302 lines
40 KiB
XML
302 lines
40 KiB
XML
<document id="4B7CF1E4949F29C76EE9DAB027DE00A1" ENCODING="UTF-8" ID-GBIF-Dataset="bd3ade24-f935-4ef2-8998-0f76c0be1431" ModsDocID="z01067p049" checkinTime="1247054979857" checkinUser="thomas" docAuthor="Jie Li, Xianglin Chen & Bosco P. - L. Chan" docDate="2005" docId="C7644B8DBB9E37A8980111BDEE492B3A" docLanguage="en" docName="2005_Li_Chen_Chan_gg2.xml" docOrigin="Zootaxa 1067" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:F5B656F8-34C2-47A6-A0AB-11D2E1E8337C" docTitle="Pseudobagrus gracilis Li, Chen & Chan, 2005, sp. nov." docType="treatment" docVersion="12" lastPageNumber="56" masterDocId="484FE46A76D8FF7E64FAAE4C6E6D6A52" masterDocTitle="A new species of Pseudobagrus (Teleostei: Siluriformes: Bagridae) from southern China." masterLastPageNumber="57" masterPageNumber="49" pageNumber="50" updateTime="1701314227365" updateUser="plazi">
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<mods:mods id="F62DDF707A22938986B9A84B0EB9D5D3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="38B3929E337DA6BEFC7251FEFDD2FCDE">
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<mods:title id="C051DC549819AD1151DDDAD37B6E5525">A new species of Pseudobagrus (Teleostei: Siluriformes: Bagridae) from southern China.</mods:title>
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</mods:titleInfo>
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<mods:name id="EC513C9012892C4D9CFEC44C8D3607D1" type="personal">
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<mods:role id="9957E8B9D67B9B4551E9B86143F50D7F">
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<mods:roleTerm id="3DD2B7D2B2462809D3BAA49AF9ED4343">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F3C161D7808E46DABA28FE2ECE64A203">Jie Li</mods:namePart>
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</mods:name>
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<mods:name id="A0401ECD8CC5925A51183A5EDB3AD05D" type="personal">
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<mods:role id="4590F955138E3EE7382E015B9BC35495">
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<mods:roleTerm id="B08017C3663E042F30484587FE32214B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1E7BD60E1F00E95D155033CEA34514E7">Xianglin Chen</mods:namePart>
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</mods:name>
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<mods:name id="2DA0CA14415AD17ABAECE571E20E4900" type="personal">
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<mods:role id="649BB30E1922FED2E2DCD6803FF4BC2C">
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<mods:roleTerm id="E42DE42D28100588CC1A67CF2F63F604">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="60A0F1BB7886CEA0EA171E3F11B3C247">Bosco P. - L. Chan</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="558847343A375E292B96E488AB3D055B">text</mods:typeOfResource>
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<mods:relatedItem id="828FD43BA3CC0857D9054FEB72B2F9A7" type="host">
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<mods:titleInfo id="24412D61B2AEA2AB7A5ECB86671FDA80">
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<mods:title id="07FB550B8885078564EA991A623A1EA2">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="8E7DE012D432F3AF896CC9822EE1E970">
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<mods:date id="ECCA50C8EA34DECFDB95AA3C05E34771">2005</mods:date>
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<mods:detail id="7F4F528EEED2F0C833C6A8B3757E1FCD" type="volume">
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<mods:number id="CF77F72BBA779F64DC74572D223081BB">1067</mods:number>
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</mods:detail>
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<mods:extent id="0029367CEF0BD0F814CD128841BC1A18" unit="page">
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<mods:start id="B1DFB8410B5399B9B25EC33C2B892AB3">49</mods:start>
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<mods:end id="13445F112784CAFDB3D5573BDBB951F2">57</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="BF8E22CA6CA694A70732DDA6D09CCED1">
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<mods:url id="400C87EF46706A6FFDF7A0753C61B282">http://www.zoobank.org/urn:lsid:zoobank.org:pub:F5B656F8-34C2-47A6-A0AB-11D2E1E8337C</mods:url>
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</mods:location>
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<mods:classification id="828D68DBC5B737CFEA2B33891F259E13">journal article</mods:classification>
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<mods:identifier id="5A9BD4BAA797AEB7273633CB7FC43FE4" type="Plazi-Custom">z01067p049</mods:identifier>
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</mods:mods>
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<treatment id="C7644B8DBB9E37A8980111BDEE492B3A" ID-DOI="http://doi.org/10.5281/zenodo.6267283" ID-GBIF-Taxon="125136392" ID-Zenodo-Dep="6267283" LSID="urn:lsid:plazi:treatment:C7644B8DBB9E37A8980111BDEE492B3A" httpUri="http://treatment.plazi.org/id/C7644B8DBB9E37A8980111BDEE492B3A" lastPageNumber="56" pageNumber="50">
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<subSubSection id="1D7EB776C45DDEF6A5BAFC6F00D0BEA7" type="nomenclature">
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<paragraph id="CE88444B38CBC72EC66159B96CB3FDDC" pageNumber="50">
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<taxonomicName id="1579C6A340E93716A20B0071C22F195D" ID-CoL="4NPDB" LSID-ZBK="urn:lsid:zoobank.org:act:540277B6-207B-4119-8CAF-333AE24C632A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus gracilis Li, Chen & Chan 2005" order="Siluriformes" pageNumber="50" phylum="Chordata" rank="species" species="gracilis">Pseudobagrus gracilis</taxonomicName>
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<taxonomicNameLabel id="86B8C59389B50C257E2FAFE3C51C8C13" rank="species">sp. nov.</taxonomicNameLabel>
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(Fig. 1)
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</paragraph>
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</subSubSection>
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<subSubSection id="1B9E2D9F1F153089C384C8F0C55688A4" type="materials_examined">
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<paragraph id="5CF7E454D105D80BFF6AD85A26ECA62D" pageNumber="50">Measurements were taken on 16 specimens (62.80-203.76mm TL, 54.98-173.07mm SL) collected from Wushui drainage, Shaoguan City, Guangdong Province and Lijiang drainage, Guilin City, Guangxi Province; tributaries of the Beijiang and Xijiang rivers, respectively. Both rivers are tributaries of the Zhujiang, the largest catchment basin in southern China draining into the South China Sea. Collecting localities are shown in Fig. 2.</paragraph>
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</subSubSection>
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<subSubSection id="758CAD807D0104DC49D1832E93166D5A" type="nomenclature">
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<paragraph id="35AC1311D4C3154438A7A6578AE4A6EF" pageNumber="51">
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<materialsCitation id="50BF40947B315B33C9CDBA4D54C38CA4" ID-GBIF-Occurrence="1416639112" collectingDate="2004-10-26" collectionCode="SCNU" collectorName="Li Jie" country="China" latitude="29.923056" location="Guilin City, Zhujiang basin, Lijiang drainage" longitude="118.47" specimenCode="SCNU 0410002" stateProvince="Guangxi Province" typeStatus="Holotype">
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<typeStatus id="5DD6036290A3FEBBEC87CBF27DB03D76">Holotype</typeStatus>
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:
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<specimenCode id="64436060075FAC6A93799B8B6744164F">
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<collectionCode id="FD377076595CD0867825B2E39B77D01A">SCNU</collectionCode>
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0410002
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</specimenCode>
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, 203.76mm TL, 173.07mm SL;
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<collectingCountry id="5361749E784F6037F8EB838ADF278435">China</collectingCountry>
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:
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<collectingRegion id="8990529810F47187BA8EDE433C6FF84D">Guangxi Province</collectingRegion>
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,
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<location id="515AEB457AA02350F640548C7DB1D29A" LSID="urn:lsid:plazi:treatment:C7644B8DBB9E37A8980111BDEE492B3A:515AEB457AA02350F640548C7DB1D29A" country="China" name="Guilin City, Zhujiang basin, Lijiang drainage" stateProvince="Guangxi Province">Guilin City, Zhujiang basin, Lijiang drainage</location>
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;
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<geoCoordinate id="BACAFBAAEC16960B9D2C5550D2709780">29° 55' 23 N</geoCoordinate>
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<geoCoordinate id="860F7989CE9EF684BA8C15BA96ED9195">118° 28' 12 E</geoCoordinate>
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;
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<collectorName id="802E9CAF4864DB71BD3C972D77830506">Li Jie</collectorName>
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,
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<date id="B3BB05D9571476E991D35B6268C84F6B" day="26" month="10" value="2004-10-26" year="2004">
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<collectingDate id="7694B44713C55CC8B64197A62D76D4F2" day="26" month="10" value="2004-10-26" year="2004">26 October 2004</collectingDate>
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</date>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="9DA0BAE8F91EDEF2AFF2F83E6316DBF7" pageNumber="51">
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<materialsCitation id="5F94E63644BF40758D7244EF4E6DB7FB" ID-GBIF-Occurrence="1416639108" collectionCode="SCNU" collectorName="Li Jie" country="China" latitude="29.923056" location="Guilin City, Zhujiang basin, Lijiang drainage" longitude="118.47" specimenCode="SCNU 0410007-0410009" stateProvince="Guangxi Province" typeStatus="Paratypes">
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<typeStatus id="AFBBC3AACCFA409A5C9777AEE0B0E506">Paratypes</typeStatus>
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:
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<specimenCode id="19BEBBE0E3BFBED1BE2F166AAE76F460">
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<collectionCode id="677730561FDC78820031E104D17C6BBE">SCNU</collectionCode>
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0410007-0410009
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</specimenCode>
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(3), 101.91-125.41mm TL, 86.85-110.31 mm SL; collection locality and collector as holotype.
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</materialsCitation>
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<materialsCitation id="99C3CDD2257321CB67732466EE18F7C7" ID-GBIF-Occurrence="1416639111" collectingDate="2004-11-07" collectionCode="SCNU, UF" collectorName="Li Jie" country="China" latitude="24.8" location="Shaoguan City, Zhujiang basin, Wushui drainage" longitude="113.583336" specimenCode="SCNU 0411008, UF 149607, SCNU 0411010" stateProvince="Guangdong Province" typeStatus="Paratypes">
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<specimenCode id="FD5B846CC6C05A1491E343BED58692FF">
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<collectionCode id="64AFD63E16193D901A4448E17DE7821F">SCNU</collectionCode>
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0411008
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</specimenCode>
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,
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<specimenCode id="CFC36DB2423A94A2E6748CCB491046C2">
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<collectionCode id="0B172C7F55CFE224C928C94BD315E985">UF</collectionCode>
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149607
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</specimenCode>
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(115.22mm TL, 97.22mm SL) and
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<specimenCode id="0E8A243B3347D18E604224CEC1A13C7F">
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<collectionCode id="254A299E65E8C60C5894094B4D06F724">SCNU</collectionCode>
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0411010
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</specimenCode>
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(3), 78.92-131.23mm TL, 67.46-92.55 mm SL;
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<collectingCountry id="27C1F364A1268B688037D75D7C1DCC8B">China</collectingCountry>
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:
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<collectingRegion id="5088F5CF073F69EA505FC6794BE0DD15">Guangdong Province</collectingRegion>
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,
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<location id="75616BD8901F1E0FD7B583483BD0247E" LSID="urn:lsid:plazi:treatment:C7644B8DBB9E37A8980111BDEE492B3A:75616BD8901F1E0FD7B583483BD0247E" country="China" latitude="24.8" longitude="113.583336" name="Shaoguan City, Zhujiang basin, Wushui drainage" stateProvince="Guangdong Province">Shaoguan City, Zhujiang basin, Wushui drainage</location>
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;
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<geoCoordinate id="AD2F3EF7298375252E355D29F81BC547" direction="north" orientation="latitude" value="24.8">24°48' N</geoCoordinate>
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<geoCoordinate id="ABF9BE9669608FD5DB8E13D51A855293" direction="east" orientation="longitude" value="113.583336">113° 35' E</geoCoordinate>
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;
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<collectorName id="5DED582AEB910B85D90896273B197959">Li Jie</collectorName>
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,
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<date id="8A66C0ECB3C893A317F14307C998AA09" day="07" month="11" value="2004-11-07" year="2004">
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<collectingDate id="87F742332CC09BD7B5837D7A68B3A384" day="07" month="11" value="2004-11-07" year="2004">7 November 2004</collectingDate>
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</date>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="4A7F197D9DBA4415455FA4DBC4040087" type="diagnosis">
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<paragraph id="54371D0F2F35A792DB7AC22A19CD2875" pageNumber="51">
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Diagnosis. Among southern Chinese congeners,
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<taxonomicName id="52CC1643D7F217A3BE9C7B90F179447E" LSID-ZBK="urn:lsid:zoobank.org:act:540277B6-207B-4119-8CAF-333AE24C632A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus gracilis Li, Chen & Chan 2005" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="gracilis">Pseudobagrus gracilis</taxonomicName>
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most closely resembles
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<taxonomicName id="717C8274CB360376EC5D1EFBB7E61A36" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
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and
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<taxonomicName id="A934AB9041538DBDD7A4D4E43A6CEC88" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="ussuriensis">P. ussuriensis</taxonomicName>
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, but differs from topotypes of
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<taxonomicName id="97B34951B379112E0AAD4C3639AEC40D" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
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and
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<taxonomicName id="A6B21C12E3BE14F29596705A8D2ABFCE" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="ussuriensis">P. ussuriensis</taxonomicName>
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in possessing a larger elliptical eye (19.8-24.4% HL vs. 11.5-12.7% HL in
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<taxonomicName id="8C1132514A7A71F27268FA6690934F5E" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
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and 12.4-13.9% HL in
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<taxonomicName id="6C04C9D1C1DDAA0D47ACEE76D3E97766" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="ussuriensis">P. ussuriensis</taxonomicName>
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), lower body depth (11.2-16.4% SL vs. 14.2-16.1% SL in
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<taxonomicName id="264BFD5C88CBB22B3280561E4758818B" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
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and 14.8-18.4% SL in
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<taxonomicName id="9762B034F5FCB03C812DC9085BDBB8BA" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="ussuriensis">P. ussuriensis</taxonomicName>
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), lower caudal peduncle depth (5.2-8.0% SL vs. 6.9-7.9% SL in
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<taxonomicName id="D6C5681974646E14874A94B6D1A212DC" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
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and 6.6-8.7% SL in
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<taxonomicName id="BB2A0703A1D8822FF45B94642D3EE8E6" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="ussuriensis">P. ussuriensis</taxonomicName>
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), fewer serrae on the posterior edge of the pectoral fin spine (10-11 vs. 14 in
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<taxonomicName id="2DC59571A8FA6F5955F40A110C8E1E59" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
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and 12-14 in
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<taxonomicName id="15958E692F6E3C39AE6809ACD4725919" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="ussuriensis">P. ussuriensis</taxonomicName>
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), and fewer vertebrae (5+42-43 vs. 5+46-47 in
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<taxonomicName id="E3A65E10714C43BF48F2476070422AB3" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
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and 5+44 in
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<taxonomicName id="096903F362D53F7734685A0E1C074BFD" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="ussuriensis">P. ussuriensis</taxonomicName>
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).
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</paragraph>
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<paragraph id="E20509E7F176CCC10D72C2DF727D4C23" pageNumber="51">
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Additionally, the new species shows the following differences from
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<taxonomicName id="D7AC4098BD06EDEB01414B0637D15412" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
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: posterior margin of second dorsal spine lightly serrated (vs. smooth) and subequal caudal lobes, with upper lobe slighter longer (vs. symmetrical lobes of equal length). It also differs from
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<taxonomicName id="CDFBF2B0E248483331CE889C9A2CDF4A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="ussuriensis">P. ussuriensis</taxonomicName>
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in having more gill rakers (11-13 vs. 10-11) and a caudal fin without black margin (vs. with prominent black margin). The new species also has a wider distance between the supraoccipital process and the first dorsal spine than does
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<taxonomicName id="1A547FB3AFB61FA4E49E9BA3E0F98B14" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
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and
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<taxonomicName id="93488F6D5D8F51E525027434C3F85D18" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="ussuriensis">P. ussuriensis</taxonomicName>
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. Major diagnostic features of
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<taxonomicName id="7FFA4F0E2225D0DDC2E432C5E30F225D" LSID-ZBK="urn:lsid:zoobank.org:act:540277B6-207B-4119-8CAF-333AE24C632A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus gracilis Li, Chen & Chan 2005" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="gracilis">Pseudobagrus gracilis</taxonomicName>
|
|
in comparison with
|
|
<taxonomicName id="708A68EA3B705E46D715739C17DEC7A9" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
|
|
and
|
|
<taxonomicName id="4889A7DBC6ED891F361E96629A1ECC46" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="51" phylum="Chordata" rank="species" species="ussuriensis">P. ussuriensis</taxonomicName>
|
|
are summarized in Table 1.
|
|
</paragraph>
|
|
<paragraph id="046A774480B42E037A9452DC2FBC302A" pageNumber="52">
|
|
<taxonomicName id="02F8B0BFC2DCFB658FF42BC6E1BD7E87" LSID-ZBK="urn:lsid:zoobank.org:act:540277B6-207B-4119-8CAF-333AE24C632A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus gracilis Li, Chen & Chan 2005" order="Siluriformes" pageNumber="52" phylum="Chordata" rank="species" species="gracilis">Pseudobagrus gracilis</taxonomicName>
|
|
can be readily distinguished from other species of
|
|
<taxonomicName id="DBD468655E1DBAACE6FF90D4EEE3D5BA" LSID-ZBK="urn:lsid:zoobank.org:act:BD503EEA-AE89-4240-A85C-3EE5D3AE890B" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus Bleeker 1859:257" order="Siluriformes" pageNumber="52" phylum="Chordata" rank="genus">Pseudobagrus</taxonomicName>
|
|
of southeastern China as follows: from
|
|
<taxonomicName id="0BF2A8CA24BC5F3BBC9CC0D6F51D0D9F" LSID-ZBK="urn:lsid:zoobank.org:act:3BC08BB0-2816-4EC2-979D-363EF4C370AC" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus ondon Shaw 1930:111" order="Siluriformes" pageNumber="52" phylum="Chordata" rank="species" species="ondon">P. ondon Shaw</taxonomicName>
|
|
in having the anterior edge of the pectoral spine smooth (vs. serrated); from
|
|
<taxonomicName id="01224F8940B61C531C3CBCFF8AAD5D4C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="52" phylum="Chordata" rank="species" species="pratti">P. pratti (Gunther)</taxonomicName>
|
|
in having a slightly emarginate caudal fin (vs. moderately forked); from
|
|
<taxonomicName id="4D907B84AE2E02E386C8DE53C023212E" LSID-ZBK="urn:lsid:zoobank.org:act:37CF73C6-CFDC-498C-89FF-30665A59AAF4" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Macrones Pseudobagrus tenuis Günther 1873:244" order="Siluriformes" pageNumber="52" phylum="Chordata" rank="species" species="tenuis">P. tenuis (Gunther)</taxonomicName>
|
|
in having an emarginate caudal fin (vs. round fin with prominent white margin); from
|
|
<taxonomicName id="0C553DBF7327E831670ADE34440BA289" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="52" phylum="Chordata" rank="species" species="truncatus">P. truncatus (Regan)</taxonomicName>
|
|
in having more vertebrae (5+42-43 vs. 5+39-40), posterior edge of second dorsal spine slightly serrated (vs. smooth), more gill rakers (11-13 vs. 8-10), and gonopore slightly closer to origin of anal fin than to distal origin of pelvic fin (vs. slightly closer to distal origin of pelvic fin than to origin of anal fin). Figure 3 depicts the different caudal fin shapes of various Chinese
|
|
<taxonomicName id="C0057DE56D769674BC9F631A2171015E" LSID-ZBK="urn:lsid:zoobank.org:act:BD503EEA-AE89-4240-A85C-3EE5D3AE890B" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus Bleeker 1859:257" order="Siluriformes" pageNumber="52" phylum="Chordata" rank="genus">Pseudobagrus</taxonomicName>
|
|
species.
|
|
</paragraph>
|
|
<paragraph id="3443436BD0048846A67162665219A1B3" pageNumber="53">
|
|
In comparison with the morphologically most similar
|
|
<taxonomicName id="2B56D45DA84EB001B5AAE8669D82BD41" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="53" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
|
|
, the new species has the following distinctive osteological characters (Fig. 4): the neck of the mesethmoid is elongated (vs. short and stout in
|
|
<taxonomicName id="E0C85ACA05250BFD3236176B7DAFFA85" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="53" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
|
|
); frontal edge smooth (vs. rough); both the anterior and posterior fontanelle well developed (vs. not well developed, with a longer anterior fontanelle and a fused posterior fontanelle); supraoccipital process short, distally not forked, and far from the first dorsal spine (vs. supraoccipital process long, distally forked and close to the first dorsal spine).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="AA0501D697BC08B4B9C66F1402B12A35" type="description">
|
|
<paragraph id="3DFDBB71E7F0F1C9E24EACB7EDF48C18" pageNumber="54">Description. Body elongated and cylindrical, laterally compressed posterior to the pelvic fin. Head broad and wide, dorsally flattened, covered by smooth skin. Snout rounded. Upper jaw protruded, mouth subterminal; large, fleshy lips present, joined at corner of mouth. Upper jaw teeth villiform, forming dentary patches. Eyes large, elliptical, located on anterior half of head; visible dorsally, but not ventrally. Nostrils widely separated, tubular anterior pair located anterolaterally on snout tip; fleshy posterior pair located dorsally immediately before eye. Four pairs of barbels relatively short and white; nasal barbels nearly reaching to anterior edge of eye; maxillary barbels reaching to posterior edge of eye, but not the branchiostegal membrane; outer mandibular barbels reaching to center of eye; inner mandibular barbels approximately half the length of the outer mandibular barbel. Gill opening wide, gill membranes fused, not connected with gular fold.</paragraph>
|
|
<paragraph id="6CCB1FB174DEF2F5C918D5E57E533C44" pageNumber="54">Body naked, lateral line complete and straight. Dorsal fin short, origin of dorsal fin between pectoral and pelvic fins, closer to tip of snout than to origin of adipose fin. First dorsal spine tiny and subcutaneous, second dorsal spine long, almost equal to body height, anterior edge smooth, posterior edge slightly serrated. Adipose fin moderately long with a rounded distal margin separated from body. Pectoral fin spine slightly shorter than second dorsal spine, anterior edge smooth but with serrations on posterior edge. Pelvic fin much closer to anal fin than to pectoral fin, but distal margin not reaching anal fin. Origin of anal fin behind origin of adipose fin, the length of the anal fin shorter than the length of the adipose fin. Caudal fin slightly emarginated, both lobes of juveniles having equal length, but the upper lobe is slightly longer in the adult. Some individuals have white on the tips of the lobes. Gonopore slightly closer to anal fin than to pelvic fin; urogenital papillae not reaching anal fin.</paragraph>
|
|
<paragraph id="506316D05AB169D0B74B3E25A5EFA9C0" pageNumber="54">
|
|
Proportional measurements. Morphometric data of
|
|
<taxonomicName id="3F413ECF56C1898B450098BDA2C83787" LSID-ZBK="urn:lsid:zoobank.org:act:540277B6-207B-4119-8CAF-333AE24C632A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus gracilis Li, Chen & Chan 2005" order="Siluriformes" pageNumber="54" phylum="Chordata" rank="species" species="gracilis">Pseudobagrus gracilis</taxonomicName>
|
|
are listed in Table 2.
|
|
</paragraph>
|
|
<paragraph id="0A1B3DB742EA9182A3D467B5331B68F4" pageNumber="54">Counts (n = 10). Branchiostegal rays: 8-12. Gill rakers: 11-13. Fin ray counts: dorsal II -7; pectoral I-7; pelvic i-5; anal 17-21 (average 19). Vertebral counts: 5+42-43. Swimbladder with one chamber.</paragraph>
|
|
<paragraph id="4D9B718C613017BDFA0658E8DE893A8A" pageNumber="54">Coloration The dorsum is dark grey, and the underside is whitish in live specimens. Alcohol-preserved specimens are greyish-brown dorsally, and flanks have a lighter tinge.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E117AED210A42846F712624E0CF6882C" type="distribution">
|
|
<paragraph id="AB54FEC7C4893A750AD2AE5AE43DA1AB" pageNumber="54">
|
|
Distribution.
|
|
<taxonomicName id="66478E3156C33D12E826A98225956EC4" LSID-ZBK="urn:lsid:zoobank.org:act:540277B6-207B-4119-8CAF-333AE24C632A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus gracilis Li, Chen & Chan 2005" order="Siluriformes" pageNumber="54" phylum="Chordata" rank="species" species="gracilis">Pseudobagrus gracilis</taxonomicName>
|
|
has been collected from the Wushui drainage of the Beijiang drainage in Guangdong Province and the Lijiang drainge of the Xijiang drainage in Guangxi Province; there are also literature records (misidentified as
|
|
<taxonomicName id="1AE03DE00E893A623C1BC1F183CCC37C" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="54" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
|
|
) from the Dongjiang drainage in Guangdong Province (Ye, 1991). These drainages are all part of the Zhujiang catchment basin in southern China, which discharges into the South China Sea.
|
|
<taxonomicName id="5DD385C6F4575515C299AA502B36C5DF" LSID-ZBK="urn:lsid:zoobank.org:act:540277B6-207B-4119-8CAF-333AE24C632A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus gracilis Li, Chen & Chan 2005" order="Siluriformes" pageNumber="54" phylum="Chordata" rank="species" species="gracilis">Pseudobagrus gracilis</taxonomicName>
|
|
appears to be a Zhujiang endemic.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="96649BBC4C261F19219730BC746D8628" type="biology_ecology">
|
|
<paragraph id="D8440EE94EED8B5333CEFAE1FAD02A47" pageNumber="54">
|
|
Ecology.
|
|
<taxonomicName id="196148C77C5CFB38ABF3B8348FBE4DD2" LSID-ZBK="urn:lsid:zoobank.org:act:540277B6-207B-4119-8CAF-333AE24C632A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus gracilis Li, Chen & Chan 2005" order="Siluriformes" pageNumber="54" phylum="Chordata" rank="species" species="gracilis">Pseudobagrus gracilis</taxonomicName>
|
|
is found in medium to large rivers in lowland areas. These waterways have a rocky substrate with moderate or fast current, and water clarity is generally good. Sampling data suggests that
|
|
<taxonomicName id="CCC1A4D81DE3294C39749B51C3A77751" LSID-ZBK="urn:lsid:zoobank.org:act:540277B6-207B-4119-8CAF-333AE24C632A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus gracilis Li, Chen & Chan 2005" order="Siluriformes" pageNumber="54" phylum="Chordata" rank="species" species="gracilis">P. gracilis</taxonomicName>
|
|
may be nocturnal as are many other catfishes, but its ecology and biology are otherwise unknown.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="2B670FF3D3FDF18897BC2B85FDCFC3F8" type="etymology">
|
|
<paragraph id="B434603D46B9389B30F8BF6F5BE02ABD" pageNumber="55">Etymology Gracilis is Latin for slender. The species is named for its elongated and thin body form (see Fig. 1).</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="ABCDD6BF528B731D3D5C31CFD23AF7D7" type="discussion">
|
|
<paragraph id="A5887BAB4A628D9787F8A17A460D619C" pageNumber="55">Discussion</paragraph>
|
|
<paragraph id="5E5C46362801367B42034D669A3157C9" pageNumber="55">
|
|
The new species has been confused or misidentified with two other
|
|
<taxonomicName id="72D34123C399CAF98DB5DDE9E7C5EE2B" LSID-ZBK="urn:lsid:zoobank.org:act:BD503EEA-AE89-4240-A85C-3EE5D3AE890B" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus Bleeker 1859:257" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="genus">Pseudobagrus</taxonomicName>
|
|
species since records have been made for freshwater fishes in South China. Yue (1981) recognised
|
|
<taxonomicName id="B9041CEF6BAB67B8A99DAC135B9303AA" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="species" species="pratti" subGenus="Leiocassis">Pseudobagrus (Leiocassis) pratti</taxonomicName>
|
|
in Guangxi Province, but from the description and line drawing the species seems to be
|
|
<taxonomicName id="478BE097762FDE79E582D82C713306BC" LSID-ZBK="urn:lsid:zoobank.org:act:540277B6-207B-4119-8CAF-333AE24C632A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus gracilis Li, Chen & Chan 2005" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="species" species="gracilis">P. gracilis</taxonomicName>
|
|
. Pan (1984) collected an elongated bagrid from the Beijiang drainage of the Zhujiang basin and also called it
|
|
<taxonomicName id="728EAA14A3F69199C9C2405B84FAF839" family="Bagridae" genus="Leiocassis" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="species" species="pratti">Leiocassis pratti</taxonomicName>
|
|
, but did not provide descriptions or figures. Cheng and Zheng (1987) and Zhu (1995) listed
|
|
<taxonomicName id="47BAAF696F5F76761A3AA86CF5A60D94" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
|
|
as native to southeastern China, from Danshui River of Taiwan, Lingjiang and Oujiang of Zhejiang Province, Xianjiang of Hunan Province, and the Zhujiang basin. Ye (1991) also reported
|
|
<taxonomicName id="524FF1619F41BFBC9DABA3BC4ED2937A" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
|
|
from the Dongjiang drainage of the Zhujiang basin. Gao (1991) and Zheng and Dai (1999) likewise failed to recognise these cryptic species. It is likely that the authors failed to distinguish these species within this large geographic area.
|
|
</paragraph>
|
|
<paragraph id="B02A50EDB722F092D2AF124C3529F1CF" pageNumber="55">
|
|
We examined the literature cited above and examined specimens of
|
|
<taxonomicName id="C0D7F8251DB059B1FB2341F10D67F219" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="species" species="pratti">P. pratti</taxonomicName>
|
|
Pan (1984) collected in the Beijiang and topotypes of
|
|
<taxonomicName id="F79F47EF127E4157272004B09711A941" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis Oshima</taxonomicName>
|
|
, collected in Taiwan. Evidence presented here suggests that most information in the literature referring to
|
|
<taxonomicName id="99A5BB48CC00372528764D35F3FAA5C9" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="species" species="pratti">P. pratti</taxonomicName>
|
|
and
|
|
<taxonomicName id="7100713156D754DA97A52279D58D4357" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
|
|
from South China likely pertains to
|
|
<taxonomicName id="BDDA4E470F556CFC5365FB83C8D144AC" LSID-ZBK="urn:lsid:zoobank.org:act:540277B6-207B-4119-8CAF-333AE24C632A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus gracilis Li, Chen & Chan 2005" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="species" species="gracilis">P. gracilis</taxonomicName>
|
|
, which apparently is endemic to the Zhujiang basin.
|
|
<taxonomicName id="EE93E6EFCE47685A23A6D44DDA3D14FC" LSID-ZBK="urn:lsid:zoobank.org:act:540277B6-207B-4119-8CAF-333AE24C632A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus gracilis Li, Chen & Chan 2005" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="species" species="gracilis">P. gracilis</taxonomicName>
|
|
and
|
|
<taxonomicName id="E7E9A33B220D5379691C8CD80A4851AD" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="species" species="pratti">P. pratti</taxonomicName>
|
|
are two distinct species, easily separable by caudal fin shape (emarginate in
|
|
<taxonomicName id="E626CD602CE9DCE4024B63534BC07D77" LSID-ZBK="urn:lsid:zoobank.org:act:540277B6-207B-4119-8CAF-333AE24C632A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus gracilis Li, Chen & Chan 2005" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="species" species="gracilis">P. gracilis</taxonomicName>
|
|
vs. deeply forked in
|
|
<taxonomicName id="404A7BDB0197455ECF1B9A7804E6FDE9" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="species" species="pratti">P. pratti</taxonomicName>
|
|
, see Fig. 3) and distribution (Zhujiang vs. upper Yangtze), and distinct from
|
|
<taxonomicName id="96458838972F02F77BED9CFA4C4AC451" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="55" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
|
|
by meristic counts, morphometrics, osteology (see Diagnosis and Description), and distribution (vs. Danshui River, Taiwan Island).
|
|
</paragraph>
|
|
<paragraph id="332A037AAD3E980B7D332F2C5A6E430B" pageNumber="56">Systematic position</paragraph>
|
|
<paragraph id="AC6ED92B39325F1F648C7FB8E92F9033" pageNumber="56">
|
|
The development of fontanelle may well be indicative of an evoluntionary trend in the genus
|
|
<taxonomicName id="0722920C35FE4CA4F86D288779EFD6B6" LSID-ZBK="urn:lsid:zoobank.org:act:BD503EEA-AE89-4240-A85C-3EE5D3AE890B" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus Bleeker 1859:257" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="genus">Pseudobagrus</taxonomicName>
|
|
, from two well-developed fontanelle to the disappearance of the posterior fontanelle.
|
|
<taxonomicName id="EC3E676EEF07E7A0B73C70CCBB7F76CE" LSID-ZBK="urn:lsid:zoobank.org:act:540277B6-207B-4119-8CAF-333AE24C632A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus gracilis Li, Chen & Chan 2005" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="gracilis">P. gracilis</taxonomicName>
|
|
has well-developed fontanelle indicating that it is probably a primitive member of the genus whereas
|
|
<taxonomicName id="B4E318291F57FA34AF04612888D876DE" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
|
|
has a reduced posterior fontanelle indicating evolutionary advancement. Another putative evoluntionary trend is the increase in the length of the basal bone of the dorsal spine.
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</paragraph>
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|
<paragraph id="57A9303B7E0EBB11D29A7B02398E25B4" pageNumber="56">
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|
Osteological examination of various
|
|
<taxonomicName id="7FEDB40A911D70C9355523F73D0A0F2F" LSID-ZBK="urn:lsid:zoobank.org:act:BD503EEA-AE89-4240-A85C-3EE5D3AE890B" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus Bleeker 1859:257" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="genus">Pseudobagrus</taxonomicName>
|
|
species suggests that the new species has many primitive characters most comparable to those of
|
|
<taxonomicName id="5826604EDF44D003D63D1674398CA443" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="albomarginatus">P. albomarginatus (Rendahl)</taxonomicName>
|
|
, including the possession of well-developed fontanelle and moderately developed supraoccipital bone.
|
|
<taxonomicName id="D03CEF93DA4D959BE3CD686F27EA376F" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
|
|
, on the other hand, has a diminutive posterior fontanelle and a well-developed supraoccipital bone, with a pronounced process along the median line. More work is needed to ascertain whether the insular
|
|
<taxonomicName id="68E4166D1A971D9DA129F94B25172E83" LSID-ZBK="urn:lsid:zoobank.org:act:2AE4F2AD-6150-4A8C-BF3C-B08CADFD2E8C" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus adiposalis Oshima 1919:181" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="adiposalis">P. adiposalis</taxonomicName>
|
|
evolved from
|
|
<taxonomicName id="CD00274BC4DBF82D55E357D9F2708ADF" LSID-ZBK="urn:lsid:zoobank.org:act:540277B6-207B-4119-8CAF-333AE24C632A" family="Bagridae" genus="Pseudobagrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Pseudobagrus gracilis Li, Chen & Chan 2005" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="gracilis">P. gracilis</taxonomicName>
|
|
on the mainland.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
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</document> |