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<document id="97DF21187055CE15B9D7336E9C4E74A7" ID-CLB-Dataset="27502" ID-DOI="10.11646/zootaxa.4550.1.1" ID-GBIF-Dataset="291701d4-652e-440e-adab-f6b0201c0972" ID-ISSN="1175-5326" ID-Zenodo-Dep="2625030" ID-ZooBank="2F7D1EC4-D4ED-4FAE-B227-CF7B79EAE833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554274879827" checkinUser="plazi" docAuthor="Huber, Bernhard A." docDate="2019" docId="CA3B104CFFB8FFB4FF3DFCDFFC8FE356" docLanguage="en" docName="zootaxa.4550.1.1.pdf" docOrigin="Zootaxa 4550 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wanniyala mapalena Huber 2019, sp. n." docType="treatment" docVersion="14" lastPageNumber="48" masterDocId="36026834FF95FF9BFFAAFFA1FFB6E16A" masterDocTitle="The pholcid spiders of Sri Lanka (Araneae: Pholcidae)" masterLastPageNumber="57" masterPageNumber="1" pageNumber="46" updateTime="1698716853299" updateUser="ExternalLinkService">
<mods:mods id="B68B5FB33A39B3132D07AC17A77CA32E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4B84AB36E36B691400110A92E6F827CC">
<mods:title id="80CB69460F927689667B47B9FC6A66CB">The pholcid spiders of Sri Lanka (Araneae: Pholcidae)</mods:title>
</mods:titleInfo>
<mods:name id="37BE68658D768F4D431E8373E041D6EA" type="personal">
<mods:role id="E7E8FD44E51290B2C68489394092103A">
<mods:roleTerm id="CDA59059C00A26B93839853061CDEEFF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="84BD55C95DB62C2E5FEA1FB74C1C0562">Huber, Bernhard A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C66683A7713A9B92125433D1D1875F03">text</mods:typeOfResource>
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<mods:title id="E24AA9069E90B1A54F20F56AC71E2B54">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4B64DC816402439A5C6539200B28F241">
<mods:date id="6E6FDDB2F48FA2F0FEF20FDC80940D29">2019</mods:date>
<mods:detail id="522CB616E0E665D1EEF3C5A68083E71A" type="pubDate">
<mods:number id="E4FF52F4A88CCF70BCC4E9E53D8AE8E0">2019-01-23</mods:number>
</mods:detail>
<mods:detail id="F6EF215E779C2B8DC3C8B348E9ABE83A" type="volume">
<mods:number id="DC0F630A9D28278FDBB32E0F223A1FCF">4550</mods:number>
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<mods:detail id="17294E9C352E21A3FCC58F6E924A7D86" type="issue">
<mods:number id="BFDA00412656428F52C6DF12758574F6">1</mods:number>
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<mods:start id="3BFFF39CC243EAF3B6F538686CABC8B4">1</mods:start>
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<mods:classification id="B7B665712058E174036512FC92576F46">journal article</mods:classification>
<mods:identifier id="0ECA13DB37004F38ECC15A62E8156F1A" type="CLB-Dataset">27502</mods:identifier>
<mods:identifier id="89FDF0D523457F7224D51F0745FD5AFF" type="DOI">10.11646/zootaxa.4550.1.1</mods:identifier>
<mods:identifier id="954D1DC0A06E52842B4FB158D1ACDAFE" type="GBIF-Dataset">291701d4-652e-440e-adab-f6b0201c0972</mods:identifier>
<mods:identifier id="CAB0C300CB2F21F27D3F840A108B9353" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6DCFCA9056198B3B1387894D27B26B84" type="Zenodo-Dep">2625030</mods:identifier>
<mods:identifier id="E5A0878934E4861CD4B1B7EDCB72B195" type="ZooBank">2F7D1EC4-D4ED-4FAE-B227-CF7B79EAE833</mods:identifier>
</mods:mods>
<treatment id="CA3B104CFFB8FFB4FF3DFCDFFC8FE356" ID-DOI="http://doi.org/10.5281/zenodo.4581693" ID-GBIF-Taxon="154534095" ID-Zenodo-Dep="4581693" LSID="urn:lsid:plazi:treatment:CA3B104CFFB8FFB4FF3DFCDFFC8FE356" httpUri="http://treatment.plazi.org/id/CA3B104CFFB8FFB4FF3DFCDFFC8FE356" lastPageId="47" lastPageNumber="48" pageId="45" pageNumber="46">
<subSubSection id="0A88F2D1FFB8FFB6FF3DFCDFFE56E2F2" box="[151,480,894,920]" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="422DA15AFFB8FFB6FF3DFCDFFE56E2F2" blockId="45.[151,508,894,954]" box="[151,480,894,920]" pageId="45" pageNumber="46">
<heading id="19651636FFB8FFB6FF3DFCDFFE56E2F2" bold="true" box="[151,480,894,920]" fontSize="11" level="1" pageId="45" pageNumber="46" reason="1">
<emphasis id="70E67D48FFB8FFB6FF3DFCDFFE56E2F2" bold="true" box="[151,480,894,920]" pageId="45" pageNumber="46">
<taxonomicName id="8592DAD9FFB8FFB6FF3DFCDFFE21E2F2" ID-CoL="7G5T8" authority="Huber, 2019" authorityName="Huber" authorityYear="2019" box="[151,407,894,920]" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="mapalena" status="sp. nov.">
<emphasis id="70E67D48FFB8FFB6FF3DFCDFFE21E2F2" bold="true" box="[151,407,894,920]" italics="true" pageId="45" pageNumber="46">Wanniyala mapalena</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6BD5C033FFB8FFB6FE34FCDFFE56E2F2" box="[414,480,894,920]" pageId="45" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FFB8FFB6FF3DFC00FE4AE2D0" box="[151,508,929,954]" pageId="45" pageNumber="46" type="description">
<paragraph id="422DA15AFFB8FFB6FF3DFC00FE4AE2D0" blockId="45.[151,508,894,954]" box="[151,508,929,954]" pageId="45" pageNumber="46">
<figureCitation id="DAA9BDDFFFB8FFB6FF3DFC00FEE0E2D0" box="[151,342,929,954]" captionStart="FIGURES 138–149" captionStartId="34.[151,264,1387,1410]" captionTargetBox="[151,1436,195,1366]" captionTargetId="figure@34.[151,1436,195,1366]" captionTargetPageId="34" captionText="FIGURES 138–149. Wanniyala, live specimens, part 2. 138–140. W. ohiya sp. n., males and female with egg-sac from Ohiya (Fig. 139 shows a male in typical resting position). 141–143. W. viharekele sp. n., males from Viharekele and Kottawa, female from Kottawa. 144–146. W. mapalena sp. n., male and females from between Ratnapura and Mapalena Ella. 147–149. W. labugama sp. n., males and female with egg-sac from Labugama." httpUri="https://zenodo.org/record/2625076/files/figure.png" pageId="45" pageNumber="46">Figures 144–146</figureCitation>
,
<figureCitation id="DAA9BDDFFFB8FFB6FEC8FC00FE71E2D0" box="[354,455,929,954]" captionStart="FIGURES 197–205" captionStartId="46.[151,264,1840,1863]" captionTargetBox="[152,1436,228,1785]" captionTargetId="figure@46.[151,1436,228,1785]" captionTargetPageId="46" captionText="FIGURES 197–205. Wanniyala mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 197–198. Male chelicerae, clypeus and ocular area, frontal and lateral views. 199. Left male palpal trochanter and femur, retrolateral view. 200. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 201–202. Left procursus, prolateral and retrolateral views (arrow points at distinctive process on retrolateral sclerite of procursus). 203–204. Epigynum, ventral and lateral views. 205. Cleared female genitalia, dorsal view (arrow points at V-shaped element). Scale lines: 0.3 mm (197–202, 205), 0.5 mm (203–204)." httpUri="https://zenodo.org/record/2625092/files/figure.png" pageId="45" pageNumber="46">197–205</figureCitation>
,
<figureCitation id="DAA9BDDFFFB8FFB6FE78FC03FE4AE2D0" box="[466,508,930,954]" captionStart="FIGURES 213–219" captionStartId="49.[151,264,1720,1743]" captionTargetBox="[151,1436,412,1657]" captionTargetId="figure@49.[151,1436,412,1657]" captionTargetPageId="49" captionText="FIGURES 213–219. Wanniyala spp., cleared female genitalia in dorsal views. 213. W. mudita sp. n., from Kandy (ZFMK Ar 20077). 214. W. orientalis sp. n., from Dimbulagala (ZFMK Ar 20079). 215. W. upekkha sp. n., from Dematagala (ZFMK Ar 20086). 216. W. ohiya sp. n., from Ohiya (ZFMK Ar 20089). 217. W. viharekele sp. n., from Viharekele (ZFMK Ar 20092). 218. W. mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 219. W. labugama sp. n., from Labugama (ZFMK Ar 20097). At various scales." httpUri="https://zenodo.org/record/2625096/files/figure.png" pageId="45" pageNumber="46">218</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FFB8FFB6FF3DFC48FDBFE5DC" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="422DA15AFFB8FFB6FF3DFC48FDBFE5DC" blockId="45.[151,1437,1001,1998]" pageId="45" pageNumber="46">
<emphasis id="70E67D48FFB8FFB6FF3DFC48FEBCE568" bold="true" box="[151,266,1001,1026]" pageId="45" pageNumber="46">Diagnosis</emphasis>
. Males are distinguished from congeners by details of palp (retrolateral sclerite of procursus with distinctive process between dorsal process and main branch; distinctive processes of palpal trochanter, and small curved bulbal apophysis;
<figureCitation id="DAA9BDDFFFB8FFB6FE10FB90FDE1E520" box="[442,599,1073,1098]" captionStart="FIGURES 197–205" captionStartId="46.[151,264,1840,1863]" captionTargetBox="[152,1436,228,1785]" captionTargetId="figure@46.[151,1436,228,1785]" captionTargetPageId="46" captionText="FIGURES 197–205. Wanniyala mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 197–198. Male chelicerae, clypeus and ocular area, frontal and lateral views. 199. Left male palpal trochanter and femur, retrolateral view. 200. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 201–202. Left procursus, prolateral and retrolateral views (arrow points at distinctive process on retrolateral sclerite of procursus). 203–204. Epigynum, ventral and lateral views. 205. Cleared female genitalia, dorsal view (arrow points at V-shaped element). Scale lines: 0.3 mm (197–202, 205), 0.5 mm (203–204)." httpUri="https://zenodo.org/record/2625092/files/figure.png" pageId="45" pageNumber="46">Figs 199–202</figureCitation>
) and by modification of clypeus (pair of large processes at rim and small median pointed process;
<figureCitation id="DAA9BDDFFFB8FFB6FE04FBF4FDFDE504" box="[430,587,1109,1134]" captionStart="FIGURES 197–205" captionStartId="46.[151,264,1840,1863]" captionTargetBox="[152,1436,228,1785]" captionTargetId="figure@46.[151,1436,228,1785]" captionTargetPageId="46" captionText="FIGURES 197–205. Wanniyala mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 197–198. Male chelicerae, clypeus and ocular area, frontal and lateral views. 199. Left male palpal trochanter and femur, retrolateral view. 200. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 201–202. Left procursus, prolateral and retrolateral views (arrow points at distinctive process on retrolateral sclerite of procursus). 203–204. Epigynum, ventral and lateral views. 205. Cleared female genitalia, dorsal view (arrow points at V-shaped element). Scale lines: 0.3 mm (197–202, 205), 0.5 mm (203–204)." httpUri="https://zenodo.org/record/2625092/files/figure.png" pageId="45" pageNumber="46">Figs 197–198</figureCitation>
). Females are possibly indistinguishable externally from
<taxonomicName id="8592DAD9FFB8FFB6FB60FBF7FAEBE504" authorityName="Huber" authorityYear="2019" box="[1226,1373,1109,1134]" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="labugama">
<emphasis id="70E67D48FFB8FFB6FB60FBF7FAEBE504" box="[1226,1373,1109,1134]" italics="true" pageId="45" pageNumber="46">W. labugama</emphasis>
</taxonomicName>
; they differ from
<taxonomicName id="8592DAD9FFB8FFB6FEB6FBDBFE07E5F8" authorityName="Huber" authorityYear="2019" box="[284,433,1145,1170]" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="labugama">
<emphasis id="70E67D48FFB8FFB6FEB6FBDBFE07E5F8" box="[284,433,1145,1170]" italics="true" pageId="45" pageNumber="46">W. labugama</emphasis>
</taxonomicName>
and other congeners by details of internal genitalia (distinctive pair of membranous, Vshaped elements (
<figureCitation id="DAA9BDDFFFB8FFB6FEF4FB3CFE72E5DC" box="[350,452,1181,1206]" captionStart="FIGURES 197–205" captionStartId="46.[151,264,1840,1863]" captionTargetBox="[152,1436,228,1785]" captionTargetId="figure@46.[151,1436,228,1785]" captionTargetPageId="46" captionText="FIGURES 197–205. Wanniyala mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 197–198. Male chelicerae, clypeus and ocular area, frontal and lateral views. 199. Left male palpal trochanter and femur, retrolateral view. 200. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 201–202. Left procursus, prolateral and retrolateral views (arrow points at distinctive process on retrolateral sclerite of procursus). 203–204. Epigynum, ventral and lateral views. 205. Cleared female genitalia, dorsal view (arrow points at V-shaped element). Scale lines: 0.3 mm (197–202, 205), 0.5 mm (203–204)." httpUri="https://zenodo.org/record/2625092/files/figure.png" pageId="45" pageNumber="46">Figs 205</figureCitation>
,
<figureCitation id="DAA9BDDFFFB8FFB6FE65FB3FFE4BE5DC" box="[463,509,1182,1206]" captionStart="FIGURES 213–219" captionStartId="49.[151,264,1720,1743]" captionTargetBox="[151,1436,412,1657]" captionTargetId="figure@49.[151,1436,412,1657]" captionTargetPageId="49" captionText="FIGURES 213–219. Wanniyala spp., cleared female genitalia in dorsal views. 213. W. mudita sp. n., from Kandy (ZFMK Ar 20077). 214. W. orientalis sp. n., from Dimbulagala (ZFMK Ar 20079). 215. W. upekkha sp. n., from Dematagala (ZFMK Ar 20086). 216. W. ohiya sp. n., from Ohiya (ZFMK Ar 20089). 217. W. viharekele sp. n., from Viharekele (ZFMK Ar 20092). 218. W. mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 219. W. labugama sp. n., from Labugama (ZFMK Ar 20097). At various scales." httpUri="https://zenodo.org/record/2625096/files/figure.png" pageId="45" pageNumber="46">218</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FFB8FFB6FF6DFB60FB37E5B0" box="[199,1153,1217,1242]" pageId="45" pageNumber="46" type="etymology">
<paragraph id="422DA15AFFB8FFB6FF6DFB60FB37E5B0" blockId="45.[151,1437,1001,1998]" box="[199,1153,1217,1242]" pageId="45" pageNumber="46">
<emphasis id="70E67D48FFB8FFB6FF6DFB60FEF0E5B0" bold="true" box="[199,326,1217,1242]" pageId="45" pageNumber="46">Etymology</emphasis>
. The specific name is derived from the
<typeStatus id="9D291FF8FFB8FFB6FCAEFB63FC82E5B0" box="[772,820,1218,1242]" pageId="45" pageNumber="46">type</typeStatus>
locality (noun in apposition).
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FFB8FFB6FF6DFB44FDE3E400" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="422DA15AFFB8FFB6FF6DFB44FBA5E448" blockId="45.[151,1437,1001,1998]" pageId="45" pageNumber="46">
<emphasis id="70E67D48FFB8FFB6FF6DFB44FED9E594" bold="true" box="[199,367,1253,1278]" pageId="45" pageNumber="46">Type material</emphasis>
.
<materialsCitation id="F2FAAB07FFB8FFB6FED5FB47FBB9E448" ID-GBIF-Occurrence="2234226249" collectingDate="2017-03-19" collectionCode="ZFMK" collectorName="B. A. Huber" country="Sri Lanka" elevation="100" latitude="6.762" location="Ar" longLatPrecision="78" longitude="80.428" pageId="45" pageNumber="46" specimenCount="1" specimenCount-male="1" stateProvince="Sabaragamuwa" typeStatus="holotype">
<collectingCountry id="3A85E1CAFFB8FFB6FED5FB47FDADE594" box="[383,539,1254,1278]" name="Sri Lanka" pageId="45" pageNumber="46">SRI LANKA</collectingCountry>
:
<specimenCount id="54946AD3FFB8FFB6FD8CFB47FD8DE597" box="[550,571,1254,1277]" pageId="45" pageNumber="46" type="male">♂</specimenCount>
<typeStatus id="9D291FF8FFB8FFB6FDEEFB44FD1FE594" box="[580,681,1253,1278]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
,
<collectionCode id="2483399FFFB8FFB6FD1CFB47FCB2E597" box="[694,772,1254,1277]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="45" pageNumber="46" type="Museum">ZFMK</collectionCode>
(
<location id="474DF781FFB8FFB6FCBEFB47FC85E594" LSID="urn:lsid:plazi:treatment:CA3B104CFFB8FFB4FF3DFCDFFC8FE356:474DF781FFB8FFB6FCBEFB47FC85E594" box="[788,819,1254,1278]" country="Sri Lanka" latitude="6.762" longLatPrecision="78" longitude="80.428" name="Ar" pageId="45" pageNumber="46" stateProvince="Sabaragamuwa">Ar</location>
20094),
<collectingRegion id="80566FB8FFB8FFB6FC36FB44FB01E594" box="[924,1207,1253,1278]" country="Sri Lanka" name="Sabaragamuwa" pageId="45" pageNumber="46">Sabaragamuwa Province</collectingRegion>
,
<locationDeviation id="539FB878FFB8FFB6FB69FB44FEDFE448" firstLocation="Ratnapura" pageId="45" pageNumber="46" secondLocation="Mapalena Ella">between Ratnapura and Mapalena Ella</locationDeviation>
(
<geoCoordinate id="27A6C79DFFB8FFB6FED2FAA8FE6DE448" box="[376,475,1289,1314]" direction="north" orientation="latitude" pageId="45" pageNumber="46" precision="55" value="6.762">6.762°N</geoCoordinate>
,
<geoCoordinate id="27A6C79DFFB8FFB6FE4CFAABFDE5E448" box="[486,595,1290,1314]" direction="east" orientation="longitude" pageId="45" pageNumber="46" precision="55" value="80.428">80.428°E</geoCoordinate>
),
<elevation id="C9BF4669FFB8FFB6FDCCFAABFD5CE448" box="[614,746,1289,1314]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="45" pageNumber="46" unit="m" value="100.0">
<quantity id="856A0CBFFFB8FFB6FDCCFAABFD1BE44B" box="[614,685,1290,1314]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="45" pageNumber="46" unit="m" value="100.0">100 m</quantity>
a.s.l.
</elevation>
,
<date id="362C879AFFB8FFB6FD5CFAA8FCC6E448" box="[758,880,1289,1314]" pageId="45" pageNumber="46" value="2017-03-19">
<collectingDate id="26687E72FFB8FFB6FD5CFAA8FCC6E448" box="[758,880,1289,1314]" pageId="45" pageNumber="46" value="2017-03-19">19.iii.2017</collectingDate>
</date>
(
<collectorName id="EF67C48CFFB8FFB6FC2AFAABFBB1E448" box="[896,1031,1289,1314]" pageId="45" pageNumber="46">B.A. Huber</collectorName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="422DA15AFFB8FFB6FF6DFA8CFDE3E400" blockId="45.[151,1437,1001,1998]" pageId="45" pageNumber="46">
<emphasis id="70E67D48FFB8FFB6FF6DFA8CFE4FE42C" bold="true" box="[199,505,1325,1350]" pageId="45" pageNumber="46">Other material examined</emphasis>
.
<materialsCitation id="F2FAAB07FFB8FFB6FDA3FA8FFBDFE42C" ID-GBIF-Occurrence="2234226171" box="[521,1129,1325,1350]" collectionCode="ZFMK" country="Sri Lanka" location="SRI LANKA" pageId="45" pageNumber="46" specimenCount="10" specimenCount-female="6" specimenCount-male="3">
<collectingCountry id="3A85E1CAFFB8FFB6FDA3FA8FFD13E42C" box="[521,677,1326,1350]" name="Sri Lanka" pageId="45" pageNumber="46">SRI LANKA</collectingCountry>
:
<specimenCount id="54946AD3FFB8FFB6FD1BFA8FFD63E42C" box="[689,725,1326,1350]" pageId="45" pageNumber="46" type="male">3♂</specimenCount>
<specimenCount id="54946AD3FFB8FFB6FD77FA8CFCB6E42C" box="[733,768,1325,1350]" pageId="45" pageNumber="46" type="female">6♀</specimenCount>
<specimenCount id="54946AD3FFB8FFB6FCA0FA8FFCFBE42C" box="[778,845,1325,1350]" pageId="45" pageNumber="46" type="generic">1 juv.</specimenCount>
,
<collectionCode id="2483399FFFB8FFB6FCF0FA8FFC1EE42F" box="[858,936,1326,1349]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="45" pageNumber="46" type="Museum">ZFMK</collectionCode>
(Ar 20095), and
</materialsCitation>
<materialsCitation id="F2FAAB07FFB8FFB6FBD9FA8FFDE7E400" ID-GBIF-Occurrence="2234226166" collectingDate="2017-03-19" collectionCode="ZFMK" collectorName="B. A. Huber" country="Sri Lanka" elevation="100" latitude="6.762" location="Ar" longLatPrecision="78" longitude="80.428" pageId="45" pageNumber="46" specimenCode="SL165" specimenCount="4" specimenCount-female="3" stateProvince="Sabaragamuwa">
<specimenCount id="54946AD3FFB8FFB6FBD9FA8FFB20E42C" box="[1139,1174,1326,1350]" pageId="45" pageNumber="46" type="female">3♀</specimenCount>
<specimenCount id="54946AD3FFB8FFB6FB35FA8FFB56E42C" box="[1183,1248,1325,1350]" pageId="45" pageNumber="46" type="generic">1 juv.</specimenCount>
in pure ethanol,
<collectionCode id="2483399FFFB8FFB6FF3DFAF3FF53E403" box="[151,229,1362,1385]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="45" pageNumber="46" type="Museum">ZFMK</collectionCode>
(
<specimenCode id="12340921FFB8FFB6FF5EFAF0FEF5E400" box="[244,323,1361,1386]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="45" pageNumber="46" type="Museum">SL165</specimenCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FFB8FFB4FF6DFAD4FAA1E372" lastPageId="47" lastPageNumber="48" pageId="45" pageNumber="46" type="description">
<paragraph id="422DA15AFFB8FFB6FF6DFAD4FB16E4BC" blockId="45.[151,1437,1001,1998]" pageId="45" pageNumber="46">
<emphasis id="70E67D48FFB8FFB6FF6DFAD4FE28E4E4" bold="true" box="[199,414,1397,1422]" pageId="45" pageNumber="46">Description. Male</emphasis>
(
<typeStatus id="9D291FF8FFB8FFB6FE04FAD4FDA0E4E4" box="[430,534,1397,1422]" pageId="45" pageNumber="46" type="holotype">holotype</typeStatus>
). MEASUREMENTS. Total body length 1.6, carapace width 0.70. Distance PME- PME 80 µm, diameter PME 80 µm, distance PME-ALE 30 µm; AME absent. Sternum width/length: 0.50/0.40. Leg 1: 12.4 (2.9 + 0.3 + 3.0 + 4.7 + 1.5), tibia 2: 1.6, tibia 3: 1.2, tibia 4: 1.7; tibia 1 L/d: 50.
</paragraph>
<paragraph id="422DA15AFFB8FFB6FF6DFA43FCF7E70F" blockId="45.[151,1437,1001,1998]" pageId="45" pageNumber="46">COLOR (in ethanol). Carapace ochre yellow with median and lateral dark bands; ocular area posteriorly dark; clypeus with pair of dark bands below eye triads; sternum mostly black, with light median band; legs ochre-yellow, with dark rings on femora (subdistally) and tibiae (proximally and subdistally); abdomen ochre-gray with dark marks dorsally and laterally, ventrally also with dark pattern.</paragraph>
<paragraph id="422DA15AFFB8FFB6FF6DF9D3FE96E7B8" blockId="45.[151,1437,1001,1998]" pageId="45" pageNumber="46">
BODY. Habitus as in
<figureCitation id="DAA9BDDFFFB8FFB6FE1DF9D0FDACE7E0" box="[439,538,1649,1674]" captionStart="FIGURES 138–149" captionStartId="34.[151,264,1387,1410]" captionTargetBox="[151,1436,195,1366]" captionTargetId="figure@34.[151,1436,195,1366]" captionTargetPageId="34" captionText="FIGURES 138–149. Wanniyala, live specimens, part 2. 138–140. W. ohiya sp. n., males and female with egg-sac from Ohiya (Fig. 139 shows a male in typical resting position). 141–143. W. viharekele sp. n., males from Viharekele and Kottawa, female from Kottawa. 144–146. W. mapalena sp. n., male and females from between Ratnapura and Mapalena Ella. 147–149. W. labugama sp. n., males and female with egg-sac from Labugama." httpUri="https://zenodo.org/record/2625076/files/figure.png" pageId="45" pageNumber="46">Fig. 144</figureCitation>
; ocular area slightly raised; carapace with shallow but distinct median furrow; clypeus with pair of relatively large processes at rim and small median pointed process (
<figureCitation id="DAA9BDDFFFB8FFB6FB26F934FA9AE7C4" box="[1164,1324,1685,1710]" captionStart="FIGURES 197–205" captionStartId="46.[151,264,1840,1863]" captionTargetBox="[152,1436,228,1785]" captionTargetId="figure@46.[151,1436,228,1785]" captionTargetPageId="46" captionText="FIGURES 197–205. Wanniyala mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 197–198. Male chelicerae, clypeus and ocular area, frontal and lateral views. 199. Left male palpal trochanter and femur, retrolateral view. 200. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 201–202. Left procursus, prolateral and retrolateral views (arrow points at distinctive process on retrolateral sclerite of procursus). 203–204. Epigynum, ventral and lateral views. 205. Cleared female genitalia, dorsal view (arrow points at V-shaped element). Scale lines: 0.3 mm (197–202, 205), 0.5 mm (203–204)." httpUri="https://zenodo.org/record/2625092/files/figure.png" pageId="45" pageNumber="46">Figs 197–198</figureCitation>
); sternum unmodified.
</paragraph>
<paragraph id="422DA15AFFB8FFB6FF6DF97FFF79E654" blockId="45.[151,1437,1001,1998]" pageId="45" pageNumber="46">
CHELICERAE. As in
<figureCitation id="DAA9BDDFFFB8FFB6FE17F97CFDD6E79C" box="[445,608,1757,1782]" captionStart="FIGURES 197–205" captionStartId="46.[151,264,1840,1863]" captionTargetBox="[152,1436,228,1785]" captionTargetId="figure@46.[151,1436,228,1785]" captionTargetPageId="46" captionText="FIGURES 197–205. Wanniyala mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 197–198. Male chelicerae, clypeus and ocular area, frontal and lateral views. 199. Left male palpal trochanter and femur, retrolateral view. 200. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 201–202. Left procursus, prolateral and retrolateral views (arrow points at distinctive process on retrolateral sclerite of procursus). 203–204. Epigynum, ventral and lateral views. 205. Cleared female genitalia, dorsal view (arrow points at V-shaped element). Scale lines: 0.3 mm (197–202, 205), 0.5 mm (203–204)." httpUri="https://zenodo.org/record/2625092/files/figure.png" pageId="45" pageNumber="46">Figs 197–198</figureCitation>
, with pair of small apophyses proximally laterally and pair of long apophyses directed forwards, tips of apophyses distally directed downwards, distance between tips of apophyses: 0.51.
</paragraph>
<paragraph id="422DA15AFFB8FFB6FF6DF8EBFDA6E6A4" blockId="45.[151,1437,1001,1998]" pageId="45" pageNumber="46">
PALPS. In general very similar to known congeners (cf.
<figureCitation id="DAA9BDDFFFB8FFB6FCF4F8E8FBB7E608" box="[862,1025,1865,1890]" captionStart="FIGURES 150–155" captionStartId="36.[151,264,1893,1916]" captionTargetBox="[218,1340,196,1875]" captionTargetId="figure@36.[202,1387,195,1876]" captionTargetPageId="36" captionText="FIGURES 150–155. Wanniyala left male palps in prolateral and retrolateral views. 150–151. W. upekkha sp. n. from Dematagala (ZFMK Ar 20086). 152–153. W. viharekele sp. n. from Viharekele (ZFMK Ar 20091). 154–155. W. labugama sp. n. from Labugama (ZFMK Ar 20097). Abbreviations: b, genital bulb; ba, bulbal apophysis; c, coxa; e, embolus; f, femur; p, procursus; pa, patella; pbs, proximal bulbal sclerite; ta, tarsus; ti, tibia; tr, trochanter. Scale lines: 0.5 mm." httpUri="https://zenodo.org/record/2625078/files/figure.png" pageId="45" pageNumber="46">Figs 150–155</figureCitation>
); coxa unmodified; trochanter with distinctive processes (
<figureCitation id="DAA9BDDFFFB8FFB6FE3BF8CCFE43E6EC" box="[401,501,1901,1926]" captionStart="FIGURES 197–205" captionStartId="46.[151,264,1840,1863]" captionTargetBox="[152,1436,228,1785]" captionTargetId="figure@46.[151,1436,228,1785]" captionTargetPageId="46" captionText="FIGURES 197–205. Wanniyala mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 197–198. Male chelicerae, clypeus and ocular area, frontal and lateral views. 199. Left male palpal trochanter and femur, retrolateral view. 200. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 201–202. Left procursus, prolateral and retrolateral views (arrow points at distinctive process on retrolateral sclerite of procursus). 203–204. Epigynum, ventral and lateral views. 205. Cleared female genitalia, dorsal view (arrow points at V-shaped element). Scale lines: 0.3 mm (197–202, 205), 0.5 mm (203–204)." httpUri="https://zenodo.org/record/2625092/files/figure.png" pageId="45" pageNumber="46">Fig. 199</figureCitation>
); femur with small dorsal apophysis proximally; tibia very long (0.68); procursus with several distinctive elements (
<figureCitation id="DAA9BDDFFFB8FFB6FDB9F830FD04E6C0" box="[531,690,1937,1962]" captionStart="FIGURES 197–205" captionStartId="46.[151,264,1840,1863]" captionTargetBox="[152,1436,228,1785]" captionTargetId="figure@46.[151,1436,228,1785]" captionTargetPageId="46" captionText="FIGURES 197–205. Wanniyala mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 197–198. Male chelicerae, clypeus and ocular area, frontal and lateral views. 199. Left male palpal trochanter and femur, retrolateral view. 200. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 201–202. Left procursus, prolateral and retrolateral views (arrow points at distinctive process on retrolateral sclerite of procursus). 203–204. Epigynum, ventral and lateral views. 205. Cleared female genitalia, dorsal view (arrow points at V-shaped element). Scale lines: 0.3 mm (197–202, 205), 0.5 mm (203–204)." httpUri="https://zenodo.org/record/2625092/files/figure.png" pageId="45" pageNumber="46">Figs 201–202</figureCitation>
); bulb with simple membranous embolus and short, weakly curved and pointed apophysis (
<figureCitation id="DAA9BDDFFFB8FFB6FE0BF814FDB2E6A4" box="[417,516,1973,1998]" captionStart="FIGURES 197–205" captionStartId="46.[151,264,1840,1863]" captionTargetBox="[152,1436,228,1785]" captionTargetId="figure@46.[151,1436,228,1785]" captionTargetPageId="46" captionText="FIGURES 197–205. Wanniyala mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 197–198. Male chelicerae, clypeus and ocular area, frontal and lateral views. 199. Left male palpal trochanter and femur, retrolateral view. 200. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 201–202. Left procursus, prolateral and retrolateral views (arrow points at distinctive process on retrolateral sclerite of procursus). 203–204. Epigynum, ventral and lateral views. 205. Cleared female genitalia, dorsal view (arrow points at V-shaped element). Scale lines: 0.3 mm (197–202, 205), 0.5 mm (203–204)." httpUri="https://zenodo.org/record/2625092/files/figure.png" pageId="45" pageNumber="46">Fig. 200</figureCitation>
).
</paragraph>
<caption id="16EDF1D2FFBBFFB5FF3DF891FDB2E68A" httpUri="https://zenodo.org/record/2625092/files/figure.png" pageId="46" pageNumber="47" startId="46.[151,264,1840,1863]" targetBox="[152,1436,228,1785]" targetPageId="46">
<paragraph id="422DA15AFFBBFFB5FF3DF891FDB2E68A" blockId="46.[151,1437,1840,2016]" pageId="46" pageNumber="47">
<emphasis id="70E67D48FFBBFFB5FF3DF891FEDEE62C" bold="true" box="[151,360,1840,1863]" pageId="46" pageNumber="47">FIGURES 197–205</emphasis>
.
<taxonomicName id="8592DAD9FFBBFFB5FEDCF890FDF8E62D" authority="Huber, 2019" authorityName="Huber" authorityYear="2019" box="[374,590,1841,1863]" class="Arachnida" family="Pholcidae" genus="Wanniyala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="mapalena" status="sp. nov.">
<emphasis id="70E67D48FFBBFFB5FEDCF890FDF8E62D" box="[374,590,1841,1863]" italics="true" pageId="46" pageNumber="47">Wanniyala mapalena</emphasis>
</taxonomicName>
<emphasis id="70E67D48FFBBFFB5FDFFF890FD26E62C" bold="true" box="[597,656,1841,1862]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="6BD5C033FFBBFFB5FDFFF890FD26E62C" box="[597,656,1841,1862]" pageId="46" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 197–198. Male chelicerae, clypeus and ocular area, frontal and lateral views. 199. Left male palpal trochanter and femur, retrolateral view. 200. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 201–202. Left procursus, prolateral and retrolateral views (arrow points at distinctive process on retrolateral sclerite of procursus). 203–204. Epigynum, ventral and lateral views. 205. Cleared female genitalia, dorsal view (arrow points at V-shaped element). Scale lines: 0.3 mm (197–202, 205), 0.5 mm (203–204).
</paragraph>
</caption>
<paragraph id="422DA15AFFBAFFB4FF6DFF39FD6EE192" blockId="47.[151,1436,151,573]" pageId="47" pageNumber="48">LEGS. Without spines and curved hairs; with short vertical hairs in higher than usual density on all metatarsi (especially proximally); retrolateral trichobothrium on tibia 1 at 11%; prolateral trichobothrium absent on tibia 1; tarsus 1 with ~20 pseudosegments, distally distinct.</paragraph>
<paragraph id="422DA15AFFBAFFB4FF6DFEA5FAE7E076" blockId="47.[151,1436,151,573]" box="[199,1361,260,285]" pageId="47" pageNumber="48">
<emphasis id="70E67D48FFBAFFB4FF6DFEA5FEB2E077" bold="true" box="[199,260,260,285]" pageId="47" pageNumber="48">Male</emphasis>
(variation). Tibia
<quantity id="856A0CBFFFBAFFB4FE7AFEA5FE4DE077" box="[464,507,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="47" pageNumber="48" unit="in" value="1.0">1 in</quantity>
three other males: 2.7, 2.9, 3.0. One male with irregular pattern on sternum.
</paragraph>
<paragraph id="422DA15AFFBAFFB4FF6DFE86FE34E09F" blockId="47.[151,1436,151,573]" pageId="47" pageNumber="48">
<emphasis id="70E67D48FFBAFFB4FF6DFE86FEA8E02A" bold="true" box="[199,286,295,320]" pageId="47" pageNumber="48">Female</emphasis>
. In general similar to male (
<figureCitation id="DAA9BDDFFFBAFFB4FDF6FE86FD4EE02A" box="[604,760,295,320]" captionStart="FIGURES 138–149" captionStartId="34.[151,264,1387,1410]" captionTargetBox="[151,1436,195,1366]" captionTargetId="figure@34.[151,1436,195,1366]" captionTargetPageId="34" captionText="FIGURES 138–149. Wanniyala, live specimens, part 2. 138–140. W. ohiya sp. n., males and female with egg-sac from Ohiya (Fig. 139 shows a male in typical resting position). 141–143. W. viharekele sp. n., males from Viharekele and Kottawa, female from Kottawa. 144–146. W. mapalena sp. n., male and females from between Ratnapura and Mapalena Ella. 147–149. W. labugama sp. n., males and female with egg-sac from Labugama." httpUri="https://zenodo.org/record/2625076/files/figure.png" pageId="47" pageNumber="48">Figs 145–146</figureCitation>
) but clypeus unmodified and legs with usual low number of short vertical hairs; light median band on sternum in some females barely visible. Tibia
<quantity id="856A0CBFFFBAFFB4FB3BFEEDFB09E00F" box="[1169,1215,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="47" pageNumber="48" unit="in" value="1.0">1 in</quantity>
<specimenCount id="54946AD3FFBAFFB4FB62FEEDFA8DE00F" box="[1224,1339,332,357]" pageId="47" pageNumber="48" type="female">6 females</specimenCount>
: 2.1–2.4 (mean 2.3). Epigynum as in
<figureCitation id="DAA9BDDFFFBAFFB4FE52FECEFD2CE0E2" box="[504,666,367,392]" captionStart="FIGURES 197–205" captionStartId="46.[151,264,1840,1863]" captionTargetBox="[152,1436,228,1785]" captionTargetId="figure@46.[151,1436,228,1785]" captionTargetPageId="46" captionText="FIGURES 197–205. Wanniyala mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 197–198. Male chelicerae, clypeus and ocular area, frontal and lateral views. 199. Left male palpal trochanter and femur, retrolateral view. 200. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 201–202. Left procursus, prolateral and retrolateral views (arrow points at distinctive process on retrolateral sclerite of procursus). 203–204. Epigynum, ventral and lateral views. 205. Cleared female genitalia, dorsal view (arrow points at V-shaped element). Scale lines: 0.3 mm (197–202, 205), 0.5 mm (203–204)." httpUri="https://zenodo.org/record/2625092/files/figure.png" pageId="47" pageNumber="48">Figs 203–204</figureCitation>
, weakly protruding and rounded in lateral view; posterior plate apparently reduced(?) to two lateral sclerites mostly hidden behind epigynal plate and poorly visible in ventral view; internal genitalia as in
<figureCitation id="DAA9BDDFFFBAFFB4FE42FE16FDF9E0BA" box="[488,591,439,464]" captionStart="FIGURES 197–205" captionStartId="46.[151,264,1840,1863]" captionTargetBox="[152,1436,228,1785]" captionTargetId="figure@46.[151,1436,228,1785]" captionTargetPageId="46" captionText="FIGURES 197–205. Wanniyala mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 197–198. Male chelicerae, clypeus and ocular area, frontal and lateral views. 199. Left male palpal trochanter and femur, retrolateral view. 200. Left genital bulb (proximal sclerite, bulbal apophysis and embolus), prolateral view. 201–202. Left procursus, prolateral and retrolateral views (arrow points at distinctive process on retrolateral sclerite of procursus). 203–204. Epigynum, ventral and lateral views. 205. Cleared female genitalia, dorsal view (arrow points at V-shaped element). Scale lines: 0.3 mm (197–202, 205), 0.5 mm (203–204)." httpUri="https://zenodo.org/record/2625092/files/figure.png" pageId="47" pageNumber="48">Figs 205</figureCitation>
,
<figureCitation id="DAA9BDDFFFBAFFB4FDF7FE19FD3CE0BA" box="[605,650,440,464]" captionStart="FIGURES 213–219" captionStartId="49.[151,264,1720,1743]" captionTargetBox="[151,1436,412,1657]" captionTargetId="figure@49.[151,1436,412,1657]" captionTargetPageId="49" captionText="FIGURES 213–219. Wanniyala spp., cleared female genitalia in dorsal views. 213. W. mudita sp. n., from Kandy (ZFMK Ar 20077). 214. W. orientalis sp. n., from Dimbulagala (ZFMK Ar 20079). 215. W. upekkha sp. n., from Dematagala (ZFMK Ar 20086). 216. W. ohiya sp. n., from Ohiya (ZFMK Ar 20089). 217. W. viharekele sp. n., from Viharekele (ZFMK Ar 20092). 218. W. mapalena sp. n., from between Ratnapura and Mapalena Ella (ZFMK Ar 20095). 219. W. labugama sp. n., from Labugama (ZFMK Ar 20097). At various scales." httpUri="https://zenodo.org/record/2625096/files/figure.png" pageId="47" pageNumber="48">218</figureCitation>
, with distinctive pair of v-shaped membranous elements and pair of roundish pore plates.
</paragraph>
<paragraph id="422DA15AFFBAFFB4FF6DFE5EFAA1E372" blockId="47.[151,1436,151,573]" box="[199,1303,511,536]" pageId="47" pageNumber="48">
<emphasis id="70E67D48FFBAFFB4FF6DFE5EFECBE372" bold="true" box="[199,381,511,536]" pageId="47" pageNumber="48">Natural history</emphasis>
. The spiders were found in small webs among small exposed roots on the ground.
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FFBAFFB4FF6DFD85FC8FE356" box="[199,825,548,573]" pageId="47" pageNumber="48" type="distribution">
<paragraph id="422DA15AFFBAFFB4FF6DFD85FC8FE356" blockId="47.[151,1436,151,573]" box="[199,825,548,573]" pageId="47" pageNumber="48">
<emphasis id="70E67D48FFBAFFB4FF6DFD85FEEFE357" bold="true" box="[199,345,548,573]" pageId="47" pageNumber="48">Distribution</emphasis>
. Known from
<typeStatus id="9D291FF8FFBAFFB4FE54FD84FD98E357" box="[510,558,549,573]" pageId="47" pageNumber="48">type</typeStatus>
locality only (
<figureCitation id="DAA9BDDFFFBAFFB4FD7BFD85FC84E356" box="[721,818,548,573]" captionStart="FIGURES 226–227" captionStartId="53.[151,264,898,921]" captionTargetBox="[151,1436,183,885]" captionTargetId="figure@53.[151,1436,183,885]" captionTargetPageId="53" captionText="FIGURES 226–227. Known distribution of Wanniyala." httpUri="https://zenodo.org/record/2625104/files/figure.png" pageId="47" pageNumber="48">Fig. 227</figureCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>
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