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<mods:title id="5A24DF37E78793B6BF50A590856EE11B">An updated synthesis of the Geophilomorpha (Chilopoda) of Asian Russia</mods:title>
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<mods:affiliation id="B052BBAB7F72E293D0FD77F1C1D225AF">Dipartimento di Biologia, Universita di Padova, via U. Bassi 58 b, 35131 Padova, Italy</mods:affiliation>
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30.
<taxonomicName id="FF76AAC8413152789E81384B4200C31E" ID-CoL="3BGL9" LSID="C71816E0-2517-590B-B5F0-0F7ECC61766F" authority="Takakuwa, 1937 *" authorityName="Takakuwa" authorityYear="1937" class="Chilopoda" family="Schendylidae" genus="Escaryus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escaryus koreanus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="koreanus">Escaryus koreanus Takakuwa, 1937 *</taxonomicName>
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<taxonomicName id="9D50870562C671BA50B34C083CC69DBB" authorityName="Takakuwa" authorityYear="1937" class="Chilopoda" family="Schendylidae" genus="Escaryus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escaryus koreanus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="koreanus">Escaryus koreanus</taxonomicName>
<bibRefCitation id="CE7C2FB02C3825DD341CDB2837AB9E7B" author="Takakuwa, Y" journalOrPublisher="Zoological Magazine" pageId="0" pageNumber="17" pagination="297 - 299" refId="B95" refString="Takakuwa, Y, 1937a. Eine neue Escaryus -Art aus Korea. Zoological Magazine 49: 297 - 299" title="Eine neue Escaryus - Art aus Korea." volume="49" year="1937 a">Takakuwa 1937a</bibRefCitation>
: 297.
</paragraph>
<paragraph id="CB1E96AFDFA723B60D95CCE6E59A2863" pageId="0" pageNumber="17">
<taxonomicName id="F895FC554081E164667E08AAD0579485" authorityName="Takakuwa" authorityYear="1937" class="Chilopoda" family="Schendylidae" genus="Escaryus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escaryus koreanus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="koreanus">Escaryus koreanus</taxonomicName>
-
<bibRefCitation id="56F086B726B46AD276915CF2B92DBEF4" author="Takakuwa, Y" journalOrPublisher="Zoological Magazine" pageId="0" pageNumber="17" refId="B99" refString="Takakuwa, Y, 1940. Geophilomorpha. In: Okada Y et al. (Eds) Fauna Nipponica, 9, fasc. 8(1), Sanseido, Tokyo, 1-156." title="Geophilomorpha. In: Okada Y et al. (Eds) Fauna Nipponica, 9, fasc. 8 (1), Sanseido, Tokyo, 1 - 156." year="1940">Takakuwa 1940</bibRefCitation>
: 39;
<bibRefCitation id="DBACA5237E9E05FF691D0FE27B876FF7" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Materials of the 4 th All-Union Conference. Baku, 1972. Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="135 - 136" refId="B103" refString="Titova, LP, 1972. Pattern of the distribution of the genus Escaryus (Chilopoda) in the USSR. In: Ghilarov, MS, Ed., Problemy pochvennoi zoologii. Materials of the 4th All-Union Conference. Baku, 1972. Nauka Publisher, Moscow: 135 - 136" title="Pattern of the distribution of the genus Escaryus (Chilopoda) in the USSR." volumeTitle="Problemy pochvennoi zoologii." year="1972">Titova 1972</bibRefCitation>
: 135;
<bibRefCitation id="EF83179701F388F1D0BAF504A52A391A" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="94 - 119" refId="B104" refString="Titova, LP, 1973. New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda). In: Ghilarov, MS, Ed., Ekologiya pochvennykh bespozvonochnykh. Nauka Publisher, Moscow: 94 - 119" title="New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda)." volumeTitle="Ekologiya pochvennykh bespozvonochnykh." year="1973">1973</bibRefCitation>
: 112;
<bibRefCitation id="EE4783A7CA89EE36286EA0B4A7B4C25C" author="Ghilarov, MS" editor="Ghilarov, MS" journalOrPublisher="Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="40 - 59" refId="B52" refString="Ghilarov, MS, Perel, TS, 1973. Complexes of soil invertebrates of coniferous-deciduous forests of Far East as indicator of soil type. In: Ghilarov, MS, Ed., Ekologiya pochvennykh bespozvonochnykh. Nauka Publisher, Moscow: 40 - 59" title="Complexes of soil invertebrates of coniferous-deciduous forests of Far East as indicator of soil type." volumeTitle="Ekologiya pochvennykh bespozvonochnykh." year="1973">Ghilarov and Perel 1973</bibRefCitation>
: 46;
<bibRefCitation id="FD69C9A30697E302240806DB04D21C60" author="Kurcheva, GF" journalOrPublisher="Nauka Publisher, Moscow" pageId="0" pageNumber="17" refId="B59" refString="Kurcheva, GF, 1977. Soil invertebrates of Soviet Far East. Nauka Publisher, Moscow" title="Soil invertebrates of Soviet Far East." year="1977">Kurcheva 1977</bibRefCitation>
: 45;
<bibRefCitation id="F5FFED46F3BD53BA438F85B3D15CFFE5" author="Ganin, GN" journalOrPublisher="Dalnauka, Vladivostok-Khabarovsk" pageId="0" pageNumber="17" refId="B48" refString="Ganin, GN, 1997. Soil Invertebrates of the Ussuri Taiga. Dalnauka, Vladivostok-Khabarovsk" title="Soil Invertebrates of the Ussuri Taiga." year="1997">Ganin 1997</bibRefCitation>
: 105, 109, 112, 114, 121, 124, 126, 128;
<bibRefCitation id="8F65A42FAE84AE307D95031EF47E3B5D" DOI="https://doi.org/10.1134/S1062359006050128" author="Ganin, GN" journalOrPublisher="Biology Bulletin of the Russian Academy of Sciences" pageId="0" pageNumber="17" pagination="498 - 507" refId="B49" refString="Ganin, GN, 2006. Some rules of soil invertebrate community organization (by the Example of Amur Basin Mesofauna). Biology Bulletin of the Russian Academy of Sciences 33 (5): 498 - 507, DOI: https://doi.org/10.1134/S1062359006050128" title="Some rules of soil invertebrate community organization (by the Example of Amur Basin Mesofauna)." url="https://doi.org/10.1134/S1062359006050128" volume="33" year="2006">2006</bibRefCitation>
: 501;
<bibRefCitation id="6EEBC73E69DB407A418BC9C23412F2AE" author="Rybalov, LB" journalOrPublisher="Russian Entomological Journal" pageId="0" pageNumber="17" pagination="77 - 86" refId="B78" refString="Rybalov, LB, 2002. Zonal and landscape changes in soil invertebrate populations in a near-Yenisei River region of middle Siberia and the role of temperature adaptations in the meridional (zonal) distribution of invertebrates. Russian Entomological Journal 11 (1): 77 - 86" title="Zonal and landscape changes in soil invertebrate populations in a near-Yenisei River region of middle Siberia and the role of temperature adaptations in the meridional (zonal) distribution of invertebrates." volume="11" year="2002">Rybalov 2002</bibRefCitation>
: 83;
<bibRefCitation id="243FA63CD746478D6A5579846880056C" author="Vorobiova, IG" journalOrPublisher="Materialy II (XII) Vserossiiskogo soveshchaniya po pochvennoi zoologii. KMK Press, Moscow" pageId="0" pageNumber="17" refId="B115" refString="Vorobiova, IG, Rybalov, LB, Rossolimo, TE, Zalesskaja, NT, 2002. Zonal and landscape distribution of the myriapod fauna and populations (Myriapoda) in the Yenisei River basin. In: Izuchenie, sokhranenie i vosstanovlenie bioraznoobraziya ekosistem na Yeniseiskom ekologicheskom transekte: Zhivotnyi mir, etno-ekologicheskie issledovniya, 2. IEE RAS Publisher, Moscow, 60-71. [In Russian]" title="Zonal and landscape distribution of the myriapod fauna and populations (Myriapoda) in the Yenisei River basin. In: Izuchenie, sokhranenie i vosstanovlenie bioraznoobraziya ekosistem na Yeniseiskom ekologicheskom transekte: Zhivotnyi mir, etno-ekologicheskie issledovniya, 2. IEE RAS Publisher, Moscow, 60 - 71. [In Russian]" year="2002">Vorobiova et al. 2002</bibRefCitation>
: 62;
<bibRefCitation id="A56B076E25B4B520B8A98D0E63C46827" DOI="https://doi.org/10.15298/arthsel.26.1.02" author="Nefediev, PS" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="17" pagination="8 - 14" refId="B69" refString="Nefediev, PS, Tuf, IH, Farzalieva, GS, 2017a. Centipedes from urban areas in southwestern Siberia, Russia (Chilopoda). Part 2. Geophilomorpha. Arthropoda Selecta 26 (1): 8 - 14, DOI: https://doi.org/10.15298/arthsel.26.1.02" title="Centipedes from urban areas in southwestern Siberia, Russia (Chilopoda). Part 2. Geophilomorpha." url="https://doi.org/10.15298/arthsel.26.1.02" volume="26" year="2017 a">Nefediev et al. 2017a</bibRefCitation>
: 11;
<bibRefCitation id="90AB6EF9024370771408AC7430627D97" DOI="https://doi.org/10.15298/arthsel.26.3.02" author="Nefediev, PS" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="17" pagination="217 - 224" refId="B71" refString="Nefediev, PS, Farzalieva, GS, Tuf, IH, 2017c. A preliminary review of the centipede fauna of the Altai State Nature Biosphere Reserve, southwestern Siberia, Russia (Chilopoda: Lithobiomorpha, Geophilomorpha). Arthropoda Selecta 26 (3): 217 - 224, DOI: https://doi.org/10.15298/arthsel.26.3.02" title="A preliminary review of the centipede fauna of the Altai State Nature Biosphere Reserve, southwestern Siberia, Russia (Chilopoda: Lithobiomorpha, Geophilomorpha)." url="https://doi.org/10.15298/arthsel.26.3.02" volume="26" year="2017 c">2017c</bibRefCitation>
: 222;
<bibRefCitation id="5569242D735FDA15205443274E8384BC" DOI="https://doi.org/10.3897/zookeys.741.21936" author="Nefediev, PS" journalOrPublisher="ZooKeys" pageId="0" pageNumber="17" pagination="219 - 254" refId="B72" refString="Nefediev, PS, Farzalieva, GS, Tuf, IH, Nedoev, KK, Niyazov, ST, 2018. Millipede and centipede assemblages on the northern and southern slopes of the lowland Altais, southwestern Siberia, Russia (Diplopoda, Chilopoda). ZooKeys 741: 219 - 254, DOI: https://doi.org/10.3897/zookeys.741.21936" title="Millipede and centipede assemblages on the northern and southern slopes of the lowland Altais, southwestern Siberia, Russia (Diplopoda, Chilopoda)." url="https://doi.org/10.3897/zookeys.741.21936" volume="741" year="2018">2018</bibRefCitation>
: 238;
<bibRefCitation id="69214D2D3BFBA67FAD5495CE7E6E0144" DOI="https://doi.org/10.15421/2017_141" author="Dyachkov, YuV" journalOrPublisher="Ukrainian Journal of Ecology" pageId="0" pageNumber="17" pagination="453 - 456" refId="B31" refString="Dyachkov, YuV, 2017. The first data on centipede (Chilopoda: Geophilomorpha; Lithobiomorpha) fauna of the Katunskiy Biosphere State Nature Reserve, Altai Mts. Ukrainian Journal of Ecology 7 (4): 453 - 456, DOI: https://doi.org/10.15421/2017_141" title="The first data on centipede (Chilopoda: Geophilomorpha; Lithobiomorpha) fauna of the Katunskiy Biosphere State Nature Reserve, Altai Mts." url="https://doi.org/10.15421/2017_141" volume="7" year="2017">Dyachkov 2017</bibRefCitation>
: 454;
<bibRefCitation id="D80A637FB1E5B88724AEE2B5791A9BC7" DOI="https://doi.org/10.15298/arthsel.27.4.04" author="Dyachkov, YuV" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="17" pagination="293 - 299" refId="B40" refString="Dyachkov, YuV, Tuf, IH, 2018. New data on the genus Escaryus Cook et Collins, 1891 (Chilopoda: Geophilomorpha: Schendylidae) from Kazakhstan. Arthropoda Selecta 26 (4): 293 - 299, DOI: https://doi.org/10.15298/arthsel.27.4.04" title="New data on the genus Escaryus Cook et Collins, 1891 (Chilopoda: Geophilomorpha: Schendylidae) from Kazakhstan." url="https://doi.org/10.15298/arthsel.27.4.04" volume="26" year="2018">Dyachkov and Tuf 2018</bibRefCitation>
: 296;
<bibRefCitation id="D4AC367E975D7FD06218EDDAB09657AB" DOI="https://doi.org/10.25221/fee.380.4" author="Nefediev, PS" journalOrPublisher="Far Eastern Entomologist = Dal'nevostochnyi Entomolog" pageId="0" pageNumber="17" pagination="23 - 28" refId="B68" refString="Nefediev, PS, 2019. New records of geophilomorph centipedes (Chilopoda: Geophilomorpha) from natural and anthropogenic habitats of Siberia. Far Eastern Entomologist = Dal'nevostochnyi Entomolog 380: 23 - 28, DOI: https://doi.org/10.25221/fee.380.4" title="New records of geophilomorph centipedes (Chilopoda: Geophilomorpha) from natural and anthropogenic habitats of Siberia." url="https://doi.org/10.25221/fee.380.4" volume="380" year="2019">Nefediev 2019</bibRefCitation>
: 26.
</paragraph>
</subSubSection>
<subSubSection id="64D8AAB79581847650DEB7DDD37A24C7" pageId="0" pageNumber="17" type="type locality">
<paragraph id="EDA767A76A31351D40F1055357A8C30B" pageId="0" pageNumber="17">Type locality.</paragraph>
<paragraph id="2B5DAF73B975C0C32514CA3DC1FFA6D4" pageId="0" pageNumber="17">
North Korea:
<normalizedToken id="48D0EC01C61F46DF9BC53B839EC4D635" originalValue="“Husenzan”">&quot;Husenzan&quot;</normalizedToken>
(
<bibRefCitation id="1578AE81B797C216D2F783A2B9DC0CA2" author="Takakuwa, Y" journalOrPublisher="Zoological Magazine" pageId="0" pageNumber="17" pagination="297 - 299" refId="B95" refString="Takakuwa, Y, 1937a. Eine neue Escaryus -Art aus Korea. Zoological Magazine 49: 297 - 299" title="Eine neue Escaryus - Art aus Korea." volume="49" year="1937 a">Takakuwa 1937a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4A31AB068E6FC642ED390796387FB360" pageId="0" pageNumber="17" type="type series">
<paragraph id="E0369F037598455856AC5A2D4DAFC470" pageId="0" pageNumber="17">Type series.</paragraph>
<paragraph id="E87BF13CF28976D5ED58A41404374934" pageId="0" pageNumber="17">
<emphasis id="7D4634199ACD88EAD3187DDDEA182DB7" bold="true" italics="true" pageId="0" pageNumber="17">Syntypes</emphasis>
: unknown number of specimens, both sexes. Depository unknown.
</paragraph>
</subSubSection>
<subSubSection id="C1EE9D642117DC3B782737B5A90E913F" pageId="0" pageNumber="17" type="diagnosis">
<paragraph id="5D3239A6DE81FCA1C8C0AC7F3A7D6276" pageId="0" pageNumber="17">Diagnosis.</paragraph>
<paragraph id="3ADA3ADC6A6DA8A1012F1D9DB964EE68" pageId="0" pageNumber="17">
An
<taxonomicName id="E9930758B4BF69600EB004459CF643F1" authorityName="Cook &amp; Collins" authorityYear="1891" class="Chilopoda" family="Schendylidae" genus="Escaryus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escaryus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="98C17869A7936AE4A850C670AB49E8EF" italics="true" pageId="0" pageNumber="17">Escaryus</emphasis>
</taxonomicName>
species with body length reaching ≥ 6.5 cm; clypeus with large plagulae; labral arc relatively shallow, with long denticles, the middle denticles obtuse, the lateral ones pointed; first maxillae with one pair of lappets; forcipular trochanteroprefemur with an small obtuse denticle, femur and tibia without denticles, tarsungulum with a small basal bulge; 43-55 leg-bearing segments; metasternites with relatively dense setae; metasternite of the ultimate leg-bearing segment rectangular, ~ 2
<normalizedToken id="1188898C5DDD821A0FEA7A45B282DC6C" originalValue="×">x</normalizedToken>
as long as wide; coxal pores of different size, on both ventral and lateral sides of coxopleura, including a pair of much larger pores on each coxopleuron; anal pores present.
</paragraph>
</subSubSection>
<subSubSection id="92F0F2FA89ECF036AC447B36902FE98F" pageId="0" pageNumber="17" type="distribution">
<paragraph id="AE32EBC67D7C65D157B578106AA8B96E" pageId="0" pageNumber="17">Distribution.</paragraph>
<paragraph id="C375EE7B58C91F6277C9A3FC17DBB340" pageId="0" pageNumber="17">
Western Siberia: Altai krai, republics of Altai and Khakassia, Kemerovo, Novosibirsk, and Tomsk oblasts (
<bibRefCitation id="5E5DB81F1513D27EA2C61E6B692B5415" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Materials of the 4 th All-Union Conference. Baku, 1972. Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="135 - 136" refId="B103" refString="Titova, LP, 1972. Pattern of the distribution of the genus Escaryus (Chilopoda) in the USSR. In: Ghilarov, MS, Ed., Problemy pochvennoi zoologii. Materials of the 4th All-Union Conference. Baku, 1972. Nauka Publisher, Moscow: 135 - 136" title="Pattern of the distribution of the genus Escaryus (Chilopoda) in the USSR." volumeTitle="Problemy pochvennoi zoologii." year="1972">Titova 1972</bibRefCitation>
,
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). Far East: Maritime and Khabarovsk krais, Amur Oblast (
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;
<bibRefCitation id="8282D7FE41E2BA86C87690DD025E9422" author="Ghilarov, MS" editor="Ghilarov, MS" journalOrPublisher="Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="40 - 59" refId="B52" refString="Ghilarov, MS, Perel, TS, 1973. Complexes of soil invertebrates of coniferous-deciduous forests of Far East as indicator of soil type. In: Ghilarov, MS, Ed., Ekologiya pochvennykh bespozvonochnykh. Nauka Publisher, Moscow: 40 - 59" title="Complexes of soil invertebrates of coniferous-deciduous forests of Far East as indicator of soil type." volumeTitle="Ekologiya pochvennykh bespozvonochnykh." year="1973">Ghilarov and Perel 1973</bibRefCitation>
;
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). Outside Asian Russia: Eastern Kazakhstan (
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), Japan, and North Korea (
<bibRefCitation id="1ECFAFE78D0E4709A55D18BFF29887C2" author="Takakuwa, Y" journalOrPublisher="Zoological Magazine" pageId="0" pageNumber="17" pagination="297 - 299" refId="B95" refString="Takakuwa, Y, 1937a. Eine neue Escaryus -Art aus Korea. Zoological Magazine 49: 297 - 299" title="Eine neue Escaryus - Art aus Korea." volume="49" year="1937 a">Takakuwa 1937a</bibRefCitation>
,
<bibRefCitation id="DD13A02393561D708D30553F4B3F5C6F" author="Takakuwa, Y" journalOrPublisher="Zoological Magazine" pageId="0" pageNumber="17" refId="B99" refString="Takakuwa, Y, 1940. Geophilomorpha. In: Okada Y et al. (Eds) Fauna Nipponica, 9, fasc. 8(1), Sanseido, Tokyo, 1-156." title="Geophilomorpha. In: Okada Y et al. (Eds) Fauna Nipponica, 9, fasc. 8 (1), Sanseido, Tokyo, 1 - 156." year="1940">1940</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3E203BEB946C8AA411A11D03A7538605" pageId="0" pageNumber="17" type="remarks">
<paragraph id="A986AFB02E22E9A330E3031999F761F6" pageId="0" pageNumber="17">Remarks.</paragraph>
<paragraph id="A3CB680B7403BD76BEDACF6CEAFA5893" pageId="0" pageNumber="17">
<bibRefCitation id="E451976504F1A89A0826EBFCFF124005" author="Titova, LP" editor="Ghilarov, MS" journalOrPublisher="Nauka Publisher, Moscow" pageId="0" pageNumber="17" pagination="94 - 119" refId="B104" refString="Titova, LP, 1973. New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda). In: Ghilarov, MS, Ed., Ekologiya pochvennykh bespozvonochnykh. Nauka Publisher, Moscow: 94 - 119" title="New species of the genus Escaryus Cook et Collins (Schendylidae, Chilopoda)." volumeTitle="Ekologiya pochvennykh bespozvonochnykh." year="1973">Titova (1973</bibRefCitation>
: 113) suggested that
<taxonomicName id="CBB05A6E3CA867A6888996961BEBB233" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="E. koreanus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="koreanus">
<emphasis id="D2352AF88DA1D4678D0E84508021BAD1" italics="true" pageId="0" pageNumber="17">E. koreanus</emphasis>
</taxonomicName>
can be a junior synonym of
<taxonomicName id="F498CA44ED65DC2D5E5FA52551825B1B" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="E. sibiricus" order="Geophilomorpha" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="sibiricus">
<emphasis id="B96E434D6D0E708530C6A53F07CDDA08" italics="true" pageId="0" pageNumber="17">E. sibiricus</emphasis>
</taxonomicName>
Cook, 1899.
</paragraph>
</subSubSection>
</treatment>
</document>