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<document id="9EE1D8BCC0506652CF73D9B8CF2B5B35" ID-CLB-Dataset="35979" ID-DOI="http://dx.doi.org/10.3897/zookeys.427.7618" ID-GBIF-Dataset="7ff4e90a-d601-4933-9e0b-fc7377e63dbc" ID-PMC="PMC4137312" ID-Pensoft-Pub="1313-2970-427-21" ID-PubMed="25147463" ID-ZooBank="FE1E99BF29D84751BB92F460E06BD84A" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-427-21" ModsDocOrigin="ZooKeys 427" ModsDocTitle="Revision of the East Mediterranean Orthomus (Coleoptera, Carabidae, Pterostichini), with description of Parorthomus gen. n. socotranus sp. n. from Socotra Island and key to the Old World genera of subtribe Euchroina" checkinTime="1451245594833" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav, Wrase, David W. &amp; Farkac, Jan" docDate="2014" docId="CA9D04061FD0D30F83923132B2B5C89D" docLanguage="en" docName="ZooKeys 427: 21-57" docOrigin="ZooKeys 427" docSource="http://dx.doi.org/10.3897/zookeys.427.7618" docTitle="Orthomus longulus Reiche &amp; Saulcy 1855" docType="treatment" docVersion="6" lastPageNumber="35" masterDocId="FFFCFFA6D31DFFBF3719FA19D2613A34" masterDocTitle="Revision of the East Mediterranean Orthomus (Coleoptera, Carabidae, Pterostichini), with description of Parorthomus gen. n. socotranus sp. n. from Socotra Island and key to the Old World genera of subtribe Euchroina" masterLastPageNumber="57" masterPageNumber="21" pageNumber="34" updateTime="1701385333588" updateUser="ExternalLinkService">
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<mods:title id="0CEA89D4C63840AC8CD3F7BFDE783E4D">Revision of the East Mediterranean Orthomus (Coleoptera, Carabidae, Pterostichini), with description of Parorthomus gen. n. socotranus sp. n. from Socotra Island and key to the Old World genera of subtribe Euchroina</mods:title>
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<mods:namePart id="5284EF332D5B8C9AAC89ACDE425150A2">Gueorguiev, Borislav</mods:namePart>
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<mods:namePart id="8B08A78943C7C68620DF581673240ED2">Wrase, David W.</mods:namePart>
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<mods:namePart id="8D294D444ED4BF9FF0FA633CB8E42CF8">Farkac, Jan</mods:namePart>
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<mods:date id="C7A343093A8CB6F4DE8DA67BB6CFC82F">2014</mods:date>
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<mods:identifier id="0F240EB3D3456A374F406DFDB86AAB06" type="DOI">http://dx.doi.org/10.3897/zookeys.427.7618</mods:identifier>
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<treatment id="CA9D04061FD0D30F83923132B2B5C89D" ID-GBIF-Taxon="152054444" LSID="urn:lsid:plazi:treatment:CA9D04061FD0D30F83923132B2B5C89D" httpUri="http://treatment.plazi.org/id/CA9D04061FD0D30F83923132B2B5C89D" lastPageId="14" lastPageNumber="35" pageId="13" pageNumber="34">
<subSubSection id="2BBA57136528AEC8373233944DBF7F90" pageId="13" pageNumber="34" type="multiple">
<paragraph id="EFC50A6319A5288128DEC4BDB3457400" pageId="13" pageNumber="34">Taxon classification Animalia Coleoptera Carabidae</paragraph>
</subSubSection>
<subSubSection id="9A4E21EFAECC9B1CF2C8BF678FF19A3D" pageId="13" pageNumber="34" type="nomenclature">
<paragraph id="420300663FAA789291D506B273B72F11" pageId="13" pageNumber="34">
<taxonomicName id="BEAFE89BED153D3013E655F8AF25D3A2" ID-CoL="758QC" authority="Reiche &amp; Saulcy, 1855" authorityName="Reiche &amp; Saulcy" authorityYear="1855" class="Insecta" family="Carabidae" genus="Orthomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomus longulus" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longulus">Orthomus longulus (Reiche &amp; Saulcy, 1855)</taxonomicName>
Figs 3, 6, 10, 14
</paragraph>
</subSubSection>
<subSubSection id="163500F5CFD977400D4766E9117B7A49" pageId="13" pageNumber="34" type="reference_group">
<paragraph id="2D1C761A24FB849F0EF2E8BC6E076BA0" pageId="13" pageNumber="34">
<taxonomicName id="0F76BD5158D7349682EE773600868545" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia (Argutor) longula" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longula" subGenus="Argutor">Feronia (Argutor) longula</taxonomicName>
Reiche &amp; Saulcy, 1855: 616 (type locality: &quot;De Beyrouth&quot;), part
</paragraph>
<paragraph id="DBBBEA7BA9EFCCC24E79985BC909E58F" pageId="13" pageNumber="34">
<taxonomicName id="DE3C32EF59E4573AC2C457ED91AC2AEC" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia (Argutor) berytensis" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="berytensis" subGenus="Argutor">Feronia (Argutor) berytensis</taxonomicName>
Reiche &amp; Saulcy, 1855: 618, part
</paragraph>
<paragraph id="C9A4F39F4F697BB0AAAD06553F5E8864" pageId="13" pageNumber="34">
<taxonomicName id="4727B805572A129672A3BEDC2C2742B3" class="Insecta" family="Carabidae" genus="Orthomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomus longior" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longior">Orthomus longior</taxonomicName>
Chaudoir, 1873: 105, part
</paragraph>
<paragraph id="33661EE9A8A01F655FCA3239CFD4E2F2" pageId="13" pageNumber="34">
<taxonomicName id="1D08B9F993E51EF50B4A72793C26ABD9" class="Insecta" family="Carabidae" genus="Orthomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomus longulus" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longulus">Orthomus longulus</taxonomicName>
s.str.:
<bibRefCitation id="DA0345367D6FD6DE96D7BD2F31641E91" author="Mateu, J" journalOrPublisher="Eos, Revista Espanola de Entomologia" pageId="28" pageNumber="49" pagination="53 - 85" title="Notas sobre los Orthomus Chaudoir (segunda nota)." volume="31" year="1955">Mateu 1955</bibRefCitation>
: 56, 60
</paragraph>
</subSubSection>
<subSubSection id="48A3D3358CCB13004F6B8102D0749C1D" pageId="13" pageNumber="34" type="type material">
<paragraph id="9138CB35B342FE0DB3341B8E2F577057" pageId="13" pageNumber="34">Type material.</paragraph>
<paragraph id="7D82D606EF7902350FB50C935525D37C" pageId="13" pageNumber="34">
<taxonomicName id="BACB16C5F2FF5ED92D1F9B7E6275AF55" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia longula" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longula">Feronia longula</taxonomicName>
Reiche &amp; Saulcy, 1855 (specimens belonging to
<taxonomicName id="1BB84C718ED7E061BBDA2C41EB3D34E9" class="Insecta" family="Carabidae" genus="Orthomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomus longulus" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longulus">Orthomus longulus</taxonomicName>
). The type series of
<taxonomicName id="2ECFB94B8BED67167943C088C7288FCE" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia longula" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longula">Feronia longula</taxonomicName>
consists of 11 syntypes, 4 ♂♂, 6 ♀♀ of them in MHNG and 1 ♂ in MNHP. The specimens in MHNG are placed under a
<normalizedToken id="717737347F81BCB817ABF849C6637EF4" originalValue="Melly’s">Melly's</normalizedToken>
taxa label &quot;longulus Reiche., Egypte, Syrie&quot; [handwritten by Melly with pen] pinned to the bottom of the drawer, which thus are to be considered as pertaining collectively to all specimens. Our revision revealed that the specimens in MHNG are not conspecific but belong to three distinct species. Three males and one female belong to
<taxonomicName id="BEAEED12D4882A1D40A06234DD5B4406" class="Insecta" family="Carabidae" genus="Orthomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomus longior" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longior">Orthomus longior</taxonomicName>
(for these specimens see under
<taxonomicName id="10F6CF0E13A5E37D7D417438B0AF0374" class="Insecta" family="Carabidae" genus="Orthomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomus longior" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longior">Orthomus longior</taxonomicName>
), while another male and another female belong to
<taxonomicName id="F8EC84F22F515242B2E326C1B16307E1" class="Insecta" family="Carabidae" genus="Orthomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomus berytensis" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="berytensis">Orthomus berytensis</taxonomicName>
(for these specimens see under
<taxonomicName id="12D1139142F3BFAA11378BB4993F3E9E" class="Insecta" family="Carabidae" genus="Orthomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomus berytensis" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="berytensis">Orthomus berytensis</taxonomicName>
). Only four females are representatives of
<taxonomicName id="BCE5D663FE85746C47A608855729D2AD" class="Insecta" family="Carabidae" genus="Orthomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomus longulus" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longulus">Orthomus longulus</taxonomicName>
, labelled individually as follow. One female selected for lectotype:
<normalizedToken id="D3FAF30E967BEF14AAB5FBBFA05072D0" originalValue="“…”">&quot;...&quot;</normalizedToken>
[handwritten remnant on small quadratic yellow label],
<normalizedToken id="3C800FAF9B190F82254B4236EE7A3524" originalValue="“Beyrouth”">&quot;Beyrouth&quot;</normalizedToken>
[Reiche's handwriting on brown label], &quot;Coll. Reiche&quot; [black print on white label], &quot;
<taxonomicName id="A3242AEA8FC94B130E4B22564F34E287" class="Insecta" family="Carabidae" genus="Orthomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomus longulus" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longulus">Orthomus longulus</taxonomicName>
Reiche Label MHNG 2010&quot; [black print on white label by Cuccodoro], &quot;Lectotype
<taxonomicName id="F2BCECDB7A6DCC6A1919E17CF41B6E78" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia longula" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longula">Feronia longula</taxonomicName>
Reiche &amp; Saulcy, 1855 B.
<normalizedToken id="69261F8AB54EBE445556FD848783DF5D" originalValue="Guéorguiev">Gueorguiev</normalizedToken>
&amp; D.W.Wrase des. 2012&quot; [black print on red label]; 3 ♀♀, &quot;Coll. Reiche&quot; [black print on white label], &quot;
<taxonomicName id="8A2AA024042283026A394945FF38CC84" class="Insecta" family="Carabidae" genus="Orthomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomus longulus" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longulus">Orthomus longulus</taxonomicName>
Reiche 'Egypte, Syrie.' Label MHNG 2010&quot; [black print on white label by Cuccodoro], &quot;Paralectotype
<taxonomicName id="BB1665C919B072C038A2A11FCBE82EEB" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia longula" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longula">Feronia longula</taxonomicName>
Reiche &amp; Saulcy, 1855 B.
<normalizedToken id="4FEC073826404E328B488C348D351075" originalValue="Guéorguiev">Gueorguiev</normalizedToken>
&amp; D.W.Wrase des. 2012&quot; [black print on red label]. The specimen from MNHP is a male, with previously extracted genitalia and glued to a separate card pinned beneath the specimen. The most part of its aedeagus is destroyed, certainly by a species of the genus
<taxonomicName id="72E0188DD1F0E191DFD98C109B6B0A3E" class="Insecta" family="Dermestidae" genus="Anthrenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthrenus" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="genus">Anthrenus</taxonomicName>
Geoffroy, 1762, but the apical lamella is still preserved. This male is conspecific with the above last four females from MHNG and it is designated as paralectotype, too: &quot;longulus type Reiche&quot; [handwritten on white label by Mateu], &quot;Paralectotype
<taxonomicName id="DEDD2E76F3C5DE1D8DED2B7CC62E6801" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia longula" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longula">Feronia longula</taxonomicName>
Reiche &amp; Saulcy, 1855 B.
<normalizedToken id="665677147AE95E34087A2ED0A0175385" originalValue="Guéorguiev">Gueorguiev</normalizedToken>
&amp; D.W.Wrase des. 2012&quot; [black print on red label]. All five specimens, pertaining to the true longulus, with a label, subsequently added: &quot;
<taxonomicName id="E14F3C1C240431A2A4A7CA4A1F9272EF" class="Insecta" family="Carabidae" genus="Orthomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomus longulus" order="Coleoptera" pageId="13" pageNumber="34" phylum="Arthropoda" rank="species" species="longulus">Orthomus longulus</taxonomicName>
(Reiche &amp; Saulcy) det. B.
<normalizedToken id="0F543C1FF44DE994422A4D1F95BF71BB" originalValue="Guéorguiev”">Gueorguiev&quot;</normalizedToken>
[black print on white label].
</paragraph>
</subSubSection>
<subSubSection id="EC0C36DAA053A9D402EDCD501121C9C2" pageId="14" pageNumber="35" type="other material studied">
<paragraph id="15D8D2C3ABF9E7DAAB55E620323C659B" pageId="14" pageNumber="35">
<pageBreakToken id="7F79997813A2215A7CF0C51F613720FA" pageId="14" pageNumber="35" start="start">Other</pageBreakToken>
material studied.
</paragraph>
<paragraph id="5D7B96CB6D1D809CF34BC7F7A2F02C47" pageId="14" pageNumber="35">
ISRAEL: - Northern District: 1 ♂, &quot;Palestine.
<normalizedToken id="D1545FE7E201090E5768726C9B32CF19" originalValue="…">...</normalizedToken>
Galilee XII.1924 O. Theodor.&quot; (BMNH); 1 spec., &quot;Nazaret, 17.3.-3.4.87, Kfar
<normalizedToken id="8BD4EF5A9AB3FF899B2E78D6420B520F" originalValue="….">....</normalizedToken>
L. Blumenthal leg.&quot; (NMW); 3 ♂♂, 5 ♀♀, Megiddo, Ein
<normalizedToken id="A633C43705CB3336BFAFD3E222BDDD58" originalValue="Ha’emek">Ha'emek</normalizedToken>
, 230 m, Getreideacker, 08.V.1996, Schnitter &amp; Staven leg. (DWBG); 1 ♂, 4 ♀♀, &quot;Merom Golan, 12.VI.2000, V. Chikatunov leg.&quot; (TAU); 2 ♂♂, 3 ♀♀, Upper Galilee, N. sea shore of Sea of Galilee, Ein Sheva (Tabkha), -192 m,
<geoCoordinate id="9D69461DA66EEEDD4A0072D94F1C5005" direction="north" orientation="latitude" precision="1" value="32.874218">32°52.453'N</geoCoordinate>
,
<geoCoordinate id="03C261CF4CA202A05F120DFB9BF8CF56" direction="east" orientation="longitude" precision="1" value="35.545433">035°32.726'E</geoCoordinate>
(stony and loamy pasture), 25.IV.2006, D.W.Wrase leg. (DWBG); 1 ♂, Lower Galilee, ca 4 km W Tamra, (route 70),
<geoCoordinate id="77F1ADA12784ED41FC963B194FE53873" direction="north" orientation="latitude" precision="1" value="32.863316">32°51.799'N</geoCoordinate>
,
<geoCoordinate id="3562E240881633E778DCA75B56355F48" direction="east" orientation="longitude" precision="1" value="35.17153">035°10.292'E</geoCoordinate>
(loamy field edge), 25 m, 25.IV.2006 D.W.Wrase leg. (DWBG); 1 ♂, 6 ♀♀, Upper Galilee, Ha Khula Valley,
<normalizedToken id="3B464E2F88D004297269EB8FFAD8A546" originalValue="Ma’agar">Ma'agar</normalizedToken>
Einan lake, 73 m,
<geoCoordinate id="C9D99D9F0DB43E5B550DEC94C2CCACFA" direction="north" orientation="latitude" precision="1" value="33.085617">33°05.137'N</geoCoordinate>
,
<geoCoordinate id="2289B5144A61BFDC97039FA294C87FA8" direction="east" orientation="longitude" precision="1" value="35.578835">035°34.730'E</geoCoordinate>
(toe of dam, in moist loamy soil), 1./2.V.2006 D.W.Wrase leg. (DWBG, JSAG); 1 ♂, Bir el Maksur,
<geoCoordinate id="4FDF7F41550A41087C4EFD26B19E39BB" direction="north" orientation="latitude" precision="1" value="32.76502">32°45.901'N</geoCoordinate>
,
<geoCoordinate id="1507B2D5AEB937DB3B6C71901DA50DD1" direction="east" orientation="longitude" precision="1" value="35.231384">035°13.883'E</geoCoordinate>
, 23.II.2005, W. Starke leg. (DWBG); 1 ♀, Nazareth, Kfar?Hochbreeh, 17.III.-3.IV.1987, Blumenthal leg. (DWBG). - Haifa District: 2 ♂♂, 2 ♀♀, Haifa [&quot;Syrien Haifa Reitter&quot;] (BMNH, MHNG, MIZ, NMW); 1 ♀, Mount Carmel, 23.XII.25 (MIZ); 4 ♀♀, Haifa, 15.XII.1941 / 8.I.1942 / 4.XII.1954 Bytinski-Salz (TAU); 1 ♀, Nahal Oren, Mount Carmel, 18.3.1996,
<normalizedToken id="7582CA5EBEDD52BEE3CAA26AD59672E8" originalValue="Pavliček">Pavlicek</normalizedToken>
&amp; Chikatunov leg. (TAU); 1 ♀, Haifa, Check Post, 8.II.2000, V. Chikatunov &amp; T.
<normalizedToken id="71B9DA2E27841D62B44A78AE5269C138" originalValue="Pavliček">Pavlicek</normalizedToken>
leg. (TAU); 1 ♂, Mount Carmel,
<normalizedToken id="18E187A372967C58DE3F951F1CB071D6" originalValue="Ya’ar">Ya'ar</normalizedToken>
ha-
<normalizedToken id="9291EE239F9BB4E8AF6320FBA9DF2B12" originalValue="Ya’aramin">Ya'aramin</normalizedToken>
ca. 500 m (under stones), 30.III.2008, D.W.Wrase leg. (DWBG).
</paragraph>
</subSubSection>
<subSubSection id="581476CECEDFD03391D1D799BFA97255" pageId="14" pageNumber="35" type="wrong locality">
<paragraph id="E8807A9643712B80FE511A31305F5183" pageId="14" pageNumber="35">Wrong locality.</paragraph>
<paragraph id="3CA9A9149EE2D5AC9BE4D2D4BA3B505C" pageId="14" pageNumber="35">
1 ♂, &quot;42 St.&quot; / &quot;
<taxonomicName id="4D3A842F826DB62BFC0CBA38FF7051A1" class="Insecta" family="Carabidae" genus="Orthomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomus berytensis" order="Coleoptera" pageId="14" pageNumber="35" phylum="Arthropoda" rank="species" species="berytensis">Orthomus berytensis</taxonomicName>
Reich. Portugal Dr Stierlin (above), 2. b. (underneath)&quot; (MHNG).
</paragraph>
</subSubSection>
<subSubSection id="FB2BA910B8D541C6F9DA03FEC04CCAEB" pageId="14" pageNumber="35" type="male genitalia">
<paragraph id="CB95D20B40A73C8764F5C36C4E530C00" pageId="14" pageNumber="35">Male genitalia</paragraph>
<paragraph id="6F286D65A96B09B5BAD42B753CCC42B7" pageId="14" pageNumber="35">(9 specimens examined).</paragraph>
</subSubSection>
<subSubSection id="6A67A5E468D536E6E32B271893C7887A" pageId="14" pageNumber="35" type="distribution">
<paragraph id="738EFA3A0CDB2F25248F07160A918651" pageId="14" pageNumber="35">Distribution.</paragraph>
<paragraph id="5B2DC0416FCA432CC64C41F30EA47C87" pageId="14" pageNumber="35">North Israel (Northern District; Haifa District), Lebanon (Beyrouth, type material).</paragraph>
</subSubSection>
</treatment>
</document>
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