261 lines
39 KiB
XML
261 lines
39 KiB
XML
<document id="A0C9A5B2BA7DF75D19F51755D5E3FCEA" ID-DOI="10.11646/zootaxa.3915.3.6" ID-GBIF-Dataset="f0d85ec3-6831-4512-abea-52d8e50f4a32" ID-ISSN="1175-5326" ID-Zenodo-Dep="232721" ID-ZooBank="248111C9-0C68-4F1C-A9B0-B5F1E0DAB54B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460972527156" checkinUser="plazi" docAuthor="Kundrata, Robin, Baena, Manuel & Bocak, Ladislav" docDate="2015" docId="CB0A287AFFCCEC31CC9FFCBDFDB73FC7" docLanguage="en" docName="zt03915p422.pdf" docOrigin="Zootaxa 3915 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paradrilus opacus Kiesenwetter 1865" docType="treatment" docVersion="7" lastPageNumber="419" masterDocId="37335002FFC8EC37CC08FF87FFCE3D0A" masterDocTitle="Classification of Omalisidae based on molecular data and morphology, with description of Paradrilinae subfam. nov. (Coleoptera: Elateroidea)" masterLastPageNumber="422" masterPageNumber="413" pageNumber="417" updateTime="1698559276336" updateUser="plazi">
|
||
<mods:mods id="C9E89FACC9A8DD3722417EF5C1B073B3" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="A5CFF70AB5683FECE2A176D721261FD2">
|
||
<mods:title id="2F474ED1691E9F46A6B242BFE87FFCC7">Classification of Omalisidae based on molecular data and morphology, with description of Paradrilinae subfam. nov. (Coleoptera: Elateroidea)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="B4FA905E0C6F235E303AE2FADCD13BE2" type="personal">
|
||
<mods:role id="1E7B935DB9662F390C3C507F29D998EB">
|
||
<mods:roleTerm id="093951BE90509956CA34D5E79129912A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="09B46CD8A4D2B9187E9A3A6FABB668DC">Kundrata, Robin</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="C0AF8B1174ECA43250DB1A8E00339B5E" type="personal">
|
||
<mods:role id="FF3BDFB437F46B9BD462F7E8E1CFCAAB">
|
||
<mods:roleTerm id="E7D400D458ADE3AC93804089242A8301">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D5ED0961DAFE87668227CBC3B2A52FA9">Baena, Manuel</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D895E7C6430D530305DA112402566E56" type="personal">
|
||
<mods:role id="B831A3C8D0A50656DF51AC4827588692">
|
||
<mods:roleTerm id="D25B586E872D1B9F8996968B0F282B12">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9261992987A8E3C891AEDDD60353D30B">Bocak, Ladislav</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="F010FB8FB779A543011B861820B1E9D0">text</mods:typeOfResource>
|
||
<mods:relatedItem id="29A289588BE04EED34ADB11CE0D1CB52" type="host">
|
||
<mods:titleInfo id="E113F075176D4CBE60881EC82C74044D">
|
||
<mods:title id="E032616723E8501B91549606AF2C953F">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="0A212E3438606A1E31E3CCD44D7A0769">
|
||
<mods:date id="6C3166AAE85CAA25FBEAF445B009FB83">2015</mods:date>
|
||
<mods:detail id="F336E0BBAE6F8F6EC177236CCFD079E3" type="volume">
|
||
<mods:number id="05D45629E1A400B5AC517A7C5055B42F">3915</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="275782D90A7528EF676DFD963E4518EA" type="issue">
|
||
<mods:number id="D210796FF95A8CB02618D5117A179178">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="EB5DB61C8616E7DBA396D733AF89835C" unit="page">
|
||
<mods:start id="8EBED1BBCC45E7950540FDA63C11B5C9">413</mods:start>
|
||
<mods:end id="70AF199C0EE6A0AC006BAF28CF9B8CDC">422</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="C528BAA229DEB6E6C4AF9433E43A240E">journal article</mods:classification>
|
||
<mods:identifier id="68EBA0A19D9C85E4F23C03AA628CB1A5" type="DOI">10.11646/zootaxa.3915.3.6</mods:identifier>
|
||
<mods:identifier id="0814A504CE7640945365F2C2EA0E454E" type="GBIF-Dataset">f0d85ec3-6831-4512-abea-52d8e50f4a32</mods:identifier>
|
||
<mods:identifier id="1F8A2A38065BE85C2797B9AD3D96641A" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="023FCB7024E831718D54ECD045AC3C7F" type="Zenodo-Dep">232721</mods:identifier>
|
||
<mods:identifier id="7E735AA2AF12E614FFE60077F3852D28" type="ZooBank">248111C9-0C68-4F1C-A9B0-B5F1E0DAB54B</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="CB0A287AFFCCEC31CC9FFCBDFDB73FC7" ID-DOI="http://doi.org/10.5281/zenodo.5661133" ID-GBIF-Taxon="119594015" ID-Zenodo-Dep="5661133" LSID="urn:lsid:plazi:treatment:CB0A287AFFCCEC31CC9FFCBDFDB73FC7" httpUri="http://treatment.plazi.org/id/CB0A287AFFCCEC31CC9FFCBDFDB73FC7" lastPageId="6" lastPageNumber="419" pageId="4" pageNumber="417">
|
||
<subSubSection id="0BB9CAE7FFCCEC33CC9FFCBDFEFA3E7C" pageId="4" pageNumber="417" type="nomenclature">
|
||
<paragraph id="431C996CFFCCEC33CC9FFCBDFDA83E5E" blockId="4.[151,614,826,886]" box="[151,614,826,852]" pageId="4" pageNumber="417">
|
||
<heading id="18542E00FFCCEC33CC9FFCBDFDA83E5E" bold="true" box="[151,614,826,852]" fontSize="11" level="1" pageId="4" pageNumber="417" reason="1">
|
||
<taxonomicName id="84A3E2EFFFCCEC33CC9FFCBDFDA83E5E" authority="Kiesenwetter, 1865" authorityName="Kiesenwetter" authorityYear="1865" box="[151,614,826,852]" class="Insecta" family="Drilidae" genus="Paradrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="417" phylum="Arthropoda" rank="species" species="opacus">
|
||
<emphasis id="71D7457EFFCCEC33CC9FFCBDFDA83E5E" bold="true" box="[151,614,826,852]" pageId="4" pageNumber="417">
|
||
<emphasis id="71D7457EFFCCEC33CC9FFCBDFEBA3E5E" bold="true" box="[151,372,826,852]" italics="true" pageId="4" pageNumber="417">Paradrilus opacus</emphasis>
|
||
<bibRefCitation id="2732E49DFFCCEC33CD72FCBDFDA83E5E" author="Kiesenwetter" box="[378,614,826,852]" pageId="4" pageNumber="417" refString="Kiesenwetter, E. A. H. (1865) Eine entomologische Excursion nach Spanien in Sommer 1865. Berliner Entomologische Zeitschrift, 9, 369 - 375. http: // dx. doi. org / 10.1002 / mmnd. 18650090316" type="journal article" year="1865">Kiesenwetter, 1865</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="431C996CFFCCEC33CC9FFCDAFEFA3E7C" blockId="4.[151,614,826,886]" box="[151,308,861,886]" pageId="4" pageNumber="417">
|
||
(
|
||
<figureCitation id="DB9885E9FFCCEC33CC97FCDAFF293E7C" box="[159,231,861,886]" captionStart="FIGURES 1 – 7. 1" captionStartId="3.[151,264,1748,1770]" captionTargetBox="[166,1428,776,1720]" captionTargetId="figure@3.[151,1436,772,1724]" captionTargetPageId="3" captionText="FIGURES 1 – 7. 1 — The maximum likelihood (ML) Elateroidea phylogenetic tree of 557 taxa aligned by BlastAlign and inferred from RAxML; 2 — Omalisidae phylogeny inferred from RAxML. The values at branches represent bootstrap values inferred from ML analyses of datasets aligned by BlastAlign and Mafft, respectively. 3 — Euanoma marketae Kundrata & Bocak, 2007 (holotype); 4 — Pseudeuanoma sp.; 5 — Paradrilus opacus Kiesenwetter, 1865 (lectotype); 6 — Phaeopterus unicolor Costa, 1857 (holotype); 7 — Omalisus fontisbellaquei Geoffroy, 1762. Scale 2 mm (Figs 3 – 7)." httpUri="https://zenodo.org/record/232722/files/figure.png" pageId="4" pageNumber="417">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="DB9885E9FFCCEC33CCFBFCD9FEE33E7C" box="[243,301,862,886]" captionStart="FIGURES 8 – 18" captionStartId="5.[151,264,1629,1651]" captionTargetBox="[162,1421,634,1597]" captionTargetId="figure@5.[151,1436,625,1608]" captionTargetPageId="5" captionText="FIGURES 8 – 18. Paradrilus opacus Kiesenwetter, 1865; 8 — antenna; 9 — mandible; 10 — maxilla; 11 — labium; 12 — hypopharynx; 13 — pronotum; 14 — prosternum; 15 — wing; 16 — leg; 17 — last abdominal segments; 18 — aedeagus. Scale 0.05 mm (Fig. 11), 0.1 mm (Figs 9 – 10, 12, 17 – 18), 0.2 mm (Figs 13 – 14, 16), 0.5 mm (Figs 8, 15)." httpUri="https://zenodo.org/record/232723/files/figure.png" pageId="4" pageNumber="417">8–18</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB9CAE7FFCCEC33CC9FFC22FD993EB1" box="[151,599,933,955]" pageId="4" pageNumber="417" type="reference_group">
|
||
<paragraph id="431C996CFFCCEC33CC9FFC22FD993EB1" blockId="4.[151,599,933,955]" box="[151,599,933,955]" pageId="4" pageNumber="417">
|
||
<treatmentCitationGroup id="63B3BE42FFCCEC33CC9FFC22FD993EB1" box="[151,599,933,955]" pageId="4" pageNumber="417">
|
||
<taxonomicName id="84A3E2EFFFCCEC33CC9FFC22FD9D3EB1" authority="Kiesenwetter, 1865: 369" authorityName="Kiesenwetter" authorityPageNumber="369" authorityYear="1865" box="[151,595,933,955]" class="Insecta" family="Drilidae" genus="Paradrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="417" phylum="Arthropoda" rank="species" species="opacus">
|
||
<emphasis id="71D7457EFFCCEC33CC9FFC22FE9C3EB1" box="[151,338,933,955]" italics="true" pageId="4" pageNumber="417">Paradrilus opacus</emphasis>
|
||
<treatmentCitation id="C202BF7DFFCCEC33CD51FC22FD9D3EB1" author="Kiesenwetter" box="[345,595,933,955]" page="369" pageId="4" pageNumber="417" year="1865">
|
||
<bibRefCitation id="2732E49DFFCCEC33CD51FC22FDEE3EB1" author="Kiesenwetter" box="[345,544,933,955]" pageId="4" pageNumber="417" refString="Kiesenwetter, E. A. H. (1865) Eine entomologische Excursion nach Spanien in Sommer 1865. Berliner Entomologische Zeitschrift, 9, 369 - 375. http: // dx. doi. org / 10.1002 / mmnd. 18650090316" type="journal article" year="1865">Kiesenwetter, 1865</bibRefCitation>
|
||
: 369
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB9CAE7FFCCEC33CC9FFC60FE1D39D2" pageId="4" pageNumber="417" type="materials_examined">
|
||
<paragraph id="431C996CFFCCEC33CC9FFC60FE153942" blockId="4.[151,1437,999,2033]" pageId="4" pageNumber="417">
|
||
<emphasis id="71D7457EFFCCEC33CC9FFC60FEF1390A" bold="true" box="[151,319,999,1024]" pageId="4" pageNumber="417">
|
||
<typeStatus id="9C1827CEFFCCEC33CC9FFC60FF1C390A" box="[151,210,999,1024]" pageId="4" pageNumber="417">Type</typeStatus>
|
||
material
|
||
</emphasis>
|
||
.
|
||
<typeStatus id="9C1827CEFFCCEC33CD46FC6FFE70390A" box="[334,446,1000,1024]" pageId="4" pageNumber="417" type="lectotype">Lectotype</typeStatus>
|
||
[here designated], male, "Andalusia, Kiesenwetter, 326. 33." [invalidly designated as a
|
||
<typeStatus id="9C1827CEFFCCEC33CC9FFB8AFF39392F" box="[151,247,1037,1061]" pageId="4" pageNumber="417" type="paratype">paratype</typeStatus>
|
||
by a local curator];
|
||
<typeStatus id="9C1827CEFFCCEC33CDD5FB8BFDBB392F" box="[477,629,1036,1061]" pageId="4" pageNumber="417" type="paralectotype">Paralectotype</typeStatus>
|
||
[here designated], male, the same data as
|
||
<typeStatus id="9C1827CEFFCCEC33C85BFB8BFB74392F" box="[1107,1210,1036,1061]" pageId="4" pageNumber="417" type="lectotype">lectotype</typeStatus>
|
||
(Hungarian Natural History Museum, Budapest).
|
||
</paragraph>
|
||
<paragraph id="431C996CFFCCEC33CCCFFBD3FE1D39D2" blockId="4.[151,1437,999,2033]" pageId="4" pageNumber="417">
|
||
<emphasis id="71D7457EFFCCEC33CCCFFBD3FE353967" bold="true" box="[199,507,1108,1133]" pageId="4" pageNumber="417">Other material examined</emphasis>
|
||
.
|
||
<specimenCount id="55A552E5FFCCEC33CE04FBD3FDA63967" box="[524,616,1108,1133]" pageId="4" pageNumber="417" type="male">2 males</specimenCount>
|
||
, "Andalusien" (Naturhistorisches Museum, Basel),
|
||
<specimenCount id="55A552E5FFCCEC33C8B3FBD3FAD93967" box="[1211,1303,1108,1133]" pageId="4" pageNumber="417" type="male">2 males</specimenCount>
|
||
, "Andalus., Kratz, 64." (Muséum national d'Histoire naturelle, Paris),
|
||
<specimenCount id="55A552E5FFCCEC33CF32FBFFFC42399A" box="[826,908,1143,1168]" pageId="4" pageNumber="417" type="male">1 male</specimenCount>
|
||
, "
|
||
<collectingCountry id="3BB4D9FCFFCCEC33CFACFBF0FC27399A" box="[932,1001,1143,1168]" name="Spain" pageId="4" pageNumber="417">Spain</collectingCountry>
|
||
, Córdoba, Santa María de Trassierra, Arroyo del Molino,
|
||
<date id="371DBFACFFCCEC33CD75FB1BFDFE39BE" box="[381,560,1180,1205]" pageId="4" pageNumber="417" value="2012-05-20" valueMax="2012-05-25" valueMin="2012-05-20">20.–25. v. 2012</date>
|
||
, M. Baena leg." (
|
||
<collectionCode id="25B201A9FFCCEC33CEF1FB1BFCF639BE" LSID="urn:lsid:biocol.org:col:15536" box="[761,824,1180,1204]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="4" pageNumber="417">DNA</collectionCode>
|
||
voucher specimen RK0626; Laboratory of Molecular Systematics, UP, Olomouc).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB9CAE7FFCCEC32CCCFFB63FEF13AC2" lastPageId="5" lastPageNumber="418" pageId="4" pageNumber="417" type="description">
|
||
<paragraph id="431C996CFFCCEC33CCCFFB63FB06384F" blockId="4.[151,1437,999,2033]" pageId="4" pageNumber="417">
|
||
<emphasis id="71D7457EFFCCEC33CCCFFB63FEA339F7" bold="true" box="[199,365,1252,1277]" pageId="4" pageNumber="417">Redescription</emphasis>
|
||
. Adult, male. Body slender,
|
||
<quantity id="845B3489FFCCEC33CEB0FB63FC8F39F6" box="[696,833,1252,1276]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="4" pageNumber="417" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.2–2.7 mm</quantity>
|
||
long, 3.7 times longer than width at humeri, weakly dorso-ventrally flattened, parallel-sided (
|
||
<figureCitation id="DB9885E9FFCCEC33CE7DFA80FD71382A" box="[629,703,1287,1312]" captionStart="FIGURES 1 – 7. 1" captionStartId="3.[151,264,1748,1770]" captionTargetBox="[166,1428,776,1720]" captionTargetId="figure@3.[151,1436,772,1724]" captionTargetPageId="3" captionText="FIGURES 1 – 7. 1 — The maximum likelihood (ML) Elateroidea phylogenetic tree of 557 taxa aligned by BlastAlign and inferred from RAxML; 2 — Omalisidae phylogeny inferred from RAxML. The values at branches represent bootstrap values inferred from ML analyses of datasets aligned by BlastAlign and Mafft, respectively. 3 — Euanoma marketae Kundrata & Bocak, 2007 (holotype); 4 — Pseudeuanoma sp.; 5 — Paradrilus opacus Kiesenwetter, 1865 (lectotype); 6 — Phaeopterus unicolor Costa, 1857 (holotype); 7 — Omalisus fontisbellaquei Geoffroy, 1762. Scale 2 mm (Figs 3 – 7)." httpUri="https://zenodo.org/record/232722/files/figure.png" pageId="4" pageNumber="417">Fig. 5</figureCitation>
|
||
); moderately sclerotized, cuticle of elytra and abdomen soft, flexible; dark brown to black colored, surface mat, finely punctured, with dense, short vestiture.
|
||
</paragraph>
|
||
<paragraph id="431C996CFFCCEC33CCCFFAC8FDC73AA2" blockId="4.[151,1437,999,2033]" pageId="4" pageNumber="417">
|
||
Head small, hypognathous, slightly narrower than prothorax, exposed from prothorax, with fully developed mouthparts; cranium prolonged anteriorly, with inconspicuous antennal sockets, antennal insertions fronto-lateral, separated by narrow bridge, about 0.6 times width of maximum diameter of antennal cavity. Eyes small, moderately prominent, eye diameter 1.4 times their frontal distance. Antennae filiform, 11-segmented, antennomeres round in cross section (
|
||
<figureCitation id="DB9885E9FFCCEC33CE4AFA58FD4938F2" box="[578,647,1503,1528]" captionStart="FIGURES 8 – 18" captionStartId="5.[151,264,1629,1651]" captionTargetBox="[162,1421,634,1597]" captionTargetId="figure@5.[151,1436,625,1608]" captionTargetPageId="5" captionText="FIGURES 8 – 18. Paradrilus opacus Kiesenwetter, 1865; 8 — antenna; 9 — mandible; 10 — maxilla; 11 — labium; 12 — hypopharynx; 13 — pronotum; 14 — prosternum; 15 — wing; 16 — leg; 17 — last abdominal segments; 18 — aedeagus. Scale 0.05 mm (Fig. 11), 0.1 mm (Figs 9 – 10, 12, 17 – 18), 0.2 mm (Figs 13 – 14, 16), 0.5 mm (Figs 8, 15)." httpUri="https://zenodo.org/record/232723/files/figure.png" pageId="4" pageNumber="417">Fig. 8</figureCitation>
|
||
); scape robust, parallel-sided, constricted at base, apex asymmetrical; pedicel slightly triangular, robust, constricted at base, widely rounded apically; antennomere 3 shortest, triangular; antennomere 4 much longer; following antennomeres gradually shortened except terminal one. All antennomeres with moderately dense, erect pubescence. Fronto-clypeal suture absent; anterior edge of clypeus concave; labrum membranous, hypopharynx T-shaped, sclerotized, lateral processes long, suboesophaginal ring absent (
|
||
<figureCitation id="DB9885E9FFCCEC33C930F9E8FA413B82" box="[1336,1423,1647,1672]" captionStart="FIGURES 8 – 18" captionStartId="5.[151,264,1629,1651]" captionTargetBox="[162,1421,634,1597]" captionTargetId="figure@5.[151,1436,625,1608]" captionTargetPageId="5" captionText="FIGURES 8 – 18. Paradrilus opacus Kiesenwetter, 1865; 8 — antenna; 9 — mandible; 10 — maxilla; 11 — labium; 12 — hypopharynx; 13 — pronotum; 14 — prosternum; 15 — wing; 16 — leg; 17 — last abdominal segments; 18 — aedeagus. Scale 0.05 mm (Fig. 11), 0.1 mm (Figs 9 – 10, 12, 17 – 18), 0.2 mm (Figs 13 – 14, 16), 0.5 mm (Figs 8, 15)." httpUri="https://zenodo.org/record/232723/files/figure.png" pageId="4" pageNumber="417">Fig. 12</figureCitation>
|
||
). Mandibles slender apically, robust at base, slightly curved; incisor without teeth (
|
||
<figureCitation id="DB9885E9FFCCEC33C82BF913FBA63BA6" box="[1059,1128,1684,1709]" captionStart="FIGURES 8 – 18" captionStartId="5.[151,264,1629,1651]" captionTargetBox="[162,1421,634,1597]" captionTargetId="figure@5.[151,1436,625,1608]" captionTargetPageId="5" captionText="FIGURES 8 – 18. Paradrilus opacus Kiesenwetter, 1865; 8 — antenna; 9 — mandible; 10 — maxilla; 11 — labium; 12 — hypopharynx; 13 — pronotum; 14 — prosternum; 15 — wing; 16 — leg; 17 — last abdominal segments; 18 — aedeagus. Scale 0.05 mm (Fig. 11), 0.1 mm (Figs 9 – 10, 12, 17 – 18), 0.2 mm (Figs 13 – 14, 16), 0.5 mm (Figs 8, 15)." httpUri="https://zenodo.org/record/232723/files/figure.png" pageId="4" pageNumber="417">Fig. 9</figureCitation>
|
||
). Maxilla with small cardo, stipes well developed, plate-shaped; galea and lacinia fused in setose, mala basally sclerotized; palpifer triangular, robust; maxillary palpi four-segmented, palpomere 1 small, transverse, palpomeres 2 and 3 robust, about as long as wide, apical palpomere twice longer than width at base, slender apically, apex widely rounded (
|
||
<figureCitation id="DB9885E9FFCCEC33C8D2F978FAE13A12" box="[1242,1327,1791,1816]" captionStart="FIGURES 8 – 18" captionStartId="5.[151,264,1629,1651]" captionTargetBox="[162,1421,634,1597]" captionTargetId="figure@5.[151,1436,625,1608]" captionTargetPageId="5" captionText="FIGURES 8 – 18. Paradrilus opacus Kiesenwetter, 1865; 8 — antenna; 9 — mandible; 10 — maxilla; 11 — labium; 12 — hypopharynx; 13 — pronotum; 14 — prosternum; 15 — wing; 16 — leg; 17 — last abdominal segments; 18 — aedeagus. Scale 0.05 mm (Fig. 11), 0.1 mm (Figs 9 – 10, 12, 17 – 18), 0.2 mm (Figs 13 – 14, 16), 0.5 mm (Figs 8, 15)." httpUri="https://zenodo.org/record/232723/files/figure.png" pageId="4" pageNumber="417">Fig. 10</figureCitation>
|
||
). Labium small, praementum short, wide, reduced to convex plate with extensive palpal cavities, projected and pointed apically, without ligula or apical setae; postmentum fused with gula, sclerotized, plate-like, about as long as width; labial palpi tiny, two-segmented, basal palpomere wide, transverse, apical palpomere slender, almost parallel-sided, weakly pointed at apex (
|
||
<figureCitation id="DB9885E9FFCCEC33CDA1F808FE353AA2" box="[425,507,1935,1960]" captionStart="FIGURES 8 – 18" captionStartId="5.[151,264,1629,1651]" captionTargetBox="[162,1421,634,1597]" captionTargetId="figure@5.[151,1436,625,1608]" captionTargetPageId="5" captionText="FIGURES 8 – 18. Paradrilus opacus Kiesenwetter, 1865; 8 — antenna; 9 — mandible; 10 — maxilla; 11 — labium; 12 — hypopharynx; 13 — pronotum; 14 — prosternum; 15 — wing; 16 — leg; 17 — last abdominal segments; 18 — aedeagus. Scale 0.05 mm (Fig. 11), 0.1 mm (Figs 9 – 10, 12, 17 – 18), 0.2 mm (Figs 13 – 14, 16), 0.5 mm (Figs 8, 15)." httpUri="https://zenodo.org/record/232723/files/figure.png" pageId="4" pageNumber="417">Fig. 11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="431C996CFFCCEC32CCCFF833FE9C3F37" blockId="4.[151,1437,999,2033]" lastBlockId="5.[151,1437,151,573]" lastPageId="5" lastPageNumber="418" pageId="4" pageNumber="417">
|
||
Prothorax about as long as width (pronotum length/width ratio=0.9), anterior edge straight, frontal angles obtuse, anterior foramen widely rounded; posterior angles obtuse, posterior margin weakly emarginate in middle; lateral edges widely convex, widest in frontal fourth, straight posteriorly, pronotum narrower basally than elytral humeri, disc convex, lateral margins bent downwards, with inconspicuous pleura anteriorly, disc without any carinae or keel, mat, finely structured, with punctures at frontal and posterior corners (
|
||
<figureCitation id="DB9885E9FFCDEC32C8A0FF58FACF3DF2" box="[1192,1281,223,248]" captionStart="FIGURES 8 – 18" captionStartId="5.[151,264,1629,1651]" captionTargetBox="[162,1421,634,1597]" captionTargetId="figure@5.[151,1436,625,1608]" captionTargetPageId="5" captionText="FIGURES 8 – 18. Paradrilus opacus Kiesenwetter, 1865; 8 — antenna; 9 — mandible; 10 — maxilla; 11 — labium; 12 — hypopharynx; 13 — pronotum; 14 — prosternum; 15 — wing; 16 — leg; 17 — last abdominal segments; 18 — aedeagus. Scale 0.05 mm (Fig. 11), 0.1 mm (Figs 9 – 10, 12, 17 – 18), 0.2 mm (Figs 13 – 14, 16), 0.5 mm (Figs 8, 15)." httpUri="https://zenodo.org/record/232723/files/figure.png" pageId="5" pageNumber="418">Fig. 13</figureCitation>
|
||
). Prosternum moderately long, with wide parallel-sided prosternal process bearing divergent short and robust branches at apex (
|
||
<figureCitation id="DB9885E9FFCDEC32CC97FEA0FF3A3C4A" box="[159,244,295,320]" captionStart="FIGURES 8 – 18" captionStartId="5.[151,264,1629,1651]" captionTargetBox="[162,1421,634,1597]" captionTargetId="figure@5.[151,1436,625,1608]" captionTargetPageId="5" captionText="FIGURES 8 – 18. Paradrilus opacus Kiesenwetter, 1865; 8 — antenna; 9 — mandible; 10 — maxilla; 11 — labium; 12 — hypopharynx; 13 — pronotum; 14 — prosternum; 15 — wing; 16 — leg; 17 — last abdominal segments; 18 — aedeagus. Scale 0.05 mm (Fig. 11), 0.1 mm (Figs 9 – 10, 12, 17 – 18), 0.2 mm (Figs 13 – 14, 16), 0.5 mm (Figs 8, 15)." httpUri="https://zenodo.org/record/232723/files/figure.png" pageId="5" pageNumber="418">Fig. 14</figureCitation>
|
||
). Scutellum bilobate. Mesoventrite wide, coxae in extensive cavities laterally. Metasternum long, pleural part connected to metasternum with membrane, epimeron and episternum slender. Elytra very slender, tapering to apex, with humeral edge and pleura present in basal third, posterior half of elytra flat, elytra without longitudinal costae, surface with fine papillae bearing apical setae; elytral apexes separately rounded (
|
||
<figureCitation id="DB9885E9FFCDEC32C898FE13FB183CA7" box="[1168,1238,404,429]" captionStart="FIGURES 1 – 7. 1" captionStartId="3.[151,264,1748,1770]" captionTargetBox="[166,1428,776,1720]" captionTargetId="figure@3.[151,1436,772,1724]" captionTargetPageId="3" captionText="FIGURES 1 – 7. 1 — The maximum likelihood (ML) Elateroidea phylogenetic tree of 557 taxa aligned by BlastAlign and inferred from RAxML; 2 — Omalisidae phylogeny inferred from RAxML. The values at branches represent bootstrap values inferred from ML analyses of datasets aligned by BlastAlign and Mafft, respectively. 3 — Euanoma marketae Kundrata & Bocak, 2007 (holotype); 4 — Pseudeuanoma sp.; 5 — Paradrilus opacus Kiesenwetter, 1865 (lectotype); 6 — Phaeopterus unicolor Costa, 1857 (holotype); 7 — Omalisus fontisbellaquei Geoffroy, 1762. Scale 2 mm (Figs 3 – 7)." httpUri="https://zenodo.org/record/232722/files/figure.png" pageId="5" pageNumber="418">Fig. 5</figureCitation>
|
||
). Wing as in
|
||
<figureCitation id="DB9885E9FFCDEC32C978FE13FF783CDA" captionStart="FIGURES 8 – 18" captionStartId="5.[151,264,1629,1651]" captionTargetBox="[162,1421,634,1597]" captionTargetId="figure@5.[151,1436,625,1608]" captionTargetPageId="5" captionText="FIGURES 8 – 18. Paradrilus opacus Kiesenwetter, 1865; 8 — antenna; 9 — mandible; 10 — maxilla; 11 — labium; 12 — hypopharynx; 13 — pronotum; 14 — prosternum; 15 — wing; 16 — leg; 17 — last abdominal segments; 18 — aedeagus. Scale 0.05 mm (Fig. 11), 0.1 mm (Figs 9 – 10, 12, 17 – 18), 0.2 mm (Figs 13 – 14, 16), 0.5 mm (Figs 8, 15)." httpUri="https://zenodo.org/record/232723/files/figure.png" pageId="5" pageNumber="418">Fig. 15</figureCitation>
|
||
. Legs slender, compressed (
|
||
<figureCitation id="DB9885E9FFCDEC32CE1FFE30FDBB3CDA" box="[535,629,439,464]" captionStart="FIGURES 8 – 18" captionStartId="5.[151,264,1629,1651]" captionTargetBox="[162,1421,634,1597]" captionTargetId="figure@5.[151,1436,625,1608]" captionTargetPageId="5" captionText="FIGURES 8 – 18. Paradrilus opacus Kiesenwetter, 1865; 8 — antenna; 9 — mandible; 10 — maxilla; 11 — labium; 12 — hypopharynx; 13 — pronotum; 14 — prosternum; 15 — wing; 16 — leg; 17 — last abdominal segments; 18 — aedeagus. Scale 0.05 mm (Fig. 11), 0.1 mm (Figs 9 – 10, 12, 17 – 18), 0.2 mm (Figs 13 – 14, 16), 0.5 mm (Figs 8, 15)." httpUri="https://zenodo.org/record/232723/files/figure.png" pageId="5" pageNumber="418">Fig. 16</figureCitation>
|
||
); coxae separated, elongate; trochanters slender, moderately long, transversally cut at apex; femora moderately robust; tibiae slender, with pair of small spurs apically; tarsi with five slender tarsomeres, tarsomeres 1–3 subequal, tarsomere 4 short, tarsomere 5 slender, long; pulvilli absent; claws simple (
|
||
<figureCitation id="DB9885E9FFCDEC32CCF9FDA3FE8A3F37" box="[241,324,548,573]" captionStart="FIGURES 8 – 18" captionStartId="5.[151,264,1629,1651]" captionTargetBox="[162,1421,634,1597]" captionTargetId="figure@5.[151,1436,625,1608]" captionTargetPageId="5" captionText="FIGURES 8 – 18. Paradrilus opacus Kiesenwetter, 1865; 8 — antenna; 9 — mandible; 10 — maxilla; 11 — labium; 12 — hypopharynx; 13 — pronotum; 14 — prosternum; 15 — wing; 16 — leg; 17 — last abdominal segments; 18 — aedeagus. Scale 0.05 mm (Fig. 11), 0.1 mm (Figs 9 – 10, 12, 17 – 18), 0.2 mm (Figs 13 – 14, 16), 0.5 mm (Figs 8, 15)." httpUri="https://zenodo.org/record/232723/files/figure.png" pageId="5" pageNumber="418">Fig. 16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="17DCC9E4FFCDEC32CC9FF9DAFBB53BBA" httpUri="https://zenodo.org/record/232723/files/figure.png" pageId="5" pageNumber="418" targetBox="[162,1421,634,1597]" targetPageId="5">
|
||
<paragraph id="431C996CFFCDEC32CC9FF9DAFBB53BBA" blockId="5.[151,1436,1629,1712]" pageId="5" pageNumber="418">
|
||
<emphasis id="71D7457EFFCDEC32CC9FF9DAFE963B79" bold="true" box="[151,344,1629,1651]" pageId="5" pageNumber="418">FIGURES 8–18.</emphasis>
|
||
<taxonomicName id="84A3E2EFFFCDEC32CD78F9DAFCE33B79" authority="Kiesenwetter, 1865" authorityName="Kiesenwetter" authorityYear="1865" box="[368,813,1629,1651]" class="Insecta" family="Drilidae" genus="Paradrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="418" phylum="Arthropoda" rank="species" species="opacus">
|
||
<emphasis id="71D7457EFFCDEC32CD78F9DAFDF33B79" box="[368,573,1629,1651]" italics="true" pageId="5" pageNumber="418">Paradrilus opacus</emphasis>
|
||
Kiesenwetter, 1865
|
||
</taxonomicName>
|
||
; 8—antenna; 9—mandible; 10—maxilla; 11—labium; 12—hypopharynx; 13—pronotum; 14—prosternum; 15—wing; 16—leg; 17—last abdominal segments; 18—aedeagus. Scale 0.05 mm (Fig. 11), 0.1 mm (Figs 9–10, 12, 17–18), 0.2 mm (Figs 13–14, 16), 0.5 mm (Figs 8, 15).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="431C996CFFCDEC32CCCFF95FFE713A56" blockId="5.[151,1436,1752,1993]" pageId="5" pageNumber="418">
|
||
Abdomen with eight visible ventrites, basal ventrite membranous in middle part, terminal male segments as in
|
||
<figureCitation id="DB9885E9FFCDEC32CC9FF97BFF273A1E" box="[151,233,1788,1813]" captionStart="FIGURES 8 – 18" captionStartId="5.[151,264,1629,1651]" captionTargetBox="[162,1421,634,1597]" captionTargetId="figure@5.[151,1436,625,1608]" captionTargetPageId="5" captionText="FIGURES 8 – 18. Paradrilus opacus Kiesenwetter, 1865; 8 — antenna; 9 — mandible; 10 — maxilla; 11 — labium; 12 — hypopharynx; 13 — pronotum; 14 — prosternum; 15 — wing; 16 — leg; 17 — last abdominal segments; 18 — aedeagus. Scale 0.05 mm (Fig. 11), 0.1 mm (Figs 9 – 10, 12, 17 – 18), 0.2 mm (Figs 13 – 14, 16), 0.5 mm (Figs 8, 15)." httpUri="https://zenodo.org/record/232723/files/figure.png" pageId="5" pageNumber="418">Fig. 17</figureCitation>
|
||
. Aedeagus of trilobate
|
||
<typeStatus id="9C1827CEFFCDEC32CDE5F97AFDEE3A1F" box="[493,544,1789,1813]" pageId="5" pageNumber="418">type</typeStatus>
|
||
, with long parameres; apices of parameres simple, partly membranous; internal sac inconspicuous, membranous; phallobase plate-like, v-shaped, with strengthened lateral margins and posterolateral processes (
|
||
<figureCitation id="DB9885E9FFCDEC32CD56F8C3FE7C3A56" box="[350,434,1860,1885]" captionStart="FIGURES 8 – 18" captionStartId="5.[151,264,1629,1651]" captionTargetBox="[162,1421,634,1597]" captionTargetId="figure@5.[151,1436,625,1608]" captionTargetPageId="5" captionText="FIGURES 8 – 18. Paradrilus opacus Kiesenwetter, 1865; 8 — antenna; 9 — mandible; 10 — maxilla; 11 — labium; 12 — hypopharynx; 13 — pronotum; 14 — prosternum; 15 — wing; 16 — leg; 17 — last abdominal segments; 18 — aedeagus. Scale 0.05 mm (Fig. 11), 0.1 mm (Figs 9 – 10, 12, 17 – 18), 0.2 mm (Figs 13 – 14, 16), 0.5 mm (Figs 8, 15)." httpUri="https://zenodo.org/record/232723/files/figure.png" pageId="5" pageNumber="418">Fig. 18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="431C996CFFCDEC32CCCFF8EFFEF13AC2" blockId="5.[151,1436,1752,1993]" pageId="5" pageNumber="418">
|
||
<emphasis id="71D7457EFFCDEC32CCCFF8EFFE383A8B" bold="true" box="[199,502,1896,1921]" pageId="5" pageNumber="418">
|
||
Measurements (
|
||
<typeStatus id="9C1827CEFFCDEC32CD81F8EFFE383A8B" box="[393,502,1896,1921]" pageId="5" pageNumber="418" type="lectotype">lectotype</typeStatus>
|
||
</emphasis>
|
||
). Body length
|
||
<quantity id="845B3489FFCDEC32CEA2F8EFFCDA3A8A" box="[682,788,1896,1920]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="5" pageNumber="418" unit="mm" value="2.7">2.70 mm</quantity>
|
||
, width at humeri
|
||
<quantity id="845B3489FFCDEC32CFE1F8EFFB9A3A8A" box="[1001,1108,1896,1920]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="5" pageNumber="418" unit="mm" value="0.73">0.73 mm</quantity>
|
||
, pronotum length
|
||
<quantity id="845B3489FFCDEC32C925F8EFFA563A8A" box="[1325,1432,1896,1921]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="418" unit="mm" value="0.5">0.50 mm</quantity>
|
||
; pronotum width
|
||
<quantity id="845B3489FFCDEC32CD50F80BFE0F3AAE" box="[344,449,1932,1957]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="5" pageNumber="418" unit="mm" value="0.57">0.57 mm</quantity>
|
||
, minimum frontal distance between eyes
|
||
<quantity id="845B3489FFCDEC32CFADF80BFBC13AAE" box="[933,1039,1932,1956]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="418" unit="mm" value="0.3">0.30 mm</quantity>
|
||
, maximum eye diameter in lateral view
|
||
<quantity id="845B3489FFCDEC32CCDCF837FEF53AC2" box="[212,315,1968,1992]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="418" unit="mm" value="0.21">0.21 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB9CAE7FFCEEC31CCCFFF10FDB03DDF" pageId="6" pageNumber="419" type="distribution">
|
||
<paragraph id="431C996CFFCEEC31CCCFFF10FDB03DDF" blockId="6.[151,1437,151,717]" pageId="6" pageNumber="419">
|
||
<emphasis id="71D7457EFFCEEC31CCCFFF10FE973DBA" bold="true" box="[199,345,151,176]" pageId="6" pageNumber="419">Distribution</emphasis>
|
||
. Southern
|
||
<collectingCountry id="3BB4D9FCFFCEEC31CDDEFF10FDD83DBA" box="[470,534,151,176]" name="Spain" pageId="6" pageNumber="419">Spain</collectingCountry>
|
||
(Andalusia: Córdoba, Huelva, Jaén, and Sevilla provinces;
|
||
<bibRefCitation id="2732E49DFFCEEC31C8C9FF10FA563DBA" author="Kiesenwetter" box="[1217,1432,151,176]" pageId="6" pageNumber="419" refString="Kiesenwetter, E. A. H. (1866) Beitrage zur Kaferfauna Spaniens. Erstes Stuck. Malacodermata, Melyridae. Berliner Entomologische Zeitschrift, 10, 241 - 274. http: // dx. doi. org / 10.1002 / mmnd. 18660100117" type="journal article" year="1866">Kiesenwetter 1866</bibRefCitation>
|
||
, Bahillo de la Puebla & López Colón 2005).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB9CAE7FFCEEC31CCCFFF58FD2D3C16" pageId="6" pageNumber="419" type="biology_ecology">
|
||
<paragraph id="431C996CFFCEEC31CCCFFF58FD2D3C16" blockId="6.[151,1437,151,717]" pageId="6" pageNumber="419">
|
||
<emphasis id="71D7457EFFCEEC31CCCFFF58FE7D3DF2" bold="true" box="[199,435,223,248]" pageId="6" pageNumber="419">Biology and ecology</emphasis>
|
||
. The specimens were collected by sweeping of the herbal stratum in a forested place near Santa María de Trassierra, Córdoba,
|
||
<collectingCountry id="3BB4D9FCFFCEEC31CE3BFE83FDBD3C17" box="[563,627,260,285]" name="Spain" pageId="6" pageNumber="419">Spain</collectingCountry>
|
||
(
|
||
<figureCitation id="DB9885E9FFCEEC31CE8AFE83FD1B3C16" box="[642,725,260,285]" captionStart="FIGURE 19" captionStartId="6.[151,250,1775,1797]" captionTargetBox="[151,1436,772,1754]" captionTargetId="figure@6.[151,1436,772,1754]" captionTargetPageId="6" captionText="FIGURE 19. Habitat of Paradrilus opacus Kiesenwetter, 1865 — Arroyo del Molino, Santa María de Trassierra, Córdoba, Spain." httpUri="https://zenodo.org/record/232724/files/figure.png" pageId="6" pageNumber="419">Fig. 19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0BB9CAE7FFCEEC31CCCFFEA0FDB73FC7" pageId="6" pageNumber="419" type="discussion">
|
||
<paragraph id="431C996CFFCEEC31CCCFFEA0FDB83C82" blockId="6.[151,1437,151,717]" pageId="6" pageNumber="419">
|
||
<emphasis id="71D7457EFFCEEC31CCCFFEA0FEFA3C4A" bold="true" box="[199,308,295,320]" pageId="6" pageNumber="419">Remarks</emphasis>
|
||
. The female and larva are unknown.
|
||
<collectingCountry id="3BB4D9FCFFCEEC31CEFBFEAFFCDC3C4A" box="[755,786,296,320]" name="American Samoa" pageId="6" pageNumber="419">As</collectingCountry>
|
||
only males are present in all examined collections, we suppose that the females of
|
||
<taxonomicName id="84A3E2EFFFCEEC31CE01FECBFD4D3C6F" box="[521,643,332,357]" class="Insecta" family="Drilidae" genus="Paradrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="71D7457EFFCEEC31CE01FECBFD4D3C6F" box="[521,643,332,357]" italics="true" pageId="6" pageNumber="419">Paradrilus</emphasis>
|
||
</taxonomicName>
|
||
do not complete metamorphosis similarly to other
|
||
<taxonomicName id="84A3E2EFFFCEEC31C914FECBFA523C6F" box="[1308,1436,332,357]" class="Insecta" family="Omalisidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="419" phylum="Arthropoda" rank="family">Omalisidae</taxonomicName>
|
||
(
|
||
<bibRefCitation id="2732E49DFFCEEC31CC96FEE8FEB83C82" author="Kiesenwetter" box="[158,374,367,392]" pageId="6" pageNumber="419" refString="Kiesenwetter, E. A. H. (1866) Beitrage zur Kaferfauna Spaniens. Erstes Stuck. Malacodermata, Melyridae. Berliner Entomologische Zeitschrift, 10, 241 - 274. http: // dx. doi. org / 10.1002 / mmnd. 18660100117" type="journal article" year="1866">Kiesenwetter 1866</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="2732E49DFFCEEC31CD89FEE8FDA43C82" author="Bocak" box="[385,618,367,392]" pageId="6" pageNumber="419" refString="Bocak, L. & Brlik, M. (2008) Revision of the family Omalisidae (Coleoptera, Elateroidea). Insect Systematics & Evolution, 39, 189 - 212. http: // dx. doi. org / 10.1163 / 187631208788784101" type="journal article" year="2008">Bocak & Brlik 2008</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="431C996CFFCEEC31CCCFFE13FDB73FC7" blockId="6.[151,1437,151,717]" pageId="6" pageNumber="419">
|
||
The
|
||
<typeStatus id="9C1827CEFFCEEC31CCF6FE12FEE03CA7" box="[254,302,405,429]" pageId="6" pageNumber="419">type</typeStatus>
|
||
material is not present in the Kiesenwetter collection in Zoologische Staatssammlung München,
|
||
<collectingCountry id="3BB4D9FCFFCEEC31CC9FFE3FFF313CDA" box="[151,255,440,464]" name="Germany" pageId="6" pageNumber="419">Germany</collectingCountry>
|
||
(K. Neven, personal communication) and might be distributed in several museums. We were able to locate
|
||
<typeStatus id="9C1827CEFFCEEC31CCECFE5AFED13CFF" box="[228,287,477,501]" pageId="6" pageNumber="419">types</typeStatus>
|
||
only in the Budapest museum and several conspecific individuals elsewhere. The specimens in the Budapest collection were designated by a curator as
|
||
<typeStatus id="9C1827CEFFCEEC31CEECFD87FC9C3F12" box="[740,850,512,536]" pageId="6" pageNumber="419" type="paratype">paratypes</typeStatus>
|
||
, but there is no indication in the original description that
|
||
<typeStatus id="9C1827CEFFCEEC31CCC0FDA3FEE43F37" box="[200,298,548,573]" pageId="6" pageNumber="419" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="9C1827CEFFCEEC31CD69FDA2FE023F37" box="[353,460,549,573]" pageId="6" pageNumber="419" type="paratype">paratypes</typeStatus>
|
||
were designated and all
|
||
<typeStatus id="9C1827CEFFCEEC31CEEAFDA2FCD33F37" box="[738,797,549,573]" pageId="6" pageNumber="419">types</typeStatus>
|
||
must be considered as
|
||
<typeStatus id="9C1827CEFFCEEC31C82BFDA2FB4B3F37" box="[1059,1157,549,573]" pageId="6" pageNumber="419" type="syntype">syntypes</typeStatus>
|
||
(
|
||
<bibRefCitation id="2732E49DFFCEEC31C89CFDA3FAD43F36" author="ICZN" box="[1172,1306,548,572]" pageId="6" pageNumber="419" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 305 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
|
||
). We prefer to designate
|
||
<typeStatus id="9C1827CEFFCEEC31CD27FDC0FE5B3F6A" box="[303,405,583,608]" pageId="6" pageNumber="419" type="lectotype">lectotype</typeStatus>
|
||
as the beetles with limited dispersal propensity tend to produce separate, evolutionary isolated populations in a limited geographical space (
|
||
<bibRefCitation id="2732E49DFFCEEC31CEE2FDEBFBDE3F8E" author="Malohlava" box="[746,1040,620,645]" pageId="6" pageNumber="419" refString="Malohlava, V. & Bocak, L. (2010) Evidence of extreme habitat stability in a Southeast Asian biodiversity hotspot based on the evolutionary analysis of neotenic net-winged beetles. Molecular Ecology, 19, 4800 - 4811. http: // dx. doi. org / 10.1111 / j. 1365 - 294 X. 2010.04850. x" type="journal article" year="2010">Malohlava & Bocak 2010</bibRefCitation>
|
||
) and the original series can include several species. The here fixed morphological concept of
|
||
<taxonomicName id="84A3E2EFFFCEEC31CF5EFD08FC013FA2" box="[854,975,655,680]" class="Insecta" family="Drilidae" genus="Paradrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="71D7457EFFCEEC31CF5EFD08FC013FA2" box="[854,975,655,680]" italics="true" pageId="6" pageNumber="419">Paradrilus</emphasis>
|
||
</taxonomicName>
|
||
follows that represented by identified specimens in several European collections.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |