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<document id="24BEFC82EC874B08DCE95DF78CEF8DA6" ID-DOI="http://dx.doi.org/10.3897/zookeys.734.22774" ID-GBIF-Dataset="5ce053ba-431c-4fc5-83ea-1b8c389fe55f" ID-PMC="PMC5904335" ID-Pensoft-Pub="1313-2970-734-121" ID-PubMed="29674857" ID-ZooBank="2FD0E47C9C5C4022ABF78482872414A0" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-734-121" ModsDocOrigin="ZooKeys 734" ModsDocTitle="Review of the genus Coccus Linnaeus from Korea, with description of a new species (Hemiptera, Coccomorpha, Coccidae)" checkinTime="1517868730254" checkinUser="pensoft" docAuthor="Choi, Jinyeong &amp; Lee, Seunghwan" docDate="2018" docId="CB0CEC6B298E399CB11FE216BC455EA3" docLanguage="en" docName="ZooKeys 734: 121-135" docOrigin="ZooKeys 734" docSource="http://dx.doi.org/10.3897/zookeys.734.22774" docTitle="Coccus malloti Takahashi 1956" docType="treatment" docVersion="5" lastPageNumber="128" masterDocId="E70DD842700F3542B635FFA00601D609" masterDocTitle="Review of the genus Coccus Linnaeus from Korea, with description of a new species (Hemiptera, Coccomorpha, Coccidae)" masterLastPageNumber="135" masterPageNumber="121" pageNumber="126" updateTime="1701316762108" updateUser="plazi">
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<mods:title id="AFE62085D572D12C9BB9BE291841D510">Review of the genus Coccus Linnaeus from Korea, with description of a new species (Hemiptera, Coccomorpha, Coccidae)</mods:title>
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<mods:namePart id="7E02390A1EA6EE8469DC2EF64A75CEEF">Choi, Jinyeong</mods:namePart>
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<mods:namePart id="0C22F5C1611F20EE9DF43AAFA312F149">Lee, Seunghwan</mods:namePart>
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<mods:classification id="47E70E1E9668CD03F13506EBB5648BC5">journal article</mods:classification>
<mods:identifier id="12A638973114AB0A7B676C4E954290D7" type="DOI">http://dx.doi.org/10.3897/zookeys.734.22774</mods:identifier>
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<treatment id="CB0CEC6B298E399CB11FE216BC455EA3" ID-GBIF-Taxon="141129076" LSID="urn:lsid:plazi:treatment:CB0CEC6B298E399CB11FE216BC455EA3" httpUri="http://treatment.plazi.org/id/CB0CEC6B298E399CB11FE216BC455EA3" lastPageId="7" lastPageNumber="128" pageId="5" pageNumber="126">
<subSubSection id="9A44765D8BEE40DE4462170D15D12E3E" pageId="5" pageNumber="126" type="nomenclature">
<paragraph id="6ED0102196F44D053AF070032DF9A531" pageId="5" pageNumber="126">
<taxonomicName id="80F050AF200F8F3E4505D6164EE128B1" ID-CoL="5ZHNF" authority="Takahashi, 1956" authorityName="Takahashi" authorityYear="1956" class="Insecta" family="Coccidae" genus="Coccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccus malloti" order="Hemiptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="malloti">Coccus malloti (Takahashi, 1956)</taxonomicName>
Figs 4
<normalizedToken id="1258778594A661BB458447FEB7C8F3B4" originalValue="AB">A-B</normalizedToken>
, 5
<normalizedToken id="83F25A80A86630B4C34FD718E9F31057" originalValue="AQ">A-Q</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection id="71FE97F1AC67FD056AD188F6E9D78664" pageId="5" pageNumber="126" type="reference_group">
<paragraph id="6C31464879D9738F2B377C0EEAE3364C" pageId="5" pageNumber="126">
<taxonomicName id="70915DD0E8CF28A6801E142C6A7383E3" class="Insecta" family="Coccidae" genus="Pulvinaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulvinaria malloti" order="Hemiptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="malloti">Pulvinaria malloti</taxonomicName>
Takahashi, 1956: 25.
</paragraph>
</subSubSection>
<subSubSection id="B2E95B712C07301237D2C85D198839F8" pageId="5" pageNumber="126" type="materials_examined">
<paragraph id="9317BDEF8C69C23BD1A8EDB4D7FF2ECF" pageId="5" pageNumber="126">Material examined.</paragraph>
<paragraph id="8E65F4370025A4E841DEDF796532F44F" pageId="5" pageNumber="126">
Adult female: Jeollanam-do, Gwangyang-si, Ongnyong-myeon, Chusan-ri, 28.v.2015, coll. J.Y. Choi, on
<taxonomicName id="343E28BF41F73FE288476526C61D3347" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ilex cornuta" order="Aquifoliales" pageId="5" pageNumber="126" phylum="Tracheophyta" rank="species" species="cornuta">Ilex cornuta</taxonomicName>
Lindl. (
<taxonomicName id="1CB2F6026B7E481A66CB17E48F2CCB79" family="Aquifoliaceae" lsidName="" pageId="5" pageNumber="126" rank="family">Aquifoliaceae</taxonomicName>
), 9♀♀; Jeju-do, Seogwipo-si, Andeok-myeon, Gamsan-ri, 27.iv.2016, coll. J.Y. Choi, on
<taxonomicName id="0BFBC42536AB08F1F2D812DCABD05164" class="Magnoliopsida" family="Cannabaceae" genus="Aphananthe" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aphananthe aspera" order="Rosales" pageId="5" pageNumber="126" phylum="Tracheophyta" rank="species" species="aspera">Aphananthe aspera</taxonomicName>
(Thunb.) (
<taxonomicName id="0EA282BCFD20CB57B809E2B776A8DB6E" family="Cannabaceae" lsidName="" pageId="5" pageNumber="126" rank="family">Cannabaceae</taxonomicName>
), 1♀.
</paragraph>
</subSubSection>
<subSubSection id="D9361E8E157FDA78C427C84BD7B60CC0" pageId="5" pageNumber="126" type="diagnosis">
<paragraph id="DF7918DC11A622C4023A442A4D1909E4" pageId="5" pageNumber="126">Diagnosis.</paragraph>
<paragraph id="48C49A1C4876011CE2414BC344A6E905" pageId="5" pageNumber="126">
Adult females in life (Fig. 4
<normalizedToken id="099846F0834F23BB8F5302AFB33226A3" originalValue="AB">A-B</normalizedToken>
) with a reticulated pattern of black stripes and a longitudinal band medially on dorsum; dermal areolations present but small; dorsal tubercles present; dorsal tubular ducts absent; dorsal setae sharply spinose; marginal setae mostly with simple pointed apices; multilocular disc-pores usually with ten loculi; ventral tubular ducts of three types: Type I with a broad inner ductule, densely present on posterior medial area of head; frequent on anterior medial area of prothorax, extending to inner submarginal area of thorax; and also sparsely scattered on inner submarginal area of abdomen; Type II with a narrow inner ductule and a large flower-shaped terminal gland, rarely present on inner submarginal area and posterior medial area of abdomen; Type III with a long filamentous inner ductule and a quite small terminal gland, moderately present on submarginal area between anal clefts and each posterior spiracular furrow; all types of ventral tubular ducts absent on medial area of meso-, metathorax and anterior abdomen, and submarginal area of head; antennae each eight segments; legs each with a tibio-tarsal sclerosis on the articulation.
</paragraph>
<caption id="A8C11CDBEF90D636233CE194E6309AA7" pageId="5" pageNumber="126">
<paragraph id="6226FBD76E7641C4E05E865C16706A56" pageId="5" pageNumber="126">
Figure 4.
<taxonomicName id="33A17F485B760078E0CE0F3E59551928" class="Insecta" family="Coccidae" genus="Coccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccus malloti" order="Hemiptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="malloti">Coccus malloti</taxonomicName>
(Takahashi, 1956). A adult female on
<taxonomicName id="0DAF31474570EDC5FF8EF877E69C7662" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ilex cornuta" order="Aquifoliales" pageId="5" pageNumber="126" phylum="Tracheophyta" rank="species" species="cornuta">Ilex cornuta</taxonomicName>
B adult female on
<taxonomicName id="031DEF36FB2F9AC8C40B7E3ACBE3826C" class="Magnoliopsida" family="Cannabaceae" genus="Aphananthe" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aphananthe aspera" order="Rosales" pageId="5" pageNumber="126" phylum="Tracheophyta" rank="species" species="aspera">Aphananthe aspera</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="A03A7585CF2B5D75E1CFC9C118BE8E46" lastPageId="7" lastPageNumber="128" pageId="6" pageNumber="127" type="redescription">
<paragraph id="CEBCBDF9E35E78A6B72A03F1DF388BC3" pageId="6" pageNumber="127">
<pageBreakToken id="28A0025D6EEB8B6D65BEA2D652693685" pageId="6" pageNumber="127" start="start">Redescription</pageBreakToken>
.
</paragraph>
<paragraph id="EEDD56384259036D2E085A6419DB820A" pageId="6" pageNumber="127">
Adult female. Living appearance (Figs 4
<normalizedToken id="993AC851149A98AF5DC7957D332F202D" originalValue="AB">A-B</normalizedToken>
). Body elongate oval, flattened, or slightly convex. Young adult females yellowish to dark brownish, with a reticulated pattern of brown or black stripes, getting darker at maturity. Eggs reddish in color, stored beneath venter.
</paragraph>
<paragraph id="89C3FF9231F79A153B9B868DFF2B2BD7" pageId="6" pageNumber="127">
Slide-mounted material (Fig. 5
<normalizedToken id="BC25D669D4E85F50E37C14CBEB5473E2" originalValue="AQ">A-Q</normalizedToken>
). Body elongate oval, 3.6-4.8 mm long, 2.2-3.3 mm wide, with shallow to deep stigmatic cleft; anal clefts approximately 1/6 of body length.
</paragraph>
<caption id="48843491D6D99084E26A64B6FF7A16D3" pageId="6" pageNumber="127">
<paragraph id="4B8AB202D3F5DF960D6ED0A0CF686D51" pageId="6" pageNumber="127">
Figure 5.
<taxonomicName id="F6A74FDBCC48D131DB9D49C49EAB9ED2" class="Insecta" family="Coccidae" genus="Coccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccus malloti" order="Hemiptera" pageId="6" pageNumber="127" phylum="Arthropoda" rank="species" species="malloti">Coccus malloti</taxonomicName>
(Takahashi, 1956), adult female. A dorsal microduct B dorsal tubercle C dermal areolations D dorsal seta E stigmatic spines F marginal seta G anal plates H preopercular pore I multilocular disc-pores J ventral tubular duct (Type II) K leg L ventral tubular duct (Type III) M ventral tubular ducts (Type I) N ventral seta O spiracular pore P ventral microduct Q antenna. Scale bars: 200
<normalizedToken id="7FD5861C806D50F6BA695078F24B64CA" originalValue="μm">μm</normalizedToken>
(K, Q); 100μm (G); 50
<normalizedToken id="B79C5341006EA42F523103E232436D48" originalValue="μm">μm</normalizedToken>
(C, E); others = 10
<normalizedToken id="A0C160564FF5BFD64BC9150153DCC268" originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
<paragraph id="4E8E5C491A93DFF61FF6E8012F13CA7F" pageId="6" pageNumber="127">
Dorsum. Derm membranous. Dermal areolations oval and small, each with a microduct. Dorsal tubercles normally convex, present on submarginal area, 1-5 in total on each side: one or two between apex of head and anterior stigmatic cleft, zero to two between anterior and posterior stigmatic clefts, and zero or one between posterior stigmatic cleft and anal cleft. Dorsal setae sharply spinose, short, stout, each 6-9
<normalizedToken id="28CB05D948F9476FFF6142ADB2552172" originalValue="μm">μm</normalizedToken>
long, moderately distributed on dorsum. Dorsal tubular ducts absent. Dorsal microducts evenly scattered over entire dorsum. Preopercular pores round and small, 3-5
<normalizedToken id="7206AD6D183DC389B1356553719BFD67" originalValue="μm">μm</normalizedToken>
wide, set in a small group of approximately 8 in front of anal plates. Anal plates quadrate, 217-249
<normalizedToken id="08AB76299283DFCBE99C90E5D5F26E32" originalValue="μm">μm</normalizedToken>
long, 169-198
<normalizedToken id="3E381AA2C2458F513AB1195265123A5F" originalValue="μm">μm</normalizedToken>
wide, each with slightly concaved posterolateral margin, usually posterolateral margin quite longer than anterolateral margin; anterolateral margin 123-143
<normalizedToken id="505DD993128854D9F383E5D9936486E3" originalValue="μm">μm</normalizedToken>
long, posterolateral margin 150-169
<normalizedToken id="CBE0854FC5579ECE443343B0CFBFEA6B" originalValue="μm">μm</normalizedToken>
long; each plate with four apical setae. Ano-genital fold with two pairs of anterior and three pairs of lateral margin setae. Anal ring with six long setae.
</paragraph>
<paragraph id="9AB009C5866CEF1D5C3793C35FFBCF3D" pageId="6" pageNumber="127">
Margin. Marginal setae spinose, straight or slightly bent, each 14-22
<normalizedToken id="646C90252F3EF051770955ABF41EEDA5" originalValue="μm">μm</normalizedToken>
long, mostly with simple pointed apices; with 30-43 present between anterior stigmatic clefts; 13-17 laterally present between anterior and posterior stigmatic clefts, 22-38 present between posterior stigmatic cleft and anal clefts. Stigmatic clefts shallow to deep, each with three stigmatic spines, median spine nearly twice as long as lateral spine: medians 60-75μm long, laterals 24-38
<normalizedToken id="A9B97307876C31E5C6CBFA46E7AEAC5D" originalValue="μm">μm</normalizedToken>
long. Eyespots located near margin.
</paragraph>
<paragraph id="7B141B5AD621A07808EB62F13AAC84A5" lastPageId="7" lastPageNumber="128" pageId="6" pageNumber="127">
Venter. Derm membranous. Multilocular disc-pores 6-7
<normalizedToken id="E059FBCDC6C9D374DAD086101064CF4B" originalValue="μm">μm</normalizedToken>
wide, each with 8-10 loculi, mostly with ten loculi, abundant around vulvar area; one or two transverse rows on each abdominal segments; and also small groups present laterad of each metacoxa and mesocoxa, but not observed around procoxa. Spiracular pores 4-5
<normalizedToken id="164197645807CA35C4EE770043AA20F8" originalValue="μm">μm</normalizedToken>
wide, each with five loculi, in a narrow band 2-4 pores wide between each spiracle and stigmatic cleft. Ventral tubular ducts of three types: Type I each with 33-39
<normalizedToken id="763C977667C7FE3A45646518BB06CA7B" originalValue="μm">μm</normalizedToken>
long, with a developed outer ductule and a moderately broad inner ductule, straight or slightly curved, with a flower-shaped terminal gland, densely present on posterior medial area of head; frequent on anterior medial area of prothorax, extending to inner submarginal area of thorax; and also sparsely scattered on inner submarginal area of abdomen; Type II each with 16-24
<normalizedToken id="B08266713D27D0E0D25CCB1FC8B70B3E" originalValue="μm">μm</normalizedToken>
long, with a narrow inner ductule and a large flower-shaped terminal gland, rarely present on inner submarginal area and posterior medial area of abdomen; Type III each with 15-24
<normalizedToken id="0AAB6C72C39E41A36000B7BFB0E93692" originalValue="μm">μm</normalizedToken>
long, with a long filamentous inner ductule and a quite small terminal gland, moderately present on submarginal area between anal clefts and each posterior spiracular furrow; all types of ventral tubular ducts absent on medial area of meso- and metathorax and anterior abdomen, and submarginal area of head. Ventral microducts present on entire venter, especially frequent on sub
<pageBreakToken id="C1FFFDF8C0AF5037058DEB23AAD92F6E" pageId="7" pageNumber="128" start="start">margin</pageBreakToken>
. Ventral setae with three pairs of long pregenital setae; approximately three or four pairs of long and short setae between antennae; other setae sharply spinose, each 5-10
<normalizedToken id="2965F27F9BE91FE50EC90F9F010C050E" originalValue="μm">μm</normalizedToken>
long, sparsely distributed over entire venter. Legs well developed, each with a tibio-tarsal articulation and an articulatory sclerosis; total length of each metathoracic leg 614-769
<normalizedToken id="48A7A431D187ED2DF778D35893B6BE78" originalValue="μm">μm</normalizedToken>
long: each coxa 141-189
<normalizedToken id="8F887813522CA22F602EEB1BBD5CD610" originalValue="μm">μm</normalizedToken>
long, trochanter+femer 209-257
<normalizedToken id="591E1513AA3FC0D586870F5B5E0A73D6" originalValue="μm">μm</normalizedToken>
long, tibia+tarsus 243-311
<normalizedToken id="F25534E98653334E7EB44E7A581E6AD9" originalValue="μm">μm</normalizedToken>
long, claw 17-24
<normalizedToken id="4CCA838279A07812A797AF71EC041C7C" originalValue="μm">μm</normalizedToken>
long. Tarsal digitules thinner and longer than claw digitules. Spiracles normal, mostly posterior peritreme broader than anterior: anterior peritremes each 37-52
<normalizedToken id="26A37B69897F9AF333D9717978EDC8C4" originalValue="μm">μm</normalizedToken>
wide, posterior peritremes each 45-61
<normalizedToken id="276BBF33FDE27B1E480768B7228ACBA0" originalValue="μm">μm</normalizedToken>
wide. Antenna 8-segmented, each 339-399
<normalizedToken id="71ED6FE6F6CEDE2F2DAAF7E6DBC84734" originalValue="μm">μm</normalizedToken>
long. Clypeolabral shield 138-157
<normalizedToken id="2B0C006646D9D0311B6426FE5687BB21" originalValue="μm">μm</normalizedToken>
wide.
</paragraph>
</subSubSection>
<subSubSection id="8384B3EE5A72422C45889BDA2886B56C" pageId="7" pageNumber="128" type="host">
<paragraph id="DA538D15E7EEF8B273C8BB734C3A6CF3" pageId="7" pageNumber="128">Host plant.</paragraph>
<paragraph id="B95A69DC36AB51C8E9A4D99F17C6C4EC" pageId="7" pageNumber="128">
Recorded from six genera in six families (
<bibRefCitation id="A73A2E5629285F5D7623FC4040A35A9A" author="Garcia-Morales, M" journalOrPublisher="Bulletin of Entomological Research" pageId="10" pageNumber="131" url="http://scalenet.info" year="2016">
<normalizedToken id="77F47B66DB0F276627DB41DE42BC7BEE" originalValue="García-Morales">Garcia-Morales</normalizedToken>
et al. 2016
</bibRefCitation>
). In Korea, it was found on
<taxonomicName id="808D4A3862DF2FD6B28A0BC308E2521C" class="Magnoliopsida" family="Cannabaceae" genus="Aphananthe" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aphananthe aspera" order="Rosales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="aspera">Aphananthe aspera</taxonomicName>
(
<taxonomicName id="3C2C1C2B9FA9F9A596C0E9D367379543" family="Cannabaceae" lsidName="" pageId="7" pageNumber="128" rank="family">Cannabaceae</taxonomicName>
) and
<taxonomicName id="929878F00C0779A661D88581FFAC48FB" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ilex cornuta" order="Aquifoliales" pageId="7" pageNumber="128" phylum="Tracheophyta" rank="species" species="cornuta">Ilex cornuta</taxonomicName>
(
<taxonomicName id="0447C00CDC1AFF7964E01D1DCC73B79A" family="Aquifoliaceae" lsidName="" pageId="7" pageNumber="128" rank="family">Aquifoliaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="99BF310A4D27CC915587C3F7FF3AAE54" pageId="7" pageNumber="128" type="distribution">
<paragraph id="6F3CC79FEB915B874962634825222ACE" pageId="7" pageNumber="128">Distribution.</paragraph>
<paragraph id="31E7E21BBAD173EB5D338C7C59439AF5" pageId="7" pageNumber="128">
Only known from Japan (
<bibRefCitation id="7ACAA80D40D3F4C05BF07F3EF3A24A01" author="Takahashi, R" journalOrPublisher="Kontyu" pageId="11" pageNumber="132" pagination="23 - 30" title="Pulvinaria of Japan (Coccidae, Homoptera)." volume="24" year="1956">Takahashi 1956</bibRefCitation>
); first record for Korea.
</paragraph>
</subSubSection>
<subSubSection id="15DAC495C50225F5E248411FA0AC2CA2" pageId="7" pageNumber="128" type="comments">
<paragraph id="C2AB80AAF5806BEB932219B9ACD0FA0B" pageId="7" pageNumber="128">Comments.</paragraph>
<paragraph id="CBF942597C643B40A9739E44AEB5D7AD" pageId="7" pageNumber="128">
The above description based on Korean specimens agrees well with that of
<bibRefCitation id="D30695F6B21DC1BFE46111A55CF6ACA4" author="Takahashi, R" journalOrPublisher="Kontyu" pageId="11" pageNumber="132" pagination="23 - 30" title="Pulvinaria of Japan (Coccidae, Homoptera)." volume="24" year="1956">Takahashi (1956)</bibRefCitation>
, except that variation in the number of dorsal tubercles and marginal setae, and exact distributions of each type of ventral tubular ducts are newly provided in this study.
<taxonomicName id="6BC92641D9D8F98D0AC01DE99C8071E0" class="Insecta" family="Coccidae" genus="Coccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccus malloti" order="Hemiptera" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="malloti">Coccus malloti</taxonomicName>
probably has intermediate morphological characters between the tribes
<taxonomicName id="9BD5A8C301646BAA644A7C2EDC129290" lsidName="" pageId="7" pageNumber="128" rank="tribe" tribe="Coccini">Coccini</taxonomicName>
and
<taxonomicName id="D9AF4825732A27371A36A5D14E0B57EA" lsidName="" pageId="7" pageNumber="128" rank="tribe" tribe="Pulvinariini">Pulvinariini</taxonomicName>
. However, the woolly test, known as ovisac and one of the typical characters of the
<taxonomicName id="F084C6D1D80C886C80E580425478AA71" lsidName="" pageId="7" pageNumber="128" rank="tribe" tribe="Pulvinariini">Pulvinariini</taxonomicName>
, is not observed in the species. In addition, some slide-mounted specimens of
<taxonomicName id="EAAD9EA4ED7CDE45C2CC93998C6FDE32" lsidName="C. malloti" pageId="7" pageNumber="128" rank="species" species="malloti">C. malloti</taxonomicName>
contain eggs and nymphs in their body, which indirectly indicates that they would not produce an ovisac for oviposition. Although
<taxonomicName id="9ACE55C188D436E4EFBBC24CC59CF9E9" class="Insecta" family="Coccidae" genus="Coccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coccus malloti" order="Hemiptera" pageId="7" pageNumber="128" phylum="Arthropoda" rank="species" species="malloti">Coccus malloti</taxonomicName>
would be retained in the tribe
<taxonomicName id="0D2F1276252CDAAF812257BC2D610092" lsidName="" pageId="7" pageNumber="128" rank="tribe" tribe="Coccini">Coccini</taxonomicName>
, it needs to be reviewed with its type materials to clarify the exact generic position of the species.
</paragraph>
</subSubSection>
</treatment>
</document>