171 lines
19 KiB
XML
171 lines
19 KiB
XML
<document id="A45904C9BEAC2EDF500EF00729B3F8E0" ID-CLB-Dataset="27502" ID-DOI="10.11646/zootaxa.4550.1.1" ID-GBIF-Dataset="291701d4-652e-440e-adab-f6b0201c0972" ID-ISSN="1175-5326" ID-Zenodo-Dep="2625030" ID-ZooBank="2F7D1EC4-D4ED-4FAE-B227-CF7B79EAE833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554274879827" checkinUser="plazi" docAuthor="Huber, Bernhard A." docDate="2019" docId="CA3B104CFF92FF9CFF3DFE35FC59E2A8" docLanguage="en" docName="zootaxa.4550.1.1.pdf" docOrigin="Zootaxa 4550 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana benjamini Huber 2005" docType="treatment" docVersion="14" lastPageNumber="8" masterDocId="36026834FF95FF9BFFAAFFA1FFB6E16A" masterDocTitle="The pholcid spiders of Sri Lanka (Araneae: Pholcidae)" masterLastPageNumber="57" masterPageNumber="1" pageNumber="8" updateTime="1698716853299" updateUser="ExternalLinkService">
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<mods:titleInfo id="0CC05070B936789C25C6A5AAF51114DF">
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<mods:title id="00A6BE7E73EF5CFA9E33B6E0C92C152A">The pholcid spiders of Sri Lanka (Araneae: Pholcidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="2F1253202660A8B464431A68A7A1DEC9" type="personal">
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<mods:namePart id="4A8CB14BC614A5CBA353FC1D17DC58C6">Huber, Bernhard A.</mods:namePart>
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<mods:typeOfResource id="609F90C3866ACC1B9F2B7357455AF3C4">text</mods:typeOfResource>
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<mods:title id="24EBA7E92FFD24D4555E160FE9D7A1D8">Zootaxa</mods:title>
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<mods:part id="5D91985AD7B777BDC46BB3A9F8A13308">
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<mods:date id="29C497E857FC7DDBED6C1EAAE1672079">2019</mods:date>
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<mods:number id="ADEBF50EAA7BAFF1FE3547673BC40F64">2019-01-23</mods:number>
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<mods:number id="B368C83B79E73D788478E6C8FF440C24">4550</mods:number>
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<mods:identifier id="67040A34A22F9A3AFA1E485E3BA5FF4D" type="DOI">10.11646/zootaxa.4550.1.1</mods:identifier>
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<treatment id="CA3B104CFF92FF9CFF3DFE35FC59E2A8" ID-DOI="http://doi.org/10.5281/zenodo.4581582" ID-GBIF-Taxon="154534127" ID-Zenodo-Dep="4581582" LSID="urn:lsid:plazi:treatment:CA3B104CFF92FF9CFF3DFE35FC59E2A8" httpUri="http://treatment.plazi.org/id/CA3B104CFF92FF9CFF3DFE35FC59E2A8" lastPageNumber="8" pageId="7" pageNumber="8">
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<subSubSection id="0A88F2D1FF92FF9CFF3DFE35FD94E0C4" box="[151,546,404,430]" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="422DA15AFF92FF9CFF3DFE35FD94E0C4" blockId="7.[151,546,404,463]" box="[151,546,404,430]" pageId="7" pageNumber="8">
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<heading id="19651636FF92FF9CFF3DFE35FD94E0C4" bold="true" box="[151,546,404,430]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
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<taxonomicName id="8592DAD9FF92FF9CFF3DFE35FD94E0C4" ID-CoL="5WH9K" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[151,546,404,430]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="benjamini">
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<emphasis id="70E67D48FF92FF9CFF3DFE35FD94E0C4" bold="true" box="[151,546,404,430]" pageId="7" pageNumber="8">
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<emphasis id="70E67D48FF92FF9CFF3DFE35FE34E0C4" bold="true" box="[151,386,404,430]" italics="true" pageId="7" pageNumber="8">Belisana benjamini</emphasis>
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Huber, 2005
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="0A88F2D1FF92FF9CFF3DFE17FED4E0A5" box="[151,354,438,463]" pageId="7" pageNumber="8" type="description">
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<paragraph id="422DA15AFF92FF9CFF3DFE17FED4E0A5" blockId="7.[151,546,404,463]" box="[151,354,438,463]" pageId="7" pageNumber="8">
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<figureCitation id="DAA9BDDFFF92FF9CFF3DFE17FEA6E0A5" box="[151,272,438,463]" captionStart="FIGURES 17–24" captionStartId="5.[151,264,1673,1696]" captionTargetBox="[174,1406,476,1654]" captionTargetId="figure@5.[173,1412,476,1658]" captionTargetPageId="5" captionText="FIGURES 17–24. 17–22. Belisana badulla sp. n. (ZFMK Ar 20002). 17–19. Left procursus, prolateral, dorsal, and retrolateral views. 20–21. Male chelicerae, frontal and lateral views. 22. Left genital bulb, prolateral view. 23–24. B. keyti Huber, 2005 (ZFMK Ar 20004) and B. benjamini Huber, 2005 (ZFMK Ar 20005), left genital bulbs, prolateral views (all genital bulbs at same scale). Abbreviations: ba, bulbal apophysis; e, embolus. Scale lines: 0.2 mm." httpUri="https://zenodo.org/record/2625036/files/figure.png" pageId="7" pageNumber="8">Figures 24</figureCitation>
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,
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<figureCitation id="DAA9BDDFFF92FF9CFEB6FE16FED4E0A5" box="[284,354,439,463]" captionStart="FIGURES 25–30" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[177,1411,435,1882]" captionTargetId="figure@6.[177,1411,435,1882]" captionTargetPageId="6" captionText="FIGURES 25–30. Cleared female genitalia in ventral and dorsal views, all at same scale. 25–26. Belisana badulla sp. n. (ZFMK Ar 20002). 27–28. B. keyti Huber, 2005 (ZFMK Ar 20004); arrows point at distinctive pair of internal folds. 29–30. B. benjamini Huber, 2005 (ZFMK Ar 20005); arrows point at median folds. Abbreviations: po, pocket; pp, pore plate. Scale line: 0.2 mm." httpUri="https://zenodo.org/record/2625038/files/figure.png" pageId="7" pageNumber="8">29–30</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0A88F2D1FF92FF9CFF3DFE5FFC15E37E" box="[151,931,510,532]" pageId="7" pageNumber="8" type="reference_group">
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<paragraph id="422DA15AFF92FF9CFF3DFE5FFC15E37E" blockId="7.[151,931,510,532]" box="[151,931,510,532]" pageId="7" pageNumber="8">
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<taxonomicName id="8592DAD9FF92FF9CFF3DFE5FFDA5E37E" ID-CoL="5WH9K" authority="Huber, 2005: 112" authorityName="Huber" authorityPageNumber="112" authorityYear="2005" box="[151,531,510,532]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="benjamini">
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<emphasis id="70E67D48FF92FF9CFF3DFE5FFEEDE37E" box="[151,347,510,532]" italics="true" pageId="7" pageNumber="8">Belisana benjamini</emphasis>
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Huber, 2005: 112
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</taxonomicName>
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, figs 37–38, 621–625 (
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<specimenCount id="54946AD3FF92FF9CFD54FE5FFCA3E37E" box="[766,789,510,532]" pageId="7" pageNumber="8" type="male">♂</specimenCount>
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<specimenCount id="54946AD3FF92FF9CFCBFFE5FFC93E37E" box="[789,805,510,532]" pageId="7" pageNumber="8" type="female">♀</specimenCount>
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,
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<collectingCountry id="3A85E1CAFF92FF9CFC9AFE5FFC2EE37E" box="[816,920,510,532]" name="Sri Lanka" pageId="7" pageNumber="8">Sri Lanka</collectingCountry>
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).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0A88F2D1FF92FF9CFF3DFDE0FB9CE380" pageId="7" pageNumber="8" type="diagnosis">
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||
<paragraph id="422DA15AFF92FF9CFF3DFDE0FB9CE380" blockId="7.[151,1437,577,962]" pageId="7" pageNumber="8">
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||
<emphasis id="70E67D48FF92FF9CFF3DFDE0FEBCE330" bold="true" box="[151,266,577,602]" pageId="7" pageNumber="8">Diagnosis</emphasis>
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(amendments; see Huber 2005). Easily distinguished from most congeners by curvature of procursus (towards ventral; Huber 2005: fig. 622); from two Sri Lankan species with similar procursus (
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<taxonomicName id="8592DAD9FF92FF9CFB10FDC7FA99E314" authorityName="Huber" authorityYear="2019" box="[1210,1327,613,638]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="badulla">
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<emphasis id="70E67D48FF92FF9CFB10FDC7FA99E314" box="[1210,1327,613,638]" italics="true" pageId="7" pageNumber="8">B. badulla</emphasis>
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||
</taxonomicName>
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,
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<taxonomicName id="8592DAD9FF92FF9CFA94FDC7FA25E314" authorityName="Huber" authorityYear="2005" box="[1342,1427,613,638]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="keyti">
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<emphasis id="70E67D48FF92FF9CFA94FDC7FA25E314" box="[1342,1427,613,638]" italics="true" pageId="7" pageNumber="8">B. keyti</emphasis>
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||
</taxonomicName>
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) by absence of bulbal apophysis (
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<figureCitation id="DAA9BDDFFF92FF9CFDA5FD28FDD0E3C8" box="[527,614,649,674]" captionStart="FIGURES 17–24" captionStartId="5.[151,264,1673,1696]" captionTargetBox="[174,1406,476,1654]" captionTargetId="figure@5.[173,1412,476,1658]" captionTargetPageId="5" captionText="FIGURES 17–24. 17–22. Belisana badulla sp. n. (ZFMK Ar 20002). 17–19. Left procursus, prolateral, dorsal, and retrolateral views. 20–21. Male chelicerae, frontal and lateral views. 22. Left genital bulb, prolateral view. 23–24. B. keyti Huber, 2005 (ZFMK Ar 20004) and B. benjamini Huber, 2005 (ZFMK Ar 20005), left genital bulbs, prolateral views (all genital bulbs at same scale). Abbreviations: ba, bulbal apophysis; e, embolus. Scale lines: 0.2 mm." httpUri="https://zenodo.org/record/2625036/files/figure.png" pageId="7" pageNumber="8">Fig. 24</figureCitation>
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). Females are difficult to distinguish externally from similar congeners; internal genitalia smaller and with distinctive median folds and elongated pore plates (not consisting of round lateral part and long narrow elongation as in
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<taxonomicName id="8592DAD9FF92FF9CFD27FD73FCB7E380" authorityName="Huber" authorityYear="2019" box="[653,769,721,746]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="badulla">
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<emphasis id="70E67D48FF92FF9CFD27FD73FCB7E380" box="[653,769,721,746]" italics="true" pageId="7" pageNumber="8">B. badulla</emphasis>
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||
</taxonomicName>
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||
and
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<emphasis id="70E67D48FF92FF9CFC92FD73FC23E380" box="[824,917,721,746]" italics="true" pageId="7" pageNumber="8">
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||
<taxonomicName id="8592DAD9FF92FF9CFC92FD73FC38E380" authorityName="Huber" authorityYear="2005" box="[824,910,721,746]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="keyti">B. keyti</taxonomicName>
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;
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</emphasis>
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<figureCitation id="DAA9BDDFFF92FF9CFC36FD70FBA9E380" box="[924,1055,721,746]" captionStart="FIGURES 25–30" captionStartId="6.[151,264,1904,1927]" captionTargetBox="[177,1411,435,1882]" captionTargetId="figure@6.[177,1411,435,1882]" captionTargetPageId="6" captionText="FIGURES 25–30. Cleared female genitalia in ventral and dorsal views, all at same scale. 25–26. Belisana badulla sp. n. (ZFMK Ar 20002). 27–28. B. keyti Huber, 2005 (ZFMK Ar 20004); arrows point at distinctive pair of internal folds. 29–30. B. benjamini Huber, 2005 (ZFMK Ar 20005); arrows point at median folds. Abbreviations: po, pocket; pp, pore plate. Scale line: 0.2 mm." httpUri="https://zenodo.org/record/2625038/files/figure.png" pageId="7" pageNumber="8">Figs 29–30</figureCitation>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="0A88F2D1FF92FF9CFF6DFD54FE5DE2F4" pageId="7" pageNumber="8" type="description">
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<paragraph id="422DA15AFF92FF9CFF6DFD54FF44E258" blockId="7.[151,1437,577,962]" pageId="7" pageNumber="8">
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<emphasis id="70E67D48FF92FF9CFF6DFD54FEE7E264" bold="true" box="[199,337,757,782]" pageId="7" pageNumber="8">Description</emphasis>
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||
(amendments; see Huber 2005). Tibia
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<quantity id="856A0CBFFF92FF9CFCA4FD57FC8CE264" box="[782,826,757,782]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="8" unit="in" value="1.0">1 in</quantity>
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two newly examined males: 2.6, 2.9; in two females: 1.9, 2.3.
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||
</paragraph>
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<paragraph id="422DA15AFF92FF9CFF6DFC9CFE5DE2F4" blockId="7.[151,1437,577,962]" pageId="7" pageNumber="8">
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<emphasis id="70E67D48FF92FF9CFF6DFC9CFEE7E23C" bold="true" box="[199,337,829,854]" pageId="7" pageNumber="8">New record</emphasis>
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.
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<materialsCitation id="F2FAAB07FF92FF9CFECBFC9FFCDAE23C" ID-GBIF-Occurrence="2234226232" box="[353,876,829,854]" collectionCode="ZFMK" collectorName="Ar" country="Sri Lanka" location="SRI LANKA" pageId="7" pageNumber="8" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
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<collectingCountry id="3A85E1CAFF92FF9CFECBFC9FFE4CE23C" box="[353,506,830,854]" name="Sri Lanka" pageId="7" pageNumber="8">SRI LANKA</collectingCountry>
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:
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<specimenCount id="54946AD3FF92FF9CFDAFFC9FFD9FE23C" box="[517,553,830,854]" pageId="7" pageNumber="8" type="male">2♂</specimenCount>
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,
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<collectionCode id="2483399FFF92FF9CFDC8FC9FFD06E23F" box="[610,688,830,853]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="7" pageNumber="8" type="Museum">ZFMK</collectionCode>
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(
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<collectorName id="EF67C48CFF92FF9CFD15FC9FFD68E23C" box="[703,734,830,854]" pageId="7" pageNumber="8">Ar</collectorName>
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20005), and
|
||
</materialsCitation>
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||
<materialsCitation id="F2FAAB07FF92FF9CFCDEFC9FFE53E2F4" ID-GBIF-Occurrence="2234226285" collectingDate="2009-12-02" collectionCode="ZFMK" collectorName="S. P. Benjamin & S. Batuwita" country="Sri Lanka" county="Benj" elevation="1000" latitude="7.528" location="Riverstone" longLatPrecision="78" longitude="80.738" municipality="Matale District" pageId="7" pageNumber="8" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Central">
|
||
<specimenCount id="54946AD3FF92FF9CFCDEFC9FFC21E23C" box="[884,919,830,854]" pageId="7" pageNumber="8" type="male">1♂</specimenCount>
|
||
<specimenCount id="54946AD3FF92FF9CFC35FC9FFC74E23C" box="[927,962,830,854]" pageId="7" pageNumber="8" type="female">2♀</specimenCount>
|
||
in pure ethanol,
|
||
<collectionCode id="2483399FFF92FF9CFB28FC9FFB66E23F" box="[1154,1232,830,853]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="7" pageNumber="8" type="Museum">ZFMK</collectionCode>
|
||
(
|
||
<collectingCounty id="AB4CD9D6FF92FF9CFB4AFC9CFAAFE23C" box="[1248,1305,829,854]" pageId="7" pageNumber="8">Benj</collectingCounty>
|
||
53),
|
||
<collectingRegion id="80566FB8FF92FF9CFAE0FC9CFF48E210" country="Sri Lanka" name="Central" pageId="7" pageNumber="8">Central Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="A2493B20FF92FF9CFEA4FCC0FE76E210" box="[270,448,865,890]" pageId="7" pageNumber="8">Matale District</collectingMunicipality>
|
||
,
|
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<location id="474DF781FF92FF9CFE7BFCC0FDFBE210" LSID="urn:lsid:plazi:treatment:CA3B104CFF92FF9CFF3DFE35FC59E2A8:474DF781FF92FF9CFE7BFCC0FDFBE210" box="[465,589,865,890]" country="Sri Lanka" county="Benj" latitude="7.528" longLatPrecision="78" longitude="80.738" municipality="Matale District" name="Riverstone" pageId="7" pageNumber="8" stateProvince="Central">Riverstone</location>
|
||
,
|
||
<location id="474DF781FF92FF9CFDF4FCC0FCAAE210" LSID="urn:lsid:plazi:treatment:CA3B104CFF92FF9CFF3DFE35FC59E2A8:474DF781FF92FF9CFDF4FCC0FCAAE210" box="[606,796,865,890]" country="Sri Lanka" county="Benj" latitude="7.528" longLatPrecision="78" longitude="80.738" municipality="Matale District" name="Knuckles Range" pageId="7" pageNumber="8" stateProvince="Central">Knuckles Range</location>
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||
(
|
||
<geoCoordinate id="27A6C79DFF92FF9CFC9BFCC0FC22E210" box="[817,916,865,890]" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="55" value="7.528">7.528°N</geoCoordinate>
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,
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<geoCoordinate id="27A6C79DFF92FF9CFC0FFCC3FBA4E210" box="[933,1042,866,890]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="55" value="80.738">80.738°E</geoCoordinate>
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), ~
|
||
<elevation id="C9BF4669FF92FF9CFB90FCC3FB6EE210" box="[1082,1240,865,890]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="m" value="1000.0">
|
||
<quantity id="856A0CBFFF92FF9CFB90FCC3FB20E213" box="[1082,1174,866,890]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="m" value="1000.0">1000 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="362C879AFF92FF9CFB40FCC0FAEAE210" box="[1258,1372,865,890]" pageId="7" pageNumber="8" value="2009-12-02">
|
||
<collectingDate id="26687E72FF92FF9CFB40FCC0FAEAE210" box="[1258,1372,865,890]" pageId="7" pageNumber="8" value="2009-12-02">2.xii.2009</collectingDate>
|
||
</date>
|
||
(
|
||
<collectorName id="EF67C48CFF92FF9CFAD8FCC3FEB1E2F4" pageId="7" pageNumber="8">S.P. Benjamin</collectorName>
|
||
,
|
||
<collectorName id="EF67C48CFF92FF9CFEB8FC27FE21E2F4" box="[274,407,901,926]" pageId="7" pageNumber="8">S. Batuwita</collectorName>
|
||
,
|
||
<emphasis id="70E67D48FF92FF9CFE08FC27FE6DE2F4" box="[418,475,901,926]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A88F2D1FF92FF9CFF6DFC08FC59E2A8" box="[199,1007,937,962]" pageId="7" pageNumber="8" type="distribution">
|
||
<paragraph id="422DA15AFF92FF9CFF6DFC08FC59E2A8" blockId="7.[151,1437,577,962]" box="[199,1007,937,962]" pageId="7" pageNumber="8">
|
||
<emphasis id="70E67D48FF92FF9CFF6DFC08FEEFE2A8" bold="true" box="[199,345,937,962]" pageId="7" pageNumber="8">Distribution</emphasis>
|
||
. Known from two localities in Knuckles Range (
|
||
<figureCitation id="DAA9BDDFFF92FF9CFC2AFC08FC55E2A8" box="[896,995,937,962]" captionStart="FIGURES 220–221" captionStartId="51.[151,264,919,942]" captionTargetBox="[151,1436,195,898]" captionTargetId="figure@51.[151,1436,195,898]" captionTargetPageId="51" captionText="FIGURES 220–221. Known distribution of Belisana in Sri Lanka." httpUri="https://zenodo.org/record/2625098/files/figure.png" pageId="7" pageNumber="8">Fig. 220</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |