206 lines
34 KiB
XML
206 lines
34 KiB
XML
<document ID-CLB-Dataset="6712" ID-DOI="http://dx.doi.org/10.3897/dez.65.21000" ID-GBIF-Dataset="03cce405-5887-4474-b040-35de18e8d020" ID-Pensoft-Pub="1860-1324-1-1" ID-Pensoft-UUID="77D7AEC9294554E5BBDCABA37901022A" ID-Zenodo-Dep="1166397" ID-ZooBank="73DEE0F32BB04A21B4455E168FE50F54" ModsDocID="1860-1324-65-1-1" checkinTime="1621372489409" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2018" docId="CB0F8B6C344341BF29928F5D4AD9E56B" docLanguage="en" docName="DeutEntomolZeit 65(1): 1-63" docOrigin="Deutsche Entomologische Zeitschrift 65 (1)" docPubDate="2018-01-18" docSource="http://dx.doi.org/10.3897/dez.65.21000" docTitle="Mecyclothorax mouensis Moore & Liebherr 2018, sp. n." docType="treatment" docUuid="572EF2F8-5BF7-4663-8F53-74100E27DF5E" docUuidSource="ZooBank" docVersion="5" id="77D7AEC9294554E5BBDCABA37901022A" lastPageNumber="37" masterDocId="77D7AEC9294554E5BBDCABA37901022A" masterDocTitle="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel" masterLastPageNumber="63" masterPageNumber="1" pageNumber="37" updateTime="1732929906348" updateUser="ExternalLinkService">
|
||
<mods:mods id="8A046FDB11D8D5AB8798B9649C760D0D" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="9FE54DE8D1AAE1C8D766E4E454A8F029">
|
||
<mods:title id="E5E22370C1872CB23E07266E7933BC5B">Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="620F14A6B80CF0798160FC6878A6048F" type="personal">
|
||
<mods:role id="ADBE1BB4B9A584E5758117B8FCBE9E0C">
|
||
<mods:roleTerm id="6AC89657FE79C7D1D87EC5484FB055BD">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="554823D34C3B045CF48608FFE311F17E">Liebherr, James K.</mods:namePart>
|
||
<mods:nameIdentifier id="E644EE0F5489E6647B772F13D8831227" type="ORCID">https://orcid.org/0000-0001-9831-884X</mods:nameIdentifier>
|
||
<mods:affiliation id="C3164EC181B168328A4292CE79B1AF35">Cornell University Insect Collection, John H. and Anna B. Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
|
||
<mods:nameIdentifier id="24A420033CF0D05C65E15C2E024D515E" type="email">jkl5@cornell.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="F8A4EDC94289C6384EAE0001FC04E566">text</mods:typeOfResource>
|
||
<mods:relatedItem id="BE7EABFF09493B6EBA646060B1B1FCFE" type="host">
|
||
<mods:titleInfo id="08958B449C646976601885A6EC37978D">
|
||
<mods:title id="1A06049887A36A1DC4E40A4E40D21924">Deutsche Entomologische Zeitschrift</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="501B3A6E9F660261632B7E4CB5BBEA04">
|
||
<mods:date id="E6035BED7EEE0F32199D38D5677E556B">2018</mods:date>
|
||
<mods:detail id="289ECD6290E6A3E74A6649A92655A2A0" type="pubDate">
|
||
<mods:number id="5F60771A57345288F6377D48B8650F64">2018-01-18</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="8691C23C7074462E2A06B1B28ABE474C" type="volume">
|
||
<mods:number id="768579488AB0D01249C60B9620672F9D">65</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="3A7CF85D0C37C8231D64F60C7BE363B3" type="issue">
|
||
<mods:number id="2954D765BCABAA2608CB394A14F98A20">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="C9387ED02199A8C582CDDEE4A9A9BEE4" unit="page">
|
||
<mods:start id="A6E9E57B933E080EB21B0C815E95AB86">1</mods:start>
|
||
<mods:end id="10670CDC474CF13C12C27830304F89FA">63</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="1200AEB1B0D499DF29C39BAEAC493B7B">
|
||
<mods:url id="2C15F5AC9D9E08AC97BD564D4563D0D5">http://dx.doi.org/10.3897/dez.65.21000</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="F71808EB6A95FA904EDB3F545D525DD5">journal article</mods:classification>
|
||
<mods:identifier id="5294A82B82256AD14B578505494CB5C4" type="DOI">http://dx.doi.org/10.3897/dez.65.21000</mods:identifier>
|
||
<mods:identifier id="CB46151218950955F155D69C2DCFBB29" type="Pensoft-Pub">1860-1324-1-1</mods:identifier>
|
||
<mods:identifier id="9B8C0BF7C3C27A5C1DF081FA8A5E4141" type="ZooBank">73DEE0F32BB04A21B4455E168FE50F54</mods:identifier>
|
||
<mods:identifier id="08BF9CFD1CDC93430CB29FCEB38619E2" type="Pensoft-UUID">77D7AEC9294554E5BBDCABA37901022A</mods:identifier>
|
||
<mods:identifier id="E73A4C7D13EBE81A6DC8E36E4694CEAF" type="Zenodo-Dep">1166397</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="CB0F8B6C344341BF29928F5D4AD9E56B" ID-GBIF-Taxon="182223665" LSID="urn:lsid:zoobank.org:act:572EF2F8-5BF7-4663-8F53-74100E27DF5E" httpUri="http://treatment.plazi.org/id/CB0F8B6C344341BF29928F5D4AD9E56B" lastPageNumber="37" pageId="36" pageNumber="37" scope_family="Carabidae" scope_order="Coleoptera" scope_tribe="Moriomorphini">
|
||
<subSubSection id="AE85EBD0356F3B2B346FC2222014350E" pageId="36" pageNumber="37" type="nomenclature">
|
||
<paragraph id="02024102DB6F253AD0F980560898B399" pageId="36" pageNumber="37">
|
||
<pageBreakToken id="E4A73ABF2F3620017284A781961480E4" pageId="36" pageNumber="37" start="start">13</pageBreakToken>
|
||
.
|
||
<taxonomicName id="E60C6C10B40BE799DCAF07EB4B87ECD4" ID-CoL="8M6Z6" LSID="http://zoobank.org/572EF2F8-5BF7-4663-8F53-74100E27DF5E" authority="Moore & Liebherr" authorityName="Moore & Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax mouensis" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="mouensis" status="sp. n.">Mecyclothorax mouensis Moore & Liebherr</taxonomicName>
|
||
<taxonomicNameLabel id="1A02DB8A4FBC194CBFD8CC86766319A8" pageId="36" pageNumber="37">sp. n.</taxonomicNameLabel>
|
||
<figureCitation id="D65094A9C506B5F3424C148442C03CAB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Thoracic structures of Mecyclothorax and Amblytelus spp. A-B. Pronotum, dorsal view: A, M. laterorectus; B, M. mouensis. C-F. Prosternum, ventral view: C, M. fleutiauxi; D, M. curtus; E, M. blackburni; F, M. moorei. G-J. Elytral apices, dorsal view: G, M. fleutiauxi; H, M. laterorectus; I, M. lissus; J, A. curtus. K-L. Elytral lateral margin showing eighth stria, lateral view: K, M. ambiguus; L, M. blackburni. Abbreviations include: aag, anteapical groove; acc, accessory elytral dorsal setae in interval 5; as, apical elytral seta; ca, carina laterad stria 7; ls, lateral elytral setae; ri, ridge mesad stria 8; sa, subapical elytral seta; sas, subapical sinuation of elytron; scs, supracarinal elytral setae; 8, medial portion of stria 8." figureDoi="10.3897/dez.65.21000.figure2" httpUri="https://binary.pensoft.net/fig/179958" pageId="36" pageNumber="37">Figures 2B</figureCitation>
|
||
<figureCitation id="A9F40832C2E6AC4AA60CF284F131120B" captionStart="Figure 17" captionStartId="F18" captionText="Figure 17. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. plurisetosus, right view, dorsal view in situ (Aoupinie); C-D, M. megalovatulus, right view, dorsal view of righ sclerite (Mt. Panie); E-F, M. octavius, right view, dorsal view in situ (Me Maoya); G-H, M. laterovatulus, right view, dorsal view in situ (Aoupinie); I-J, M. manautei, right view, dorsal view in situ (Mt. Humboldt, 1400 m); K-L, M. paniensis, right view, dorsal view in situ (Mt. Panie); M-N, M. mouensis, right view, dorsal view in situ (Mt. Mou). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure17" httpUri="https://binary.pensoft.net/fig/179973" pageId="36" pageNumber="37">, 17M-N</figureCitation>
|
||
<figureCitation id="D9CB75B11E0D58573199E9A647E3745E" captionStart="Figure 18" captionStartId="F19" captionText="Figure 18. Paired left (above) and right (below) parameres of Mecyclothorax (Phacothorax) spp. (ectal view): A, M. plurisetosus; B, M. megalovatulus; C, M. octavius; D, M. laterovatulus; E, M. manautei; F, M. paniensis; G, M. mouensis." figureDoi="10.3897/dez.65.21000.figure18" httpUri="https://binary.pensoft.net/fig/179974" pageId="36" pageNumber="37">, 18G</figureCitation>
|
||
<figureCitation id="7D6E52DCB8F4577F27DE8097ED300953" captionStart="Figure 20" captionStartId="F21" captionText="Figure 20. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Humboldt, 580 m); E, M. kanak (Mt. Humboldt, 630 m); F, M. kanak (Mt. Dzumac); G, M. kanak (Col de Yate); H, M. picdupinsensis." figureDoi="10.3897/dez.65.21000.figure20" httpUri="https://binary.pensoft.net/fig/179976" pageId="36" pageNumber="37">, 20C</figureCitation>
|
||
<figureCitation id="EB49F6E947C633469BEE7A0A8611DC2A" captionStart="Figure 23" captionStartId="F24" captionText="Figure 23. Female reproductive tract, gonocoxites and associated laterotergites, Mecyclothorax (Phacothorax) spp: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Dzumac; common oviduct artificially darkened to show configuration); E, M. kanak (Pic du Grand Kaori); F, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure23" httpUri="https://binary.pensoft.net/fig/179980" pageId="36" pageNumber="37">, 23C</figureCitation>
|
||
<figureCitation id="AB2B25A6544B5A4C375B3625117BB267" captionStart="Figure 24" captionStartId="F25" captionText="Figure 24. Left female gonocoxa of Mecyclothorax (Phacothorax) spp., ventral view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak; E, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure24" httpUri="https://binary.pensoft.net/fig/179981" pageId="36" pageNumber="37">, 24C</figureCitation>
|
||
<figureCitation id="D79443A1B242DAD9EC6F10DEF75129C0" captionStart="Figure 25" captionStartId="F26" captionText="Figure 25. Geographical distributions of Mecyclothorax (Phacothorax) spp. (asterisked locality attributed to M. mouensis based on non-type female specimen)." figureDoi="10.3897/dez.65.21000.figure25" httpUri="https://binary.pensoft.net/fig/179982" pageId="36" pageNumber="37">, 25</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EA221BDC6305A391A5DD6463B3D1D36B" pageId="36" pageNumber="37" type="diagnosis">
|
||
<paragraph id="4BD7E6E63D60482AE5BDD201828B0E66" pageId="36" pageNumber="37">Diagnosis.</paragraph>
|
||
<paragraph id="A62FB2618CF16941EDDF9733EE64574F" pageId="36" pageNumber="37">
|
||
This species, along with
|
||
<taxonomicName id="D6CA2BD53D32FCBD5B1678FE078D156B" lsidName="M. kanak" pageId="36" pageNumber="37" rank="species" species="kanak">
|
||
<emphasis id="DE2B5F2A4282E7A77BCFFFBCB62421E9" italics="true" pageId="36" pageNumber="37">M. kanak</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="0BAB5961113927E97684E76CBF308BE9" lsidName="M. picdupinsensis" pageId="36" pageNumber="37" rank="species" species="picdupinsensis">
|
||
<emphasis id="94062FB85FEE19E14C252FCE7F14C41B" italics="true" pageId="36" pageNumber="37">M. picdupinsensis</emphasis>
|
||
</taxonomicName>
|
||
, comprises a triplet of cryptic sibling species best determined using the male aedeagal median lobe. To that end, this species is characterized by a median lobe with the apex dorsally and ventrally subparallel beyond the ostial opening, and with the
|
||
<normalizedToken id="2D83FF14DFBBDA4745D15CC6734FB741" originalValue="lobe’s">lobe's</normalizedToken>
|
||
apical face narrowly flattened (Fig.
|
||
<figureCitation id="4D21831E7190A73C14E1CA29BEDB1ED4" captionStart="Figure 17" captionStartId="F18" captionText="Figure 17. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. plurisetosus, right view, dorsal view in situ (Aoupinie); C-D, M. megalovatulus, right view, dorsal view of righ sclerite (Mt. Panie); E-F, M. octavius, right view, dorsal view in situ (Me Maoya); G-H, M. laterovatulus, right view, dorsal view in situ (Aoupinie); I-J, M. manautei, right view, dorsal view in situ (Mt. Humboldt, 1400 m); K-L, M. paniensis, right view, dorsal view in situ (Mt. Panie); M-N, M. mouensis, right view, dorsal view in situ (Mt. Mou). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure17" httpUri="https://binary.pensoft.net/fig/179973" pageId="36" pageNumber="37">17M</figureCitation>
|
||
). All three species share broadly hemiovoid elytra, with this species exhibiting a broadly rounded basal groove across the humerus. Individuals of this species can be best diagnosed externally from those of the following two species by the broader elytral lateral marginal depression outside the anterior series of lateral elytral setae (Fig.
|
||
<figureCitation id="A9532227365BB37CB4B71906EC243A16" captionStart="Figure 20" captionStartId="F21" captionText="Figure 20. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Humboldt, 580 m); E, M. kanak (Mt. Humboldt, 630 m); F, M. kanak (Mt. Dzumac); G, M. kanak (Col de Yate); H, M. picdupinsensis." figureDoi="10.3897/dez.65.21000.figure20" httpUri="https://binary.pensoft.net/fig/179976" pageId="36" pageNumber="37">20C</figureCitation>
|
||
). The anterior lateral elytral setae are situated in the lateral depression in beetles of this species, whereas they lie on the upraised lateral reaches of the elytral disc in
|
||
<taxonomicName id="4A408798F1291DEEC4A7386C81CA2893" lsidName="M. kanak" pageId="36" pageNumber="37" rank="species" species="kanak">
|
||
<emphasis id="29ACC4B9E5D1F3DCA5FF463E341B8985" italics="true" pageId="36" pageNumber="37">M. kanak</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="6A8733C03EF9B5FECFED73E4885D0EFC" lsidName="M. picdupinsensis" pageId="36" pageNumber="37" rank="species" species="picdupinsensis">
|
||
<emphasis id="28DCEBD063654D4EF51713E35223B62D" italics="true" pageId="36" pageNumber="37">M. picdupinsensis</emphasis>
|
||
</taxonomicName>
|
||
. Also the sutural stria is deep and smooth on the disc and striae 2-3 are evident and traceable to nearly obsolete in
|
||
<taxonomicName id="1E2A57CEEB4810989CEE4408549CB612" lsidName="M. mouensis" pageId="36" pageNumber="37" rank="species" species="mouensis">
|
||
<emphasis id="EF384C96340C637D9137EE7772DD2ED7" italics="true" pageId="36" pageNumber="37">M. mouensis</emphasis>
|
||
</taxonomicName>
|
||
. This differs from the sutural stria in beetles of the following two species where the sutural stria is crenulate to indistinctly punctate on the disc. Standardized body length 2.8-3.4 mm. Chaetotaxy +/+//+/-//+/2/+/+.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FDBDE27C7AE58ECF576C78B6D830C721" pageId="36" pageNumber="37" type="description">
|
||
<paragraph id="E4DDB075293943F022BA0F5544098A29" pageId="36" pageNumber="37">Description</paragraph>
|
||
<paragraph id="B7CEF44DCC1EF5F77E27743D2C9F9FB0" pageId="36" pageNumber="37">
|
||
(n = 5). Head capsule elongate, eyes small, very convex, ocular lobe-genal juncture evenly curved, a very shallow groove indicating limit of ocular lobe; 11 ommatidia along horizontal diameter of eye; ocular ratio 1.43-1.48, ocular lobe ratio 0.73-0.77, EyL/EyD = 2.17-2.38; frontal grooves narrowly incised, sinuously convergent to pit at frontoclypeal suture, briefly extended onto clypeus; mandibles moderately elongate, mandibular ratio 1.73; ligula narrowed to a moderately broad, slightly convex anterior margin, the two ligular setae separated by one setal diameter; paraglossae thin, extended twice as far beyond ligular margin as their basal length to margin, apex visibly spiculate (100
|
||
<normalizedToken id="A1E369E7A5E9330A43193060FCF39E75" originalValue="×">x</normalizedToken>
|
||
); antennae moderately elongate, antennomere 9 length 2.0
|
||
<normalizedToken id="4A28090A7457479E8FB16B211AD1520F" originalValue="×">x</normalizedToken>
|
||
maximal breadth; antennomere 3 glabrous except for apical ring of setae. Pronotum very transverse, distinctly cordate, lateral margins sinuately concave anterad obtuse-rounded hind angles (Figs
|
||
<figureCitation id="9F9406B1BBF18E6CA914E130893F4CD0" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Thoracic structures of Mecyclothorax and Amblytelus spp. A-B. Pronotum, dorsal view: A, M. laterorectus; B, M. mouensis. C-F. Prosternum, ventral view: C, M. fleutiauxi; D, M. curtus; E, M. blackburni; F, M. moorei. G-J. Elytral apices, dorsal view: G, M. fleutiauxi; H, M. laterorectus; I, M. lissus; J, A. curtus. K-L. Elytral lateral margin showing eighth stria, lateral view: K, M. ambiguus; L, M. blackburni. Abbreviations include: aag, anteapical groove; acc, accessory elytral dorsal setae in interval 5; as, apical elytral seta; ca, carina laterad stria 7; ls, lateral elytral setae; ri, ridge mesad stria 8; sa, subapical elytral seta; sas, subapical sinuation of elytron; scs, supracarinal elytral setae; 8, medial portion of stria 8." figureDoi="10.3897/dez.65.21000.figure2" httpUri="https://binary.pensoft.net/fig/179958" pageId="36" pageNumber="37">2B</figureCitation>
|
||
,
|
||
<figureCitation id="324CE1ABE9BD05D5143A8EFF125BC1CE" captionStart="Figure 20" captionStartId="F21" captionText="Figure 20. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Humboldt, 580 m); E, M. kanak (Mt. Humboldt, 630 m); F, M. kanak (Mt. Dzumac); G, M. kanak (Col de Yate); H, M. picdupinsensis." figureDoi="10.3897/dez.65.21000.figure20" httpUri="https://binary.pensoft.net/fig/179976" pageId="36" pageNumber="37">20C</figureCitation>
|
||
), MPW/BPW = 2.43-2.82, MPW/PL = 1.35-1.39; front angles slightly protruded, obtuse, APW/BPW = 1.42-1.50; median base unmargined basally, trapezoidally depressed relative to disc, the median longitudinal impression divided basally into two transverse impressions that isolate disc from base, each transverse basal impression terminated laterally in a short laterally arcuate laterobasal depression; median longitudinal impression finely incised on disc, terminated anteriorly at anterior transverse impression, a narrow fovea at posterior juncture with basal transverse impressions; anterior transverse impression shallow, broad, traceable to front angles; proepisternum separated from prosternum by a shallow groove anteriorly, and deep, distinct groove ventrally; smooth anteapical impression well developed laterally, continuous though shallower ventrally; prosternal process broadly, shallowly depressed between procoxae, the shallow depression extended 1/2 distance toward anterior prothoracic margin. Elytra broadly hemiovoid, humeri well-extended laterally before lateral margins curve posteriorly; basal elytral groove posteriorly curved laterad obtuse humeral angle; MEW/EL = 0.85-0.91; sutural stria well developed in basal 2/3, striae 2-3 traceable on disc near positions of dorsal elytral setae, stria 4 indicated by an series of intermittent longitudinal depressions; only stria 8 evident on elytral apex; elytra appressed and conjoined apically, the sutural margin upraised at apex. Pterothoracic mesepisternal anterior furrow with five to six depressions in one to two irregular rows; mesosternal-mesepisternal suture complete (as in Fig.
|
||
<figureCitation id="711F906C625570F447824D5A9D3F3DB3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Thoracic structures of Mecyclothorax spp. A-B. Lateral view of elytral epipleuron and meso-and metapleurites: A, M. laterorectus; B, M. fleutiauxi. C-D. Metathoracic flight wing: C, M. punctipennis, showing fully developed flight wing folded under elytron; D, M. lissus, stenopterous wing rudiment that extends beyond metanotum, rudimentary wing venation present at base of strap. Abbreviations include: mms, mesepisternal-mesosternal suture; msp, mesepimeron; mst, mesepisternum; mtp, metepimeron; mtt, metepisternum." figureDoi="10.3897/dez.65.21000.figure3" httpUri="https://binary.pensoft.net/fig/179959" pageId="36" pageNumber="37">3a</figureCitation>
|
||
); metepisternum length and breadth subequal; metepisternal-metepimeral suture complete though fine laterally. Abdomen with elongate crescent-shaped depression along suture between first and second ventrite, second ventrite only slightly depressed within crescent; suture between second and third ventrites reduced though traceable laterally; ventrites 3-6 with broad, shallow, linear plaques near lateral margin. Microsculpture of frons reduced, surface glossy, indistinct transverse mesh-sculpticell breadth 2
|
||
<normalizedToken id="1BE28CFA291B0A61E109A9A6DA9824D0" originalValue="×">x</normalizedToken>
|
||
length-visible over portions of vertex; pronotal disc glossy but with transverse sculpticells visible outside areas of reflection, trapezoidal median base glossy with indistinct transverse mesh anterad laterobasal depression; elytra opalescent, disc covered with very finely separated transverse lines, elytral apex covered with elongate transverse sculpticells and lines.
|
||
</paragraph>
|
||
<paragraph id="7F34657436214EE16203C8BD23972E62" pageId="36" pageNumber="37">
|
||
<emphasis id="15FD477B0C748487CDB50DBC27A7A953" italics="true" pageId="36" pageNumber="37">Male genitalia</emphasis>
|
||
(n = 8). Antecostal margin of mediotergite IX angulate, not extended (Fig.
|
||
<figureCitation id="179F2D0AFF56AB62A0ABD3FDA05C2682" captionStart="Figure 17" captionStartId="F18" captionText="Figure 17. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. plurisetosus, right view, dorsal view in situ (Aoupinie); C-D, M. megalovatulus, right view, dorsal view of righ sclerite (Mt. Panie); E-F, M. octavius, right view, dorsal view in situ (Me Maoya); G-H, M. laterovatulus, right view, dorsal view in situ (Aoupinie); I-J, M. manautei, right view, dorsal view in situ (Mt. Humboldt, 1400 m); K-L, M. paniensis, right view, dorsal view in situ (Mt. Panie); M-N, M. mouensis, right view, dorsal view in situ (Mt. Mou). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure17" httpUri="https://binary.pensoft.net/fig/179973" pageId="36" pageNumber="37">17N</figureCitation>
|
||
); right paramere narrow, with basal cuff articulating with elongate, whiplike apical extension (Fig.
|
||
<figureCitation id="446D4DB23B127AA1CDEC3EA31A995C36" captionStart="Figure 18" captionStartId="F19" captionText="Figure 18. Paired left (above) and right (below) parameres of Mecyclothorax (Phacothorax) spp. (ectal view): A, M. plurisetosus; B, M. megalovatulus; C, M. octavius; D, M. laterovatulus; E, M. manautei; F, M. paniensis; G, M. mouensis." figureDoi="10.3897/dez.65.21000.figure18" httpUri="https://binary.pensoft.net/fig/179974" pageId="36" pageNumber="37">18G</figureCitation>
|
||
), six setae along ventral margin to complement to longer apical setae; left paramere narrow basally, evenly narrowed to whiplike apex; aedeagal median lobe robust, broadest medially basad apex of ostial opening, apex subparallel dorsoventrally beyond ostium, with apical face somewhat flattened (Fig.
|
||
<figureCitation id="048EA739430B1AFDB8370B05475F77AA" captionStart="Figure 17" captionStartId="F18" captionText="Figure 17. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. plurisetosus, right view, dorsal view in situ (Aoupinie); C-D, M. megalovatulus, right view, dorsal view of righ sclerite (Mt. Panie); E-F, M. octavius, right view, dorsal view in situ (Me Maoya); G-H, M. laterovatulus, right view, dorsal view in situ (Aoupinie); I-J, M. manautei, right view, dorsal view in situ (Mt. Humboldt, 1400 m); K-L, M. paniensis, right view, dorsal view in situ (Mt. Panie); M-N, M. mouensis, right view, dorsal view in situ (Mt. Mou). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure17" httpUri="https://binary.pensoft.net/fig/179973" pageId="36" pageNumber="37">17M</figureCitation>
|
||
); aedeagal median lobe internal sac with broad, basal dorsal plate and a large field of macrospicules apicad the plate, but flagellum and flagellar sheath not present.
|
||
</paragraph>
|
||
<paragraph id="6B613445AB8A26DBF14799D9C0AFB3C8" pageId="36" pageNumber="37">
|
||
<emphasis id="163A994E8EA4F1A7F066DA4D69507C7D" italics="true" pageId="36" pageNumber="37">Female reproductive tract</emphasis>
|
||
(n = 1). Bursa copulatrix length subequal to circumference, its surface thin, translucent, slightly wrinkled (Fig.
|
||
<figureCitation id="4C6E1BDC005472625365E1A76DD1E24B" captionStart="Figure 23" captionStartId="F24" captionText="Figure 23. Female reproductive tract, gonocoxites and associated laterotergites, Mecyclothorax (Phacothorax) spp: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Dzumac; common oviduct artificially darkened to show configuration); E, M. kanak (Pic du Grand Kaori); F, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure23" httpUri="https://binary.pensoft.net/fig/179980" pageId="36" pageNumber="37">23C</figureCitation>
|
||
); spermathecal duct entering near bursa-common oviduct juncture with duct oriented toward right side of bursa; elongate laminar helminthoid sclerite present at base of spermathecal duct; spermatheca of only slightly greater circumference than spermathecal duct, spermathecal gland duct entering at base of spermathecal reservoir; ligular apophysis present on common oviduct; basal gonocoxite 1 with apical fringe of two setae laterally, a very small setae may be present immediately mesad, and several smaller setae along medial margin (Fig.
|
||
<figureCitation id="D2ED6F674C74D6E1CD4BE55E97791F2A" captionStart="Figure 24" captionStartId="F25" captionText="Figure 24. Left female gonocoxa of Mecyclothorax (Phacothorax) spp., ventral view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak; E, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure24" httpUri="https://binary.pensoft.net/fig/179981" pageId="36" pageNumber="37">24C</figureCitation>
|
||
); medial margin bearing a dense field of microtrichia basally, this field about 3-4 microtrichia broad and appearing like the hooklike surface of a Velcro® closure; gonocoxite 2 moderately broad basally, basal width 0.6
|
||
<normalizedToken id="BAFF3F36EEFC1B93B88DB437F3C82306" originalValue="×">x</normalizedToken>
|
||
medial length; two gracile lateral ensiform setae of moderate length present.
|
||
</paragraph>
|
||
<paragraph id="39C54259D72F563B3E4660D853EF52DE" pageId="36" pageNumber="37">
|
||
<emphasis id="06597CFA0F15524B90B86B2E8ED56A52" italics="true" pageId="36" pageNumber="37">Types</emphasis>
|
||
- Holotype male (MNHN): NEW CALEDONIA / Mt Mou summit / 24 May 1984 / G. Monteith & D. Cook // Q.M. Berlesate 659 / 22.04S X 166.21E / Rainforest, 1200 n / Litter // QUEENSLAND / MUSEUM LOAN / DATE: Nov. 2003 / No. LEN-1686 (green label) // HOLOTYPE /
|
||
<taxonomicName id="012CD35BFDEE3965DF6FA4AA3CA35A96" authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
|
||
/
|
||
<taxonomicName id="CD021066B6ED4FC9CF52D5B128B0A3AC" authorityName="Moore & Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax mouensis" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="mouensis">Mecyclothorax mouensis</taxonomicName>
|
||
/ B.P. Moore & J.K. / Liebherr 2017 (black-bordered red label).
|
||
</paragraph>
|
||
<paragraph id="C3D9075228C188F7D8AC8DB9E3E31FE9" pageId="36" pageNumber="37">
|
||
Paratypes (33 specimens). NEW CALEDONIA: Mts. Koghis, [code PA 58],
|
||
<geoCoordinate id="90E6E2DB640C58BE8BB6680A0059D43F" degrees="22" direction="south" minutes="11" orientation="latitude" precision="925" value="-22.183332">22°11'S</geoCoordinate>
|
||
<geoCoordinate id="2045D9D9FD86FA7641D94358A8925D2B" degrees="166" direction="east" minutes="31" orientation="longitude" precision="925" value="166.51666">166°31'E</geoCoordinate>
|
||
, 30-viii-1970, Franz (NHMW, 3), ~5 km N Noumea, 550 m el.,
|
||
<geoCoordinate id="750E20EFDD087DF469F46BC9DD9682C8" degrees="22" direction="south" minutes="10.5" orientation="latitude" precision="92" value="-22.175">22 10.5'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="15EC21BCE4BFE76439A56EC533F7E50F" degrees="166" direction="east" minutes="30.3" orientation="longitude" precision="92" value="166.505">166 30.3'E</geoCoordinate>
|
||
, forest litter, 25-xi-2009, Schuh (NHMW, 1), forest, 520 m el.,
|
||
<geoCoordinate id="70E04F58ED672BE073F5B41F355C69E4" degrees="22" direction="south" minutes="10.7" orientation="latitude" precision="92" value="-22.178333">22°10.7'S</geoCoordinate>
|
||
<geoCoordinate id="22BF967B79EEA73FD5B3F59FFC2CAEED" degrees="166" direction="east" minutes="30.3" orientation="longitude" precision="92" value="166.505">166°30.3'E</geoCoordinate>
|
||
, sifted litter, 25-x-2008, Wanat (MNHW, 9), La Roussette, 600 m el.,
|
||
<geoCoordinate id="A0AEBC96DC3B245E04262E01AB8DEB34" degrees="22" direction="south" minutes="10.8" orientation="latitude" precision="92" value="-22.18">22°10.8'S</geoCoordinate>
|
||
<geoCoordinate id="9856A9B37C0FA1D5ABE3ADCD42DAEE0C" degrees="166" direction="east" minutes="30.7" orientation="longitude" precision="92" value="166.51167">166°30.7'E</geoCoordinate>
|
||
, sifted litter, 27-x-2008, Wanat (MNHW, 4), track entrance, 500 m el.,
|
||
<geoCoordinate id="7FC888114F499E25A5E4A7B84DF65E04" degrees="22" direction="south" minutes="11" orientation="latitude" precision="925" value="-22.183332">22°11'S</geoCoordinate>
|
||
<geoCoordinate id="3A383BE2790D3FC2E9275173FE1237CB" degrees="166" direction="east" minutes="31" orientation="longitude" precision="925" value="166.51666">166°31'E</geoCoordinate>
|
||
, berlesate, 06-v-2005, lot 12264, Monteith (QMB, 4), 500 m el.,
|
||
<geoCoordinate id="6B08F498738B1E87C770A91648DFD24E" degrees="22" direction="south" minutes="10" orientation="latitude" precision="925" value="-22.166666">22°10'S</geoCoordinate>
|
||
<geoCoordinate id="0CD280A21E77CD7C73FA0A88C6305A89" degrees="166" direction="east" minutes="31" orientation="longitude" precision="925" value="166.51666">166°31'E</geoCoordinate>
|
||
, 26-viii-1978, S. & J. Peck (CNC, 1), 400-500 m el.,
|
||
<geoCoordinate id="26669B978A5885011B2D03A12687CEC2" degrees="22" direction="south" minutes="11" orientation="latitude" precision="925" value="-22.183332">22°11'S</geoCoordinate>
|
||
<geoCoordinate id="03EAA6195F5876B3E2A340D64726FE22" degrees="166" direction="east" minutes="31" orientation="longitude" precision="925" value="166.51666">166°31'E</geoCoordinate>
|
||
, primary forest, 18-19-x-1998,
|
||
<normalizedToken id="C2D364BA063F19A429F93872DFE01CEA" originalValue="Löbl">Loebl</normalizedToken>
|
||
(MBC, 1; MHNG, 6); Mt. Mou, summit, 1200 m el.,
|
||
<geoCoordinate id="2E9116004C2B65AD011FFFB21A7DF6F4" degrees="22" direction="south" minutes="04" orientation="latitude" precision="925" value="-22.066668">22°04'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="799A749600341700BE342F73D62A9AFB" degrees="166" direction="east" minutes="21" orientation="longitude" precision="925" value="166.35">166°21'E</geoCoordinate>
|
||
, rainforest litter, 24-v-1984, Monteith & Cook (QMB, 2), top camp, 1150 m el.,
|
||
<geoCoordinate id="DF8AEFE5D2130F7491B54B8119DDC693" degrees="22" direction="south" minutes="04" orientation="latitude" precision="925" value="-22.066668">22°04'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="96CD5B6D5A27DCF1CB5106A80CC694A9" degrees="166" direction="east" minutes="21" orientation="longitude" precision="925" value="166.35">166°21'E</geoCoordinate>
|
||
, berlesate, sieved litter, 27-xii-2004, lot 12013, Monteith (QMB, 2). Non-type female specimen:
|
||
<normalizedToken id="DCC04CD2BD1D022A5CD7B281B5027AD9" originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken id="ADF8D3FC0ADDB60862926E79E8D18FA5" originalValue="Cachée">Cachee</normalizedToken>
|
||
, 250 m el.,
|
||
<geoCoordinate id="B940FBC8F6C5DB2922CB1131C176CEB3" degrees="22" direction="south" minutes="11.5" orientation="latitude" precision="92" value="-22.191668">22°11.5'S</geoCoordinate>
|
||
<geoCoordinate id="163F9FFF6627FC6151BF9F588A4A8529" degrees="166" direction="east" minutes="47.2" orientation="longitude" precision="92" value="166.78667">166°47.2'E</geoCoordinate>
|
||
, sifted litter, 26-x-2008, Wanat (MNHW, 1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CE2E9E454F601D7A56CFC7169237ECC2" pageId="36" pageNumber="37" type="etymology">
|
||
<paragraph id="26068BFA8318BB1482F4415B77D03C7B" pageId="36" pageNumber="37">Etymology.</paragraph>
|
||
<paragraph id="932D9EA1A0ED7DA2EED245D51D4794AB" pageId="36" pageNumber="37">
|
||
As in the species above, this species epithet elides the type locality, Mt. Mou, with the genitive, adjectival ending -ensis. As Dr. Barry P. Moore had both diagnosed this species from the following
|
||
<taxonomicName id="95DC41D2084FBF34841F95E37F890D7F" lsidName="M. kanak" pageId="36" pageNumber="37" rank="species" species="kanak">
|
||
<emphasis id="628251BD612AC462FEF962C3DD0EC3F4" italics="true" pageId="36" pageNumber="37">M. kanak</emphasis>
|
||
</taxonomicName>
|
||
through the use of male genitalic characters, and had chosen a holotype specimen, he is given senior author status for this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7AE4322DAAD328DD7ED1FBA1E468CEF3" pageId="36" pageNumber="37" type="distribution">
|
||
<paragraph id="4D44A5D44498E433510037BAA06DD3D9" pageId="36" pageNumber="37">Distribution and habitat.</paragraph>
|
||
<paragraph id="B6D8FC54F8C2D3FA7F861129B4D3A228" pageId="36" pageNumber="37">
|
||
This species is restricted to the southern portion of Grande Terre, from Mt. Mou on the west, Mts. Koghis north of Noumea, and with a easternmost record from
|
||
<normalizedToken id="D640F452DF70633BEAD2CBF23D550B5B" originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken id="C4186151CDBC9328D27E7F85D04C7276" originalValue="Cachée">Cachee</normalizedToken>
|
||
based on a single non-type female specimen (Fig.
|
||
<figureCitation id="4C2D01D06AEBE016AE409E03C21C029B" captionStart="Figure 25" captionStartId="F26" captionText="Figure 25. Geographical distributions of Mecyclothorax (Phacothorax) spp. (asterisked locality attributed to M. mouensis based on non-type female specimen)." figureDoi="10.3897/dez.65.21000.figure25" httpUri="https://binary.pensoft.net/fig/179982" pageId="36" pageNumber="37">25</figureCitation>
|
||
). Collection localities range 250-1200 m elevation. Given that 24 of the 35 specimens are labeled from Berlese samples or sieved litter, and another 7 specimens (1, S. & J. Peck, CNC; 6,
|
||
<normalizedToken id="B75EC10F9B8DBA3AC81CFDE6809F4201" originalValue="Löbl">Loebl</normalizedToken>
|
||
, MHNG) were collected by coleopterists studying the microcoleopteran litter fauna, it can be concluded that this species occupies the ground-level litter layer.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |