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<document id="9C592F1729E9DD62E3B31EEF422ADD11" ID-CLB-Dataset="26128" ID-DOI="10.11646/zootaxa.4646.2.9" ID-GBIF-Dataset="5e229ea9-be13-4238-aef6-13a32e4eeba8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3349497" ID-ZooBank="D32B0AE3-773E-4244-99C0-2FB82D41D12D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1563947575391" checkinUser="plazi" docAuthor="Roxo, Fabio F., Messias, Felipe L. &amp; Silva, Gabriel S. C." docDate="2019" docId="C94673767577FFF2FF04A5B3850A0D77" docLanguage="en" docName="zootaxa.4646.2.9.pdf" docOrigin="Zootaxa 4646 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parotocinclus pentakelis Roxo &amp; Messias &amp; Silva 2019, new species" docType="treatment" docVersion="9" lastPageNumber="350" masterDocId="357F0B0E7576FFF6FF93A76A86030E60" masterDocTitle="A new species of Parotocinclus (Loricariidae: Hypoptopomatinae) from Rio Tocantins basin, Brazil" masterLastPageNumber="356" masterPageNumber="346" pageNumber="347" updateTime="1698747211264" updateUser="ExternalLinkService">
<mods:mods id="4F31D77EC1CD095A592808FCE8A0F6A1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C23EEF200B1AFDED645CB79F9A028ECF">
<mods:title id="6F4CF0828D4884D928753BCC1B50E9B4">A new species of Parotocinclus (Loricariidae: Hypoptopomatinae) from Rio Tocantins basin, Brazil</mods:title>
</mods:titleInfo>
<mods:name id="5FE2AEC2B95DC6154D834F698791F0BB" type="personal">
<mods:role id="24410BAAB6827CDAE1B586044CCBB7BD">
<mods:roleTerm id="52D0ED5753C5E9C5DC674B13AA9407AB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D411CE8211A690856EAF67F8E574413A">Roxo, Fabio F.</mods:namePart>
</mods:name>
<mods:name id="E870D4C1215779A14A752040C1B3F866" type="personal">
<mods:role id="668EB0494AE263496E4B8EC5DF513FBA">
<mods:roleTerm id="29E0A90EE69D0652D176D2AC403AB8B0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0AB72550798F56B380EFC77098FD249F">Messias, Felipe L.</mods:namePart>
</mods:name>
<mods:name id="0218A5DC0FB771375B51776A0B8B5FFE" type="personal">
<mods:role id="4CEAACA8225CBA359EE8348892D56801">
<mods:roleTerm id="7B684285B3D48B7CEA664728B54348CA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5AE5D90D4D64B1CD25EE6F480B5998B8">Silva, Gabriel S. C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="9141160EB72BE48E07770878C7117390">text</mods:typeOfResource>
<mods:relatedItem id="92A3AE15953D548D8ABB7D46AFB6BE86" type="host">
<mods:titleInfo id="535ED3DA905E68FEC22CDB18F543DC69">
<mods:title id="B908EAA853FBC410DBB2C39CD596DF69">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="57361A1D765AB2BF1120AD474D04B44E">
<mods:date id="755F19B553203DC781A0609BF312B90C">2019</mods:date>
<mods:detail id="B4CC0B1BFB9F2BCD9E90E413B357AF01" type="pubDate">
<mods:number id="EEB2822FA38B6734EB8A950149E6FF62">2019-07-24</mods:number>
</mods:detail>
<mods:detail id="0BC7EA265C0C64314E92FFB549B5D691" type="volume">
<mods:number id="A7F14F5182515602129825B4FAD2B162">4646</mods:number>
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<mods:detail id="4DEB355E02103C740BE644E88D74958F" type="issue">
<mods:number id="F7F28A1F6F931508030ADEFBF62F54E4">2</mods:number>
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<mods:start id="BA3CBD2EDC3EA729D746A2457EB82AAA">346</mods:start>
<mods:end id="413F3AC9BCEAF67CDDC9D8D09CB1EA19">356</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="15B05BB513C8C91AD1FFA76B338FAB82">journal article</mods:classification>
<mods:identifier id="DD1C05D96AC59B15BC933604E22623D4" type="CLB-Dataset">26128</mods:identifier>
<mods:identifier id="760144DA46DBF3118464DA79D665DF98" type="DOI">10.11646/zootaxa.4646.2.9</mods:identifier>
<mods:identifier id="83AE0BAEEED618CA14D25F9EF28E6031" type="GBIF-Dataset">5e229ea9-be13-4238-aef6-13a32e4eeba8</mods:identifier>
<mods:identifier id="1503FC47DBFA5B522D6F7F0042EA60E1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C67317A8DD81EEAAA70602DA3179E356" type="Zenodo-Dep">3349497</mods:identifier>
<mods:identifier id="B0C671016B11FAA0BF4181E34C6219B5" type="ZooBank">D32B0AE3-773E-4244-99C0-2FB82D41D12D</mods:identifier>
</mods:mods>
<treatment id="C94673767577FFF2FF04A5B3850A0D77" ID-GBIF-Taxon="158852402" LSID="urn:lsid:plazi:treatment:C94673767577FFF2FF04A5B3850A0D77" httpUri="http://treatment.plazi.org/id/C94673767577FFF2FF04A5B3850A0D77" lastPageId="4" lastPageNumber="350" pageId="1" pageNumber="347">
<subSubSection id="09F591EB7577FFF7FF04A5B3845C0C93" box="[151,607,729,756]" pageId="1" pageNumber="347" type="nomenclature">
<paragraph id="4150C2607577FFF7FF04A5B3845C0C93" blockId="1.[151,607,729,792]" box="[151,607,729,756]" pageId="1" pageNumber="347">
<heading id="1A18750C7577FFF7FF04A5B3845C0C93" bold="true" box="[151,607,729,756]" fontSize="11" level="1" pageId="1" pageNumber="347" reason="1">
<emphasis id="739B1E727577FFF7FF04A5B3845C0C93" bold="true" box="[151,607,729,756]" pageId="1" pageNumber="347">
<taxonomicName id="86EFB9E37577FFF7FF04A5B387C30C94" authority="Roxo &amp; Messias &amp; Silva, 2019" authorityName="Roxo &amp; Messias &amp; Silva" authorityYear="2019" box="[151,448,729,756]" class="Actinopterygii" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="347" phylum="Chordata" rank="species" species="pentakelis" status="sp. nov.">
<emphasis id="739B1E727577FFF7FF04A5B387C30C94" bold="true" box="[151,448,729,756]" italics="true" pageId="1" pageNumber="347">Parotocinclus pentakelis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="68A8A3097577FFF7FE5CA5B0845C0C93" box="[463,607,729,756]" pageId="1" pageNumber="347" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="09F591EB7577FFF7FF04A597875F0D78" box="[151,348,765,792]" pageId="1" pageNumber="347" type="description">
<paragraph id="4150C2607577FFF7FF04A597875F0D78" blockId="1.[151,607,729,792]" box="[151,348,765,792]" pageId="1" pageNumber="347">
<figureCitation id="D9D4DEE57577FFF7FF04A59786DA0D77" box="[151,217,765,791]" captionStart="FIGURE 1" captionStartId="2.[151,250,1737,1762]" captionTargetBox="[151,1436,417,1718]" captionTargetId="figure@2.[151,1436,417,1718]" captionTargetPageId="2" captionText="FIGURE 1. Parotocinclus pentakelis, MZUSP 124900, holotype, 22.8 mm SL, rio Palmas, rio Tocantins basin." httpUri="https://zenodo.org/record/3349499/files/figure.png" pageId="1" pageNumber="347">Fig. 1</figureCitation>
and Table I
</paragraph>
</subSubSection>
<subSubSection id="09F591EB7577FFF2FF04A42C850A0D77" lastPageId="4" lastPageNumber="350" pageId="1" pageNumber="347" type="materials_examined">
<paragraph id="4150C2607577FFF7FF04A42C839A0DE3" blockId="1.[151,1437,837,2016]" pageId="1" pageNumber="347">
<materialsCitation id="F187C83D7577FFF7FF04A42C839A0DE3" ID-GBIF-Occurrence="2294326647" collectingDate="2017-07-02" collectionCode="MZUSP" collectorName="R. Devide &amp; Devide, G. S. C &amp; Silva, C &amp; B. F. Melo" country="Brazil" county="Lavandeira" latitude="-12.801389" location="Rio" longLatPrecision="21" longitude="-46.477222" municipality="Rio Palmas" pageId="1" pageNumber="347" specimenCode="MZUSP 124900" specimenCount="1" stateProvince="Tocantins" typeStatus="holotype">
<emphasis id="739B1E727577FFF7FF04A42C87080D00" bold="true" box="[151,267,838,864]" pageId="1" pageNumber="347">
<typeStatus id="9E547CC27577FFF7FF04A42C87050D00" box="[151,262,838,864]" pageId="1" pageNumber="347" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="11496A1B7577FFF7FE82A42F87C90D3F" box="[273,458,837,863]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="347">MZUSP 124900</specimenCode>
,
<quantity id="86176F857577FFF7FE47A42F84350D00" box="[468,566,837,864]" metricMagnitude="-2" metricUnit="m" metricValue="2.28" pageId="1" pageNumber="347" unit="mm" value="22.8">22.8 mm</quantity>
SL, female,
<collectingCountry id="39F882F07577FFF7FD51A42F850A0D3F" box="[706,777,837,863]" name="Brazil" pageId="1" pageNumber="347">Brazil</collectingCountry>
,
<collectingRegion id="832B0C827577FFF7FC80A42F85C30D3F" box="[787,960,837,863]" country="Brazil" name="Tocantins" pageId="1" pageNumber="347">Tocantins State</collectingRegion>
, municipality
<collectingCounty id="A831BAEC7577FFF7FBCEA42F82DD0D3F" box="[1117,1246,837,863]" pageId="1" pageNumber="347">Lavandeira</collectingCounty>
,
<collectingMunicipality id="A134581A7577FFF7FB7BA42F83690D3F" box="[1256,1386,837,863]" pageId="1" pageNumber="347">Rio Palmas</collectingMunicipality>
,
<location id="443094BB7577FFF7FAE7A42F839F0D3F" LSID="urn:lsid:plazi:treatment:C94673767577FFF2FF04A5B3850A0D77:443094BB7577FFF7FAE7A42F839F0D3F" box="[1396,1436,837,863]" country="Brazil" county="Lavandeira" latitude="-12.801389" longLatPrecision="21" longitude="-46.477222" municipality="Rio Palmas" name="Rio" pageId="1" pageNumber="347" stateProvince="Tocantins">Rio</location>
<collectingRegion id="832B0C827577FFF7FF04A40387000DE3" box="[151,259,873,899]" country="Brazil" name="Tocantins" pageId="1" pageNumber="347">Tocantins</collectingRegion>
basin;
<geoCoordinate id="24DBA4A77577FFF7FEC6A40387EB0DE3" box="[341,488,872,900]" degrees="12" direction="south" minutes="48" orientation="latitude" pageId="1" pageNumber="347" precision="15" seconds="05" value="-12.801389">
12
<superScript id="B69A6F287577FFF7FEE2A402877A0D16" attach="left" box="[369,377,872,886]" fontSize="6" pageId="1" pageNumber="347">o</superScript>
48’ 05” S
</geoCoordinate>
;
<geoCoordinate id="24DBA4A77577FFF7FE66A40384910DE4" box="[501,658,872,900]" degrees="46" direction="west" minutes="28" orientation="longitude" pageId="1" pageNumber="347" precision="15" seconds="38" value="-46.477222">
46
<superScript id="B69A6F287577FFF7FD82A402841A0D16" attach="left" box="[529,537,872,886]" fontSize="6" pageId="1" pageNumber="347">o</superScript>
28’ 38” W
</geoCoordinate>
; col.
<collectorName id="EC1AA7B67577FFF7FD41A40085450DE3" box="[722,838,873,900]" pageId="1" pageNumber="347">R. Devidé</collectorName>
, G. S. C.
<collectorName id="EC1AA7B67577FFF7FC21A403820C0DE3" box="[946,1039,873,899]" pageId="1" pageNumber="347">Silva, C</collectorName>
. Arraya &amp;
<collectorName id="EC1AA7B67577FFF7FB19A40083070DE3" box="[1162,1284,873,900]" pageId="1" pageNumber="347">B. F. Melo</collectorName>
,
<collectingDate id="25151D487577FFF7FA9CA40383950DE3" box="[1295,1430,873,899]" pageId="1" pageNumber="347" value="2017-07-02">
<date id="3551E4A07577FFF7FA9CA40383950DE3" box="[1295,1430,873,899]" pageId="1" pageNumber="347" value="2017-07-02">2 July 2017</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="4150C2607577FFF7FF54A4E485B50A1F" blockId="1.[151,1437,837,2016]" pageId="1" pageNumber="347">
<materialsCitation id="F187C83D7577FFF7FF54A4E482CF0DC7" ID-GBIF-Occurrence="2294326643" box="[199,1228,909,936]" collectionCode="LBP" pageId="1" pageNumber="347" specimenCount="2" typeStatus="paratype">
<emphasis id="739B1E727577FFF7FF54A4E487450DC8" bold="true" box="[199,326,910,936]" pageId="1" pageNumber="347">
<typeStatus id="9E547CC27577FFF7FF54A4E487420DC8" box="[199,321,910,936]" pageId="1" pageNumber="347" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="27FE5AA57577FFF7FEC2A4E487800DC8" box="[337,387,910,936]" pageId="1" pageNumber="347">LBP</collectionCode>
25358 (two specimens
<quantity id="86176F857577FFF7FD0DA4E785460DC8" box="[670,837,909,936]" metricMagnitude="-2" metricUnit="m" metricValue="1.425" metricValueMax="1.53" metricValueMin="1.3199999999999998" pageId="1" pageNumber="347" unit="mm" value="14.25" valueMax="15.3" valueMin="13.2">13.2–15.3 mm</quantity>
SL), collected with the
<typeStatus id="9E547CC27577FFF7FBF5A4E782CF0DC7" box="[1126,1228,909,935]" pageId="1" pageNumber="347" type="holotype">holotype</typeStatus>
</materialsCitation>
.
<materialsCitation id="F187C83D7577FFF7FB49A4E4828E0DAB" ID-GBIF-Occurrence="2294326644" collectionCode="LBP" pageId="1" pageNumber="347" specimenCount="1" typeStatus="holotype">
<collectionCode id="27FE5AA57577FFF7FB49A4E4830F0DC8" box="[1242,1292,910,936]" pageId="1" pageNumber="347">LBP</collectionCode>
26866 (two specimens
<quantity id="86176F857577FFF7FE81A4DB87B70DAC" box="[274,436,945,972]" metricMagnitude="-2" metricUnit="m" metricValue="2.23" metricValueMax="2.28" metricValueMin="2.18" pageId="1" pageNumber="347" unit="mm" value="22.3" valueMax="22.8" valueMin="21.8">21.8–22.8 mm</quantity>
SL, one c&amp;s specimen
<quantity id="86176F857577FFF7FD29A4DB851D0DAC" box="[698,798,945,972]" metricMagnitude="-2" metricUnit="m" metricValue="2.31" pageId="1" pageNumber="347" unit="mm" value="23.1">23.1 mm</quantity>
SL), collected with the
<typeStatus id="9E547CC27577FFF7FBB4A4DB828E0DAB" box="[1063,1165,945,971]" pageId="1" pageNumber="347" type="holotype">holotype</typeStatus>
</materialsCitation>
.
<materialsCitation id="F187C83D7577FFF7FB05A4D884F50D8F" ID-GBIF-Occurrence="2294326648" collectionCode="NUP" pageId="1" pageNumber="347" specimenCount="1" typeStatus="holotype">
<collectionCode id="27FE5AA57577FFF7FB05A4D882CE0DAC" box="[1174,1229,946,972]" pageId="1" pageNumber="347">NUP</collectionCode>
21298 (two specimens
<quantity id="86176F857577FFF7FF49A4BF877D0D90" box="[218,382,981,1008]" metricMagnitude="-2" metricUnit="m" metricValue="2.28" metricValueMax="2.35" metricValueMin="2.21" pageId="1" pageNumber="347" unit="mm" value="22.8" valueMax="23.5" valueMin="22.1">22.1–23.5 mm</quantity>
SL), collected with the
<typeStatus id="9E547CC27577FFF7FD03A4BF84F50D8F" box="[656,758,981,1007]" pageId="1" pageNumber="347" type="holotype">holotype</typeStatus>
</materialsCitation>
.
<materialsCitation id="F187C83D7577FFF7FC93A4BF84960A57" ID-GBIF-Occurrence="2294326645" collectingDate="1920-07-02" collectingDateMax="2002-11-24" collectingDateMin="1920-07-02" collectionCode="MZUSP" collectorName="C. R. Moreira &amp; J. C. Nolasco" country="Brazil" county="Lavandeira" latitude="-12.801389" location="Rio" longLatPrecision="21" longitude="-46.477222" municipality="Rio Palmas" pageId="1" pageNumber="347" specimenCode="MZUSP 84141" specimenCount="1" stateProvince="Tocantins" typeStatus="paratype">
<specimenCode id="11496A1B7577FFF7FC93A4BF85A80D8F" box="[768,939,981,1007]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="347">MZUSP 84141</specimenCode>
(two specimens
<collectingDate id="25151D487577FFF7FBF8A4BF82DF0D90" box="[1131,1244,981,1008]" pageId="1" pageNumber="347" value="1920-07-02" valueMax="1920-07-22" valueMin="1920-07-02">
<date id="3551E4A07577FFF7FBF8A4BF82DF0D90" box="[1131,1244,981,1008]" pageId="1" pageNumber="347" value="1920-07-02" valueMax="1920-07-22" valueMin="1920-07-02">
20 2–22
<quantity id="86176F857577FFF7FB5DA4BC82DF0D90" box="[1230,1244,982,1008]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="347" unit="mm" value="7.0">7</quantity>
</date>
</collectingDate>
mm SL),
<collectingCountry id="39F882F07577FFF7FAC2A4BF839B0D8F" box="[1361,1432,981,1007]" name="Brazil" pageId="1" pageNumber="347">Brazil</collectingCountry>
,
<collectingRegion id="832B0C827577FFF7FF04A49387460A73" box="[151,325,1017,1043]" country="Brazil" name="Tocantins" pageId="1" pageNumber="347">Tocantins State</collectingRegion>
, municipality
<collectingCounty id="A831BAEC7577FFF7FE77A49384660A73" box="[484,613,1017,1043]" pageId="1" pageNumber="347">Lavandeira</collectingCounty>
,
<collectingMunicipality id="A134581A7577FFF7FDE3A49384F00A73" box="[624,755,1017,1043]" pageId="1" pageNumber="347">Rio Palmas</collectingMunicipality>
,
<location id="443094BB7577FFF7FD6EA49385260A73" LSID="urn:lsid:plazi:treatment:C94673767577FFF2FF04A5B3850A0D77:443094BB7577FFF7FD6EA49385260A73" box="[765,805,1017,1043]" country="Brazil" county="Lavandeira" latitude="-12.801389" longLatPrecision="21" longitude="-46.477222" municipality="Rio Palmas" name="Rio" pageId="1" pageNumber="347" stateProvince="Tocantins">Rio</location>
<collectingRegion id="832B0C827577FFF7FCBFA493859B0A73" box="[812,920,1017,1043]" country="Brazil" name="Tocantins" pageId="1" pageNumber="347">Tocantins</collectingRegion>
basin;
<geoCoordinate id="24DBA4A77577FFF7FC7BA49382780A73" box="[1000,1147,1016,1044]" degrees="12" direction="south" minutes="48" orientation="latitude" pageId="1" pageNumber="347" precision="15" seconds="05" value="-12.801389">
12
<superScript id="B69A6F287577FFF7FB97A492820F0A66" attach="left" box="[1028,1036,1016,1030]" fontSize="6" pageId="1" pageNumber="347">o</superScript>
48’ 05” S
</geoCoordinate>
;
<geoCoordinate id="24DBA4A77577FFF7FB14A49383270A74" box="[1159,1316,1016,1044]" degrees="46" direction="west" minutes="28" orientation="longitude" pageId="1" pageNumber="347" precision="15" seconds="38" value="-46.477222">
46
<superScript id="B69A6F287577FFF7FB30A49282A80A66" attach="left" box="[1187,1195,1016,1030]" fontSize="6" pageId="1" pageNumber="347">o</superScript>
28’ 38” W
</geoCoordinate>
; col.
<collectorName id="EC1AA7B67577FFF7FAF1A49386F20A57" pageId="1" pageNumber="347">C. R. Moreira</collectorName>
&amp;
<collectorName id="EC1AA7B67577FFF7FE86A37487AD0A57" box="[277,430,1053,1080]" pageId="1" pageNumber="347">J. C. Nolasco</collectorName>
,
<collectingDate id="25151D487577FFF7FE2AA37784960A57" box="[441,661,1053,1079]" pageId="1" pageNumber="347" value="2002-11-24">
<date id="3551E4A07577FFF7FE2AA37784960A57" box="[441,661,1053,1079]" pageId="1" pageNumber="347" value="2002-11-24">24 November 2002</date>
</collectingDate>
</materialsCitation>
.
<materialsCitation id="F187C83D7577FFF7FD0CA37785B00A1F" ID-GBIF-Occurrence="2294326646" collectingDate="2002-11-14" collectionCode="MZUSP" collectorName="C. R. Moreira &amp; J. C. Nolasco" country="Brazil" county="Porto Alegre" latitude="-11.611389" location="Rio" longLatPrecision="21" longitude="-47.044167" municipality="Rio Manuel Alves da Natividade" pageId="1" pageNumber="347" specimenCode="MZUSP 84157" specimenCount="1" stateProvince="Tocantins" typeStatus="paratype">
<specimenCode id="11496A1B7577FFF7FD0CA377854B0A58" box="[671,840,1053,1080]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="347">MZUSP 84157</specimenCode>
(six specimens
<quantity id="86176F857577FFF7FC6FA377829C0A58" box="[1020,1183,1053,1080]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" metricValueMax="2.31" metricValueMin="1.95" pageId="1" pageNumber="347" unit="mm" value="21.3" valueMax="23.1" valueMin="19.5">19.5–23.1 mm</quantity>
SL),
<collectingCountry id="39F882F07577FFF7FB4DA37783260A57" box="[1246,1317,1053,1079]" name="Brazil" pageId="1" pageNumber="347">Brazil</collectingCountry>
,
<collectingRegion id="832B0C827577FFF7FAA3A37786D10A3B" country="Brazil" name="Tocantins" pageId="1" pageNumber="347">Tocantins State</collectingRegion>
, municipality
<collectingCounty id="A831BAEC7577FFF7FEE2A32887FD0A3B" box="[369,510,1089,1116]" pageId="1" pageNumber="347">Porto Alegre</collectingCounty>
do
<collectingRegion id="832B0C827577FFF7FDB4A32B84950A3B" box="[551,662,1089,1115]" country="Brazil" name="Tocantins" pageId="1" pageNumber="347">Tocantins</collectingRegion>
,
<collectingMunicipality id="A134581A7577FFF7FD32A32B82120A3B" box="[673,1041,1089,1115]" pageId="1" pageNumber="347">Rio Manuel Alves da Natividade</collectingMunicipality>
,
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<collectingRegion id="832B0C827577FFF7FBDAA32B82B60A3B" box="[1097,1205,1089,1115]" country="Brazil" name="Tocantins" pageId="1" pageNumber="347">Tocantins</collectingRegion>
basin;
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11
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36’ 41” S
</geoCoordinate>
;
<geoCoordinate id="24DBA4A77577FFF7FF04A30F87360AE0" box="[151,309,1124,1152]" degrees="47" direction="west" minutes="02" orientation="longitude" pageId="1" pageNumber="347" precision="15" seconds="39" value="-47.044167">
47
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02’ 39” W
</geoCoordinate>
; col.
<collectorName id="EC1AA7B67577FFF7FEE7A30F840C0A1F" box="[372,527,1125,1152]" pageId="1" pageNumber="347">C. R. Moreira</collectorName>
&amp;
<collectorName id="EC1AA7B67577FFF7FDA0A30C84CF0A1F" box="[563,716,1125,1152]" pageId="1" pageNumber="347">J. C. Nolasco</collectorName>
,
<collectingDate id="25151D487577FFF7FD44A30F85B00A1F" box="[727,947,1125,1151]" pageId="1" pageNumber="347" value="2002-11-14">
<date id="3551E4A07577FFF7FD44A30F85B00A1F" box="[727,947,1125,1151]" pageId="1" pageNumber="347" value="2002-11-14">14 November 2002</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
<paragraph id="4150C2607577FFF7FF54A3E3859F0BA4" blockId="1.[151,1437,837,2016]" pageId="1" pageNumber="347">
<emphasis id="739B1E727577FFF7FF54A3E387410AC4" bold="true" box="[199,322,1161,1188]" pageId="1" pageNumber="347">Diagnosis.</emphasis>
The new species
<taxonomicName id="86EFB9E37577FFF7FD9FA3E385210AC3" authorityName="Roxo &amp; Messias &amp; Silva" authorityYear="2019" box="[524,802,1161,1187]" class="Actinopterygii" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="347" phylum="Chordata" rank="species" species="pentakelis">
<emphasis id="739B1E727577FFF7FD9FA3E385210AC3" box="[524,802,1161,1187]" italics="true" pageId="1" pageNumber="347">Parotocinclus pentakelis</emphasis>
</taxonomicName>
can be distinguished from all congeners (except
<taxonomicName id="86EFB9E37577FFF7FADDA3E087E20AA8" authority="Roxo, Silva &amp; Oliveira, 2016" authorityName="Roxo, Silva &amp; Oliveira" authorityYear="2016" class="Actinopterygii" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="347" phylum="Chordata" rank="species" species="dani">
<emphasis id="739B1E727577FFF7FADDA3E0839F0AC3" box="[1358,1436,1161,1187]" italics="true" pageId="1" pageNumber="347">P. dani</emphasis>
<bibRefCitation id="257EBF917577FFF7FF04A3C487E20AA8" author="Roxo, F. F. &amp; Silva, G. S. C. &amp; Oliveira, C." box="[151,481,1197,1224]" pageId="1" pageNumber="347" pagination="125 - 136" refId="ref6769" refString="Roxo, F. F., Silva, G. S. C. &amp; Oliveira, C. (2016) Description of a new species of Parotocinclus (Siluriformes, Hypoptopomatinae) from the rio Tapajos basin. ZooKeys, 634, 125 - 136. https: // doi. org / 10.3897 / zookeys. 634.991 7" type="journal article" year="2016">Roxo, Silva &amp; Oliveira, 2016</bibRefCitation>
</taxonomicName>
) by a unique pattern of body coloration with five conspicuous dark bars,
<figureCitation id="D9D4DEE57577FFF7FA9EA3C7835A0AA7" box="[1293,1369,1197,1223]" captionStart="FIGURE 1" captionStartId="2.[151,250,1737,1762]" captionTargetBox="[151,1436,417,1718]" captionTargetId="figure@2.[151,1436,417,1718]" captionTargetPageId="2" captionText="FIGURE 1. Parotocinclus pentakelis, MZUSP 124900, holotype, 22.8 mm SL, rio Palmas, rio Tocantins basin." httpUri="https://zenodo.org/record/3349499/files/figure.png" pageId="1" pageNumber="347">Figs. 1</figureCitation>
,
<figureCitation id="D9D4DEE57577FFF7FAF5A3C783750AA7" box="[1382,1398,1197,1223]" captionStart="FIGURE 2" captionStartId="4.[151,250,1570,1595]" captionTargetBox="[151,1436,813,1551]" captionTargetId="figure@4.[151,1436,813,1551]" captionTargetPageId="4" captionText="FIGURE 2. Parotocinclus pentakelis, live specimen, MZUSP 124900, holotype, 22.8 mm SL." httpUri="https://zenodo.org/record/3349501/files/figure.png" pageId="1" pageNumber="347">2</figureCitation>
,
<figureCitation id="D9D4DEE57577FFF7FA11A3C7839F0AA7" box="[1410,1436,1197,1223]" captionStart="FIGURE 3" captionStartId="5.[151,250,1898,1923]" captionTargetBox="[186,1400,345,1879]" captionTargetId="figure@5.[186,1400,345,1879]" captionTargetPageId="5" captionText="FIGURE 3. Body color pattern variation between a) Parotocinclus pentakelis, paratypes, NUP 21298, 22.8 mm SL. b) P. dani, holotype, MZUSP 120737, holotype, 27.3 mm SL; c) Parotocinclus variola, ICNHHN 18685, 25.9 mm SL; d) Curculionichthys tukana, holotype, MZUSP 123010, 23.4 mm SL; e) Hisonotus alberti, holotype, MZUSP 118845, 33.5 mm SL.Arrows indicate transverse dark bars." httpUri="https://zenodo.org/record/3349503/files/figure.png" pageId="1" pageNumber="347">3a</figureCitation>
(
<emphasis id="739B1E727577FFF7FF32A3B886BC0A8B" box="[161,191,1234,1259]" italics="true" pageId="1" pageNumber="347">vs.</emphasis>
four or three in all remaining species of
<taxonomicName id="86EFB9E37577FFF7FD17A3BB85230A8B" authorityName="Eigenmann &amp; Eigenmann" authorityYear="1889" box="[644,800,1233,1259]" class="Actinopterygii" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="347" phylum="Chordata" rank="genus">
<emphasis id="739B1E727577FFF7FD17A3BB85230A8B" box="[644,800,1233,1259]" italics="true" pageId="1" pageNumber="347">Parotocinclus</emphasis>
</taxonomicName>
) and by absence of an adipose fin (
<emphasis id="739B1E727577FFF7FB34A3B882C60A8B" box="[1191,1221,1234,1259]" italics="true" pageId="1" pageNumber="347">vs.</emphasis>
presence of an adipose fin). Additionally, the new species can be distinguished from
<taxonomicName id="86EFB9E37577FFF7FCE0A39C85BC0B6F" authorityName="Roxo, Silva &amp; Oliveira" authorityYear="2016" box="[883,959,1269,1295]" class="Actinopterygii" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="347" phylum="Chordata" rank="species" species="dani">
<emphasis id="739B1E727577FFF7FCE0A39C85BC0B6F" box="[883,959,1269,1295]" italics="true" pageId="1" pageNumber="347">P. dani</emphasis>
</taxonomicName>
by absence of a small platelet at the typical adipose-fin location (
<emphasis id="739B1E727577FFF7FE14A27087A60B53" box="[391,421,1306,1331]" italics="true" pageId="1" pageNumber="347">vs.</emphasis>
presence of a small platelet at typical adipose-fin region), by absence of conspicuous dark dots broadly distributed on the body (
<emphasis id="739B1E727577FFF7FDD3A254845C0B37" box="[576,607,1342,1367]" italics="true" pageId="1" pageNumber="347">vs.</emphasis>
absence of this pigmentation pattern) and by absence of a triangular dark blotch at the anterior base of the dorsal fin (
<emphasis id="739B1E727577FFF7FD15A20884A70B1B" box="[646,676,1378,1403]" italics="true" pageId="1" pageNumber="347">vs.</emphasis>
presence of a triangular dark blotch). Additionally, the new species differs from all congeners from the coastal drainages of the Brazilian shield from south and northeastern
<collectingCountry id="39F882F07577FFF7FAA7A2EF837B0BFF" box="[1332,1400,1413,1439]" name="Brazil" pageId="1" pageNumber="347">Brazil</collectingCountry>
by presence of a
<emphasis id="739B1E727577FFF7FEA1A2C087400BA3" box="[306,323,1450,1475]" italics="true" pageId="1" pageNumber="347">V</emphasis>
-shaped spinelet (
<emphasis id="739B1E727577FFF7FD9BA2C084240BA3" box="[520,551,1450,1475]" italics="true" pageId="1" pageNumber="347">vs.</emphasis>
rounded or rectangular spinelet).
</paragraph>
<paragraph id="4150C2607577FFF7FF54A2A783890967" blockId="1.[151,1437,837,2016]" pageId="1" pageNumber="347">
<emphasis id="739B1E727577FFF7FF54A2A7875A0B88" bold="true" box="[199,345,1485,1512]" pageId="1" pageNumber="347">Description.</emphasis>
Small-sized loricariid,
<typeStatus id="9E547CC27577FFF7FDF2A2A784C00B87" box="[609,707,1485,1511]" pageId="1" pageNumber="347" type="holotype">holotype</typeStatus>
<quantity id="86176F857577FFF7FD59A2A7852C0B88" box="[714,815,1485,1512]" metricMagnitude="-2" metricUnit="m" metricValue="2.28" pageId="1" pageNumber="347" unit="mm" value="22.8">22.8 mm</quantity>
SL,
<typeStatus id="9E547CC27577FFF7FCF6A2A485D30B88" box="[869,976,1486,1512]" pageId="1" pageNumber="347" type="paratype">paratypes</typeStatus>
<quantity id="86176F857577FFF7FC4BA2A782780B88" box="[984,1147,1485,1512]" metricMagnitude="-2" metricUnit="m" metricValue="1.835" metricValueMax="2.35" metricValueMin="1.3199999999999998" pageId="1" pageNumber="347" unit="mm" value="18.35" valueMax="23.5" valueMin="13.2">13.2–23.5 mm</quantity>
SL. In dorsal view, snout tip rounded, elongated (
<quantity id="86176F857577FFF7FE33A29B8446086C" box="[416,581,1521,1548]" metricMagnitude="-2" metricUnit="m" metricValue="5.24" metricValueMax="5.52" metricValueMin="4.96" pageId="1" pageNumber="347" unit="mm" value="52.4" valueMax="55.2" valueMin="49.6">49.6–55.2 mm</quantity>
HL). Snout margin completely covered by odontodes, larger than remaining ones on head. Odontodes on head and trunk well defined, and arranged in longitudinal rows, not forming parallel series. Eye median-sized and rounded, situated dorsolaterally at midpoint length of head. No ridge between eyes and nares. Supraoccipital process not elevated and without tuft of odontodes in specimens of all sizes. Nasal aperture small. Mouth wide, ventrally-oriented; oral disk rounded with papillae randomly distributed. Lower lip larger than upper, but distant from cleithrum; its border fringed. Maxillary barbel short, slender and free distally. Ventral surface of head naked in area around mouth and between lower lip and cleithrum. Buccal papilla absent. Teeth slender and bicuspid, cusps asymmetrical, main cusp larger than lateral cusp. Premaxillary teeth 15–21. Dentary teeth 14–18.
</paragraph>
<paragraph id="4150C2607577FFF7FF54A078842C09DC" blockId="1.[151,1437,837,2016]" pageId="1" pageNumber="347">In lateral view, dorsal profile of head straight from snout tip to interorbital region; slightly convex from this point to posterior margin of parieto-supraoccipital; slightly concave from this point to dorsal fin-insertion; concave and descending from this point to origin of anteriormost caudal fin procurrent ray. Ventral profile of head concave from snout tip to opercular region; straight from this point to anal-fin insertion; concave from this point to first anteriormost caudal-fin procurrent ray.</paragraph>
<paragraph id="4150C2607577FFF4FF54A0AF86E30FEB" blockId="1.[151,1437,837,2016]" lastBlockId="2.[151,1437,153,395]" lastPageId="2" lastPageNumber="348" pageId="1" pageNumber="347">
Greatest body depth at dorsal-fin origin. Greatest body width at cleithral region (
<quantity id="86176F857577FFF7FBCAA0AF82FC0980" box="[1113,1279,1989,2016]" metricMagnitude="-2" metricUnit="m" metricValue="2.52" metricValueMax="2.62" metricValueMin="2.42" pageId="1" pageNumber="347" unit="mm" value="25.2" valueMax="26.2" valueMin="24.2">24.2–26.2 mm</quantity>
SL), progres- sively narrowing towards snout and caudal fin. Trunk and caudal peduncle almost ellipsoid in cross section. Trunk covered by bony plates, except on area around fin bases and anal-fin insertion. Mid-dorsal series poorly developed reaching vertical line at middle of dorsal fin. Lateral plate series with 21–23 plates; lateral line incomplete, with a gap of two plates along mid-length of body. Mid-ventral series short reaching vertical line at adipose-fin region. Arrector fossae partially enclosed by ventral lamina of coracoid. Abdominal region entirely covered by plates; lat- eral abdominal plate series with three to four plates; 1-3 irregular and large plates on median series; five pre-anal plates.
</paragraph>
<caption id="159092E87574FFF4FF04A1A383010882" box="[151,1282,1737,1762]" httpUri="https://zenodo.org/record/3349499/files/figure.png" pageId="2" pageNumber="348" startId="2.[151,250,1737,1762]" targetBox="[151,1436,417,1718]" targetPageId="2">
<paragraph id="4150C2607574FFF4FF04A1A383010882" blockId="2.[151,1282,1737,1762]" box="[151,1282,1737,1762]" pageId="2" pageNumber="348">
<emphasis id="739B1E727574FFF4FF04A1A3870E0882" bold="true" box="[151,269,1737,1762]" pageId="2" pageNumber="348">FIGURE 1</emphasis>
.
<taxonomicName id="86EFB9E37574FFF4FE8AA1A384170881" authorityName="Roxo &amp; Messias &amp; Silva" authorityYear="2019" box="[281,532,1737,1761]" class="Actinopterygii" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="348" phylum="Chordata" rank="species" species="pentakelis">
<emphasis id="739B1E727574FFF4FE8AA1A384170881" box="[281,532,1737,1761]" italics="true" pageId="2" pageNumber="348">Parotocinclus pentakelis</emphasis>
</taxonomicName>
, MZUSP 124900, holotype, 22.8 mm SL, rio Palmas, rio Tocantins basin.
</paragraph>
</caption>
<paragraph id="4150C2607574FFF5FF54A07A87A90EB7" blockId="2.[151,1437,1808,2014]" lastBlockId="3.[151,1437,152,539]" lastPageId="3" lastPageNumber="349" pageId="2" pageNumber="348">
Dorsal-fin rays II,7; dorsal-fin spinelet
<emphasis id="739B1E727574FFF4FDEAA07B8489094A" box="[633,650,1809,1834]" italics="true" pageId="2" pageNumber="348">V</emphasis>
-shaped; second dorsal-fin unbranched ray reaching half of anal-fin unbranched ray when depressed; dorsal-fin locking mechanism functional. Second dorsal-fin unbranched ray covered by odontodes. Pectoral-fin rays I,6; reaching more than two thirds of pelvic-fin unbranched ray when depressed; dorsal surface of pectoral spine covered by odontodes, larger on distal portion. Pectoral-fin axillary slit present and well visible. Pelvic-fin rays i,5; its posterior tip almost reaching anal-fin insertion when depressed in females and extending beyond anal-fin insertion when depressed in males; pelvic-fin spine covered by odontodes, on ventral surface. Anal-fin rays i,5; anal-fin spine covered by odontodes. Caudal-fin rays i,14,i; caudal fin forked, all rays covered with odontodes.
</paragraph>
<paragraph id="4150C2607575FFF5FF54A78B87FC0C7B" blockId="3.[151,1437,152,539]" pageId="3" pageNumber="349">Snout tip formed by a single rostral rectangular-shaped plate in dorsal view. Prenasals plates positioned posteriorly to rostral plate and formed by two large and two small triangular-shaped plates located between nares, and one small, rectangular-shaped plate. Nasal almost rectangular forming anterioromedial nostril margin, contacting frontals. Dorsal surface of head at its posterior portion composed by compound pterotic laterally, parieto-supraoccipital and frontal medially, and sphenotic. Compound pterotic on lateral portion. Postrostral plates rectangular-shaped. Infraorbital series incomplete, with only four bones, present just above postrostral series, all covered by laterosensory canal system. Preopercle elongated and rectangular, covered by laterosensory canal and situated ventral to infraorbital 3 and infraorbital 4, and upper to compound pterotic. Subocular cheek plates and opercle forming posterior lateral margin of head.</paragraph>
<caption id="159092E87575FFF5FF04A52084B40CE7" ID-Table-UUID="159092E87575FFF5FF04A52084B40CE7" httpUri="http://table.plazi.org/id/159092E87575FFF5FF04A52084B40CE7" pageId="3" pageNumber="349" targetBox="[166,1390,658,1779]" targetIsTable="true" targetPageId="3">
<paragraph id="4150C2607575FFF5FF04A52084B40CE7" blockId="3.[151,1437,586,647]" pageId="3" pageNumber="349">
<emphasis id="739B1E727575FFF5FF04A52087090C02" bold="true" box="[151,266,586,610]" pageId="3" pageNumber="349">TABLE 1.</emphasis>
Morphometrics and meristics of
<taxonomicName id="86EFB9E37575FFF5FDFFA52185700C03" authorityName="Roxo &amp; Messias &amp; Silva" authorityYear="2019" box="[620,883,587,611]" class="Actinopterygii" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="349" phylum="Chordata" rank="species" species="pentakelis">
<emphasis id="739B1E727575FFF5FDFFA52185700C03" box="[620,883,587,611]" italics="true" pageId="3" pageNumber="349">Parotocinclus pentakelis</emphasis>
</taxonomicName>
. SD = standard deviation. SL = standard length. HL = head length. N = number of analyzed specimens.
</paragraph>
</caption>
<paragraph id="4150C2607575FFF5FD60A5F883530892" pageId="3" pageNumber="349">
<table id="33EF30C075750009FF35A5F8836D0893" box="[166,1390,658,1779]" gridcols="7" gridrows="30" pageId="3" pageNumber="349">
<tr id="FFDFC02275750009FF35A5F8836D0CCB" box="[166,1390,658,683]" gridrow="0" pageId="3" pageNumber="349" rowspan-0="1">
<th id="BC0EA95E75750009FD60A5F885520CCB" box="[755,849,658,683]" gridcol="1" gridrow="0" pageId="3" pageNumber="349">Holotype</th>
<th id="BC0EA95E75750009FC30A5F885BF0CCB" box="[931,956,658,683]" gridcol="2" gridrow="0" pageId="3" pageNumber="349">N</th>
<th id="BC0EA95E75750009FC7EA5F882180CCB" box="[1005,1051,658,683]" gridcol="3" gridrow="0" pageId="3" pageNumber="349">Low</th>
<th id="BC0EA95E75750009FBC5A5F8828A0CCB" box="[1110,1161,658,683]" gridcol="4" gridrow="0" pageId="3" pageNumber="349">High</th>
<th id="BC0EA95E75750009FB55A5F883020CCB" box="[1222,1281,658,683]" gridcol="5" gridrow="0" pageId="3" pageNumber="349">Mean</th>
<th id="BC0EA95E75750009FAD1A5F8836D0CCB" box="[1346,1390,658,683]" gridcol="6" gridrow="0" pageId="3" pageNumber="349">SD</th>
</tr>
<tr id="FFDFC02275750009FF35A5D2836D0CB1" box="[166,1390,696,721]" gridrow="1" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A5D287F10CB1" box="[166,498,696,721]" gridcol="0" gridrow="1" pageId="3" pageNumber="349">SL (mm)</th>
<td id="BC0EA95E75750009FD60A5D285520CB1" box="[755,849,696,721]" gridcol="1" gridrow="1" pageId="3" pageNumber="349">22.8</td>
<td id="BC0EA95E75750009FC30A5D285BF0CB1" box="[931,956,696,721]" gridcol="2" gridrow="1" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA5D282180CB1" box="[1005,1051,696,721]" gridcol="3" gridrow="1" pageId="3" pageNumber="349">19.5</td>
<td id="BC0EA95E75750009FBC5A5D2828A0CB1" box="[1110,1161,696,721]" gridcol="4" gridrow="1" pageId="3" pageNumber="349">23.5</td>
<td id="BC0EA95E75750009FB55A5D283020CB1" box="[1222,1281,696,721]" gridcol="5" gridrow="1" pageId="3" pageNumber="349">21.8</td>
<td id="BC0EA95E75750009FAD1A5D2836D0CB1" box="[1346,1390,696,721]" gridcol="6" gridrow="1" pageId="3" pageNumber="349">1.32</td>
</tr>
<tr id="FFDFC02275750009FF35A5B4836D0C97" box="[166,1390,734,759]" gridrow="2" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A5B4836D0C97" box="[166,1390,734,759]" colspan="7" colspanRight="6" gridcol="0" gridrow="2" pageId="3" pageNumber="349">Percent of SL</th>
</tr>
<tr id="FFDFC02275750009FF35A46E836D0D7D" box="[166,1390,772,797]" gridrow="3" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A46E87F10D7D" box="[166,498,772,797]" gridcol="0" gridrow="3" pageId="3" pageNumber="349">Head length</th>
<td id="BC0EA95E75750009FD60A46E85520D7D" box="[755,849,772,797]" gridcol="1" gridrow="3" pageId="3" pageNumber="349">37.1</td>
<td id="BC0EA95E75750009FC30A46E85BF0D7D" box="[931,956,772,797]" gridcol="2" gridrow="3" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA46E82180D7D" box="[1005,1051,772,797]" gridcol="3" gridrow="3" pageId="3" pageNumber="349">35.4</td>
<td id="BC0EA95E75750009FBC5A46E828A0D7D" box="[1110,1161,772,797]" gridcol="4" gridrow="3" pageId="3" pageNumber="349">40.1</td>
<td id="BC0EA95E75750009FB55A46E83020D7D" box="[1222,1281,772,797]" gridcol="5" gridrow="3" pageId="3" pageNumber="349">37.4</td>
<td id="BC0EA95E75750009FAD1A46E836D0D7D" box="[1346,1390,772,797]" gridcol="6" gridrow="3" pageId="3" pageNumber="349">1.16</td>
</tr>
<tr id="FFDFC02275750009FF35A443836D0D22" box="[166,1390,809,834]" gridrow="4" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A44387F10D22" box="[166,498,809,834]" gridcol="0" gridrow="4" pageId="3" pageNumber="349">Predorsal length</th>
<td id="BC0EA95E75750009FD60A44385520D22" box="[755,849,809,834]" gridcol="1" gridrow="4" pageId="3" pageNumber="349">48.6</td>
<td id="BC0EA95E75750009FC30A44385BF0D22" box="[931,956,809,834]" gridcol="2" gridrow="4" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA44382180D22" box="[1005,1051,809,834]" gridcol="3" gridrow="4" pageId="3" pageNumber="349">45.2</td>
<td id="BC0EA95E75750009FBC5A443828A0D22" box="[1110,1161,809,834]" gridcol="4" gridrow="4" pageId="3" pageNumber="349">49.9</td>
<td id="BC0EA95E75750009FB55A44383020D22" box="[1222,1281,809,834]" gridcol="5" gridrow="4" pageId="3" pageNumber="349">47.4</td>
<td id="BC0EA95E75750009FAD1A443836D0D22" box="[1346,1390,809,834]" gridcol="6" gridrow="4" pageId="3" pageNumber="349">1.39</td>
</tr>
<tr id="FFDFC02275750009FF35A425836D0D08" box="[166,1390,847,872]" gridrow="5" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A42587F10D08" box="[166,498,847,872]" gridcol="0" gridrow="5" pageId="3" pageNumber="349">Dorsal-fin spine length</th>
<td id="BC0EA95E75750009FD60A42585520D08" box="[755,849,847,872]" gridcol="1" gridrow="5" pageId="3" pageNumber="349">28.9</td>
<td id="BC0EA95E75750009FC30A42585BF0D08" box="[931,956,847,872]" gridcol="2" gridrow="5" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA42582180D08" box="[1005,1051,847,872]" gridcol="3" gridrow="5" pageId="3" pageNumber="349">26.2</td>
<td id="BC0EA95E75750009FBC5A425828A0D08" box="[1110,1161,847,872]" gridcol="4" gridrow="5" pageId="3" pageNumber="349">29.6</td>
<td id="BC0EA95E75750009FB55A42583020D08" box="[1222,1281,847,872]" gridcol="5" gridrow="5" pageId="3" pageNumber="349">27.6</td>
<td id="BC0EA95E75750009FAD1A425836D0D08" box="[1346,1390,847,872]" gridcol="6" gridrow="5" pageId="3" pageNumber="349">0.95</td>
</tr>
<tr id="FFDFC02275750009FF35A41F836D0DEE" box="[166,1390,885,910]" gridrow="6" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A41F87F10DEE" box="[166,498,885,910]" gridcol="0" gridrow="6" pageId="3" pageNumber="349">Anal-fin unbranched ray length</th>
<td id="BC0EA95E75750009FD60A41F85520DEE" box="[755,849,885,910]" gridcol="1" gridrow="6" pageId="3" pageNumber="349">18.4</td>
<td id="BC0EA95E75750009FC30A41F85BF0DEE" box="[931,956,885,910]" gridcol="2" gridrow="6" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA41F82180DEE" box="[1005,1051,885,910]" gridcol="3" gridrow="6" pageId="3" pageNumber="349">17.3</td>
<td id="BC0EA95E75750009FBC5A41F828A0DEE" box="[1110,1161,885,910]" gridcol="4" gridrow="6" pageId="3" pageNumber="349">20.5</td>
<td id="BC0EA95E75750009FB55A41F83020DEE" box="[1222,1281,885,910]" gridcol="5" gridrow="6" pageId="3" pageNumber="349">18.7</td>
<td id="BC0EA95E75750009FAD1A41F836D0DEE" box="[1346,1390,885,910]" gridcol="6" gridrow="6" pageId="3" pageNumber="349">1.01</td>
</tr>
<tr id="FFDFC02275750009FF35A4F1836D0DD4" box="[166,1390,923,948]" gridrow="7" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A4F187F10DD4" box="[166,498,923,948]" gridcol="0" gridrow="7" pageId="3" pageNumber="349">Pectoral-fin spine length</th>
<td id="BC0EA95E75750009FD60A4F185520DD4" box="[755,849,923,948]" gridcol="1" gridrow="7" pageId="3" pageNumber="349">30.9</td>
<td id="BC0EA95E75750009FC30A4F185BF0DD4" box="[931,956,923,948]" gridcol="2" gridrow="7" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA4F182180DD4" box="[1005,1051,923,948]" gridcol="3" gridrow="7" pageId="3" pageNumber="349">25.6</td>
<td id="BC0EA95E75750009FBC5A4F1828A0DD4" box="[1110,1161,923,948]" gridcol="4" gridrow="7" pageId="3" pageNumber="349">33.0</td>
<td id="BC0EA95E75750009FB55A4F183020DD4" box="[1222,1281,923,948]" gridcol="5" gridrow="7" pageId="3" pageNumber="349">30.1</td>
<td id="BC0EA95E75750009FAD1A4F1836D0DD4" box="[1346,1390,923,948]" gridcol="6" gridrow="7" pageId="3" pageNumber="349">1.86</td>
</tr>
<tr id="FFDFC02275750009FF35A4AB836D0DBA" box="[166,1390,961,986]" gridrow="8" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A4AB87F10DBA" box="[166,498,961,986]" gridcol="0" gridrow="8" pageId="3" pageNumber="349">Pelvic-fin unbranched ray length</th>
<td id="BC0EA95E75750009FD60A4AB85520DBA" box="[755,849,961,986]" gridcol="1" gridrow="8" pageId="3" pageNumber="349">19.2</td>
<td id="BC0EA95E75750009FC30A4AB85BF0DBA" box="[931,956,961,986]" gridcol="2" gridrow="8" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA4AB82180DBA" box="[1005,1051,961,986]" gridcol="3" gridrow="8" pageId="3" pageNumber="349">17.7</td>
<td id="BC0EA95E75750009FBC5A4AB828A0DBA" box="[1110,1161,961,986]" gridcol="4" gridrow="8" pageId="3" pageNumber="349">23.5</td>
<td id="BC0EA95E75750009FB55A4AB83020DBA" box="[1222,1281,961,986]" gridcol="5" gridrow="8" pageId="3" pageNumber="349">20.2</td>
<td id="BC0EA95E75750009FAD1A4AB836D0DBA" box="[1346,1390,961,986]" gridcol="6" gridrow="8" pageId="3" pageNumber="349">1.76</td>
</tr>
<tr id="FFDFC02275750009FF35A48C836D0D9F" box="[166,1390,998,1023]" gridrow="9" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A48C87F10D9F" box="[166,498,998,1023]" gridcol="0" gridrow="9" pageId="3" pageNumber="349">Cleithral width</th>
<td id="BC0EA95E75750009FD60A48C85520D9F" box="[755,849,998,1023]" gridcol="1" gridrow="9" pageId="3" pageNumber="349">25.4</td>
<td id="BC0EA95E75750009FC30A48C85BF0D9F" box="[931,956,998,1023]" gridcol="2" gridrow="9" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA48C82180D9F" box="[1005,1051,998,1023]" gridcol="3" gridrow="9" pageId="3" pageNumber="349">24.2</td>
<td id="BC0EA95E75750009FBC5A48C828A0D9F" box="[1110,1161,998,1023]" gridcol="4" gridrow="9" pageId="3" pageNumber="349">26.2</td>
<td id="BC0EA95E75750009FB55A48C83020D9F" box="[1222,1281,998,1023]" gridcol="5" gridrow="9" pageId="3" pageNumber="349">25.3</td>
<td id="BC0EA95E75750009FAD1A48C836D0D9F" box="[1346,1390,998,1023]" gridcol="6" gridrow="9" pageId="3" pageNumber="349">0.60</td>
</tr>
<tr id="FFDFC02275750009FF35A366836D0A45" box="[166,1390,1036,1061]" gridrow="10" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A36687F10A45" box="[166,498,1036,1061]" gridcol="0" gridrow="10" pageId="3" pageNumber="349">Thorax length</th>
<td id="BC0EA95E75750009FD60A36685520A45" box="[755,849,1036,1061]" gridcol="1" gridrow="10" pageId="3" pageNumber="349">15.8</td>
<td id="BC0EA95E75750009FC30A36685BF0A45" box="[931,956,1036,1061]" gridcol="2" gridrow="10" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA36682180A45" box="[1005,1051,1036,1061]" gridcol="3" gridrow="10" pageId="3" pageNumber="349">13.6</td>
<td id="BC0EA95E75750009FBC5A366828A0A45" box="[1110,1161,1036,1061]" gridcol="4" gridrow="10" pageId="3" pageNumber="349">16.2</td>
<td id="BC0EA95E75750009FB55A36683020A45" box="[1222,1281,1036,1061]" gridcol="5" gridrow="10" pageId="3" pageNumber="349">15.0</td>
<td id="BC0EA95E75750009FAD1A366836D0A45" box="[1346,1390,1036,1061]" gridcol="6" gridrow="10" pageId="3" pageNumber="349">0.92</td>
</tr>
<tr id="FFDFC02275750009FF35A358836D0A2B" box="[166,1390,1074,1099]" gridrow="11" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A35887F10A2B" box="[166,498,1074,1099]" gridcol="0" gridrow="11" pageId="3" pageNumber="349">Abdominal length</th>
<td id="BC0EA95E75750009FD60A35885520A2B" box="[755,849,1074,1099]" gridcol="1" gridrow="11" pageId="3" pageNumber="349">21.9</td>
<td id="BC0EA95E75750009FC30A35885BF0A2B" box="[931,956,1074,1099]" gridcol="2" gridrow="11" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA35882180A2B" box="[1005,1051,1074,1099]" gridcol="3" gridrow="11" pageId="3" pageNumber="349">18.5</td>
<td id="BC0EA95E75750009FBC5A358828A0A2B" box="[1110,1161,1074,1099]" gridcol="4" gridrow="11" pageId="3" pageNumber="349">23.3</td>
<td id="BC0EA95E75750009FB55A35883020A2B" box="[1222,1281,1074,1099]" gridcol="5" gridrow="11" pageId="3" pageNumber="349">21.4</td>
<td id="BC0EA95E75750009FAD1A358836D0A2B" box="[1346,1390,1074,1099]" gridcol="6" gridrow="11" pageId="3" pageNumber="349">1.53</td>
</tr>
<tr id="FFDFC02275750009FF35A332836D0A11" box="[166,1390,1112,1137]" gridrow="12" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A33287F10A11" box="[166,498,1112,1137]" gridcol="0" gridrow="12" pageId="3" pageNumber="349">Caudal peduncle length</th>
<td id="BC0EA95E75750009FD60A33285520A11" box="[755,849,1112,1137]" gridcol="1" gridrow="12" pageId="3" pageNumber="349">27.0</td>
<td id="BC0EA95E75750009FC30A33285BF0A11" box="[931,956,1112,1137]" gridcol="2" gridrow="12" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA33282180A11" box="[1005,1051,1112,1137]" gridcol="3" gridrow="12" pageId="3" pageNumber="349">26.7</td>
<td id="BC0EA95E75750009FBC5A332828A0A11" box="[1110,1161,1112,1137]" gridcol="4" gridrow="12" pageId="3" pageNumber="349">31.0</td>
<td id="BC0EA95E75750009FB55A33283020A11" box="[1222,1281,1112,1137]" gridcol="5" gridrow="12" pageId="3" pageNumber="349">28.5</td>
<td id="BC0EA95E75750009FAD1A332836D0A11" box="[1346,1390,1112,1137]" gridcol="6" gridrow="12" pageId="3" pageNumber="349">1.31</td>
</tr>
<tr id="FFDFC02275750009FF35A314836D0AF7" box="[166,1390,1150,1175]" gridrow="13" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A31487F10AF7" box="[166,498,1150,1175]" gridcol="0" gridrow="13" pageId="3" pageNumber="349">Caudal peduncle depth</th>
<td id="BC0EA95E75750009FD60A31485520AF7" box="[755,849,1150,1175]" gridcol="1" gridrow="13" pageId="3" pageNumber="349">10.0</td>
<td id="BC0EA95E75750009FC30A31485BF0AF7" box="[931,956,1150,1175]" gridcol="2" gridrow="13" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA31482180AF7" box="[1005,1051,1150,1175]" gridcol="3" gridrow="13" pageId="3" pageNumber="349">9.7</td>
<td id="BC0EA95E75750009FBC5A314828A0AF7" box="[1110,1161,1150,1175]" gridcol="4" gridrow="13" pageId="3" pageNumber="349">11.6</td>
<td id="BC0EA95E75750009FB55A31483020AF7" box="[1222,1281,1150,1175]" gridcol="5" gridrow="13" pageId="3" pageNumber="349">10.4</td>
<td id="BC0EA95E75750009FAD1A314836D0AF7" box="[1346,1390,1150,1175]" gridcol="6" gridrow="13" pageId="3" pageNumber="349">0.49</td>
</tr>
<tr id="FFDFC02275750009FF35A3C9836D0ADC" box="[166,1390,1187,1212]" gridrow="14" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A3C987F10ADC" box="[166,498,1187,1212]" gridcol="0" gridrow="14" pageId="3" pageNumber="349">Preanal length</th>
<td id="BC0EA95E75750009FD60A3C985520ADC" box="[755,849,1187,1212]" gridcol="1" gridrow="14" pageId="3" pageNumber="349">64.0</td>
<td id="BC0EA95E75750009FC30A3C985BF0ADC" box="[931,956,1187,1212]" gridcol="2" gridrow="14" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA3C982180ADC" box="[1005,1051,1187,1212]" gridcol="3" gridrow="14" pageId="3" pageNumber="349">62.0</td>
<td id="BC0EA95E75750009FBC5A3C9828A0ADC" box="[1110,1161,1187,1212]" gridcol="4" gridrow="14" pageId="3" pageNumber="349">68.3</td>
<td id="BC0EA95E75750009FB55A3C983020ADC" box="[1222,1281,1187,1212]" gridcol="5" gridrow="14" pageId="3" pageNumber="349">64.2</td>
<td id="BC0EA95E75750009FAD1A3C9836D0ADC" box="[1346,1390,1187,1212]" gridcol="6" gridrow="14" pageId="3" pageNumber="349">1.67</td>
</tr>
<tr id="FFDFC02275750009FF35A3A3836D0A82" box="[166,1390,1225,1250]" gridrow="15" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A3A387F10A82" box="[166,498,1225,1250]" gridcol="0" gridrow="15" pageId="3" pageNumber="349">Base of dorsal-fin length</th>
<td id="BC0EA95E75750009FD60A3A385520A82" box="[755,849,1225,1250]" gridcol="1" gridrow="15" pageId="3" pageNumber="349">15.2</td>
<td id="BC0EA95E75750009FC30A3A385BF0A82" box="[931,956,1225,1250]" gridcol="2" gridrow="15" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA3A382180A82" box="[1005,1051,1225,1250]" gridcol="3" gridrow="15" pageId="3" pageNumber="349">11.0</td>
<td id="BC0EA95E75750009FBC5A3A3828A0A82" box="[1110,1161,1225,1250]" gridcol="4" gridrow="15" pageId="3" pageNumber="349">15.5</td>
<td id="BC0EA95E75750009FB55A3A383020A82" box="[1222,1281,1225,1250]" gridcol="5" gridrow="15" pageId="3" pageNumber="349">13.4</td>
<td id="BC0EA95E75750009FAD1A3A3836D0A82" box="[1346,1390,1225,1250]" gridcol="6" gridrow="15" pageId="3" pageNumber="349">1.59</td>
</tr>
<tr id="FFDFC02275750009FF35A385836D0B68" box="[166,1390,1263,1288]" gridrow="16" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A38587F10B68" box="[166,498,1263,1288]" gridcol="0" gridrow="16" pageId="3" pageNumber="349">Lower caudal spine length</th>
<td id="BC0EA95E75750009FD60A38585520B68" box="[755,849,1263,1288]" gridcol="1" gridrow="16" pageId="3" pageNumber="349">30.2</td>
<td id="BC0EA95E75750009FC30A38585BF0B68" box="[931,956,1263,1288]" gridcol="2" gridrow="16" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA38582180B68" box="[1005,1051,1263,1288]" gridcol="3" gridrow="16" pageId="3" pageNumber="349">24.8</td>
<td id="BC0EA95E75750009FBC5A385828A0B68" box="[1110,1161,1263,1288]" gridcol="4" gridrow="16" pageId="3" pageNumber="349">31.5</td>
<td id="BC0EA95E75750009FB55A38583020B68" box="[1222,1281,1263,1288]" gridcol="5" gridrow="16" pageId="3" pageNumber="349">29.1</td>
<td id="BC0EA95E75750009FAD1A385836D0B68" box="[1346,1390,1263,1288]" gridcol="6" gridrow="16" pageId="3" pageNumber="349">1.97</td>
</tr>
<tr id="FFDFC02275750009FF35A27F836D0B4E" box="[166,1390,1301,1326]" gridrow="17" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A27F87F10B4E" box="[166,498,1301,1326]" gridcol="0" gridrow="17" pageId="3" pageNumber="349">Body width at anal-fin insertion</th>
<td id="BC0EA95E75750009FD60A27F85520B4E" box="[755,849,1301,1326]" gridcol="1" gridrow="17" pageId="3" pageNumber="349">13.0</td>
<td id="BC0EA95E75750009FC30A27F85BF0B4E" box="[931,956,1301,1326]" gridcol="2" gridrow="17" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA27F82180B4E" box="[1005,1051,1301,1326]" gridcol="3" gridrow="17" pageId="3" pageNumber="349">12.0</td>
<td id="BC0EA95E75750009FBC5A27F828A0B4E" box="[1110,1161,1301,1326]" gridcol="4" gridrow="17" pageId="3" pageNumber="349">14.0</td>
<td id="BC0EA95E75750009FB55A27F83020B4E" box="[1222,1281,1301,1326]" gridcol="5" gridrow="17" pageId="3" pageNumber="349">13.2</td>
<td id="BC0EA95E75750009FAD1A27F836D0B4E" box="[1346,1390,1301,1326]" gridcol="6" gridrow="17" pageId="3" pageNumber="349">0.64</td>
</tr>
<tr id="FFDFC02275750009FF35A251836D0B34" box="[166,1390,1339,1364]" gridrow="18" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A25187F10B34" box="[166,498,1339,1364]" gridcol="0" gridrow="18" pageId="3" pageNumber="349">Snout-opercle length</th>
<td id="BC0EA95E75750009FD60A25185520B34" box="[755,849,1339,1364]" gridcol="1" gridrow="18" pageId="3" pageNumber="349">27.2</td>
<td id="BC0EA95E75750009FC30A25185BF0B34" box="[931,956,1339,1364]" gridcol="2" gridrow="18" pageId="3" pageNumber="349">10</td>
<td id="BC0EA95E75750009FC7EA25182180B34" box="[1005,1051,1339,1364]" gridcol="3" gridrow="18" pageId="3" pageNumber="349">26.3</td>
<td id="BC0EA95E75750009FBC5A251828A0B34" box="[1110,1161,1339,1364]" gridcol="4" gridrow="18" pageId="3" pageNumber="349">30.2</td>
<td id="BC0EA95E75750009FB55A25183020B34" box="[1222,1281,1339,1364]" gridcol="5" gridrow="18" pageId="3" pageNumber="349">27.7</td>
<td id="BC0EA95E75750009FAD1A251836D0B34" box="[1346,1390,1339,1364]" gridcol="6" gridrow="18" pageId="3" pageNumber="349">1.09</td>
</tr>
<tr id="FFDFC02275750009FF35A20A836D0B19" box="[166,1390,1376,1401]" gridrow="19" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A20A87F10B19" box="[166,498,1376,1401]" gridcol="0" gridrow="19" pageId="3" pageNumber="349">Percents of HL</th>
<td id="BC0EA95E75750009FD60A20A85520B19" box="[755,849,1376,1401]" gridcol="1" gridrow="19" pageId="3" pageNumber="349">Holotype</td>
<td id="BC0EA95E75750009FC30A20A85BF0B19" box="[931,956,1376,1401]" gridcol="2" gridrow="19" pageId="3" pageNumber="349">N</td>
<td id="BC0EA95E75750009FC7EA20A82180B19" box="[1005,1051,1376,1401]" gridcol="3" gridrow="19" pageId="3" pageNumber="349">Low</td>
<td id="BC0EA95E75750009FBC5A20A828A0B19" box="[1110,1161,1376,1401]" gridcol="4" gridrow="19" pageId="3" pageNumber="349">High</td>
<td id="BC0EA95E75750009FB55A20A83020B19" box="[1222,1281,1376,1401]" gridcol="5" gridrow="19" pageId="3" pageNumber="349">Mean</td>
<td id="BC0EA95E75750009FAD1A20A836D0B19" box="[1346,1390,1376,1401]" gridcol="6" gridrow="19" pageId="3" pageNumber="349">SD</td>
</tr>
<tr id="FFDFC02275750009FF35A2EC836D0BFF" box="[166,1390,1414,1439]" gridrow="20" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A2EC87F10BFF" box="[166,498,1414,1439]" gridcol="0" gridrow="20" pageId="3" pageNumber="349">Snout length</th>
<td id="BC0EA95E75750009FD60A2EC85520BFF" box="[755,849,1414,1439]" gridcol="1" gridrow="20" pageId="3" pageNumber="349">50.6</td>
<td id="BC0EA95E75750009FC30A2EC85BF0BFF" box="[931,956,1414,1439]" gridcol="2" gridrow="20" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA2EC82180BFF" box="[1005,1051,1414,1439]" gridcol="3" gridrow="20" pageId="3" pageNumber="349">49.6</td>
<td id="BC0EA95E75750009FBC5A2EC828A0BFF" box="[1110,1161,1414,1439]" gridcol="4" gridrow="20" pageId="3" pageNumber="349">55.2</td>
<td id="BC0EA95E75750009FB55A2EC83020BFF" box="[1222,1281,1414,1439]" gridcol="5" gridrow="20" pageId="3" pageNumber="349">51.4</td>
<td id="BC0EA95E75750009FAD1A2EC836D0BFF" box="[1346,1390,1414,1439]" gridcol="6" gridrow="20" pageId="3" pageNumber="349">1.72</td>
</tr>
<tr id="FFDFC02275750009FF35A2C6836D0BA5" box="[166,1390,1452,1477]" gridrow="21" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A2C687F10BA5" box="[166,498,1452,1477]" gridcol="0" gridrow="21" pageId="3" pageNumber="349">Orbital diameter</th>
<td id="BC0EA95E75750009FD60A2C685520BA5" box="[755,849,1452,1477]" gridcol="1" gridrow="21" pageId="3" pageNumber="349">15.1</td>
<td id="BC0EA95E75750009FC30A2C685BF0BA5" box="[931,956,1452,1477]" gridcol="2" gridrow="21" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA2C682180BA5" box="[1005,1051,1452,1477]" gridcol="3" gridrow="21" pageId="3" pageNumber="349">13.6</td>
<td id="BC0EA95E75750009FBC5A2C6828A0BA5" box="[1110,1161,1452,1477]" gridcol="4" gridrow="21" pageId="3" pageNumber="349">15.7</td>
<td id="BC0EA95E75750009FB55A2C683020BA5" box="[1222,1281,1452,1477]" gridcol="5" gridrow="21" pageId="3" pageNumber="349">15.1</td>
<td id="BC0EA95E75750009FAD1A2C6836D0BA5" box="[1346,1390,1452,1477]" gridcol="6" gridrow="21" pageId="3" pageNumber="349">0.52</td>
</tr>
<tr id="FFDFC02275750009FF35A2B8836D0B8B" box="[166,1390,1490,1515]" gridrow="22" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A2B887F10B8B" box="[166,498,1490,1515]" gridcol="0" gridrow="22" pageId="3" pageNumber="349">Interorbital width</th>
<td id="BC0EA95E75750009FD60A2B885520B8B" box="[755,849,1490,1515]" gridcol="1" gridrow="22" pageId="3" pageNumber="349">37.2</td>
<td id="BC0EA95E75750009FC30A2B885BF0B8B" box="[931,956,1490,1515]" gridcol="2" gridrow="22" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA2B882180B8B" box="[1005,1051,1490,1515]" gridcol="3" gridrow="22" pageId="3" pageNumber="349">37.2</td>
<td id="BC0EA95E75750009FBC5A2B8828A0B8B" box="[1110,1161,1490,1515]" gridcol="4" gridrow="22" pageId="3" pageNumber="349">42.4</td>
<td id="BC0EA95E75750009FB55A2B883020B8B" box="[1222,1281,1490,1515]" gridcol="5" gridrow="22" pageId="3" pageNumber="349">39.7</td>
<td id="BC0EA95E75750009FAD1A2B8836D0B8B" box="[1346,1390,1490,1515]" gridcol="6" gridrow="22" pageId="3" pageNumber="349">1.54</td>
</tr>
<tr id="FFDFC02275750009FF35A292836D0871" box="[166,1390,1528,1553]" gridrow="23" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A29287F10871" box="[166,498,1528,1553]" gridcol="0" gridrow="23" pageId="3" pageNumber="349">Head depth</th>
<td id="BC0EA95E75750009FD60A29285520871" box="[755,849,1528,1553]" gridcol="1" gridrow="23" pageId="3" pageNumber="349">51.4</td>
<td id="BC0EA95E75750009FC30A29285BF0871" box="[931,956,1528,1553]" gridcol="2" gridrow="23" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA29282180871" box="[1005,1051,1528,1553]" gridcol="3" gridrow="23" pageId="3" pageNumber="349">48.1</td>
<td id="BC0EA95E75750009FBC5A292828A0871" box="[1110,1161,1528,1553]" gridcol="4" gridrow="23" pageId="3" pageNumber="349">52.2</td>
<td id="BC0EA95E75750009FB55A29283020871" box="[1222,1281,1528,1553]" gridcol="5" gridrow="23" pageId="3" pageNumber="349">50.3</td>
<td id="BC0EA95E75750009FAD1A292836D0871" box="[1346,1390,1528,1553]" gridcol="6" gridrow="23" pageId="3" pageNumber="349">1.35</td>
</tr>
<tr id="FFDFC02275750009FF35A177836D0856" box="[166,1390,1565,1590]" gridrow="24" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A17787F10856" box="[166,498,1565,1590]" gridcol="0" gridrow="24" pageId="3" pageNumber="349">Suborbital depth</th>
<td id="BC0EA95E75750009FD60A17785520856" box="[755,849,1565,1590]" gridcol="1" gridrow="24" pageId="3" pageNumber="349">18.4</td>
<td id="BC0EA95E75750009FC30A17785BF0856" box="[931,956,1565,1590]" gridcol="2" gridrow="24" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA17782180856" box="[1005,1051,1565,1590]" gridcol="3" gridrow="24" pageId="3" pageNumber="349">17.9</td>
<td id="BC0EA95E75750009FBC5A177828A0856" box="[1110,1161,1565,1590]" gridcol="4" gridrow="24" pageId="3" pageNumber="349">22.4</td>
<td id="BC0EA95E75750009FB55A17783020856" box="[1222,1281,1565,1590]" gridcol="5" gridrow="24" pageId="3" pageNumber="349">20.1</td>
<td id="BC0EA95E75750009FAD1A177836D0856" box="[1346,1390,1565,1590]" gridcol="6" gridrow="24" pageId="3" pageNumber="349">1.60</td>
</tr>
<tr id="FFDFC02275750009FF35A129836D083C" box="[166,1390,1603,1628]" gridrow="25" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A12987F1083C" box="[166,498,1603,1628]" gridcol="0" gridrow="25" pageId="3" pageNumber="349">Head width</th>
<td id="BC0EA95E75750009FD60A1298552083C" box="[755,849,1603,1628]" gridcol="1" gridrow="25" pageId="3" pageNumber="349">62.6</td>
<td id="BC0EA95E75750009FC30A12985BF083C" box="[931,956,1603,1628]" gridcol="2" gridrow="25" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA1298218083C" box="[1005,1051,1603,1628]" gridcol="3" gridrow="25" pageId="3" pageNumber="349">57.7</td>
<td id="BC0EA95E75750009FBC5A129828A083C" box="[1110,1161,1603,1628]" gridcol="4" gridrow="25" pageId="3" pageNumber="349">66.9</td>
<td id="BC0EA95E75750009FB55A1298302083C" box="[1222,1281,1603,1628]" gridcol="5" gridrow="25" pageId="3" pageNumber="349">63.6</td>
<td id="BC0EA95E75750009FAD1A129836D083C" box="[1346,1390,1603,1628]" gridcol="6" gridrow="25" pageId="3" pageNumber="349">2.36</td>
</tr>
<tr id="FFDFC02275750009FF35A103836D08E2" box="[166,1390,1641,1666]" gridrow="26" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A10387F108E2" box="[166,498,1641,1666]" gridcol="0" gridrow="26" pageId="3" pageNumber="349">Meristics</th>
<td id="BC0EA95E75750009FD60A103855208E2" box="[755,849,1641,1666]" gridcol="1" gridrow="26" pageId="3" pageNumber="349">Holotype</td>
<td id="BC0EA95E75750009FC30A10385BF08E2" box="[931,956,1641,1666]" gridcol="2" gridrow="26" pageId="3" pageNumber="349">N</td>
<td id="BC0EA95E75750009FC7EA103821808E2" box="[1005,1051,1641,1666]" gridcol="3" gridrow="26" pageId="3" pageNumber="349">Low</td>
<td id="BC0EA95E75750009FBC5A103828A08E2" box="[1110,1161,1641,1666]" gridcol="4" gridrow="26" pageId="3" pageNumber="349">High</td>
<td id="BC0EA95E75750009FB55A103830208E2" box="[1222,1281,1641,1666]" gridcol="5" gridrow="26" pageId="3" pageNumber="349">Mode</td>
<td id="BC0EA95E75750009FAD1A103836D08E2" box="[1346,1390,1641,1666]" gridcol="6" gridrow="26" pageId="3" pageNumber="349">SD</td>
</tr>
<tr id="FFDFC02275750009FF35A1E5836D08C8" box="[166,1390,1679,1704]" gridrow="27" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A1E587F108C8" box="[166,498,1679,1704]" gridcol="0" gridrow="27" pageId="3" pageNumber="349">Premaxillary teeth</th>
<td id="BC0EA95E75750009FD60A1E5855208C8" box="[755,849,1679,1704]" gridcol="1" gridrow="27" pageId="3" pageNumber="349">17</td>
<td id="BC0EA95E75750009FC30A1E585BF08C8" box="[931,956,1679,1704]" gridcol="2" gridrow="27" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA1E5821808C8" box="[1005,1051,1679,1704]" gridcol="3" gridrow="27" pageId="3" pageNumber="349">15</td>
<td id="BC0EA95E75750009FBC5A1E5828A08C8" box="[1110,1161,1679,1704]" gridcol="4" gridrow="27" pageId="3" pageNumber="349">21</td>
<td id="BC0EA95E75750009FB55A1E5830208C8" box="[1222,1281,1679,1704]" gridcol="5" gridrow="27" pageId="3" pageNumber="349">17</td>
<td id="BC0EA95E75750009FAD1A1E5836D08C8" box="[1346,1390,1679,1704]" gridcol="6" gridrow="27" pageId="3" pageNumber="349">−</td>
</tr>
<tr id="FFDFC02275750009FF35A1DF836D08AE" box="[166,1390,1717,1742]" gridrow="28" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A1DF87F108AE" box="[166,498,1717,1742]" gridcol="0" gridrow="28" pageId="3" pageNumber="349">Dentary teeth</th>
<td id="BC0EA95E75750009FD60A1DF855208AE" box="[755,849,1717,1742]" gridcol="1" gridrow="28" pageId="3" pageNumber="349">15</td>
<td id="BC0EA95E75750009FC30A1DF85BF08AE" box="[931,956,1717,1742]" gridcol="2" gridrow="28" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA1DF821808AE" box="[1005,1051,1717,1742]" gridcol="3" gridrow="28" pageId="3" pageNumber="349">14</td>
<td id="BC0EA95E75750009FBC5A1DF828A08AE" box="[1110,1161,1717,1742]" gridcol="4" gridrow="28" pageId="3" pageNumber="349">18</td>
<td id="BC0EA95E75750009FB55A1DF830208AE" box="[1222,1281,1717,1742]" gridcol="5" gridrow="28" pageId="3" pageNumber="349">17</td>
<td id="BC0EA95E75750009FAD1A1DF836D08AE" box="[1346,1390,1717,1742]" gridcol="6" gridrow="28" pageId="3" pageNumber="349">−</td>
</tr>
<tr id="FFDFC02275750009FF35A1B0836D0893" box="[166,1390,1754,1779]" gridrow="29" pageId="3" pageNumber="349">
<th id="BC0EA95E75750009FF35A1B087F10893" box="[166,498,1754,1779]" gridcol="0" gridrow="29" pageId="3" pageNumber="349">Lateral plates</th>
<td id="BC0EA95E75750009FD60A1B085520893" box="[755,849,1754,1779]" gridcol="1" gridrow="29" pageId="3" pageNumber="349">22</td>
<td id="BC0EA95E75750009FC30A1B085BF0893" box="[931,956,1754,1779]" gridcol="2" gridrow="29" pageId="3" pageNumber="349">12</td>
<td id="BC0EA95E75750009FC7EA1B082180893" box="[1005,1051,1754,1779]" gridcol="3" gridrow="29" pageId="3" pageNumber="349">21</td>
<td id="BC0EA95E75750009FBC5A1B0828A0893" box="[1110,1161,1754,1779]" gridcol="4" gridrow="29" pageId="3" pageNumber="349">23</td>
<td id="BC0EA95E75750009FB55A1B083020893" box="[1222,1281,1754,1779]" gridcol="5" gridrow="29" pageId="3" pageNumber="349">22</td>
<td id="BC0EA95E75750009FAD1A1B0836D0893" box="[1346,1390,1754,1779]" gridcol="6" gridrow="29" pageId="3" pageNumber="349">−</td>
</tr>
</table>
</paragraph>
<paragraph id="4150C2607575FFF2FF54A04984690E9B" blockId="3.[151,1436,1827,2035]" lastBlockId="4.[151,1437,153,791]" lastPageId="4" lastPageNumber="350" pageId="3" pageNumber="349">
<emphasis id="739B1E727575FFF5FF54A049878E095E" bold="true" box="[199,397,1827,1854]" pageId="3" pageNumber="349">Color in alcohol.</emphasis>
Dorsal surface of head dark brownish, except for striking
<emphasis id="739B1E727575FFF5FB8AA04F8229095E" box="[1049,1066,1829,1854]" italics="true" pageId="3" pageNumber="349">V</emphasis>
-shaped yellowish tan mark from rostral plate passing through nares to anterior orbital margins. Background color yellowish-brown in dorsal and ventral portions of body. Five dark brownish bars, forming saddles in dorsal view, laterally most broad, sometimes forming a longitudinal stripe on sides of trunk, and crossing to ventral region: first located at dorsal-fin origin, second at end of dorsal-fin base, third at adipose-fin region, and fourth at caudal peduncle and the fifth at caudal fin origin. Dorsal, pectoral, and pelvic fins with dark chromatophores, forming irregular sets of bars: three on dorsal fin, four on pectoral fins, three on pelvic fin, and two to three on anal fin. Dark blotch of pigmentation on posterior portion of caudal peduncle that extends median portion of caudal-fin. Dark chromatophores forming irregular sets of bars at middle and at end of caudal fin.
</paragraph>
<paragraph id="4150C2607572FFF2FF54A66E820E0F23" blockId="4.[151,1437,153,791]" pageId="4" pageNumber="350">
<emphasis id="739B1E727572FFF2FF54A66E875F0F7F" bold="true" box="[199,348,260,287]" pageId="4" pageNumber="350">Color in life.</emphasis>
Similar pattern described for alcohol preserved specimens, but with dark-brown background coloration and with pronounced dark chromatophores and bars over body (
<figureCitation id="D9D4DEE57572FFF2FC29A64382030F23" box="[954,1024,297,323]" captionStart="FIGURE 2" captionStartId="4.[151,250,1570,1595]" captionTargetBox="[151,1436,813,1551]" captionTargetId="figure@4.[151,1436,813,1551]" captionTargetPageId="4" captionText="FIGURE 2. Parotocinclus pentakelis, live specimen, MZUSP 124900, holotype, 22.8 mm SL." httpUri="https://zenodo.org/record/3349501/files/figure.png" pageId="4" pageNumber="350">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="4150C2607572FFF2FF54A62687680FB4" blockId="4.[151,1437,153,791]" pageId="4" pageNumber="350">
<emphasis id="739B1E727572FFF2FF54A62687BA0F07" bold="true" box="[199,441,332,359]" pageId="4" pageNumber="350">Sexual dimorphism.</emphasis>
Males have (1) a papilla urogenital (
<emphasis id="739B1E727572FFF2FCF0A62485820F07" box="[867,897,334,359]" italics="true" pageId="4" pageNumber="350">vs.</emphasis>
papilla absent in females); (2) the posterior tip of pelvic fin beyond anal-fin origin (
<emphasis id="739B1E727572FFF2FDA6A61884570FEB" box="[565,596,370,395]" italics="true" pageId="4" pageNumber="350">vs.</emphasis>
pelvic fin not reaching anal-fin origin in females); and (3) the unbranched pelvic-fin ray supporting a dermal flap on dorsal surface (
<emphasis id="739B1E727572FFF2FCBAA6FC85440FCF" box="[809,839,406,431]" italics="true" pageId="4" pageNumber="350">vs.</emphasis>
dermal flap on proximal dorsal surface of pelvic-fin absent in females).
</paragraph>
<paragraph id="4150C2607572FFF2FF54A6B682AE0C03" blockId="4.[151,1437,153,791]" pageId="4" pageNumber="350">
<emphasis id="739B1E727572FFF2FF54A6B687620F97" bold="true" box="[199,353,476,503]" pageId="4" pageNumber="350">Distribution.</emphasis>
<taxonomicName id="86EFB9E37572FFF2FEFAA6B784830F97" authorityName="Roxo &amp; Messias &amp; Silva" authorityYear="2019" box="[361,640,477,503]" class="Actinopterygii" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="350" phylum="Chordata" rank="species" species="pentakelis">
<emphasis id="739B1E727572FFF2FEFAA6B784830F97" box="[361,640,477,503]" italics="true" pageId="4" pageNumber="350">Parotocinclus pentakelis</emphasis>
</taxonomicName>
is known from two localities of the Rio
<collectingRegion id="832B0C827572FFF2FBC0A6B782BC0F97" box="[1107,1215,477,503]" country="Brazil" name="Tocantins" pageId="4" pageNumber="350">Tocantins</collectingRegion>
basin (
<figureCitation id="D9D4DEE57572FFF2FA87A6B783580F97" box="[1300,1371,477,503]" captionStart="FIGURE 4" captionStartId="6.[151,250,1957,1982]" captionTargetBox="[171,1411,1045,1914]" captionTargetId="figure@6.[151,1436,1029,1938]" captionTargetPageId="6" captionText="FIGURE 4. Type locality of Parotocinclus pentakelis (black star) at rio Palmas, and paratypes locality at rio Manuel Alves da Natividade (black circle), rio Tocantins basin." httpUri="https://zenodo.org/record/3349505/files/figure.png" pageId="4" pageNumber="350">Fig. 4</figureCitation>
). The type-locality is at Rio Palmas (
<geoCoordinate id="24DBA4A77572FFF2FE7DA56B84810C7B" box="[494,642,512,539]" degrees="12" direction="south" minutes="48" orientation="latitude" pageId="4" pageNumber="350" precision="15" seconds="05" value="-12.801389">
12
<superScript id="B69A6F287572FFF2FD98A56A84100C6E" attach="left" box="[523,531,512,526]" fontSize="6" pageId="4" pageNumber="350">o</superScript>
48’ 05” S
</geoCoordinate>
;
<geoCoordinate id="24DBA4A77572FFF2FD1DA56B85280C7B" box="[654,811,512,539]" degrees="46" direction="west" minutes="28" orientation="longitude" pageId="4" pageNumber="350" precision="15" seconds="38" value="-46.477222">
46
<superScript id="B69A6F287572FFF2FD39A56A84B10C6E" attach="left" box="[682,690,512,526]" fontSize="6" pageId="4" pageNumber="350">o</superScript>
28’ 38” W
</geoCoordinate>
), municipality of Lavandeira, southeastern of
<collectingRegion id="832B0C827572FFF2FAA3A56B86CD0C5F" country="Brazil" name="Tocantins" pageId="4" pageNumber="350">Tocantins state</collectingRegion>
. The new species is also found in a second locality at Rio Manuel Alves da Natividade (
<geoCoordinate id="24DBA4A77572FFF2FB3EA54F83420C5F" box="[1197,1345,548,575]" degrees="11" direction="south" minutes="36" orientation="latitude" pageId="4" pageNumber="350" precision="15" seconds="41" value="-11.611389">
11
<superScript id="B69A6F287572FFF2FB5BA54E82D30C52" attach="left" box="[1224,1232,548,562]" fontSize="6" pageId="4" pageNumber="350">o</superScript>
36’ 41” S
</geoCoordinate>
;
<geoCoordinate id="24DBA4A77572FFF2FADDA54F86E30C03" degrees="47" direction="west" minutes="02" orientation="longitude" pageId="4" pageNumber="350" precision="15" seconds="39" value="-47.044167">
47
<superScript id="B69A6F287572FFF2FAF9A54E83710C52" attach="left" box="[1386,1394,548,562]" fontSize="6" pageId="4" pageNumber="350">o</superScript>
02’ 39” W
</geoCoordinate>
), a river draining directly to Rio
<collectingRegion id="832B0C827572FFF2FDC7A52384C00C03" box="[596,707,585,611]" country="Brazil" name="Tocantins" pageId="4" pageNumber="350">Tocantins</collectingRegion>
, municipality of Porto Alegre do
<collectingRegion id="832B0C827572FFF2FBAAA52382AA0C03" box="[1081,1193,585,611]" country="Brazil" name="Tocantins" pageId="4" pageNumber="350">Tocantins</collectingRegion>
.
</paragraph>
<paragraph id="4150C2607572FFF2FF54A50687330CAF" blockId="4.[151,1437,153,791]" pageId="4" pageNumber="350">
<emphasis id="739B1E727572FFF2FF54A50687290CE7" bold="true" box="[199,298,620,647]" pageId="4" pageNumber="350">Habitat.</emphasis>
The new species was found associated with marginal vegetation of median size rivers, (
<figureCitation id="D9D4DEE57572FFF2FA95A507834B0CE7" box="[1286,1352,620,647]" captionStart="FIGURE 5" captionStartId="7.[151,250,1456,1481]" captionTargetBox="[151,1436,181,1437]" captionTargetId="figure@7.[151,1436,181,1437]" captionTargetPageId="7" captionText="FIGURE 5. Type locality of Parotocinclus pentakelis at rio Palmas, rio Tocantins basin, 12o 48’ 05” S; 46o 28’ 38” W. Species were found associated with marginal vegetation." httpUri="https://zenodo.org/record/3349507/files/figure.png" pageId="4" pageNumber="350">Fig. 5</figureCitation>
) that is composed of grasses, shrubs and large trees. The waters where this species is found clear fast-flowing
<quantity id="86176F857572FFF2FA8DA5FA839F0CCB" box="[1310,1436,656,683]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="4" pageNumber="350" unit="cm" value="75.0" valueMax="100.0" valueMin="50.0">50–100 cm</quantity>
near margins.
</paragraph>
<paragraph id="4150C2607572FFF2FF54A5B3850A0D77" blockId="4.[151,1437,153,791]" pageId="4" pageNumber="350">
<emphasis id="739B1E727572FFF2FF54A5B3874D0C93" bold="true" box="[199,334,729,755]" pageId="4" pageNumber="350">Etymology.</emphasis>
The specific name “
<taxonomicName id="86EFB9E37572FFF2FDABA5B384A90C93" authorityName="Roxo &amp; Messias &amp; Silva" authorityYear="2019" box="[568,682,729,755]" class="Actinopterygii" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="350" phylum="Chordata" rank="species" species="pentakelis">
<emphasis id="739B1E727572FFF2FDABA5B384A90C93" box="[568,682,729,755]" italics="true" pageId="4" pageNumber="350">pentakelis</emphasis>
</taxonomicName>
” comes from the Greek
<emphasis id="739B1E727572FFF2FC51A5B082030C93" box="[962,1024,730,755]" italics="true" pageId="4" pageNumber="350">penta</emphasis>
(five) and
<emphasis id="739B1E727572FFF2FBEDA5B382B20C93" box="[1150,1201,729,755]" italics="true" pageId="4" pageNumber="350">kelis</emphasis>
(stain) related to the color pattern of the body with four transverse dark bars.
</paragraph>
</subSubSection>
</treatment>
</document>
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