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<document id="DBA2C1ADC5D361C0666A4D5F60E43E9B" ID-DOI="10.5281/zenodo.13244981" ID-ISSN="2345-7600" ID-Zenodo-Dep="13244981" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722970368039" checkinUser="felipe" docAuthor="Stuart Ko Sok Thy Neang, Bryan L." docDate="2006" docId="CB2487A5FFFB69032C95C77EF935F16F" docLanguage="en" docName="RafflesBZool.54.1.129-155.pdf" docOrigin="Raffles Bulletin of Zoology 54 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Rana attigua Inger, Orlov &amp; Darevsky 1999" docType="treatment" docVersion="2" lastPageNumber="140" masterDocId="371DFFDDFFF169082F30C000FB36F417" masterDocTitle="A Collection Of Amphibians And Reptiles From Hilly Eastern Cambodia" masterLastPageNumber="155" masterPageNumber="129" pageNumber="139" updateTime="1722975809971" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="210EB07D40DC8341571B80D1E056ABC9">
<mods:title id="DC6C4BDAB0D6A0D132A4203D617F74F7">A Collection Of Amphibians And Reptiles From Hilly Eastern Cambodia</mods:title>
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<mods:namePart id="430230C0AB3F34E8D6EF2D23EF0FEB93">Stuart Ko Sok Thy Neang, Bryan L.</mods:namePart>
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<mods:title id="F17D0A4DEB576D89A9EA147695EB9BE6">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="D7AAB114368B7BAF6562628D6D862E13">2006</mods:date>
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<mods:number id="8678A1AD58E802FDDA50CF0B41663CE3">2006-02-28</mods:number>
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<mods:number id="89424560358604A1BE72EE3F33917854">54</mods:number>
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<mods:identifier id="2890E07258356C2FD8FDC47C5120A27D" type="DOI">10.5281/zenodo.13244981</mods:identifier>
<mods:identifier id="FDF84790A1675318C4C5FC463354AC61" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="F47FEF8ED5928869E8442C22EC6A6620" type="Zenodo-Dep">13244981</mods:identifier>
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<treatment id="CB2487A5FFFB69032C95C77EF935F16F" LSID="urn:lsid:plazi:treatment:CB2487A5FFFB69032C95C77EF935F16F" httpUri="http://treatment.plazi.org/id/CB2487A5FFFB69032C95C77EF935F16F" lastPageId="11" lastPageNumber="140" pageId="10" pageNumber="139">
<subSubSection id="0B976538FFFB69022C95C77EFE69F382" box="[933,1375,1917,1941]" pageId="10" pageNumber="139" type="nomenclature">
<paragraph id="433236B3FFFB69022C95C77EFE69F382" blockId="10.[933,1375,1917,1941]" box="[933,1375,1917,1941]" pageId="10" pageNumber="139">
<heading id="187A81DFFFFB69022C95C77EFE69F382" bold="true" box="[933,1375,1917,1941]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="139" reason="2">
<taxonomicName id="848D4D30FFFB69022C95C77EFE69F382" ID-CoL="4RDHP" authority="Inger, Orlov &amp; Darevsky" authorityName="Inger, Orlov &amp; Darevsky" authorityYear="1999" box="[933,1375,1917,1941]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="10" pageNumber="139" phylum="Chordata" rank="species" species="attigua">
<emphasis id="71F9EAA1FFFB69022C95C77EFE69F382" bold="true" box="[933,1375,1917,1941]" pageId="10" pageNumber="139">
<emphasis id="71F9EAA1FFFB69022C95C77EFF03F382" bold="true" box="[933,1077,1917,1941]" italics="true" pageId="10" pageNumber="139">Rana attigua</emphasis>
Inger, Orlov &amp; Darevsky
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0B976538FFFB69022C73C7BDFFFEFC51" pageId="10" pageNumber="139" type="materials_examined">
<paragraph id="433236B3FFFB69022C73C7BDFFFEFC51" blockId="10.[835,1474,1981,2118]" pageId="10" pageNumber="139">
<emphasis id="71F9EAA1FFFB69022C73C7BDFF2CF3C4" bold="true" box="[835,1050,1981,2003]" italics="true" pageId="10" pageNumber="139">Material examined. –</emphasis>
<materialsCitation id="F3E53CEEFFFB69022B12C7BDFE46FC1B" collectingDate="2000-06-21" collectionCode="FMNH" elevation="650" latitude="12.537778" location="Pichrada" longLatPrecision="21" longitude="107.53333" municipality="Phnom Nam Lyr Mountain" pageId="10" pageNumber="139" specimenCode="FMNH 261920-22" specimenCount="1">
<location id="46526068FFFB69022B12C7BDFF4CF3C4" LSID="urn:lsid:plazi:treatment:CB2487A5FFFB69032C95C77EF935F16F:46526068FFFB69022B12C7BDFF4CF3C4" box="[1058,1146,1981,2003]" latitude="12.537778" longLatPrecision="21" longitude="107.53333" municipality="Phnom Nam Lyr Mountain" name="Pichrada" pageId="10" pageNumber="139">Pichrada</location>
:
<specimenCode id="132B9EC8FFFB69022BB4C7BEFE0DF3C4" box="[1156,1339,1981,2003]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="139" type="Museum">FMNH 261920-22</specimenCode>
,
<collectingMunicipality id="A356ACC9FFFB69022A77C7BDF8FAF3E7" pageId="10" pageNumber="139">Phnom Nam Lyr Mountain</collectingMunicipality>
, evergreen mixed with deciduous and bamboo, near
<geoCoordinate id="26B95074FFFB69022C73C7F7F882FC1A" box="[835,948,2039,2061]" degrees="12" direction="north" minutes="32" orientation="latitude" pageId="10" pageNumber="139" precision="15" seconds="16" value="12.537778">12°32'16&quot;N</geoCoordinate>
<geoCoordinate id="26B95074FFFB69022C8CC7F7FF0EFC1B" box="[956,1080,2039,2060]" degrees="107" direction="east" minutes="32" orientation="longitude" pageId="10" pageNumber="139" precision="15" seconds="00" value="107.53333">107°32'00&quot;E</geoCoordinate>
,
<quantity id="84759B56FFFB69022B75C7F7FF87FC1B" box="[1093,1201,2039,2061]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="10" pageNumber="139" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">
<elevation id="C8A0D180FFFB69022B75C7F7FF87FC1B" box="[1093,1201,2039,2061]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="10" pageNumber="139" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">600-700 m</elevation>
</quantity>
elev.,
<date id="37331073FFFB69022BC5C7F7FE46FC1B" box="[1269,1392,2039,2060]" pageId="10" pageNumber="139" value="2000-06-21">
<collectingDate id="2777E99BFFFB69022BC5C7F7FE46FC1B" box="[1269,1392,2039,2060]" pageId="10" pageNumber="139" value="2000-06-21">21 Jun.2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="F3E53CEEFFFB69022A4CC7F7FFF3FC51" collectingDate="2000-06-22" collectionCode="FMNH" elevation="700" latitude="12.496944" longLatPrecision="21" longitude="107.49249" pageId="10" pageNumber="139" specimenCode="FMNH 261923" specimenCount="1">
<specimenCode id="132B9EC8FFFB69022A4CC7F7F8B8FC3D" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="139" type="Museum">FMNH 261923</specimenCode>
, gallery evergreen forest along stream, near
<geoCoordinate id="26B95074FFFB69022A60C814FEF4FC3E" box="[1360,1474,2068,2089]" degrees="12" direction="north" minutes="29" orientation="latitude" pageId="10" pageNumber="139" precision="15" seconds="49" value="12.496944">12°29'49&quot;N</geoCoordinate>
<geoCoordinate id="26B95074FFFB69022C73C831F8F6FC51" box="[835,960,2097,2118]" degrees="107" direction="east" minutes="29" orientation="longitude" pageId="10" pageNumber="139" precision="15" seconds="33" value="107.49249">107°29'33&quot;E</geoCoordinate>
,
<quantity id="84759B56FFFB69022CFBC831FF3FFC51" box="[971,1033,2097,2118]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="139" unit="m" value="700.0">
<elevation id="C8A0D180FFFB69022CFBC831FF3FFC51" box="[971,1033,2097,2118]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="139" unit="m" value="700.0">700 m</elevation>
</quantity>
elev.,
<date id="37331073FFFB69022B7BC831FFF3FC51" box="[1099,1221,2097,2118]" pageId="10" pageNumber="139" value="2000-06-22">
<collectingDate id="2777E99BFFFB69022B7BC831FFF3FC51" box="[1099,1221,2097,2118]" pageId="10" pageNumber="139" value="2000-06-22">22 Jun.2000</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="17F2663BFFFB69022FB8C810F9DFFC54" ID-DOI="http://doi.org/10.5281/zenodo.13244994" ID-Zenodo-Dep="13244994" httpUri="https://zenodo.org/record/13244994/files/figure.png" pageId="10" pageNumber="139" startId="10.[136,173,2064,2086]" targetBox="[136,776,1635,2048]" targetPageId="10" targetType="figure">
<paragraph id="433236B3FFFB69022FB8C810F9DFFC54" blockId="10.[136,774,2064,2115]" pageId="10" pageNumber="139">
Fig. 8.
<taxonomicName id="848D4D30FFFB69022FE8C811FA98FC32" baseAuthorityName="Bourret" baseAuthorityYear="1942" box="[216,430,2064,2086]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="139" phylum="Chordata" rank="species" species="poilani">
<emphasis id="71F9EAA1FFFB69022FE8C811FA98FC32" box="[216,430,2064,2086]" italics="true" pageId="10" pageNumber="139">Limnonectes poilani</emphasis>
</taxonomicName>
from Phnom Nam Lyr Wildlife Sanctuary, Pichrada District, Mondolkiri Province, Cambodia.
</paragraph>
</caption>
<subSubSection id="0B976538FFFA69032F41C0C0F935F16F" pageId="11" pageNumber="140" type="discussion">
<paragraph id="433236B3FFFA69032F41C0C0F9D4F08F" blockId="11.[113,753,192,1176]" pageId="11" pageNumber="140">
<emphasis id="71F9EAA1FFFA69032F41C0C0FBDBF4CF" bold="true" box="[113,237,192,216]" italics="true" pageId="11" pageNumber="140">Remarks. –</emphasis>
<specimenCount id="558BFD3AFFFA69032FC4C0C1FA5DF4CF" box="[244,363,192,216]" count="4" pageId="11" pageNumber="140" type="male">Four males</specimenCount>
with nuptial pads (SVL 34.0 – 36.6, mean ± SD 35.5 ± 1.3, N = 4) agree with the original description (
<bibRefCitation id="271C4B42FFFA69032FC8C100FA8CF50F" author="Inger, R" box="[248,442,256,280]" pageId="11" pageNumber="140" pagination="1 - 46" refId="ref25910" refString="Inger, R. F., N. Orlov, &amp; I. Darevsky, 1999. Frogs of Vietnam: a report on new collections. Fieldiana Zoology (New Series), 92: 1 - 46." type="journal article" year="1999">Inger et al., 1999</bibRefCitation>
) and the FMNH
<typeStatus id="9C368811FFFA69032D4BC101F99FF50E" box="[635,681,257,281]" pageId="11" pageNumber="140">type</typeStatus>
series from
<collectingRegion id="8149F851FFFA69032F99C120FA6CF52F" box="[169,346,288,312]" country="Vietnam" name="Gia Lai" pageId="11" pageNumber="140">Gia Lai Province</collectingRegion>
,
<collectingCountry id="3B9A7623FFFA69032E53C120FAF6F52F" box="[355,448,288,312]" name="Vietnam" pageId="11" pageNumber="140">Vietnam</collectingCountry>
, except the Cambodian males are slightly smaller than males with nuptial pads in the
<typeStatus id="9C368811FFFA69032DF2C141F9C6F54E" box="[706,752,321,345]" pageId="11" pageNumber="140">type</typeStatus>
series (SVL 36.2 – 45.5, mean ± SE 41.3 ± 0.3, N = 43;
<bibRefCitation id="271C4B42FFFA69032D87C161FBDFF58F" author="Inger, R" pageId="11" pageNumber="140" pagination="1 - 46" refId="ref25910" refString="Inger, R. F., N. Orlov, &amp; I. Darevsky, 1999. Frogs of Vietnam: a report on new collections. Fieldiana Zoology (New Series), 92: 1 - 46." type="journal article" year="1999">Inger et al., 1999</bibRefCitation>
), and the nuptial pads have a distinct notch in the ventral margin. These discrepancies are noteworthy because two of the characteristics used by
<bibRefCitation id="271C4B42FFFA69032ECEC1C1F9E6F5CF" author="Inger, R" box="[510,720,448,473]" pageId="11" pageNumber="140" pagination="1 - 46" refId="ref25910" refString="Inger, R. F., N. Orlov, &amp; I. Darevsky, 1999. Frogs of Vietnam: a report on new collections. Fieldiana Zoology (New Series), 92: 1 - 46." type="journal article" year="1999">Inger et al. (1999)</bibRefCitation>
to distinguish
<taxonomicName id="848D4D30FFFA69032FD9C1E1FA79F5EF" authorityName="Inger, Orlov &amp; Darevsky" authorityYear="1999" box="[233,335,481,504]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="attigua">
<emphasis id="71F9EAA1FFFA69032FD9C1E1FA79F5EF" box="[233,335,481,504]" italics="true" pageId="11" pageNumber="140">R. attigua</emphasis>
</taxonomicName>
from the similar, co-occurring
<taxonomicName id="848D4D30FFFA69032DA1C1E1FBC0F60F" authority="Smith, 1921" authorityName="Smith" authorityYear="1921" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="milleti">
<emphasis id="71F9EAA1FFFA69032DA1C1E1F9C6F5EF" box="[657,752,480,504]" italics="true" pageId="11" pageNumber="140">R. milleti</emphasis>
<bibRefCitation id="271C4B42FFFA69032F41C200FBC0F60F" author="Smith, M" box="[113,246,512,536]" pageId="11" pageNumber="140" pagination="423 - 440" refId="ref27080" refString="Smith, M. A., 1921. New or little-known reptiles and batrachians from southern Annam (Indo-China). Proceedings of the Zoological Society of London, 1921: 423 - 440." type="book chapter" year="1921">Smith, 1921</bibRefCitation>
</taxonomicName>
, at the
<typeStatus id="9C368811FFFA69032E74C201FA44F60E" box="[324,370,513,537]" pageId="11" pageNumber="140">type</typeStatus>
locality in
<collectingCountry id="3B9A7623FFFA69032EDCC200F97EF60F" box="[492,584,512,536]" name="Vietnam" pageId="11" pageNumber="140">Vietnam</collectingCountry>
were body size and condition of the nuptial pad. Specifically,
<bibRefCitation id="271C4B42FFFA69032D46C221FB8EF64F" author="Inger, R" pageId="11" pageNumber="140" pagination="1 - 46" refId="ref25910" refString="Inger, R. F., N. Orlov, &amp; I. Darevsky, 1999. Frogs of Vietnam: a report on new collections. Fieldiana Zoology (New Series), 92: 1 - 46." type="journal article" year="1999">Inger et al. (1999)</bibRefCitation>
reported that
<taxonomicName id="848D4D30FFFA69032E7FC241FA87F64F" authorityName="Smith" authorityYear="1921" box="[335,433,576,600]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="milleti">
<emphasis id="71F9EAA1FFFA69032E7FC241FA87F64F" box="[335,433,576,600]" italics="true" pageId="11" pageNumber="140">R. milleti</emphasis>
</taxonomicName>
males with nuptial pads have SVL 34.2 – 40.4 (mean ± SE 37.0 ± 0.2, N = 52), but only four of
<specimenCount id="558BFD3AFFFA69032F8DC280FA27F68F" box="[189,273,640,664]" count="43" pageId="11" pageNumber="140" type="male">43 male</specimenCount>
<taxonomicName id="848D4D30FFFA69032E26C281FA4BF68F" authorityName="Inger, Orlov &amp; Darevsky" authorityYear="1999" box="[278,381,641,664]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="attigua">
<emphasis id="71F9EAA1FFFA69032E26C281FA4BF68F" box="[278,381,641,664]" italics="true" pageId="11" pageNumber="140">R. attigua</emphasis>
</taxonomicName>
were less than 39.0, and the nuptial pad in
<taxonomicName id="848D4D30FFFA69032F8FC2A1FA15F6AF" authorityName="Smith" authorityYear="1921" box="[191,291,672,696]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="milleti">
<emphasis id="71F9EAA1FFFA69032F8FC2A1FA15F6AF" box="[191,291,672,696]" italics="true" pageId="11" pageNumber="140">R. milleti</emphasis>
</taxonomicName>
has a distinct notch or constriction in the ventral margin, but that of
<taxonomicName id="848D4D30FFFA69032E98C2C1F920F6CF" authorityName="Inger, Orlov &amp; Darevsky" authorityYear="1999" box="[424,534,705,728]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="attigua">
<emphasis id="71F9EAA1FFFA69032E98C2C1F920F6CF" box="[424,534,705,728]" italics="true" pageId="11" pageNumber="140">R. attigua</emphasis>
</taxonomicName>
is straight. We reexamined specimens assigned to
<taxonomicName id="848D4D30FFFA69032EE2C2E1F902F6EF" authorityName="Smith" authorityYear="1921" box="[466,564,736,760]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="milleti">
<emphasis id="71F9EAA1FFFA69032EE2C2E1F902F6EF" box="[466,564,736,760]" italics="true" pageId="11" pageNumber="140">R. milleti</emphasis>
</taxonomicName>
(FMNH 253425- 86) by
<bibRefCitation id="271C4B42FFFA69032FF0C301FABFF70F" author="Inger, R" box="[192,393,768,793]" pageId="11" pageNumber="140" pagination="1 - 46" refId="ref25910" refString="Inger, R. F., N. Orlov, &amp; I. Darevsky, 1999. Frogs of Vietnam: a report on new collections. Fieldiana Zoology (New Series), 92: 1 - 46." type="journal article" year="1999">Inger et al. (1999)</bibRefCitation>
, and only about half of the adult males have a notch or constriction in the ventral margin of the nuptial pad, rendering this character to be non-diagnostic. The most conspicuous differences in preservative between
<taxonomicName id="848D4D30FFFA69032F41C381FBEFF78F" authorityName="Inger, Orlov &amp; Darevsky" authorityYear="1999" box="[113,217,897,920]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="attigua">
<emphasis id="71F9EAA1FFFA69032F41C381FBEFF78F" box="[113,217,897,920]" italics="true" pageId="11" pageNumber="140">R. attigua</emphasis>
</taxonomicName>
and
<taxonomicName id="848D4D30FFFA69032E3AC381FA5DF78F" authorityName="Smith" authorityYear="1921" box="[266,363,896,920]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="milleti">
<emphasis id="71F9EAA1FFFA69032E3AC381FA5DF78F" box="[266,363,896,920]" italics="true" pageId="11" pageNumber="140">R. milleti</emphasis>
</taxonomicName>
are those of colouration, as stated by
<bibRefCitation id="271C4B42FFFA69032F41C3A1FA00F7AF" author="Inger, R" box="[113,310,928,953]" pageId="11" pageNumber="140" pagination="1 - 46" refId="ref25910" refString="Inger, R. F., N. Orlov, &amp; I. Darevsky, 1999. Frogs of Vietnam: a report on new collections. Fieldiana Zoology (New Series), 92: 1 - 46." type="journal article" year="1999">Inger et al. (1999)</bibRefCitation>
: the underside of the thigh is immaculate whitish and the underside of the calf is immaculate or with only faint dark speckling in
<taxonomicName id="848D4D30FFFA69032E91C3E1F93DF7EF" authorityName="Inger, Orlov &amp; Darevsky" authorityYear="1999" box="[417,523,993,1016]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="attigua">
<emphasis id="71F9EAA1FFFA69032E91C3E1F93DF7EF" box="[417,523,993,1016]" italics="true" pageId="11" pageNumber="140">R. attigua</emphasis>
</taxonomicName>
, but the underside of the thigh has dark speckling and the underside of the calf has dark spotting in
<taxonomicName id="848D4D30FFFA69032E2FC421FAB7F02F" authorityName="Smith" authorityYear="1921" box="[287,385,1056,1080]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="milleti">
<emphasis id="71F9EAA1FFFA69032E2FC421FAB7F02F" box="[287,385,1056,1080]" italics="true" pageId="11" pageNumber="140">R. milleti</emphasis>
</taxonomicName>
; and the rear of the thigh has only a light dusting of dark pigment in
<taxonomicName id="848D4D30FFFA69032ED0C441F97CF04F" authorityName="Inger, Orlov &amp; Darevsky" authorityYear="1999" box="[480,586,1089,1112]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="attigua">
<emphasis id="71F9EAA1FFFA69032ED0C441F97CF04F" box="[480,586,1089,1112]" italics="true" pageId="11" pageNumber="140">R. attigua</emphasis>
</taxonomicName>
, but the rear of the thigh has a distinct pattern of dark brown forming a network around small light spots or blotches in
<taxonomicName id="848D4D30FFFA69032D48C481F9EDF08F" authorityName="Smith" authorityYear="1921" box="[632,731,1152,1176]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="milleti">
<emphasis id="71F9EAA1FFFA69032D48C481F9EDF08F" box="[632,731,1152,1176]" italics="true" pageId="11" pageNumber="140">R. milleti</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="433236B3FFFA69032F41C4C0F9A8F0EF" blockId="11.[113,752,1216,1272]" pageId="11" pageNumber="140">
Three were taken in a flooded grassy clearing in the forest, and one was on leaf litter
<quantity id="84759B56FFFA69032EBCC4E0FA81F0EF" box="[396,439,1248,1272]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="140" unit="m" value="2.0">2 m</quantity>
from a small stream.
</paragraph>
<paragraph id="433236B3FFFA69032F41C520F935F16F" blockId="11.[113,752,1312,1400]" pageId="11" pageNumber="140">
This is the first report of the species from
<collectingCountry id="3B9A7623FFFA69032D79C520F98CF12F" box="[585,698,1312,1336]" name="Cambodia" pageId="11" pageNumber="140">Cambodia</collectingCountry>
. The similar
<taxonomicName id="848D4D30FFFA69032FF4C541FA1FF14F" authorityName="Smith" authorityYear="1921" box="[196,297,1344,1368]" class="Amphibia" family="Ranidae" genus="Rana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="140" phylum="Chordata" rank="species" species="milleti">
<emphasis id="71F9EAA1FFFA69032FF4C541FA1FF14F" box="[196,297,1344,1368]" italics="true" pageId="11" pageNumber="140">R. milleti</emphasis>
</taxonomicName>
was recently reported from southwestern
<collectingCountry id="3B9A7623FFFA69032F41C560FBE9F16F" box="[113,223,1376,1400]" name="Cambodia" pageId="11" pageNumber="140">Cambodia</collectingCountry>
(
<bibRefCitation id="271C4B42FFFA69032FDDC560FAC1F16F" author="Chuaynkern, Y &amp; Ohler, C &amp; Inthara, P" box="[237,503,1376,1400]" pageId="11" pageNumber="140" pagination="1 - 13" refId="ref25171" refString="Chuaynkern, Y., A. Ohler, C. Inthara, P. Kumtong, &amp; A. Dubois, 2004. The recent distribution of Rana milleti Smith, 1921 in mainland Southeast Asia with the first record of Cambodia. The Natural History Journal of Chulalongkorn University, 4 (1): 1 - 13." type="journal article" year="2004">Chuaynkern et al., 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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