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<document id="FF57C5C0AB64C625C782EB3D8A6D5362" ID-DOI="10.1080/00222930500393368" ID-ISSN="1464-5262" ID-Zenodo-Dep="5220802" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629364456045" checkinUser="felipe" docAuthor="Adams, Richard J., Price, Roger D. &amp; Clayton, Dale H." docDate="2005" docId="CB3C8797C300871EFE381CBCFB36FA6F" docLanguage="en" docName="JNatHist.39.41.3545-3618.pdf" docOrigin="Journal of Natural History 39 (41)" docSource="http://dx.doi.org/10.1080/00222930500393368" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Columbicola sphenurus Tendeiro 1984" docType="treatment" docVersion="3" lastPageNumber="3602" masterDocId="3705FFEFC33A8725FFF11922FFA3FF97" masterDocTitle="Taxonomic revision of Old World members of the feather louse genus Columbicola (Phthiraptera: Ischnocera), including descriptions of eight new species" masterLastPageNumber="3618" masterPageNumber="3545" pageNumber="3601" updateTime="1699069527252" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="17498991BF5B5290B538C9B9BFAE0F78">Taxonomic revision of Old World members of the feather louse genus Columbicola (Phthiraptera: Ischnocera), including descriptions of eight new species</mods:title>
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<mods:namePart id="86A954FD539C3B751C2FE56E8CC0115E">Adams, Richard J.</mods:namePart>
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<mods:namePart id="5672717FBFE36E14D42CB004E64A5B78">Price, Roger D.</mods:namePart>
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<mods:namePart id="05721524C99FFA3C7216C0078B6EAFE3">Clayton, Dale H.</mods:namePart>
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<mods:title id="721C38DB6CD526E16A63419019E51634">Journal of Natural History</mods:title>
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<mods:date id="74EC3A5DA710BD907AB33B0F6864C454">2005</mods:date>
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<mods:number id="8892AEB7A23E272D55C5EEA4DBB78FAD">2005-12-23</mods:number>
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<mods:number id="79065D5BD6061CE5FB80EFB6E17B87C3">39</mods:number>
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<mods:number id="BDB381303A62C097D2D6E096F9F5CC6C">41</mods:number>
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<mods:identifier id="8043BA56C18E2B70AED6AC3E3E9600BB" type="ISSN">1464-5262</mods:identifier>
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<subSubSection id="0B8F650AC300871FFE381CBCFCD1FA21" box="[457,882,1438,1462]" pageId="58" pageNumber="3601" type="nomenclature">
<paragraph id="432A3681C300871FFE381CBCFCD1FA21" blockId="58.[165,1121,1438,1526]" box="[457,882,1438,1462]" pageId="58" pageNumber="3601">
<heading id="186281EDC300871FFE381CBCFCD1FA21" box="[457,882,1438,1462]" centered="true" fontSize="10" level="2" pageId="58" pageNumber="3601" reason="2">
<taxonomicName id="84954D02C300871FFE381CBCFCD1FA21" ID-CoL="5ZPZ3" authority="Tendeiro" authorityName="Tendeiro" authorityYear="1984" box="[457,882,1438,1462]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="58" pageNumber="3601" phylum="Arthropoda" rank="species" species="sphenurus">
<emphasis id="71E1EA93C300871FFE381CBCFD5DFA21" bold="true" box="[457,766,1438,1462]" italics="true" pageId="58" pageNumber="3601">Columbicola sphenurus</emphasis>
Tendeiro
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0B8F650AC300871EFDBF1C9CFB36FA6F" lastPageId="59" lastPageNumber="3602" pageId="58" pageNumber="3601" type="description">
<paragraph id="432A3681C300871FFDBF1C9CFBC2FA61" blockId="58.[165,1121,1438,1526]" pageId="58" pageNumber="3601">
(
<figureCitation id="DBAE2A04C300871FFDA71C9CFD76FA41" box="[598,725,1470,1494]" captionStart="Figures 165–189" captionStartId="64.[138,207,1455,1474]" captionTargetBox="[181,1141,133,1416]" captionTargetId="figure-1@64.[181,1143,133,1419]" captionTargetPageId="64" captionText="Figures 165–189. Columbicola davisae: (165) female ventral terminalia. C. insularis: (166) female dorsal head. C. elbeli: (167) male genitalia; (168) female ventral terminalia. C. phoenicopterae: (169) male genitalia; (170) female ventral terminalia. C. sphenurus: (171) male genitalia. C. wardi: (172) male genitalia; (173) female ventral terminalia. C. emersoni: (174) male dorsal head; (175) female dorsal head; (176) male genitalia; (177) female ventral terminalia. C. wecksteini: (178) male dorsal head; (179) female dorsal head; (180) male genitalia; (181) female ventral terminalia. C. curtus: (182) male dorsal head; (183) female dorsal head (missing antennae); (184) male genitalia. C. brygooi: (185) male genitalia; (186) female ventral terminalia. C. juliusriemeri: (187) male dorsal head; (188) male genitalia. C. obliteratus: (189) male dorsal head." figureDoi="http://doi.org/10.5281/zenodo.5220824" httpUri="https://zenodo.org/record/5220824/files/figure.png" pageId="58" pageNumber="3601">Figure 171</figureCitation>
)
<taxonomicName id="84954D02C300871FFF541CFCFD24FA61" authority="Tendeiro 1984: 92" authorityName="Tendeiro" authorityPageNumber="92" authorityYear="1984" box="[165,647,1502,1526]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="58" pageNumber="3601" phylum="Arthropoda" rank="species" species="sphenurus">
<emphasis id="71E1EA93C300871FFF541CFCFE3EFA61" box="[165,413,1502,1526]" italics="true" pageId="58" pageNumber="3601">Columbicola sphenurus</emphasis>
<bibRefCitation id="27044B70C300871FFE561CFCFD24FA61" author="Tendeiro J" box="[423,647,1502,1526]" pageId="58" pageNumber="3601" pagination="77 - 118" refId="ref39526" refString="Tendeiro J. 1984. Nouvelles etudes sur la systematique, la zoogeographie et l'ecologie du genre Columbicola Ewing, 1929 (Mallophaga, Ischnocera). Garcia de Orta, Serie Zoologia 11: 77 - 118." type="journal article" year="1984">Tendeiro 1984: 92</bibRefCitation>
</taxonomicName>
.
<typeStatus id="9C2E8823C300871FFD641CFCFD72FA61" box="[661,721,1502,1526]" pageId="58" pageNumber="3601">Type</typeStatus>
host:
<emphasis id="71E1EA93C300871FFCEA1CFCFCDEFA62" box="[795,893,1502,1525]" italics="true" pageId="58" pageNumber="3601">
<taxonomicName id="84954D02C300871FFCEA1CFCFCC0FA62" authorityName="Vieillot" authorityYear="1816" box="[795,867,1502,1525]" class="Aves" family="Columbidae" genus="Treron" kingdom="Animalia" order="Columbiformes" pageId="58" pageNumber="3601" phylum="Chordata" rank="genus">Treron</taxonomicName>
s.
</emphasis>
<taxonomicName id="84954D02C300871FFC771CFCFBFEFA61" authority="(Vigors)" baseAuthorityName="Vigors" box="[902,1117,1502,1526]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="58" pageNumber="3601" phylum="Arthropoda" rank="species" species="sphenurus">
<emphasis id="71E1EA93C300871FFC771CFCFC53FA61" box="[902,1008,1502,1526]" italics="true" pageId="58" pageNumber="3601">sphenurus</emphasis>
(Vigors)
</taxonomicName>
.
</paragraph>
<paragraph id="432A3681C300871FFF7B1F0FFEA0F9D2" blockId="58.[138,259,1581,1605]" box="[138,259,1581,1605]" pageId="58" pageNumber="3601">
<emphasis id="71E1EA93C300871FFF7B1F0FFEA0F9D2" box="[138,259,1581,1605]" italics="true" pageId="58" pageNumber="3601">Description</emphasis>
</paragraph>
<paragraph id="432A3681C300871EFF7B1F7FFBD2FEB7" blockId="58.[138,1185,1629,1685]" lastBlockId="59.[125,1172,136,288]" lastPageId="59" lastPageNumber="3602" pageId="58" pageNumber="3601">
Similar to
<taxonomicName id="84954D02C300871FFEFF1F7FFEC9F9E2" box="[270,362,1629,1653]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="58" pageNumber="3601" phylum="Arthropoda" rank="species" species="elbeli">
<emphasis id="71E1EA93C300871FFEFF1F7FFEC9F9E2" box="[270,362,1629,1653]" italics="true" pageId="58" pageNumber="3601">C. elbeli</emphasis>
</taxonomicName>
, differing in structure of male genitalia. Male with HW, 0.28–0.30 (0.290); HL, 0.55–0.59 (0.570); HL/HW, 1.96. Thorax with PW, 0.20–0.21 (0.207); MW, 0.28–0.32 (0.303). Genitalia as in
<figureCitation id="DBAE2A04C301871EFD7C19AAFCAFFF37" box="[653,780,136,160]" captionStart="Figures 165–189" captionStartId="64.[138,207,1455,1474]" captionTargetBox="[181,1141,133,1416]" captionTargetId="figure-1@64.[181,1143,133,1419]" captionTargetPageId="64" captionText="Figures 165–189. Columbicola davisae: (165) female ventral terminalia. C. insularis: (166) female dorsal head. C. elbeli: (167) male genitalia; (168) female ventral terminalia. C. phoenicopterae: (169) male genitalia; (170) female ventral terminalia. C. sphenurus: (171) male genitalia. C. wardi: (172) male genitalia; (173) female ventral terminalia. C. emersoni: (174) male dorsal head; (175) female dorsal head; (176) male genitalia; (177) female ventral terminalia. C. wecksteini: (178) male dorsal head; (179) female dorsal head; (180) male genitalia; (181) female ventral terminalia. C. curtus: (182) male dorsal head; (183) female dorsal head (missing antennae); (184) male genitalia. C. brygooi: (185) male genitalia; (186) female ventral terminalia. C. juliusriemeri: (187) male dorsal head; (188) male genitalia. C. obliteratus: (189) male dorsal head." figureDoi="http://doi.org/10.5281/zenodo.5220824" httpUri="https://zenodo.org/record/5220824/files/figure.png" pageId="59" pageNumber="3602">Figure 171</figureCitation>
; parameres elongate, thickened anteriorly, with small lateral indentations; mesosome thick, ‘‘V’’-shaped, indented anteriorly; GW, 0.113. TL, 2.24–2.32 (2.29). Female with HW, 0.29–0.31 (0.303); HL, 0.58–0.60 (0.591); HL/HW, 1.87–2.00 (1.96). Thorax with PW, 0.20–0.21 (0.207); MW, 0.29–0.31 (0.309). Subgenital plate groove ovoid, edges rough. TL, 2.36–2.59 (2.48).
</paragraph>
<paragraph id="432A3681C301871EFF8C1875FF78FEF8" blockId="59.[125,219,343,367]" box="[125,219,343,367]" pageId="59" pageNumber="3602">
<emphasis id="71E1EA93C301871EFF8C1875FF78FEF8" box="[125,219,343,367]" italics="true" pageId="59" pageNumber="3602">Material</emphasis>
</paragraph>
<paragraph id="432A3681C301871EFF8C18A5FCFDFE08" blockId="59.[125,862,391,415]" box="[125,862,391,415]" pageId="59" pageNumber="3602">
<specimenCount id="5593FD08C301871EFF8C18A5FF6FFE08" box="[125,204,391,415]" pageId="59" pageNumber="3602" type="male">1 male</specimenCount>
<typeStatus id="9C2E8823C301871EFF2418A5FE9BFE08" box="[213,312,391,415]" pageId="59" pageNumber="3602" type="paratype">paratype</typeStatus>
of
<taxonomicName id="84954D02C301871EFE9018A5FE4CFE08" box="[353,495,391,415]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="sphenurus">
<emphasis id="71E1EA93C301871EFE9018A5FE4CFE08" box="[353,495,391,415]" italics="true" pageId="59" pageNumber="3602">C. sphenurus</emphasis>
</taxonomicName>
, ex
<taxonomicName id="84954D02C301871EFDD218AAFD13FE08" box="[547,688,391,415]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="sphenurus">
<emphasis id="71E1EA93C301871EFDD218AAFD13FE08" box="[547,688,391,415]" italics="true" pageId="59" pageNumber="3602">T. sphenurus</emphasis>
</taxonomicName>
,
<collectingCountry id="3B827611C301871EFD3018A5FC88FE08" box="[705,811,391,415]" name="Thailand" pageId="59" pageNumber="3602">Thailand</collectingCountry>
(1).
</paragraph>
<paragraph id="432A3681C301871EFF8C18F5FF7FFE78" blockId="59.[125,220,471,495]" box="[125,220,471,495]" pageId="59" pageNumber="3602">
<emphasis id="71E1EA93C301871EFF8C18F5FF7FFE78" box="[125,220,471,495]" italics="true" pageId="59" pageNumber="3602">Remarks</emphasis>
</paragraph>
<paragraph id="432A3681C301871EFF8C1B25FBF5FD29" blockId="59.[125,1173,519,702]" pageId="59" pageNumber="3602">
Several of the specimens used to describe
<taxonomicName id="84954D02C301871EFDAF1B25FD4AFD88" box="[606,745,519,543]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="sphenurus">
<emphasis id="71E1EA93C301871EFDAF1B25FD4AFD88" box="[606,745,519,543]" italics="true" pageId="59" pageNumber="3602">C. sphenurus</emphasis>
</taxonomicName>
were originally part of the
<typeStatus id="9C2E8823C301871EFBEC1B25FBEDFD88" box="[1053,1102,519,543]" pageId="59" pageNumber="3602">type</typeStatus>
series for
<taxonomicName id="84954D02C301871EFF581B04FECCFDA9" box="[169,367,550,574]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="elbeli">
<emphasis id="71E1EA93C301871EFF581B04FECCFDA9" box="[169,367,550,574]" italics="true" pageId="59" pageNumber="3602">Columbicola elbeli</emphasis>
</taxonomicName>
.
<taxonomicName id="84954D02C301871EFE731B04FDB1FDA9" box="[386,530,550,574]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="sphenurus">
<emphasis id="71E1EA93C301871EFE731B04FDB1FDA9" box="[386,530,550,574]" italics="true" pageId="59" pageNumber="3602">C. sphenurus</emphasis>
</taxonomicName>
was later recognized as a separate species because of differences in the male genitalia of lice from
<taxonomicName id="84954D02C301871EFD8A1B65FCB7FDC9" baseAuthorityName="J. F. Gmelin" baseAuthorityYear="1789" box="[635,788,582,606]" class="Aves" family="Columbidae" genus="Treron" kingdom="Animalia" order="Columbiformes" pageId="59" pageNumber="3602" phylum="Chordata" rank="species" species="pompadora">
<emphasis id="71E1EA93C301871EFD8A1B65FCB7FDC9" box="[635,788,582,606]" italics="true" pageId="59" pageNumber="3602">T. pompadora</emphasis>
</taxonomicName>
, the
<typeStatus id="9C2E8823C301871EFCBE1B64FC23FDC9" box="[847,896,582,606]" pageId="59" pageNumber="3602">type</typeStatus>
host for
<taxonomicName id="84954D02C301871EFC161B64FB9DFDC9" box="[999,1086,582,606]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="elbeli">
<emphasis id="71E1EA93C301871EFC161B64FB9DFDC9" box="[999,1086,582,606]" italics="true" pageId="59" pageNumber="3602">C. elbeli</emphasis>
</taxonomicName>
, versus lice from
<taxonomicName id="84954D02C301871EFF031B45FE22FDE9" box="[242,385,614,638]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="sphenurus">
<emphasis id="71E1EA93C301871EFF031B45FE22FDE9" box="[242,385,614,638]" italics="true" pageId="59" pageNumber="3602">T. sphenurus</emphasis>
</taxonomicName>
, the
<typeStatus id="9C2E8823C301871EFE321B44FE57FDE9" box="[451,500,614,638]" pageId="59" pageNumber="3602">type</typeStatus>
host of
<taxonomicName id="84954D02C301871EFDAC1B44FD4EFDE9" box="[605,749,614,638]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="sphenurus">
<emphasis id="71E1EA93C301871EFDAC1B44FD4EFDE9" box="[605,749,614,638]" italics="true" pageId="59" pageNumber="3602">C. sphenurus</emphasis>
</taxonomicName>
.
<bibRefCitation id="27044B70C301871EFCF11B44FC66FDE9" author="Tendeiro J" box="[768,965,614,638]" pageId="59" pageNumber="3602" pagination="77 - 118" refId="ref39526" refString="Tendeiro J. 1984. Nouvelles etudes sur la systematique, la zoogeographie et l'ecologie du genre Columbicola Ewing, 1929 (Mallophaga, Ischnocera). Garcia de Orta, Serie Zoologia 11: 77 - 118." type="journal article" year="1984">Tendeiro (1984)</bibRefCitation>
also recorded
<taxonomicName id="84954D02C301871EFB881B44FF44FD09" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="sphenurus">
<emphasis id="71E1EA93C301871EFB881B44FF44FD09" italics="true" pageId="59" pageNumber="3602">C. sphenurus</emphasis>
</taxonomicName>
from
<taxonomicName id="84954D02C301871EFECD1BA5FD98FD09" authority="Swinhoe." authorityName="Swinhoe." box="[316,571,646,670]" class="Aves" family="Columbidae" genus="Treron" kingdom="Animalia" order="Columbiformes" pageId="59" pageNumber="3602" phylum="Chordata" rank="species" species="formosae">
<emphasis id="71E1EA93C301871EFECD1BA5FE62FD09" box="[316,449,646,670]" italics="true" pageId="59" pageNumber="3602">T. formosae</emphasis>
Swinhoe.
</taxonomicName>
Measurements for the description above, except genitalic width, were taken from
<specimenCount id="5593FD08C301871EFDF01B84FD24FD29" box="[513,647,678,702]" pageId="59" pageNumber="3602" type="male">three males</specimenCount>
and
<specimenCount id="5593FD08C301871EFD351B84FCC0FD29" box="[708,867,678,702]" pageId="59" pageNumber="3602" type="female">seven females</specimenCount>
in
<bibRefCitation id="27044B70C301871EFC7A1B84FBF1FD29" author="Tendeiro J" box="[907,1106,678,702]" pageId="59" pageNumber="3602" pagination="77 - 118" refId="ref39526" refString="Tendeiro J. 1984. Nouvelles etudes sur la systematique, la zoogeographie et l'ecologie du genre Columbicola Ewing, 1929 (Mallophaga, Ischnocera). Garcia de Orta, Serie Zoologia 11: 77 - 118." type="journal article" year="1984">Tendeiro (1984)</bibRefCitation>
.
</paragraph>
<paragraph id="432A3681C301871EFE4D1BC1FC89FD6C" blockId="59.[152,1043,739,827]" box="[444,810,739,763]" pageId="59" pageNumber="3602">
<taxonomicName id="84954D02C301871EFE4D1BC1FC89FD6C" authority="Tendeiro" authorityName="Tendeiro" authorityYear="1965" box="[444,810,739,763]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="wardi">
<emphasis id="71E1EA93C301871EFE4D1BC1FD14FD6C" bold="true" box="[444,695,739,763]" italics="true" pageId="59" pageNumber="3602">Columbicola wardi</emphasis>
Tendeiro
</taxonomicName>
</paragraph>
<paragraph id="432A3681C301871EFE0E1A21FBB0FCAC" blockId="59.[152,1043,739,827]" pageId="59" pageNumber="3602">
(
<figureCitation id="DBAE2A04C301871EFDF71A21FD6EFC8C" box="[518,717,771,795]" captionStart="Figures 165–189" captionStartId="64.[138,207,1455,1474]" captionTargetBox="[181,1141,133,1416]" captionTargetId="figure-1@64.[181,1143,133,1419]" captionTargetPageId="64" captionText="Figures 165–189. Columbicola davisae: (165) female ventral terminalia. C. insularis: (166) female dorsal head. C. elbeli: (167) male genitalia; (168) female ventral terminalia. C. phoenicopterae: (169) male genitalia; (170) female ventral terminalia. C. sphenurus: (171) male genitalia. C. wardi: (172) male genitalia; (173) female ventral terminalia. C. emersoni: (174) male dorsal head; (175) female dorsal head; (176) male genitalia; (177) female ventral terminalia. C. wecksteini: (178) male dorsal head; (179) female dorsal head; (180) male genitalia; (181) female ventral terminalia. C. curtus: (182) male dorsal head; (183) female dorsal head (missing antennae); (184) male genitalia. C. brygooi: (185) male genitalia; (186) female ventral terminalia. C. juliusriemeri: (187) male dorsal head; (188) male genitalia. C. obliteratus: (189) male dorsal head." figureDoi="http://doi.org/10.5281/zenodo.5220824" httpUri="https://zenodo.org/record/5220824/files/figure.png" pageId="59" pageNumber="3602">Figures 172, 173</figureCitation>
)
<taxonomicName id="84954D02C301871EFF691A01FDFEFCAC" authority="Tendeiro 1965: 310" authorityName="Tendeiro" authorityPageNumber="310" authorityYear="1965" box="[152,605,803,827]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="wardi">
<emphasis id="71E1EA93C301871EFF691A01FEC6FCAC" box="[152,357,803,827]" italics="true" pageId="59" pageNumber="3602">Columbicola wardi</emphasis>
<bibRefCitation id="27044B70C301871EFE9F1A01FDFEFCAC" author="Tendeiro J" box="[366,605,803,827]" pageId="59" pageNumber="3602" pagination="1 - 460" refId="ref39197" refString="Tendeiro J. 1965. Estudos sobre Malofagos. Revisao monografica do genero Columbicola Ewing (Ischnocera, Philopteridae). Memorias da Junta de Investigacoes do Ultramar, No. 32 (1962). p 1 - 460." type="journal article" year="1965">Tendeiro 1965: 310</bibRefCitation>
</taxonomicName>
.
<typeStatus id="9C2E8823C301871EFD9B1A01FD05FCAC" box="[618,678,803,827]" pageId="59" pageNumber="3602">Type</typeStatus>
host:
<emphasis id="71E1EA93C301871EFD011A06FC66FCAC" box="[752,965,803,827]" italics="true" pageId="59" pageNumber="3602">
<taxonomicName id="84954D02C301871EFD011A06FC9BFCAC" authorityName="Vieillot" authorityYear="1816" box="[752,824,804,827]" class="Aves" family="Columbidae" genus="Treron" kingdom="Animalia" order="Columbiformes" pageId="59" pageNumber="3602" phylum="Chordata" rank="genus">Treron</taxonomicName>
a. apicauda
</emphasis>
Blyth.
</paragraph>
<paragraph id="432A3681C301871EFF8C1A50FF56FC1D" blockId="59.[125,245,882,906]" box="[125,245,882,906]" pageId="59" pageNumber="3602">
<emphasis id="71E1EA93C301871EFF8C1A50FF56FC1D" box="[125,245,882,906]" italics="true" pageId="59" pageNumber="3602">Description</emphasis>
</paragraph>
<paragraph id="432A3681C301871EFF8C1A80FB26FBED" blockId="59.[125,1173,930,1146]" pageId="59" pageNumber="3602">
<materialsCitation id="F3FD3CDCC301871EFF8C1A80FB22FBED" collectionCode="APW, HW, PW, MW" latitude="0.152" longLatPrecision="78" longitude="-0.157" pageId="59" pageNumber="3602" specimenCount="1">
Male head with
<collectionCode id="2584AE44C301871EFEDE1A80FED2FC2D" box="[303,369,930,954]" pageId="59" pageNumber="3602">APW</collectionCode>
,
<geoCoordinate id="26A15046C301871EFE881A80FE18FC2D" box="[377,443,930,954]" degrees="0.142" direction="north" orientation="latitude" pageId="59" pageNumber="3602" precision="55" value="0.142">0.142</geoCoordinate>
<geoCoordinate id="26A15046C301871EFE4A1A80FDA8FC2D" box="[443,523,930,954]" degrees="0.157" direction="west" orientation="longitude" pageId="59" pageNumber="3602" precision="55" value="-0.157">–0.157</geoCoordinate>
(0.149);
<collectionCode id="2584AE44C301871EFD801A80FD05FC2D" box="[625,678,930,954]" pageId="59" pageNumber="3602">HW</collectionCode>
, 0.31–0.32 (0.313); HL, 0.58–0.60 (0.593); HL/
<collectionCode id="2584AE44C301871EFF5E1AE0FF41FC4D" box="[175,226,962,986]" pageId="59" pageNumber="3602">HW</collectionCode>
, 1.87–1.94 (1.89); medioposterior setae short, not reaching halfway to posterior head margin. Thorax with
<collectionCode id="2584AE44C301871EFE781AC0FE1BFC6D" box="[393,440,994,1018]" country="Thailand" name="Paleontological Collections" pageId="59" pageNumber="3602">PW</collectionCode>
, 0.25;
<collectionCode id="2584AE44C301871EFDE51AC0FDEEFC6D" box="[532,589,994,1018]" country="Netherlands" name="Museum Wasmann" pageId="59" pageNumber="3602">MW</collectionCode>
, 0.33–0.35 (0.333). Genitalia as in
<figureCitation id="DBAE2A04C301871EFBE01AC0FB33FC6D" box="[1041,1168,994,1018]" captionStart="Figures 165–189" captionStartId="64.[138,207,1455,1474]" captionTargetBox="[181,1141,133,1416]" captionTargetId="figure-1@64.[181,1143,133,1419]" captionTargetPageId="64" captionText="Figures 165–189. Columbicola davisae: (165) female ventral terminalia. C. insularis: (166) female dorsal head. C. elbeli: (167) male genitalia; (168) female ventral terminalia. C. phoenicopterae: (169) male genitalia; (170) female ventral terminalia. C. sphenurus: (171) male genitalia. C. wardi: (172) male genitalia; (173) female ventral terminalia. C. emersoni: (174) male dorsal head; (175) female dorsal head; (176) male genitalia; (177) female ventral terminalia. C. wecksteini: (178) male dorsal head; (179) female dorsal head; (180) male genitalia; (181) female ventral terminalia. C. curtus: (182) male dorsal head; (183) female dorsal head (missing antennae); (184) male genitalia. C. brygooi: (185) male genitalia; (186) female ventral terminalia. C. juliusriemeri: (187) male dorsal head; (188) male genitalia. C. obliteratus: (189) male dorsal head." figureDoi="http://doi.org/10.5281/zenodo.5220824" httpUri="https://zenodo.org/record/5220824/files/figure.png" pageId="59" pageNumber="3602">Figure 172</figureCitation>
; mesosome weakly indented anteriorly, laterally sclerotized; GW,
<geoCoordinate id="26A15046C301871EFC991D20FC0BFB8D" box="[872,936,1026,1050]" degrees="0.113" direction="north" orientation="latitude" pageId="59" pageNumber="3602" precision="55" value="0.113">0.113</geoCoordinate>
<geoCoordinate id="26A15046C301871EFC591D20FC5AFB8D" box="[936,1017,1026,1050]" degrees="0.122" direction="west" orientation="longitude" pageId="59" pageNumber="3602" precision="55" value="-0.122">–0.122</geoCoordinate>
(0.119). TL, 2.21–2.30 (2.26). Female with
<collectionCode id="2584AE44C301871EFE011D00FD91FBAD" box="[496,562,1058,1082]" pageId="59" pageNumber="3602">APW</collectionCode>
,
<geoCoordinate id="26A15046C301871EFDB11D00FD20FBAD" box="[576,643,1058,1082]" degrees="0.152" direction="north" orientation="latitude" pageId="59" pageNumber="3602" precision="55" value="0.152">0.152</geoCoordinate>
<geoCoordinate id="26A15046C301871EFD721D00FD77FBAD" box="[643,724,1058,1082]" degrees="0.157" direction="west" orientation="longitude" pageId="59" pageNumber="3602" precision="55" value="-0.157">–0.157</geoCoordinate>
;
<collectionCode id="2584AE44C301871EFD121D00FCBBFBAD" box="[739,792,1058,1082]" pageId="59" pageNumber="3602">HW</collectionCode>
, 0.33; HL, 0.61–0.63; HL/
<collectionCode id="2584AE44C301871EFBAC1D00FB33FBAD" box="[1117,1168,1058,1082]" pageId="59" pageNumber="3602">HW</collectionCode>
, 1.85–1.91. Thorax with
<collectionCode id="2584AE44C301871EFE511D60FE6CFBCD" box="[416,463,1090,1114]" country="Thailand" name="Paleontological Collections" pageId="59" pageNumber="3602">PW</collectionCode>
, 0.25;
<collectionCode id="2584AE44C301871EFDD21D60FDFFFBCD" box="[547,604,1090,1114]" country="Netherlands" name="Museum Wasmann" pageId="59" pageNumber="3602">MW</collectionCode>
, 0.34–0.37. Ventral terminalia as in
<figureCitation id="DBAE2A04C301871EFBE01D60FB33FBCD" box="[1041,1168,1090,1114]" captionStart="Figures 165–189" captionStartId="64.[138,207,1455,1474]" captionTargetBox="[181,1141,133,1416]" captionTargetId="figure-1@64.[181,1143,133,1419]" captionTargetPageId="64" captionText="Figures 165–189. Columbicola davisae: (165) female ventral terminalia. C. insularis: (166) female dorsal head. C. elbeli: (167) male genitalia; (168) female ventral terminalia. C. phoenicopterae: (169) male genitalia; (170) female ventral terminalia. C. sphenurus: (171) male genitalia. C. wardi: (172) male genitalia; (173) female ventral terminalia. C. emersoni: (174) male dorsal head; (175) female dorsal head; (176) male genitalia; (177) female ventral terminalia. C. wecksteini: (178) male dorsal head; (179) female dorsal head; (180) male genitalia; (181) female ventral terminalia. C. curtus: (182) male dorsal head; (183) female dorsal head (missing antennae); (184) male genitalia. C. brygooi: (185) male genitalia; (186) female ventral terminalia. C. juliusriemeri: (187) male dorsal head; (188) male genitalia. C. obliteratus: (189) male dorsal head." figureDoi="http://doi.org/10.5281/zenodo.5220824" httpUri="https://zenodo.org/record/5220824/files/figure.png" pageId="59" pageNumber="3602">Figure 173</figureCitation>
; subgenital plate groove rounded anteriorly, distinct expansions posteriorly. TL, 2.38–2.55
</materialsCitation>
.
</paragraph>
<paragraph id="432A3681C301871EFF8C1D93FF47FB5E" blockId="59.[125,228,1201,1225]" box="[125,228,1201,1225]" pageId="59" pageNumber="3602">
<emphasis id="71E1EA93C301871EFF8C1D93FF47FB5E" box="[125,228,1201,1225]" italics="true" pageId="59" pageNumber="3602">Materials</emphasis>
</paragraph>
<paragraph id="432A3681C301871EFF8C1DC3FEB9FA8E" blockId="59.[125,1172,1249,1305]" pageId="59" pageNumber="3602">
<materialsCitation id="F3FD3CDCC301871EFF8C1DC3FF44FA8E" collectionCode="T" country="Thailand" location="Thailand" pageId="59" pageNumber="3602" specimenCount="9" specimenCount-female="4" specimenCount-male="5" typeStatus="paratype">
<specimenCount id="5593FD08C301871EFF8C1DC3FF79FB6E" box="[125,218,1249,1273]" pageId="59" pageNumber="3602" type="male">3 males</specimenCount>
,
<specimenCount id="5593FD08C301871EFF191DC3FEF4FB6E" box="[232,343,1249,1273]" pageId="59" pageNumber="3602" type="female">2 females</specimenCount>
(including
<specimenCount id="5593FD08C301871EFE111DC3FD90FB6E" box="[480,563,1249,1273]" pageId="59" pageNumber="3602" type="male">2 male</specimenCount>
,
<specimenCount id="5593FD08C301871EFDB01DC3FD06FB6E" box="[577,677,1249,1273]" pageId="59" pageNumber="3602" type="female">2 female</specimenCount>
<typeStatus id="9C2E8823C301871EFD5F1DC3FCBFFB6E" box="[686,796,1249,1273]" pageId="59" pageNumber="3602" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="84954D02C301871EFCB71DC3FC0BFB6E" authorityName="Tendeiro" authorityYear="1965" box="[838,936,1249,1273]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="wardi">
<emphasis id="71E1EA93C301871EFCB71DC3FC0BFB6E" box="[838,936,1249,1273]" italics="true" pageId="59" pageNumber="3602">C. wardi</emphasis>
</taxonomicName>
), ex
<emphasis id="71E1EA93C301871EFC171DC0FB2FFB6E" box="[998,1164,1249,1273]" italics="true" pageId="59" pageNumber="3602">
<collectionCode id="2584AE44C301871EFC171DC0FC5FFB6E" box="[998,1020,1250,1273]" name="Tavera, Department of Geology and Geophysics" pageId="59" pageNumber="3602">T</collectionCode>
. a. apicauda
</emphasis>
,
<collectingCountry id="3B827611C301871EFF8C1C23FF44FA8E" box="[125,231,1281,1305]" name="Thailand" pageId="59" pageNumber="3602">Thailand</collectingCountry>
</materialsCitation>
(2).
</paragraph>
<paragraph id="432A3681C301871EFF8C1C73FF7FFAFE" blockId="59.[125,220,1361,1385]" box="[125,220,1361,1385]" pageId="59" pageNumber="3602">
<emphasis id="71E1EA93C301871EFF8C1C73FF7FFAFE" box="[125,220,1361,1385]" italics="true" pageId="59" pageNumber="3602">Remarks</emphasis>
</paragraph>
<paragraph id="432A3681C301871EFF8C1CA3FB36FA6F" blockId="59.[125,1173,1409,1528]" pageId="59" pageNumber="3602">
While superficially resembling
<taxonomicName id="84954D02C301871EFE191CA3FDE1FA0E" box="[488,578,1409,1433]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="elbeli">
<emphasis id="71E1EA93C301871EFE191CA3FDE1FA0E" box="[488,578,1409,1433]" italics="true" pageId="59" pageNumber="3602">C. elbeli</emphasis>
</taxonomicName>
, the male mesosomal and female subgenital plate structures distinguish
<taxonomicName id="84954D02C301871EFE841C82FE71FA2F" authorityName="Tendeiro" authorityYear="1965" box="[373,466,1440,1464]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="wardi">
<emphasis id="71E1EA93C301871EFE841C82FE71FA2F" box="[373,466,1440,1464]" italics="true" pageId="59" pageNumber="3602">C. wardi</emphasis>
</taxonomicName>
. It is interesting to note that, upon re-examining the
<typeStatus id="9C2E8823C301871EFBD11C82FBF2FA2F" box="[1056,1105,1440,1464]" pageId="59" pageNumber="3602">type</typeStatus>
series of
<emphasis id="71E1EA93C301871EFF511CE2FF79FA4F" box="[160,218,1472,1496]" italics="true" pageId="59" pageNumber="3602">C. e.</emphasis>
<taxonomicName id="84954D02C301871EFF171CE2FE57FA4F" authority=", Tendeiro (1984)" authorityName=", Tendeiro" authorityYear="1984" box="[230,500,1472,1496]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="elbeli">
<emphasis id="71E1EA93C301871EFF171CE2FEB9FA4F" box="[230,282,1472,1496]" italics="true" pageId="59" pageNumber="3602">elbeli</emphasis>
,
<bibRefCitation id="27044B70C301871EFEDF1CE2FE57FA4F" author="Tendeiro J" box="[302,500,1472,1496]" pageId="59" pageNumber="3602" pagination="77 - 118" refId="ref39526" refString="Tendeiro J. 1984. Nouvelles etudes sur la systematique, la zoogeographie et l'ecologie du genre Columbicola Ewing, 1929 (Mallophaga, Ischnocera). Garcia de Orta, Serie Zoologia 11: 77 - 118." type="journal article" year="1984">Tendeiro (1984)</bibRefCitation>
</taxonomicName>
discovered that the male specimen from
<taxonomicName id="84954D02C301871EFC1C1CE3FEB3FA6F" authority="(Temminck)" baseAuthorityName="Temminck" baseAuthorityYear="1823" class="Aves" family="Columbidae" genus="Treron" kingdom="Animalia" order="Columbiformes" pageId="59" pageNumber="3602" phylum="Chordata" rank="species" species="oxyurus">
<emphasis id="71E1EA93C301871EFC1C1CE3FB37FA4F" box="[1005,1172,1473,1496]" italics="true" pageId="59" pageNumber="3602">Treron oxyurus</emphasis>
(Temminck)
</taxonomicName>
was actually identical to
<taxonomicName id="84954D02C301871EFDD51CC2FD22FA6F" authorityName="Tendeiro" authorityYear="1965" box="[548,641,1504,1528]" class="Insecta" family="Philopteridae" genus="Columbicola" kingdom="Animalia" order="Psocodea" pageId="59" pageNumber="3602" phylum="Arthropoda" rank="species" species="wardi">
<emphasis id="71E1EA93C301871EFDD51CC2FD22FA6F" box="[548,641,1504,1528]" italics="true" pageId="59" pageNumber="3602">C. wardi</emphasis>
</taxonomicName>
, which he had described nearly 20 years earlier.
</paragraph>
</subSubSection>
</treatment>
</document>
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