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<document ID-CLB-Dataset="2989" ID-DOI="http://dx.doi.org/10.3897/phytokeys.187.75534" ID-GBIF-Dataset="23192d55-d61d-4058-b71d-8331f18000bd" ID-PMC="PMC8688405" ID-Pensoft-Pub="1314-2003-187-1" ID-Pensoft-UUID="09581AB16B5A5691BE989684EBE4ACF7" ID-PubMed="35002363" ModsDocID="1314-2003-187-1" checkinTime="1639455700515" checkinUser="pensoft" docAuthor="Domina, Gianniantonio, Astuti, Giovanni, Bacchetta, Gianluigi, Barone, Giulio, Resetnik, Ivana, Terlevic, Ana, Thiebaut, Melanie &amp; Peruzzi, Lorenzo" docDate="2021" docId="CAE1C14979A35BA7AE6BF258BCD3A9D1" docLanguage="en" docName="PhytoKeys 187: 1-14" docOrigin="PhytoKeys 187" docPubDate="2021-12-13" docSource="http://dx.doi.org/10.3897/phytokeys.187.75534" docTitle="Dianthus sylvestris Wulfen in Jacq., Coll. 1: 237. 1786. [January-September 1786" docType="treatment" docVersion="6" id="09581AB16B5A5691BE989684EBE4ACF7" lastPageNumber="1" masterDocId="09581AB16B5A5691BE989684EBE4ACF7" masterDocTitle="Typification of 14 names in the Dianthus virgineus group (Caryophyllaceae)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="1" updateTime="1732737314193" updateUser="ExternalLinkService">
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<mods:title id="65E95BD98C024DE4466C70B307AA4E71">Typification of 14 names in the Dianthus virgineus group (Caryophyllaceae)</mods:title>
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<mods:namePart id="DA25C59326CD414D7CD7C74DE85191D1">Domina, Gianniantonio</mods:namePart>
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<mods:namePart id="A08341216B82F972F4F613B000D68605">Astuti, Giovanni</mods:namePart>
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<mods:affiliation id="F0B459534FE59636C65B6A323F25C7D4">Dipartimento di Biologia, Unita di Botanica, Universita di Pisa, Via Derna 1, 56126, Pisa, Italy</mods:affiliation>
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<mods:affiliation id="2017C94AD7C09CD75138DA4566AB20E0">Dipartimento di Scienze della Vita e dell'Ambiente, Centro Conservazione Biodiversita (CCB), Universita di Cagliari, V. le S. Ignazio da Laconi 13, 09123, Cagliari, Italy</mods:affiliation>
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<mods:namePart id="6F79BA78FA1B0D1FD37512A22E515F17">Barone, Giulio</mods:namePart>
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<mods:affiliation id="721521FBE6C21D9F5D335F5B3A51939E">Department of Agriculture, Food and Forest Sciences, University of Palermo, Viale delle Scienze ed. 4, I- 90128 Palermo, Italy</mods:affiliation>
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<mods:affiliation id="52946E0708E11B7F854D5DD3098F53A3">Faculty of Science, Department of Biology, University of Zagreb, Marulicev trg 20 / II, 10000, Zagreb, Croatia</mods:affiliation>
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<mods:affiliation id="F328A8BA992327431FA2242AB097DF2D">Faculty of Science, Department of Biology, University of Zagreb, Marulicev trg 20 / II, 10000, Zagreb, Croatia</mods:affiliation>
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<mods:affiliation id="221BAE26EF340A769E1B9E0012FE72C5">Dipartimento di Biologia, Unita di Botanica, Universita di Pisa, Via Derna 1, 56126, Pisa, Italy</mods:affiliation>
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<mods:date id="509CC16FAD7D3EDA5FA409333333DBDD">2021</mods:date>
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<mods:number id="934B3EC1E45BFEB6627C327C183CC10F">187</mods:number>
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<mods:start id="4694813B09CDA83F5748D20AC0753950">1</mods:start>
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<mods:classification id="2886F42FA63BE65D8C96800058C108EE">journal article</mods:classification>
<mods:identifier id="1519188199AE890E8B988A8D573F0677" type="DOI">http://dx.doi.org/10.3897/phytokeys.187.75534</mods:identifier>
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<treatment id="CAE1C14979A35BA7AE6BF258BCD3A9D1" ID-GBIF-Taxon="191535292" LSID="urn:lsid:plazi:treatment:CAE1C14979A35BA7AE6BF258BCD3A9D1" httpUri="http://treatment.plazi.org/id/CAE1C14979A35BA7AE6BF258BCD3A9D1" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Caryophyllaceae">
<subSubSection id="BCBA68916F9FD36674C8574B8DBC3E50" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="EA6A57996EDAF885AF2CC1EF111006E5" pageId="0" pageNumber="1">
<taxonomicName id="E513BEB5832BA4BF41E6F6B5E9DD85C2" ID-CoL="35B5V" LSID="CAE1C149-79A3-5BA7-AE6B-F258BCD3A9D1" authority="Wulfen in Jacq., Coll. 1: 237. 1786. [January-September 1786]" authorityName="Wulfen in Jacq., Coll. 1: 237. 1786. [January-September" authorityYear="1786" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus sylvestris" order="Caryophyllales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sylvestris">Dianthus sylvestris Wulfen in Jacq., Coll. 1: 237. 1786. [January-September 1786]</taxonomicName>
</paragraph>
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<paragraph id="58A1510C8AE654036952C74B19B668A6" pageId="0" pageNumber="1">
<taxonomicName id="145756FEB37361BE2CD2A0DD756C1D03" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus caryophyllus subsp. sylvestris" order="Caryophyllales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="subSpecies" species="caryophyllus" subSpecies="sylvestris">Dianthus caryophyllus sylvestris</taxonomicName>
<taxonomicName id="59B8F078966B504168B2B2D139216A60" class="Magnoliopsida" family="Caryophyllaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="D. caryophyllus subsp. sylvestris" order="Caryophyllales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="subSpecies" species="caryophyllus" subSpecies="sylvestris">D. caryophyllus subsp. sylvestris</taxonomicName>
(Wulfen) Rouy &amp; Foucaud, Fl. France 3: 193. 1896. Ind. Loc.: - &quot;in montibus illis prope Ponewitsch Baronis Wolkensberg in Carniolia, tum in M. Utocsek prope Pillichgraz; in iis. Vallis Rablensis; denique &amp; in iis Vallis Canalensis &amp;c.&quot;.
</paragraph>
</subSubSection>
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<paragraph id="75B483D51D6DDE17E2452C8CD36266B7" pageId="0" pageNumber="1">Types.</paragraph>
<paragraph id="6B1D5AC9D940982D2C7F86C0FB02F86C" pageId="0" pageNumber="1">
(lectotype here designated): The water-coloured iconography published by
<bibRefCitation id="947DA6F7E917A155E801D58D5A5409DB" author="Jacquin, NJ" journalOrPublisher="Systematics and Biodiversity" pageId="0" pageNumber="1" refId="B22" refString="Jacquin, NJ, 1781-1786. Icones plantarum rariorum 1 Christianum Fridericum Wappler, Vindobonae." title="Icones plantarum rariorum 1 Christianum Fridericum Wappler, Vindobonae." year="1781 - 1786">Jacquin (1781-1786</bibRefCitation>
, t. 82, the small individual on the right).
</paragraph>
</subSubSection>
<subSubSection id="7404B4AEC389002C375A19B25259D5FA" pageId="0" pageNumber="1" type="notes">
<paragraph id="A4DE68B1CD5578EACEEE6C940B40D475" pageId="0" pageNumber="1">Note.</paragraph>
<paragraph id="092F8BD257801B91A78F04C00E081E69" pageId="0" pageNumber="1">
The iconography designated by
<bibRefCitation id="AF39775A3DD1769BEA1DDA9D3A8E6A11" DOI="https://doi.org/10.1111/j.1756-1051.2009.00459.x" author="Bacchetta, G" journalOrPublisher="Nordic Journal of Botany" pageId="0" pageNumber="1" pagination="137 - 173" refId="B5" refString="Bacchetta, G, Brullo, S, Casti, M, Giusso del Galdo, GP, 2010. Taxonomic revision of the Dianthus sylvestris group (Caryophyllaceae) in Central-Southern Italy, Sicily and Sardinia. Nordic Journal of Botany 28 (2): 137 - 173, DOI: https://doi.org/10.1111/j.1756-1051.2009.00459.x" title="Taxonomic revision of the Dianthus sylvestris group (Caryophyllaceae) in Central-Southern Italy, Sicily and Sardinia." url="https://doi.org/10.1111/j.1756-1051.2009.00459.x" volume="28" year="2010">Bacchetta et al. (2010)</bibRefCitation>
as neotype is actually part of the original material as uncited illustration (Art. 9.12 of the ICN), since
<normalizedToken id="D92AC6DDABE3C5650E8624BFDFB78E18" originalValue="Jacquins">Jacquin's</normalizedToken>
Icones and Collectanea work are interrelated. Therefore this neotypification must be corrected in lectotypification. This illustration depicts two individuals: one small with a 2 branched single-flowered stem and one large, unbranched but with multiflowered stems and basal leaves three times longer, exemplifying morphological variation in this species. In the protologue, it is clearly stated that the larger plant was seen only once in Monte Re, near Lake of Predil, NE Italy (&quot;Uno duntaxat, quod miratus sum, loco Montis regii Rablensis, giganteum inveni, caulibus cubitalibus bi- &amp; trifloris&quot;), while smaller plants are common elsewhere in Carniola. Accordingly, we can conclude that the two drawings depict plants originating from two different areas, thus belonging to two different gatherings. Consequently, the type designated by
<bibRefCitation id="15EC1256BCE3ECDABE83850AD0441CB4" DOI="https://doi.org/10.1111/j.1756-1051.2009.00459.x" author="Bacchetta, G" journalOrPublisher="Nordic Journal of Botany" pageId="0" pageNumber="1" pagination="137 - 173" refId="B5" refString="Bacchetta, G, Brullo, S, Casti, M, Giusso del Galdo, GP, 2010. Taxonomic revision of the Dianthus sylvestris group (Caryophyllaceae) in Central-Southern Italy, Sicily and Sardinia. Nordic Journal of Botany 28 (2): 137 - 173, DOI: https://doi.org/10.1111/j.1756-1051.2009.00459.x" title="Taxonomic revision of the Dianthus sylvestris group (Caryophyllaceae) in Central-Southern Italy, Sicily and Sardinia." url="https://doi.org/10.1111/j.1756-1051.2009.00459.x" volume="28" year="2010">Bacchetta et al. (2010</bibRefCitation>
: 143), neotype or lectotype, belongs to more than one gathering and cannot be accepted as a type (Art. 8.1, 8.2, 9.3 of the ICN). Thus, the name remains to be typified. No other original material for this name exists (
<bibRefCitation id="7F59392426FB1870D2BC62E013F9D14F" DOI="https://doi.org/10.2307/1220794" author="de Langen, FR" journalOrPublisher="Taxon" pageId="0" pageNumber="1" pagination="716 - 724" refId="B12" refString="de Langen, FR, Oost, EH, Jarvis, CE, 1984. Lectotypification of Dianthus caryophyllus L. and D. chinensis L. (Caryophyllaceae). Taxon 33 (4): 716 - 724, DOI: https://doi.org/10.2307/1220794" title="Lectotypification of Dianthus caryophyllus L. and D. chinensis L. (Caryophyllaceae)." url="https://doi.org/10.2307/1220794" volume="33" year="1984">de Langen et al. 1984</bibRefCitation>
), so that we select here as lectotype only the small specimen of the water-coloured iconography published by Jacquin at table 86 that better fits the description &quot;folia ... pollicari aut circiter longitudine
<normalizedToken id="459DF2893497A1AC9F3C9ABAE842FEEB" originalValue="…">...</normalizedToken>
Caulis subquinquepollicaris... Flos plerumque unicus [Leaves
<normalizedToken id="6423818DF212CEE51652BE64EBF0130E" originalValue="…">...</normalizedToken>
one inch or about one inch long, stem less than 5 inches ... flower generally single]&quot;.
</paragraph>
<paragraph id="8DDBB9D5F74FB806A0B70B93397039D5" pageId="0" pageNumber="1">
The lectotype here selected agrees with the current application of the name by numerous authors, e.g.,
<bibRefCitation id="7EE9094D63865EB1B4484E669BA77003" author="Kerguelen, M" journalOrPublisher="Naturalia, Turriers" pageId="0" pageNumber="1" refId="B26" refString="Kerguelen, M, 1993. Index synonymique de la flore de France. Museum national d'histoire naturelle, Paris." title="Index synonymique de la flore de France. Museum national d'histoire naturelle, Paris." year="1993">
<normalizedToken id="33CC8D74FBB8F65A3A5D712854572494" originalValue="Kerguélen">Kerguelen</normalizedToken>
(1993)
</bibRefCitation>
,
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,
<bibRefCitation id="671667A8D7B68E4C3E1BDB266D468704" author="Tison, J-M" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="1" refId="B37" refString="Tison, J-M, de Foucault, B, 2014. Flora Gallica. Flore de France. Biotope Editons, Meze." title="Flora Gallica. Flore de France. Biotope Editons, Meze." year="2014">Tison and de Foucault (2014)</bibRefCitation>
,
<bibRefCitation id="FE8FE8CEAED9D7F4EEBB934CE5732CB1" author="Brullo, S" editor="Pignatti, S" journalOrPublisher="2). New Business Media, Milano" pageId="0" pageNumber="1" pagination="200 - 205" refId="B8" refString="Brullo, S, Guarino, R, 2017. Complesso di D. sylvestris. In: Pignatti, S, Ed., Flora d'Italia (Vol. 2). New Business Media, Milano: 200 - 205" title="Complesso di D. sylvestris." volumeTitle="Flora d'Italia (Vol." year="2017">Brullo and Guarino (2017)</bibRefCitation>
, who consider
<taxonomicName id="76B312A5923A00238E37CEDD986F7A64" class="Magnoliopsida" family="Caryophyllaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="D. sylvestris" order="Caryophyllales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis id="DA5557782B62A4B95B617BC0C23882B2" italics="true" pageId="0" pageNumber="1">D. sylvestris</emphasis>
</taxonomicName>
as an accepted species. The overall size of the plant, and the length of the leaves are not stable characters for taxonomic discrimination. The shape and relative size of calyx and epicalyx scales are better discriminating taxonomic characters and are evident in the lectotype. These features allow to distinguish
<taxonomicName id="35CB361A9E2A0FEE2C9524B48DE84F3B" authority="subsp. sylvestris" class="Magnoliopsida" family="Caryophyllaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="D. sylvestris subsp. sylvestris" order="Caryophyllales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="subSpecies" species="sylvestris" subSpecies="sylvestris">D. sylvestris subsp. sylvestris</taxonomicName>
from
<taxonomicName id="B3CB838C3973DF13286150D3F6DD5F56" class="Magnoliopsida" family="Caryophyllaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="D. sylvestris subsp. tergestinus" order="Caryophyllales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="subSpecies" species="sylvestris" subSpecies="tergestinus">D. sylvestris subsp. tergestinus</taxonomicName>
(
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).
</paragraph>
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