treatments-xml/data/C8/0F/AB/C80FAB481D5752D182F35B3C889D61A7.xml

275 lines
33 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-CLB-Dataset="3848" ID-DOI="http://dx.doi.org/10.3897/zookeys.1062.66845" ID-GBIF-Dataset="e73f66c3-4c75-4c11-9ad1-be2def71e368" ID-PMC="PMC8530997" ID-Pensoft-Pub="1313-2970-1062-73" ID-Pensoft-UUID="2BCBF5DF56B450B1A9A88480A9F8B9CE" ID-PubMed="34720618" ID-ZooBank="310272E8F5DD4DAB84603AFCCD2E9C45" ModsDocID="1313-2970-1062-73" checkinTime="1634290158865" checkinUser="pensoft" docAuthor="Zhang, Jianshuang, Yu, Hao &amp; Li, Shuqiang" docDate="2021" docId="C80FAB481D5752D182F35B3C889D61A7" docLanguage="en" docName="ZooKeys 1062: 73-122" docOrigin="ZooKeys 1062" docPubDate="2021-10-14" docSource="http://dx.doi.org/10.3897/zookeys.1062.66845" docTitle="Matidia Thorell 1878" docType="treatment" docVersion="6" id="2BCBF5DF56B450B1A9A88480A9F8B9CE" lastPageNumber="73" masterDocId="2BCBF5DF56B450B1A9A88480A9F8B9CE" masterDocTitle="On the clubionid spiders (Araneae, Clubionidae) from Xishuangbanna, China, with descriptions of two new genera and seven new species" masterLastPageNumber="122" masterPageNumber="73" pageNumber="73" updateTime="1732931016866" updateUser="ExternalLinkService">
<mods:mods id="F91FA3C33DE3B53F3C5BC1BDBD53B1B0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CE5441723310A7F19F4D098B15945394">
<mods:title id="7FFD15CBDDAD7822EBEFF4EC1E7C94F9">On the clubionid spiders (Araneae, Clubionidae) from Xishuangbanna, China, with descriptions of two new genera and seven new species</mods:title>
</mods:titleInfo>
<mods:name id="114E0629BA7D8E7D98F52822E720E8C7" type="personal">
<mods:role id="C893420B711A1406E743373255116142">
<mods:roleTerm id="9AB041BBFBC0DF16BA7A5E834BA8BDBA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C902BF99EDAC18607E9626A10B6B8BE3">Zhang, Jianshuang</mods:namePart>
<mods:affiliation id="35DB7BA7FE5DC8595992F8D1BB760047">School of Life Sciences, Guizhou Normal University, Guiyang, Guizhou, China</mods:affiliation>
</mods:name>
<mods:name id="732D46FAB15B75871B2CFC529432658B" type="personal">
<mods:role id="B62D0B318B59BC95B848EE6D0FC3867A">
<mods:roleTerm id="A9F74018C6C3533DA28709B3D9FD42FE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="927A880BE32C3752F80D7B15F6BF9480">Yu, Hao</mods:namePart>
<mods:affiliation id="503DF9DABC15E3845A9455ADBB08A265">School of Biological Sciences, Guizhou Education University, Guiyang, Guizhou, China</mods:affiliation>
<mods:nameIdentifier id="9CC1F5200D0243AA383E705FC2302B8E" type="email">insect1986@126.com</mods:nameIdentifier>
</mods:name>
<mods:name id="46873C6669E8CAAE73CC1AB5FDFF7619" type="personal">
<mods:role id="4B137682BC517C5DA52320985DA92AD0">
<mods:roleTerm id="6C6126FF46D6F5BC9E4B26C14CDAFDF5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="46ABA38509C27EAC7C58040E7951CB60">Li, Shuqiang</mods:namePart>
<mods:nameIdentifier id="FA758A8F24F7F7F24ED75E1BE2CFD829" type="ORCID">https://orcid.org/0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation id="363EC5EE7C7708F54BB979B5363D403B">Institute of Zoology, Chinese Academy of Sciences, Beijing, China</mods:affiliation>
</mods:name>
<mods:typeOfResource id="CA4F31CE762448BE98DFE21EDA5A3299">text</mods:typeOfResource>
<mods:relatedItem id="F87D975620953E1AEFB24433E15F833B" type="host">
<mods:titleInfo id="60DFC8117417FADCC26C9CD8485445B4">
<mods:title id="C3CC4DA8A832C6A6160CA4568A11981B">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="D0A34FBBC959C5BAF7053286912481DD">
<mods:date id="E02A07C3D389C8B44A2E31E68121CE78">2021</mods:date>
<mods:detail id="C88F5956980A7A2BBE8A9A04A4FD1848" type="pubDate">
<mods:number id="F051EB449D91EFA4CFD2D3DD0D89CBD8">2021-10-14</mods:number>
</mods:detail>
<mods:detail id="64A46C4CFB793B531D00DD101036C86C" type="volume">
<mods:number id="FF098474E9D7FC88D1267907E3A566B8">1062</mods:number>
</mods:detail>
<mods:extent id="953E5241618F14DB8055C17E0CB41AEB" unit="page">
<mods:start id="C5AE3952B0BC985DA56E72012C0E9B97">73</mods:start>
<mods:end id="3212361C5911C76FE1115C4901687305">122</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="E0E7E0CFE6FC6105350D8D9D26FC250E">
<mods:url id="0DE4C7A7B1B6E474BB02CF8B1CDF9F8D">http://dx.doi.org/10.3897/zookeys.1062.66845</mods:url>
</mods:location>
<mods:classification id="E89ACF209D8E6A07C01158AFB9477FE8">journal article</mods:classification>
<mods:identifier id="9FEED4DD0B0E920DC2DBB457F7EAD465" type="DOI">http://dx.doi.org/10.3897/zookeys.1062.66845</mods:identifier>
<mods:identifier id="A00AE83B16D6AA0F8C2022366FF46439" type="Pensoft-Pub">1313-2970-1062-73</mods:identifier>
<mods:identifier id="17FE09D52F3F905E7DC0F17A6D55357A" type="ZooBank">310272E8F5DD4DAB84603AFCCD2E9C45</mods:identifier>
<mods:identifier id="B32F897ACE2D0E373EECA4F1C757FC80" type="Pensoft-UUID">2BCBF5DF56B450B1A9A88480A9F8B9CE</mods:identifier>
</mods:mods>
<subSection id="3B067D02C977AC1909A9745E57398251" pageId="0" pageNumber="73" type="taxonomy">
<treatment id="C80FAB481D5752D182F35B3C889D61A7" ID-GBIF-Taxon="189468725" LSID="urn:lsid:plazi:treatment:C80FAB481D5752D182F35B3C889D61A7" httpUri="http://treatment.plazi.org/id/C80FAB481D5752D182F35B3C889D61A7" lastPageNumber="73" pageId="0" pageNumber="73" scope_family="Clubionidae" scope_order="Araneae">
<subSubSection id="8DD6AF9E8D517C14154F4D3E4A24862B" pageId="0" pageNumber="73" type="nomenclature">
<paragraph id="26D92390E1F15CBD2B415C2C4C11500A" pageId="0" pageNumber="73">
Genus
<taxonomicName id="4BDE770913DD9E0D90A4D42C1D04D718" ID-CoL="5LZG" LSID="C80FAB48-1D57-52D1-82F3-5B3C889D61A7" authority="Thorell, 1878" authorityName="Thorell" authorityYear="1878" class="Arachnida" family="Clubionidae" genus="Matidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Matidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Matidia Thorell, 1878</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="7287252C442CF23DBA2311F6D8A0BD21" pageId="0" pageNumber="73" type="reference_group">
<paragraph id="F514F231C03C12415888E05A22923D5A" pageId="0" pageNumber="73">
<taxonomicName id="BFE9547927E625A2D93491F81654CDDC" authorityName="Thorell" authorityYear="1878" class="Arachnida" family="Clubionidae" genus="Matidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Matidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Matidia</taxonomicName>
Thorell, 1878: 182.
</paragraph>
<paragraph id="F8653F0A79C8106477148297F546E51A" pageId="0" pageNumber="73">
<taxonomicName id="E59EB5B36AF1F9A9A9C2501BC62CB05F" authorityName="Barrion &amp; Litsinger" authorityYear="1995" class="Arachnida" family="Clubionidae" genus="Kakaibanoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kakaibanoides" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Kakaibanoides</taxonomicName>
Barrion &amp; Litsinger, 1995: 149 (type
<taxonomicName id="D5E149886BFD7015B90FEC8415827A50" class="Arachnida" family="Clubionidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="K. paranga" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="paranga">K. paranga</taxonomicName>
Barrion &amp; Litsinger, 1995, considered as junior synonym of
<taxonomicName id="53114DBC0C3671582CA1485D65213EE7" authorityName="Thorell" authorityYear="1878" class="Arachnida" family="Clubionidae" genus="Matidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Matidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Matidia</taxonomicName>
by Deeleman-Reinhold, 2001: 156).
</paragraph>
</subSubSection>
<subSubSection id="BD3EFAD0C1F1F4278FF5E13AEDF2A946" pageId="0" pageNumber="73" type="type species">
<paragraph id="088C467C27F7BA2C749EE2FB2BEB67C3" pageId="0" pageNumber="73">Type species.</paragraph>
<paragraph id="F792431C9BDCAE81FAA80466C806E767" pageId="0" pageNumber="73">
<taxonomicName id="05EDC8FA617981137D7F71547394A2A3" authorityName="Thorell" authorityYear="1878" class="Arachnida" family="Clubionidae" genus="Matidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Matidia virens" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="B71F310C70D2EC19F2FF4FEF44E059A6" italics="true" pageId="0" pageNumber="73">Matidia virens</emphasis>
</taxonomicName>
Thorell, 1878 from Moluccas, Sulawesi.
</paragraph>
</subSubSection>
<subSubSection id="C57AD9AE23506447087B2DBD20E78DF4" pageId="0" pageNumber="73" type="diagnosis">
<paragraph id="98D77CD809DC6BC8B8329BE8A8225D09" pageId="0" pageNumber="73">Diagnosis.</paragraph>
<paragraph id="C66FDA070B40A2DAA38A235160611538" pageId="0" pageNumber="73">
Species in this genus differ from all other clubionids by the following: the pars cephalica is 2
<normalizedToken id="255D2DBEF096A49523B7449DAFF660A8" originalValue="×">x</normalizedToken>
narrower than the pars thoracica (Figs
<figureCitation id="FBC33026C060831D470CC01C99943F47" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Matidia spatulata, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view; path of copulatory duct marked E dorsal view F lateral view G dorsal view H ventral view. Arrow (F) point at dark ventral abdominal spot in male. Abbreviations: BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.10 mm (equal for A-D); 2 mm (equal for E, F, equal for G, H)." figureDoi="10.3897/zookeys.1062.66845.figure4" httpUri="https://binary.pensoft.net/fig/599508" pageId="0" pageNumber="73">4E, G</figureCitation>
,
<figureCitation id="70EC9FB60DDC66B850BE10A110218B15" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Matidia xieqian sp. nov., female paratype and male holotype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view; path of copulatory duct marked E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: A = epigynal atrium; BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.20 mm (equal for A-D); 1 mm (equal for E, F, equal for G, H)." figureDoi="10.3897/zookeys.1062.66845.figure6" httpUri="https://binary.pensoft.net/fig/599510" pageId="0" pageNumber="73">6E-G</figureCitation>
) (vs. wider), leg I is longest (vs. not longest), and there is a dark ventral abdominal spot in males (Fig.
<figureCitation id="A14E14E8F9A7C241523641BEF77810DE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Matidia spatulata, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view; path of copulatory duct marked E dorsal view F lateral view G dorsal view H ventral view. Arrow (F) point at dark ventral abdominal spot in male. Abbreviations: BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.10 mm (equal for A-D); 2 mm (equal for E, F, equal for G, H)." figureDoi="10.3897/zookeys.1062.66845.figure4" httpUri="https://binary.pensoft.net/fig/599508" pageId="0" pageNumber="73">4F</figureCitation>
) (vs. absent).
<taxonomicName id="9E8204EB0AA88B409AC31E5E79C99A0D" authorityName="Thorell" authorityYear="1878" class="Arachnida" family="Clubionidae" genus="Matidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Matidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="7AEEFF287B4C9ABA29F02C517561D13C" italics="true" pageId="0" pageNumber="73">Matidia</emphasis>
</taxonomicName>
resembles
<taxonomicName id="7EEFE8C9B4148D13117F63955F19E4BB" authorityName="Deeleman-Reinhold" authorityYear="2001" class="Arachnida" family="Clubionidae" genus="Malamatidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Malamatidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="E782CDEEB691085455F4D2176780ED24" italics="true" pageId="0" pageNumber="73">Malamatidia</emphasis>
</taxonomicName>
,
<taxonomicName id="0A77434D6519495C1D514C3166D933A2" authorityName="Deeleman-Reinhold" authorityYear="2001" class="Arachnida" family="Clubionidae" genus="Nusatidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusatidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="8F12C455FB1F823C09C00B1FA4B9AAAA" italics="true" pageId="0" pageNumber="73">Nusatidia</emphasis>
</taxonomicName>
, and
<taxonomicName id="9C4D117A5675F6E0132382E8DF8B017D" authorityName="Yu &amp; Li" authorityYear="2021" class="Arachnida" family="Clubionidae" genus="Ramosatidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ramosatidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="2A118E61E3FA4ABD7B916248B0616A9B" italics="true" pageId="0" pageNumber="73">Ramosatidia</emphasis>
</taxonomicName>
gen. nov. by the slender, greenish body but is consistently separable by the shape of the copulatory organs: the male palp has a ribbon-shaped embolus (Figs
<figureCitation id="B748C40FB9DF16787F6CB48EE1776A27" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male palp of Matidia spatulata A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, ventrolateral view F bulb, retrolateral view. Abbreviations: C = conductor; E = embolus; EB = embolic base; PTA = prolateral tibial apophysis; RTA = retrolateral tibial apophysis; TA = tegular apophysis. Scale bars: 0.10 mm (equal for A, B, equal for C-F)." figureDoi="10.3897/zookeys.1062.66845.figure3" httpUri="https://binary.pensoft.net/fig/599507" pageId="0" pageNumber="73">3A, C-F</figureCitation>
,
<figureCitation id="700C2F7ECBD5F492308A241E54508B3D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of the holotype of Matidia xieqian sp. nov., left palp and bulb (A, B, E, F) and flipped right bulb (C, D) A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, ventrolateral view F retrolateral view. Arrows (E) point at three processes of tegular apophysis. Abbreviations: C = conductor; E = embolus; PTA = prolateral tibial apophysis; RTA = retrolateral tibial apophysis; TA = tegular apophysis. Scale bars: 0.20 mm (equal for A, B, equal for C-F)." figureDoi="10.3897/zookeys.1062.66845.figure5" httpUri="https://binary.pensoft.net/fig/599509" pageId="0" pageNumber="73">5A, C-F</figureCitation>
) (vs. embolus not ribbon-shaped), the epigyne has one or two depressions (or an atrium) and no septum (Figs
<figureCitation id="63783CD1C97F91011C0D556399706549" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Matidia spatulata, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view; path of copulatory duct marked E dorsal view F lateral view G dorsal view H ventral view. Arrow (F) point at dark ventral abdominal spot in male. Abbreviations: BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.10 mm (equal for A-D); 2 mm (equal for E, F, equal for G, H)." figureDoi="10.3897/zookeys.1062.66845.figure4" httpUri="https://binary.pensoft.net/fig/599508" pageId="0" pageNumber="73">4A, B</figureCitation>
,
<figureCitation id="37C073AC4B12256B1A19B98CF8798F76" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Matidia xieqian sp. nov., female paratype and male holotype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view; path of copulatory duct marked E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: A = epigynal atrium; BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.20 mm (equal for A-D); 1 mm (equal for E, F, equal for G, H)." figureDoi="10.3897/zookeys.1062.66845.figure6" httpUri="https://binary.pensoft.net/fig/599510" pageId="0" pageNumber="73">6A, B</figureCitation>
) (vs. depression lacking in
<taxonomicName id="86ADC738269A864B838CFADCABB82229" authorityName="Deeleman-Reinhold" authorityYear="2001" class="Arachnida" family="Clubionidae" genus="Nusatidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusatidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="AECE3B549AF2D036C4B8DF474A646CAA" italics="true" pageId="0" pageNumber="73">Nusatidia</emphasis>
</taxonomicName>
and
<taxonomicName id="27D533F48342580226A8EB5ECB7B2E90" authorityName="Yu &amp; Li" authorityYear="2021" class="Arachnida" family="Clubionidae" genus="Ramosatidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ramosatidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="31B573E1854694B446295FA7E281C0D5" italics="true" pageId="0" pageNumber="73">Ramosatidia</emphasis>
</taxonomicName>
gen. nov., or present but with a septum in
<taxonomicName id="EAB6DA146BFE0BF8BD344A8B19C805A8" authorityName="Deeleman-Reinhold" authorityYear="2001" class="Arachnida" family="Clubionidae" genus="Malamatidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Malamatidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="D4B4A0066A20FAA60A7E026EB9DBFE53" italics="true" pageId="0" pageNumber="73">Malamatidia</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="0660A24DB50A066DC9542E5F288F275A" doi="10.3897/zookeys.1062.66845.figure3" httpUri="https://binary.pensoft.net/fig/599507" pageId="0" pageNumber="73" start="Figure 3" startId="F3">
<paragraph id="FBD5745F0F0D1E65BAA82240150A8971" pageId="0" pageNumber="73">
<emphasis id="2DAE9DC2C8453E38C4416512D43B6CFE" bold="true" pageId="0" pageNumber="73">Figure 3.</emphasis>
Male palp of
<taxonomicName id="E7FF321E76BE454542F2FD5AC6EA06BB" authorityName="Chen &amp; Huang" authorityYear="2006" class="Arachnida" family="Clubionidae" genus="Matidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Matidia spatulata" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="spatulata">
<emphasis id="1211EE9C173C097D8EE086C56A4F49B2" italics="true" pageId="0" pageNumber="73">Matidia spatulata</emphasis>
</taxonomicName>
<emphasis id="46E4575FDEF6FE54A57B4F809B47B89D" bold="true" pageId="0" pageNumber="73">A</emphasis>
prolateral view
<emphasis id="8CF0BC69228577E08A26F9590B763B36" bold="true" pageId="0" pageNumber="73">B</emphasis>
retrolateral view
<emphasis id="50BF4002B40D1E638737E7E7CC02FE5E" bold="true" pageId="0" pageNumber="73">C</emphasis>
bulb, prolateral view
<emphasis id="708D78E20DEDB4F10519E1E5ECD372E1" bold="true" pageId="0" pageNumber="73">D</emphasis>
bulb, ventral view
<emphasis id="43DF01F39AD97824CB99EB8BA174B16C" bold="true" pageId="0" pageNumber="73">E</emphasis>
bulb, ventrolateral view
<emphasis id="5D09E38C53F39C80A919E94FFF243938" bold="true" pageId="0" pageNumber="73">F</emphasis>
bulb, retrolateral view. Abbreviations: C = conductor; E = embolus; EB = embolic base; PTA = prolateral tibial apophysis; RTA = retrolateral tibial apophysis; TA = tegular apophysis. Scale bars: 0.10 mm (equal for
<emphasis id="83147F57AC42E66C244BB00D2E6C77FE" bold="true" pageId="0" pageNumber="73">A, B</emphasis>
, equal for
<emphasis id="F0DCB9A59909E8F20F460CC5070E10F6" bold="true" pageId="0" pageNumber="73">C-F</emphasis>
).
</paragraph>
</caption>
<caption id="E74A4D6773619B555DC5F9F0929DCCD9" doi="10.3897/zookeys.1062.66845.figure4" httpUri="https://binary.pensoft.net/fig/599508" pageId="0" pageNumber="73" start="Figure 4" startId="F4">
<paragraph id="FC5813E326D633A30D309B7D712007F1" pageId="0" pageNumber="73">
<emphasis id="C0F3E8E11486D4A78C720002C4CCA315" bold="true" pageId="0" pageNumber="73">Figure 4.</emphasis>
<taxonomicName id="0D090292EC39A4D4A86779C2152A9D06" authorityName="Chen &amp; Huang" authorityYear="2006" class="Arachnida" family="Clubionidae" genus="Matidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Matidia spatulata" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="spatulata">
<emphasis id="EE6CA611ECECCA61F2DE2D49894007B0" italics="true" pageId="0" pageNumber="73">Matidia spatulata</emphasis>
</taxonomicName>
, epigyne (
<emphasis id="25BE988F2F34753358FA68093A538834" bold="true" pageId="0" pageNumber="73">A-D</emphasis>
), male habitus (
<emphasis id="65EE05D3D8AF7D705F0A7F75FF56F39B" bold="true" pageId="0" pageNumber="73">E, F</emphasis>
) and female habitus (
<emphasis id="BC889DC73D069AADC843A01AA0E9EDCB" bold="true" pageId="0" pageNumber="73">G, H</emphasis>
)
<emphasis id="7764F282544E78732ED70EC0362FB57B" bold="true" pageId="0" pageNumber="73">A</emphasis>
intact, ventral view
<emphasis id="B7A64D095D70E271249E0B3147BF5B75" bold="true" pageId="0" pageNumber="73">B</emphasis>
cleared, ventral view
<emphasis id="394A1E2A0EE31DCC45BAB1A1F1A2C198" bold="true" pageId="0" pageNumber="73">C</emphasis>
cleared, dorsal view
<emphasis id="88A153CF0013A0894CCF0EB5C1B0770B" bold="true" pageId="0" pageNumber="73">D</emphasis>
cleared, dorsal view; path of copulatory duct marked
<emphasis id="CDA7CC672E14CB25CAC35F5D8C19EE7E" bold="true" pageId="0" pageNumber="73">E</emphasis>
dorsal view
<emphasis id="CB0BFB347E52E3DEBE13A30033F69A7A" bold="true" pageId="0" pageNumber="73">F</emphasis>
lateral view
<emphasis id="AD5091A1EA74563DE8323BB572283309" bold="true" pageId="0" pageNumber="73">G</emphasis>
dorsal view
<emphasis id="CC73AAF990B124DE6988557AF8E3C748" bold="true" pageId="0" pageNumber="73">H</emphasis>
ventral view. Arrow (
<emphasis id="90D789133C218E9EDEF532B0EEA944A5" bold="true" pageId="0" pageNumber="73">F</emphasis>
) point at dark ventral abdominal spot in male. Abbreviations: BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.10 mm (equal for
<emphasis id="428A69D518FA238CF5FF49499F99CCD5" bold="true" pageId="0" pageNumber="73">A-D</emphasis>
); 2 mm (equal for
<emphasis id="A8145EA530A7E9572F05516F70548ACC" bold="true" pageId="0" pageNumber="73">E, F</emphasis>
, equal for
<emphasis id="005A581BFA24BA144F24039391CEB648" bold="true" pageId="0" pageNumber="73">G, H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="5A7BBC22784282745F3F75D9337E184D" pageId="0" pageNumber="73" type="comments">
<paragraph id="1158C99FC32BBB9BD5A64921B1E61CA9" pageId="0" pageNumber="73">Comments.</paragraph>
<paragraph id="BBAE61B6F6FAA8C5A31D105AE88BD533" pageId="0" pageNumber="73">
Based on the two newly discovered species, the description should be extended from
<bibRefCitation id="C147552F4FD7A95CE295C9016DC4BDF8" DOI="https://doi.org/10.1163/9789004475588" author="Deeleman-Reinhold, CL" journalOrPublisher="Brill, Leiden" pageId="0" pageNumber="73" refId="B3" refString="Deeleman-Reinhold, CL, 2001. Forest Spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae). Brill, Leiden, DOI: https://doi.org/10.1163/9789004475588" title="Forest Spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae)." url="https://doi.org/10.1163/9789004475588" year="2001">Deeleman-Reinhold (2001)</bibRefCitation>
: the epigyne of
<taxonomicName id="44420698410DE9146B218320F2AE6F3E" lsidName="M. spatulata" pageId="0" pageNumber="73" rank="species" species="spatulata">
<emphasis id="339BA478D08CA0842F7BCCE338F7FDEF" italics="true" pageId="0" pageNumber="73">M. spatulata</emphasis>
</taxonomicName>
Chen &amp; Huang, 2006 with 2 depressions, copulatory ducts relatively long (longer than epigyne) (Fig.
<figureCitation id="4C4E93C0FB892CDAE50F9C6B8627101A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Matidia spatulata, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view; path of copulatory duct marked E dorsal view F lateral view G dorsal view H ventral view. Arrow (F) point at dark ventral abdominal spot in male. Abbreviations: BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.10 mm (equal for A-D); 2 mm (equal for E, F, equal for G, H)." figureDoi="10.3897/zookeys.1062.66845.figure4" httpUri="https://binary.pensoft.net/fig/599508" pageId="0" pageNumber="73">4A-D</figureCitation>
) (vs. one central atrium and short ducts (shorter than epigyne) in all other congeners (e.g.,
<taxonomicName id="4B58AEE947FEFD04B49EA3F9BBB7F14E" lsidName="M. xieqian" pageId="0" pageNumber="73" rank="species" species="xieqian">
<emphasis id="492A47BB1D6110B67E9CE4BB6221CAA3" italics="true" pageId="0" pageNumber="73">M. xieqian</emphasis>
</taxonomicName>
sp. nov.; Fig.
<figureCitation id="42C21A25A8CC5C23F1BFBA0BCAC46C5B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Matidia xieqian sp. nov., female paratype and male holotype, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view; path of copulatory duct marked E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: A = epigynal atrium; BS = bursa; CD = copulatory duct (dashed line showing schematic course of copulatory duct, dorsal); CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.20 mm (equal for A-D); 1 mm (equal for E, F, equal for G, H)." figureDoi="10.3897/zookeys.1062.66845.figure6" httpUri="https://binary.pensoft.net/fig/599510" pageId="0" pageNumber="73">6A-D</figureCitation>
)); an additional exceptional feature in
<taxonomicName id="E4706D4453DE24E1BA3B5EA0AA5CA3FA" lsidName="M. xieqian" pageId="0" pageNumber="73" rank="species" species="xieqian">
<emphasis id="DB2AC25A926E7D9AF66C4E45A4D1FA09" italics="true" pageId="0" pageNumber="73">M. xieqian</emphasis>
</taxonomicName>
sp. nov. is that the cheliceral promarginal teeth are farther from the fang base than the retromarginal ones (vs. promarginal teeth nearer the fang base in all other known
<taxonomicName id="218A6882CC382A2C5CCE9E37A5F8C313" authorityName="Thorell" authorityYear="1878" class="Arachnida" family="Clubionidae" genus="Matidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Matidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="10A2DFA0530951864293B114C5041C6B" italics="true" pageId="0" pageNumber="73">Matidia</emphasis>
</taxonomicName>
species). Both
<taxonomicName id="289429142244749DA317867AE3D98637" lsidName="M. spatulata" pageId="0" pageNumber="73" rank="species" species="spatulata">
<emphasis id="7620CBFCB6D06419D985D3E78E593EDF" italics="true" pageId="0" pageNumber="73">M. spatulata</emphasis>
</taxonomicName>
and
<taxonomicName id="2F515199A73F92297655035F415D4C69" lsidName="M. xieqian" pageId="0" pageNumber="73" rank="species" species="xieqian">
<emphasis id="9200DFAFE90CDC02B173219833B6E60D" italics="true" pageId="0" pageNumber="73">M. xieqian</emphasis>
</taxonomicName>
sp. nov. have anteriorly located bursae (vs. bursae located laterally or posteriorly).
</paragraph>
</subSubSection>
<subSubSection id="72FE3449D4E402434444FFC59CAB430A" pageId="0" pageNumber="73" type="notes">
<paragraph id="296576AF3B4BF0081BBD76C17F6E1512" pageId="0" pageNumber="73">Note.</paragraph>
<paragraph id="CD051ABBFFBE5C9D8EDC2CE81F491EFB" pageId="0" pageNumber="73">
<bibRefCitation id="125E2DE16E8C09E1AB153802D8057199" DOI="https://doi.org/10.1163/9789004475588" author="Deeleman-Reinhold, CL" journalOrPublisher="Brill, Leiden" pageId="0" pageNumber="73" refId="B3" refString="Deeleman-Reinhold, CL, 2001. Forest Spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae). Brill, Leiden, DOI: https://doi.org/10.1163/9789004475588" title="Forest Spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae)." url="https://doi.org/10.1163/9789004475588" year="2001">Deeleman-Reinhold (2001)</bibRefCitation>
considered
<taxonomicName id="E39041BFF78FFC00ACD16C385DEF5BC0" authorityName="Thorell" authorityYear="1878" class="Arachnida" family="Clubionidae" genus="Matidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Matidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="82A45292F62EF5B9955D0B13AF8ABDF8" italics="true" pageId="0" pageNumber="73">Matidia</emphasis>
</taxonomicName>
putatively polyphyletic. In the same work, she established three new, similar closely related genera (
<taxonomicName id="693A0FCE76396E0D1463D8B424EFB791" authorityName="Deeleman-Reinhold" authorityYear="2001" class="Arachnida" family="Clubionidae" genus="Nusatidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusatidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="6EDD97A0872F80EB0E0115C911316204" italics="true" pageId="0" pageNumber="73">Nusatidia</emphasis>
</taxonomicName>
,
<taxonomicName id="AA0825F2D329FD8065D76DC1B9115E49" class="Arachnida" family="Clubionidae" genus="Pristidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pristidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="86ED1FB763D5715BFAFE32712BE61E1F" italics="true" pageId="0" pageNumber="73">Pristidia</emphasis>
</taxonomicName>
, and
<taxonomicName id="2EBFBAE475C7B4845CDBED41CC2C68FA" authorityName="Deeleman-Reinhold" authorityYear="2001" class="Arachnida" family="Clubionidae" genus="Malamatidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Malamatidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="47A5953ABC8D56E07828489B4E05F0C2" italics="true" pageId="0" pageNumber="73">Malamatidia</emphasis>
</taxonomicName>
) to accommodate 12 new species from SE Asia, and placed them in the subfamily
<taxonomicName id="FEA3F6617087A67010AF8CD61FFEC428" lsidName="" pageId="0" pageNumber="73" rank="subFamily" subFamily="Clubioninae">Clubioninae</taxonomicName>
. She also transferred four
<taxonomicName id="BD63D752F7CC370AEF9F543A41ECF7D9" authorityName="Thorell" authorityYear="1878" class="Arachnida" family="Clubionidae" genus="Matidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Matidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="11FD3C8BEA29835DE184BE45CF2E695F" italics="true" pageId="0" pageNumber="73">Matidia</emphasis>
</taxonomicName>
species to
<taxonomicName id="E3DFECA17BF3C6D4E35C7ADE31142B7F" authorityName="Deeleman-Reinhold" authorityYear="2001" class="Arachnida" family="Clubionidae" genus="Nusatidia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nusatidia" order="Araneae" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">
<emphasis id="0413E02B674771F7B6750E732C7FE31B" italics="true" pageId="0" pageNumber="73">Nusatidia</emphasis>
</taxonomicName>
. The similar somatic characters and sympatric distribution strongly suggest close relationships between the four genera. However, the phylogenetic relationships remain unresolved (
<bibRefCitation id="7A9E2340C2674CA5A69C017557E3DF2C" author="Versteirt, V" journalOrPublisher="Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Entomologie" pageId="0" pageNumber="73" pagination="75 - 107" refId="B19" refString="Versteirt, V, Baert, L, Jocque, R, 2010. New genera and species of canopy living Clubionidae (Araneae) from Papua New Guinea. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Entomologie 80: 75 - 107" title="New genera and species of canopy living Clubionidae (Araneae) from Papua New Guinea." volume="80" year="2010">Versteirt et al. 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>