593">Figure 151</figureCitation>
; subgenital plate groove narrow, anteriorly pointed, edges uneven. TL, 2.562.84
</materialsCitation>
.
</paragraph>
<caption id="17EA6609C3098716FF8C1CD6FDD3F9E6" ID-DOI="http://doi.org/10.5281/zenodo.5220820" ID-Zenodo-Dep="5220820" httpUri="https://zenodo.org/record/5220820/files/figure.png" pageId="51" pageNumber="3594" startId="51.[125,194,1524,1543]" targetBox="[158,1139,145,1488]" targetPageId="51">
<paragraph id="432A3681C3098716FF8C1CD6FDD3F9E6" blockId="51.[125,1172,1524,1649]" pageId="51" pageNumber="3594">
Figures 131146.
<taxonomicName id="84954D02C3098716FEDD1CD6FE4AF990" baseAuthorityName="Rudow" baseAuthorityYear="1869" box="[300,489,1524,1543]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="51" pageNumber="3594" phylum="Arthropoda" rank="species" species="longiceps">
<emphasis id="71E1EA93C3098716FEDD1CD6FE4AF990" box="[300,489,1524,1543]" italics="true" pageId="51" pageNumber="3594">Columbicola longiceps</emphasis>
</taxonomicName>
: (131) male genitalia.
<taxonomicName id="84954D02C3098716FD321CD6FC80F990" authorityName="Adams &amp; Price &amp; Clayton" authorityYear="2005" box="[707,803,1524,1543]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="51" pageNumber="3594" phylum="Arthropoda" rank="species" species="mendesi">
<emphasis id="71E1EA93C3098716FD321CD6FC80F990" box="[707,803,1524,1543]" italics="true" pageId="51" pageNumber="3594">C. mendesi</emphasis>
</taxonomicName>
: (132) male dorsal head; (133) female dorsal head; (134) male genitalia; (135) female ventral terminalia.
<taxonomicName id="84954D02C3098716FD2C1F2CFCE6F9B6" baseAuthorityName="Taschenberg" baseAuthorityYear="1882" box="[733,837,1550,1569]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="51" pageNumber="3594" phylum="Arthropoda" rank="species" species="cavifrons">
<emphasis id="71E1EA93C3098716FD2C1F2CFCE6F9B6" box="[733,837,1550,1569]" italics="true" pageId="51" pageNumber="3594">C. cavifrons</emphasis>
</taxonomicName>
: (136) male dorsal head; (137) male genitalia; (138) female dorsal head; (139) female ventral terminalia.
<taxonomicName id="84954D02C3098716FD1F1F0BFCF3F9AB" box="[750,848,1577,1596]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="51" pageNumber="3594" phylum="Arthropoda" rank="species" species="sikoraae">
<emphasis id="71E1EA93C3098716FD1F1F0BFCF3F9AB" box="[750,848,1577,1596]" italics="true" pageId="51" pageNumber="3594">C. sikoraae</emphasis>
</taxonomicName>
: (140) male genitalia; (141) female ventral terminalia.
<taxonomicName id="84954D02C3098716FEDA1F61FE21F9C1" authorityName="Tendeiro" authorityYear="1967" box="[299,386,1603,1622]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="51" pageNumber="3594" phylum="Arthropoda" rank="species" species="xavieri">
<emphasis id="71E1EA93C3098716FEDA1F61FE21F9C1" box="[299,386,1603,1622]" italics="true" pageId="51" pageNumber="3594">C. xavieri</emphasis>
</taxonomicName>
: (142) male genitalia; (143) female ventral terminalia.
<taxonomicName id="84954D02C3098716FC8C1F61FC46F9C1" authorityName="Tendeiro" authorityYear="1965" box="[893,997,1603,1622]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="51" pageNumber="3594" phylum="Arthropoda" rank="species" species="harrisoni">
<emphasis id="71E1EA93C3098716FC8C1F61FC46F9C1" box="[893,997,1603,1622]" italics="true" pageId="51" pageNumber="3594">C. harrisoni</emphasis>
</taxonomicName>
: (144) male dorsal head; (145) female dorsal head; (146) male genitalia.
</paragraph>
</caption>
<paragraph id="432A3681C30E8711FF7B19ABFE81FF37" blockId="52.[138,290,136,160]" box="[138,290,136,160]" pageId="52" pageNumber="3595">
<emphasis id="71E1EA93C30E8711FF7B19ABFE81FF37" box="[138,290,136,160]" italics="true" pageId="52" pageNumber="3595">Type material</emphasis>
</paragraph>
<paragraph id="432A3681C30E8711FF7B199AFBEBFE87" blockId="52.[138,1185,184,272]" pageId="52" pageNumber="3595">
<materialsCitation id="F3FD3CDCC30E8711FF7B199AFF45FF67" collectingDate="1972-10-23" collectionCode="OSU" collectorName="E. Sepik Dist." country="Guinea-Bissau" location="Sepik Dist." pageId="52" pageNumber="3595" specimenCount="1" typeStatus="holotype">
<typeStatus id="9C2E8823C30E8711FF7B199AFF56FF47" box="[138,245,184,208]" pageId="52" pageNumber="3595" type="holotype">Holotype</typeStatus>
male at
<collectionCode id="2584AE44C30E8711FEA2199AFE31FF47" box="[339,402,184,208]" country="0" httpUri="http://grbio.org/cool/9iaf-dk63" name="Oklahoma State University, Collection of Vertebrates" pageId="52" pageNumber="3595">OSU</collectionCode>
, ex
<taxonomicName id="84954D02C30E8711FE4A199BFDEFFF47" baseAuthorityName="Temminck" baseAuthorityYear="1821" box="[443,588,184,208]" class="Aves" family="Columbidae" genus="Ptilinopus" kingdom="Animalia" order="Columbiformes" pageId="52" pageNumber="3595" phylum="Chordata" rank="species" species="magnificus">
<emphasis id="71E1EA93C30E8711FE4A199BFDEFFF47" box="[443,588,184,208]" italics="true" pageId="52" pageNumber="3595">P. magnificus</emphasis>
</taxonomicName>
, New
<collectingCountry id="3B827611C30E8711FD67199AFD4DFF47" box="[662,750,184,208]" name="Guinea-Bissau" pageId="52" pageNumber="3595">Guinea</collectingCountry>
:
<collectorName id="EE605357C30E8711FD09199AFC3BFF47" box="[760,920,184,208]" pageId="52" pageNumber="3595">
E.
<location id="464A605AC30E8711FCE6199AFC3BFF47" LSID="urn:lsid:plazi:treatment:CB3C8797C3088711FE151D3EFDD4FCDC:464A605AC30E8711FCE6199AFC3BFF47" box="[791,920,184,208]" country="Guinea-Bissau" name="Sepik Dist." pageId="52" pageNumber="3595">Sepik Dist.</location>
</collectorName>
, Wewak,
<date id="372B1041C30E8711FBF5199AFB3FFF47" box="[1028,1180,184,208]" pageId="52" pageNumber="3595" value="1972-10-23">
<collectingDate id="276FE9A9C30E8711FBF5199AFB3FFF47" box="[1028,1180,184,208]" pageId="52" pageNumber="3595" value="1972-10-23">23-Oct-1972</collectingDate>
</date>
, 101673
</materialsCitation>
.
<typeStatus id="9C2E8823C30E8711FF0219FAFEC1FF67" box="[243,354,216,240]" pageId="52" pageNumber="3595" type="paratype">Paratypes</typeStatus>
ex
<taxonomicName id="84954D02C30E8711FE6119FAFD86FF67" baseAuthorityName="Temminck" baseAuthorityYear="1821" box="[400,549,216,240]" class="Aves" family="Columbidae" genus="Ptilinopus" kingdom="Animalia" order="Columbiformes" pageId="52" pageNumber="3595" phylum="Chordata" rank="species" species="magnificus">
<emphasis id="71E1EA93C30E8711FE6119FAFD86FF67" box="[400,549,216,240]" italics="true" pageId="52" pageNumber="3595">P. magnificus</emphasis>
</taxonomicName>
at
<materialsCitation id="F3FD3CDCC30E8711FDBF19FAFB27FF67" box="[590,1156,216,240]" collectingDate="1972-10-23" collectionCode="OSU, BMNH" collectorName="E. Sepik Dist." country="Guinea-Bissau" location="Sepik Dist." pageId="52" pageNumber="3595" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectionCode id="2584AE44C30E8711FDBF19FAFD2EFF67" box="[590,653,216,240]" country="0" httpUri="http://grbio.org/cool/9iaf-dk63" name="Oklahoma State University, Collection of Vertebrates" pageId="52" pageNumber="3595">OSU</collectionCode>
,
<collectionCode id="2584AE44C30E8711FD6A19FAFD5AFF67" box="[667,761,216,240]" pageId="52" pageNumber="3595">BMNH</collectionCode>
:
<specimenCount id="5593FD08C30E8711FCF619FAFCCDFF67" box="[775,878,216,240]" pageId="52" pageNumber="3595" type="female">1 female</specimenCount>
, same data as holotype
</materialsCitation>
;
<specimenCount id="5593FD08C30E8711FB6319FAFF67FE87" pageId="52" pageNumber="3595" type="male">1 male</specimenCount>
, L. Harrison Coll.,
<emphasis id="71E1EA93C30E8711FE5919DAFE1DFE99" box="[424,446,248,270]" italics="true" pageId="52" pageNumber="3595">#</emphasis>
929;
<specimenCount id="5593FD08C30E8711FE0B19DAFDFCFE87" box="[506,607,248,272]" pageId="52" pageNumber="3595" type="female">1 female</specimenCount>
, same except
<emphasis id="71E1EA93C30E8711FD0E19DAFCB6FE99" box="[767,789,248,270]" italics="true" pageId="52" pageNumber="3595">#</emphasis>
930;
<specimenCount id="5593FD08C30E8711FCA119DAFC0FFE87" box="[848,940,248,272]" pageId="52" pageNumber="3595" type="male">2 males</specimenCount>
, L. Harrison.
</paragraph>
<paragraph id="432A3681C30E8711FF7B1867FF4AFECA" blockId="52.[138,1185,325,683]" box="[138,233,325,349]" pageId="52" pageNumber="3595">
<emphasis id="71E1EA93C30E8711FF7B1867FF4AFECA" box="[138,233,325,349]" italics="true" pageId="52" pageNumber="3595">Remarks</emphasis>
</paragraph>
<paragraph id="432A3681C30E8711FF7B1856FEB9FD3C" blockId="52.[138,1185,325,683]" pageId="52" pageNumber="3595">
We have split
<taxonomicName id="84954D02C30E8711FECC1856FE34FE1B" authorityName="Adams &amp; Price &amp; Clayton" authorityYear="2005" box="[317,407,372,396]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="52" pageNumber="3595" phylum="Arthropoda" rank="species" species="reedi">
<emphasis id="71E1EA93C30E8711FECC1856FE34FE1B" box="[317,407,372,396]" italics="true" pageId="52" pageNumber="3595">C. reedi</emphasis>
</taxonomicName>
from the
<taxonomicName id="84954D02C30E8711FDE81856FD03FE1B" authorityName="Tendeiro" authorityYear="1965" box="[537,672,372,396]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="52" pageNumber="3595" phylum="Arthropoda" rank="species" species="harrisoni">
<emphasis id="71E1EA93C30E8711FDE81856FD03FE1B" box="[537,672,372,396]" italics="true" pageId="52" pageNumber="3595">C. harrisoni</emphasis>
</taxonomicName>
<typeStatus id="9C2E8823C30E8711FD5C1856FCB3FE1B" box="[685,784,372,396]" pageId="52" pageNumber="3595" type="paratype">paratype</typeStatus>
series on the basis of consistent differences in genitalia, i.e. the arrangement of the male transverse mesosomal sclerite and the narrower subgenital plate groove of the female. The
<typeStatus id="9C2E8823C30E8711FCB11896FC07FE5B" box="[832,932,436,460]" pageId="52" pageNumber="3595" type="holotype">holotype</typeStatus>
male and associated female are noticeably smaller than the other
<taxonomicName id="84954D02C30E8711FD7F18F6FD40FE7B" authorityName="Adams &amp; Price &amp; Clayton" authorityYear="2005" box="[654,739,468,492]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="52" pageNumber="3595" phylum="Arthropoda" rank="species" species="reedi">
<emphasis id="71E1EA93C30E8711FD7F18F6FD40FE7B" box="[654,739,468,492]" italics="true" pageId="52" pageNumber="3595">C. reedi</emphasis>
</taxonomicName>
specimens. These specimens are from New
<collectingCountry id="3B827611C30E8711FF3A18D6FE80FD9B" box="[203,291,500,524]" name="Guinea-Bissau" pageId="52" pageNumber="3595">Guinea</collectingCountry>
, which is home to one of the smaller subspecies of the host,
<taxonomicName id="84954D02C30E8711FBF218D7FB3AFD9B" baseAuthorityName="Temminck" baseAuthorityYear="1821" box="[1027,1177,500,524]" class="Aves" family="Columbidae" genus="Ptilinopus" kingdom="Animalia" order="Columbiformes" pageId="52" pageNumber="3595" phylum="Chordata" rank="species" species="magnificus">
<emphasis id="71E1EA93C30E8711FBF218D7FB3AFD9B" box="[1027,1177,500,524]" italics="true" pageId="52" pageNumber="3595">P. magnificus</emphasis>
</taxonomicName>
. Geographically,
<taxonomicName id="84954D02C30E8711FEBA1B37FE42FDBB" baseAuthorityName="Temminck" baseAuthorityYear="1821" box="[331,481,532,556]" class="Aves" family="Columbidae" genus="Ptilinopus" kingdom="Animalia" order="Columbiformes" pageId="52" pageNumber="3595" phylum="Chordata" rank="species" species="magnificus">
<emphasis id="71E1EA93C30E8711FEBA1B37FE42FDBB" box="[331,481,532,556]" italics="true" pageId="52" pageNumber="3595">P. magnificus</emphasis>
</taxonomicName>
varies greatly in size (del
<bibRefCitation id="27044B70C30E8711FCEF1B36FC4FFDBB" author="Hoyo J &amp; Elliot A &amp; Sargatal J" box="[798,1004,532,556]" pageId="52" pageNumber="3595" refId="ref38095" refString="del Hoyo J, Elliot A, Sargatal J, editors. 1997. Handbook of birds of the world. Vol. 4, Sandgrouse to Cuckoos. Barcelona: Lynx Edicions. 677 p." type="book" year="1997">Hoyo et al. 1997</bibRefCitation>
) and the other specimens of
<taxonomicName id="84954D02C30E8711FED71B16FED9FDDB" authorityName="Adams &amp; Price &amp; Clayton" authorityYear="2005" box="[294,378,564,588]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="52" pageNumber="3595" phylum="Arthropoda" rank="species" species="reedi">
<emphasis id="71E1EA93C30E8711FED71B16FED9FDDB" box="[294,378,564,588]" italics="true" pageId="52" pageNumber="3595">C. reedi</emphasis>
</taxonomicName>
appear to be from 2 larger Australian subspecies. The trend shown by these few specimens of
<taxonomicName id="84954D02C30E8711FE661B76FE48FDFB" authorityName="Adams &amp; Price &amp; Clayton" authorityYear="2005" box="[407,491,596,620]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="52" pageNumber="3595" phylum="Arthropoda" rank="species" species="reedi">
<emphasis id="71E1EA93C30E8711FE661B76FE48FDFB" box="[407,491,596,620]" italics="true" pageId="52" pageNumber="3595">C. reedi</emphasis>
</taxonomicName>
suggests that this species may demonstrate Harrisons Rule, which states that parasite size correlates with host size (
<bibRefCitation id="27044B70C30E8711FCFC1B51FC1EFD1C" author="Harrison L" box="[781,957,627,651]" pageId="52" pageNumber="3595" pagination="88 - 100" refId="ref38535" refString="Harrison L. 1915. Mallophaga from Apteryx, and their significance; with a note on the genus Rallicola. Parasitology 8: 88 - 100." type="journal article" year="1915">Harrison 1915</bibRefCitation>
,
<bibRefCitation id="27044B70C30E8711FC3B1B51FBE6FD1C" author="Clay T" box="[970,1093,627,651]" pageId="52" pageNumber="3595" pagination="207 - 215" refId="ref37731" refString="Clay T. 1951. The Mallophaga as an aid to the classification of birds with special reference to the structure of feathers. Proceedings of the 10 th International Ornithological Congress. p 207 - 215." type="book chapter" year="1951">Clay 1951</bibRefCitation>
, Johson et al. 2005).
</paragraph>
<paragraph id="432A3681C30E8711FF7B1BC1FF58FD6C" blockId="52.[138,251,739,763]" box="[138,251,739,763]" pageId="52" pageNumber="3595">
<emphasis id="71E1EA93C30E8711FF7B1BC1FF58FD6C" box="[138,251,739,763]" italics="true" pageId="52" pageNumber="3595">Etymology</emphasis>
</paragraph>
<paragraph id="432A3681C30E8711FF7B1A31FDD4FCDC" blockId="52.[138,1185,787,843]" pageId="52" pageNumber="3595">
This species is named for David L. Reed, University of
<collectingRegion id="8151F863C30E8711FCFF1A31FCC5FCBC" box="[782,870,787,811]" country="United States of America" name="Florida" pageId="52" pageNumber="3595">Florida</collectingRegion>
, Gainesville, in recognition of his work on hostparasite relationships.
</paragraph>
</subSubSection>
</treatment>
</document>