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<mods:title id="7989D9C4E5A13FB70FBF33D80AF4EA11">Taxonomic revision of the geckos of the Gonatodes concinnatus complex (Squamata: Sphaerodactylidae), with description of two new species</mods:title>
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<mods:namePart id="327BCFFBFA4E7D9D1684601023947CC4">Sturaro, Marcelo José</mods:namePart>
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<heading id="145681F6FFEA9945FF75F9BCFD47EC7A" bold="true" box="[151,727,1570,1596]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<taxonomicName id="88A14D19FFEA9945FF75F9BCFD47EC7A" authority="O'Shaughnessy, 1881" authorityName="O'Shaughnessy" authorityYear="1881" box="[151,727,1570,1596]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="concinnatus">
<emphasis id="7DD5EA88FFEA9945FF75F9BCFD47EC7A" bold="true" box="[151,727,1570,1596]" pageId="6" pageNumber="7">
<emphasis id="7DD5EA88FFEA9945FF75F9BCFE23EC7A" bold="true" box="[151,435,1570,1596]" italics="true" pageId="6" pageNumber="7">Gonatodes concinnatus</emphasis>
(
<bibRefCitation id="2B304B6BFFEA9945FE20F9BCFD5FEC7A" author="O'Shaughnessy" box="[450,719,1570,1596]" pageId="6" pageNumber="7" refString="O'Shaughnessy, A. W. E. (1881) An account of the Collection of Lizards made by Mr. Buckley in Ecuador, and now in the British Museum, with descriptions of the new species. Proceedings of the Zoological Society of London, 1881, 227 - 245, Pls. 22 - 25." type="proceedings paper" year="1881">OShaughnessy, 1881</bibRefCitation>
)
</emphasis>
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<paragraph id="4F1E369AFFEA9945FF75F9DBFEC7EC1B" blockId="6.[151,727,1570,1629]" box="[151,343,1605,1629]" pageId="6" pageNumber="7">
(
<figureCitation id="D79A2A1FFFEA9945FF7DF9DBFF75EC1B" box="[159,229,1605,1629]" captionStart="FIGURE 1" captionStartId="5.[151,250,1952,1975]" captionTargetBox="[228,1363,223,1918]" captionTargetId="figure@5.[211,1387,197,1926]" captionTargetPageId="5" captionText="FIGURE 1. Dorsal and ventral color patterns in preservative in the Gonatodes concinnatus complex. (A) pattern A (MZUSP 54655); (B) pattern B (TCWC 57237); (C) pattern C (MZUSP 53674, holotype); (D) pattern D (MPEG 25596, holotype); and (E) pattern E (MZUSP 49153, holotype). Scale bar = 5 mm." httpUri="https://zenodo.org/record/277514/files/figure.png" pageId="6" pageNumber="7">Figs.1</figureCitation>
A, 4, 5, 6)
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<taxonomicName id="88A14D19FFEA9945FF75F913FD41ECE5" ID-CoL="8GQXR" authority="O'Shaughnessy, 1881: 237" authorityName="O'Shaughnessy" authorityPageNumber="237" authorityYear="1881" box="[151,721,1677,1699]" class="Squamata" family="Sphaerodactylidae" genus="Goniodactylus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="concinnatus">
<emphasis id="7DD5EA88FFEA9945FF75F913FE3FECE5" box="[151,431,1677,1699]" italics="true" pageId="6" pageNumber="7">Goniodactylus concinnatus</emphasis>
<treatmentCitation id="CE00108BFFEA9945FE5BF913FD41ECE5" author="O'Shaughnessy" box="[441,721,1677,1699]" page="237" pageId="6" pageNumber="7" year="1881">
<bibRefCitation id="2B304B6BFFEA9945FE5BF913FD0DECE5" author="O'Shaughnessy" box="[441,669,1677,1699]" pageId="6" pageNumber="7" refString="O'Shaughnessy, A. W. E. (1881) An account of the Collection of Lizards made by Mr. Buckley in Ecuador, and now in the British Museum, with descriptions of the new species. Proceedings of the Zoological Society of London, 1881, 227 - 245, Pls. 22 - 25." type="proceedings paper" year="1881">OShaughnessy, 1881</bibRefCitation>
: 237
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(
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BMNH 1046.9.7.1012, formerly BMNH 80.12.8.2931, type-locality: Canelos,
<collectingCountry id="37B6760AFFEA9945FE50F932FD98EC84" box="[434,520,1708,1730]" name="Ecuador" pageId="6" pageNumber="7">Ecuador</collectingCountry>
, collected by Buckley).
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<taxonomicName id="88A14D19FFEA9945FF75F955FD32ECA6" ID-CoL="8H6CC" authority="O'Shaughnessy, 1881: 238" authorityName="O'Shaughnessy" authorityPageNumber="238" authorityYear="1881" box="[151,674,1738,1761]" class="Squamata" family="Sphaerodactylidae" genus="Goniodactylus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="buckleyi">
<emphasis id="7DD5EA88FFEA9945FF75F955FE17ECA7" box="[151,391,1739,1761]" italics="true" pageId="6" pageNumber="7">Goniodactylus buckleyi</emphasis>
<treatmentCitation id="CE00108BFFEA9945FE6DF954FD32ECA6" author="O'Shaughnessy" box="[399,674,1738,1760]" page="238" pageId="6" pageNumber="7" year="1881">
<bibRefCitation id="2B304B6BFFEA9945FE6DF954FDE0ECA6" author="O'Shaughnessy" box="[399,624,1738,1760]" pageId="6" pageNumber="7" refString="O'Shaughnessy, A. W. E. (1881) An account of the Collection of Lizards made by Mr. Buckley in Ecuador, and now in the British Museum, with descriptions of the new species. Proceedings of the Zoological Society of London, 1881, 227 - 245, Pls. 22 - 25." type="proceedings paper" year="1881">OShaughnessy, 1881</bibRefCitation>
: 238
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</taxonomicName>
(
<typeStatus id="901A8838FFEA9945FD53F955FC9BECA6" box="[689,779,1739,1760]" pageId="6" pageNumber="7" type="syntype">syntypes</typeStatus>
BMNH 1046.9.7.1314, formerly BMNH 80.12.8.3233, typelocality: Canelos; BMNH 1946.9.7.15, formerly BMNH 80.12.8.34, type-locality: Pallatanga [probably
<emphasis id="7DD5EA88FFEA9945FAF5F974FAFAED46" box="[1303,1386,1770,1792]" italics="true" pageId="6" pageNumber="7">in error</emphasis>
, see
<treatmentCitation id="CE00108BFFEA9945FF25F896FE40ED58" author="Rivero-Blanco" box="[199,464,1800,1822]" page="105" pageId="6" pageNumber="7" year="1968">
<bibRefCitation id="2B304B6BFFEA9945FF25F896FE0FED58" author="Rivero-Blanco" box="[199,415,1800,1822]" pageId="6" pageNumber="7" refString="Rivero-Blanco, C. (1968) Un genero y dos especies de tuqueques (Sauria: Sphaerodactylinae) citados por primera vez para Venezuela, con notas sobre la distribucion de otras especies poco conocidas. Memoria de La Sociedad de Ciencias Naturales la Salle, 27, 103 - 119." type="journal article" year="1968">Rivero-Blanco, 1968</bibRefCitation>
: 105
</treatmentCitation>
, 1979: 92]; all in
<collectingCountry id="37B6760AFFEA9945FD66F896FD4AED58" box="[644,730,1800,1822]" name="Ecuador" pageId="6" pageNumber="7">Ecuador</collectingCountry>
, collected by Buckley).
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</paragraph>
<paragraph id="4F1E369AFFEA9945FF75F8B9FDF5EDF1" blockId="6.[151,1436,1677,2007]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="6FB111B4FFEA9945FF75F8B9FDF5EDF1" pageId="6" pageNumber="7">
<taxonomicName id="88A14D19FFEA9945FF75F8B9FE15ED7B" box="[151,389,1831,1853]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="concinnatus">
<emphasis id="7DD5EA88FFEA9945FF75F8B9FE15ED7B" box="[151,389,1831,1853]" italics="true" pageId="6" pageNumber="7">Gonatodes concinnatus</emphasis>
</taxonomicName>
;
<treatmentCitation id="CE00108BFFEA9945FE70F8B9FDF0ED7B" author="Boulenger" box="[402,608,1831,1853]" page="61" pageId="6" pageNumber="7" year="1885">
<bibRefCitation id="2B304B6BFFEA9945FE70F8B9FDADED7B" author="Boulenger" box="[402,573,1831,1853]" pageId="6" pageNumber="7" refString="Boulenger, G. A. (1885) Catalogue of the Lizards in the British Museum (Natural History.) Vol. I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. Trustees of the British Museum, London, xii + 436 pp., Pls. 1 - 32." type="book chapter" year="1885">Boulenger, 1885</bibRefCitation>
: 61
</treatmentCitation>
62;
<treatmentCitation id="CE00108BFFEA9945FD71F8B6FCF9ED7A" author="Burt" box="[659,873,1831,1853]" page="2" pageId="6" pageNumber="7" year="1933">
<bibRefCitation id="2B304B6BFFEA9945FD71F8B6FCC0ED7A" author="Burt" box="[659,848,1831,1853]" pageId="6" pageNumber="7" refString="Burt, C. E. &amp; Burt, M. D. (1933) A preliminary check list of the lizards of South America. Transactions of the Academy of Science of St. Louis, 28, 1 - 104." type="journal article" year="1933">Burt &amp; Burt, 1933</bibRefCitation>
: 2
</treatmentCitation>
; Vanzolini, 1955: 123, 1968: 25 (part);
<treatmentCitation id="CE00108BFFEA9945FB1AF8B9FF73ED1D" author="Wermuth" page="44" pageId="6" pageNumber="7" year="1965">
<bibRefCitation id="2B304B6BFFEA9945FB1AF8B9FA08ED7B" author="Wermuth" box="[1272,1432,1831,1853]" pageId="6" pageNumber="7" refString="Wermuth, H. (1965) Liste der rezenten Amphibien und Reptilien: Gekkonidae, Pygopodidae, Xantusiidae. Das Tierreich, 89, i - xxii, 1 - 246." type="journal article" year="1965">Wermuth, 1965</bibRefCitation>
: 44
</treatmentCitation>
;
<treatmentCitation id="CE00108BFFEA9945FF0FF8DBFE22ED1D" author="Mechler" box="[237,434,1861,1883]" page="331" pageId="6" pageNumber="7" year="1968">
<bibRefCitation id="2B304B6BFFEA9945FF0FF8DBFE13ED1D" author="Mechler" box="[237,387,1861,1883]" pageId="6" pageNumber="7" refString="Mechler, B. (1968) Les geckonides de la Colombie. Revue Suisse de Zoologie 75, 305 - 371." type="journal article" year="1968">Mechler, 1968</bibRefCitation>
: 331
</treatmentCitation>
(part);
<treatmentCitation id="CE00108BFFEA9945FE1EF8DBFC93ED1D" author="Rivero-Blanco" box="[508,771,1861,1883]" page="104" pageId="6" pageNumber="7" year="1968">
<bibRefCitation id="2B304B6BFFEA9945FE1EF8DBFD44ED1D" author="Rivero-Blanco" box="[508,724,1861,1883]" pageId="6" pageNumber="7" refString="Rivero-Blanco, C. (1968) Un genero y dos especies de tuqueques (Sauria: Sphaerodactylinae) citados por primera vez para Venezuela, con notas sobre la distribucion de otras especies poco conocidas. Memoria de La Sociedad de Ciencias Naturales la Salle, 27, 103 - 119." type="journal article" year="1968">Rivero-Blanco, 1968</bibRefCitation>
: 104
</treatmentCitation>
(part);
<treatmentCitation id="CE00108BFFEA9945FCACF8D8FB27ED1D" author="Peters" box="[846,1207,1862,1883]" page="132" pageId="6" pageNumber="7" year="1970">
<bibRefCitation id="2B304B6BFFEA9945FCACF8D8FB18ED1D" author="Peters" box="[846,1160,1862,1883]" pageId="6" pageNumber="7" refString="Peters, J. A. &amp; Donoso-Barros, R. (1970) Catalogue of the Neotropical Squamata: part II. Lizards and amphisbaenians. United States National Museum Bulletin, 297, 1 - 293." type="journal article" year="1970">Peters &amp; Donoso-Barros, 1970</bibRefCitation>
: 132
</treatmentCitation>
(part);
<treatmentCitation id="CE00108BFFEA9945FAE0F8DBFEB6ED3F" author="Dixon" page="19" pageId="6" pageNumber="7" year="1975">
<bibRefCitation id="2B304B6BFFEA9945FAE0F8DBFF6DED3C" author="Dixon" pageId="6" pageNumber="7" refString="Dixon, J. R. &amp; Soini, P. (1975) The reptiles of the upper Amazon Basin, Iquitos Region, Peru: I. Lizards and amphisbaenians. Milwaukee Public Museum, Contributions in Biology and Geology, 4, 1 - 58." type="journal article" year="1975">Dixon &amp; Soini, 1975</bibRefCitation>
: 19
</treatmentCitation>
, 1986: 23;
<treatmentCitation id="CE00108BFFEA9945FE47F8FAFD18ED3C" author="Duellman" box="[421,648,1892,1914]" page="195" pageId="6" pageNumber="7" year="1978">
<bibRefCitation id="2B304B6BFFEA9945FE47F8FAFDC1ED3F" author="Duellman" box="[421,593,1892,1914]" pageId="6" pageNumber="7" refString="Duellman, W. E. (1978) The biology of an Equatorial Herpetofauna in Amazonian Ecuador. Miscellaneous Publications, Museum of Natural History, University of Kansas, 65, 1 - 352." type="journal article" year="1978">Duellman, 1978</bibRefCitation>
: 195
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;
<treatmentCitation id="CE00108BFFEA9945FD75F8FAFCEEED3F" author="Rodrigues" box="[663,894,1892,1914]" page="313" pageId="6" pageNumber="7" year="1980">
<bibRefCitation id="2B304B6BFFEA9945FD75F8FAFCD6ED3F" author="Rodrigues" box="[663,838,1892,1914]" pageId="6" pageNumber="7" refString="Rodrigues, M. T. (1980) Descricao de uma nova especie de Gonatodes da Amazonia (Sauria, Gekkonidae). Papeis Avulsos de Zoologia, 33, 309 - 314." type="journal article" year="1980">Rodrigues, 1980</bibRefCitation>
: 313
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;
<treatmentCitation id="CE00108BFFEA9945FC6EF8FAFABDED3C" author="Duellman" box="[908,1325,1892,1914]" page="358" pageId="6" pageNumber="7" year="1995">
<bibRefCitation id="2B304B6BFFEA9945FC6EF8FAFB65ED3C" author="Duellman" box="[908,1269,1892,1914]" pageId="6" pageNumber="7" refString="Duellman, W. E. &amp; Mendelson III, J. R. (1995) Amphibians and reptiles from northern Departamento Loreto, Peru: taxonomy and biogeography. The University of Kansas Science Bulletin, 55, 329 - 376." type="journal article" year="1995">Duellman &amp; Mendelson III, 1995</bibRefCitation>
: 358
</treatmentCitation>
;
<treatmentCitation id="CE00108BFFEA9945FADEF8FBFE63EDDE" author="Moravec" page="51" pageId="6" pageNumber="7" year="2001">
<bibRefCitation id="2B304B6BFFEA9945FADEF8FBFE5EEDDE" author="Moravec" pageId="6" pageNumber="7" refString="Moravec, J., Tuanama, I. A. &amp; Burgos, A. M. (2001) Reptiles recently recorded from the surroundings of Iquitos (Departamento Loreto, Peru). C asopis Narodniho muzea, R ada p r irodov e dna, 170, 47 - 68." type="journal article" year="2001">Moravec, Tuanama &amp; Burgos, 2001</bibRefCitation>
: 51
</treatmentCitation>
;
<treatmentCitation id="CE00108BFFEA9945FE1CF81CFCB6EDDE" author="Bartlett" box="[510,806,1922,1944]" page="158" pageId="6" pageNumber="7" year="2003">
<bibRefCitation id="2B304B6BFFEA9945FE1CF81CFD67EDDE" author="Bartlett" box="[510,759,1922,1944]" pageId="6" pageNumber="7" refString="Bartlett, R. D. &amp; Bartlett, P. (2003) Reptiles and Amphibians of the Amazon: An Ecotourist's Guide. University Press of Florida, Gainesville, 291 pp." type="book" year="2003">Bartlett &amp; Bartlett, 2003</bibRefCitation>
: 158
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(part);
<treatmentCitation id="CE00108BFFEA9945FC93F81CFB28EDDE" author="Powell" box="[881,1208,1922,1944]" page="714" pageId="6" pageNumber="7" year="2005">
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: 714
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;
<treatmentCitation id="CE00108BFFEA9945FB20F81CFA07EDDE" author="Cole" box="[1218,1431,1922,1944]" page="4" pageId="6" pageNumber="7" year="2006">
<bibRefCitation id="2B304B6BFFEA9945FB20F81CFA10EDDE" author="Cole" box="[1218,1408,1922,1944]" pageId="6" pageNumber="7" refString="Cole, C. J. &amp; Kok, P. J. R. (2006) A new species of gekkonid lizard (Sphaerodactylinae: Gonatodes) from Guyana, South American. American Museum Novitates, 3524, 1 - 13." type="journal article" year="2006">Cole &amp; Kok, 2006</bibRefCitation>
: 4
</treatmentCitation>
; Gamble, Simon,
<treatmentCitation id="CE00108BFFEA9945FE90F83FFDF1EDF1" author="Colli" box="[370,609,1953,1975]" page="271" pageId="6" pageNumber="7" year="2008">
<bibRefCitation id="2B304B6BFFEA9945FE90F83FFDBEEDF1" author="Colli" box="[370,558,1953,1975]" pageId="6" pageNumber="7" refString="Gamble, T.; Simons, A. M.; Colli, G. R. &amp; Vitt, L. J. (2008) Tertiary climate change and the diversification of the Amazonian gecko genus Gonatodes (Sphaerodactylidae, Squamata). Molecular Phylogenetics and Evolution, 46, 269 - 277." type="journal article" year="2008">Colli &amp; Vitt, 2008</bibRefCitation>
: 271
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4F1E369AFFEA9945FF75F85EFCC4ED91" blockId="6.[151,1436,1677,2007]" box="[151,852,1984,2007]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="6FB111B4FFEA9945FF75F85EFCC4ED91" box="[151,852,1984,2007]" pageId="6" pageNumber="7">
<taxonomicName id="88A14D19FFEA9945FF75F85EFD95ED90" box="[151,517,1984,2006]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="subSpecies" species="concinnatus" subSpecies="concinnatus">
<emphasis id="7DD5EA88FFEA9945FF75F85EFD95ED90" box="[151,517,1984,2006]" italics="true" pageId="6" pageNumber="7">Gonatodes concinnatus concinnatus</emphasis>
</taxonomicName>
;
<treatmentCitation id="CE00108BFFEA9945FDF1F85EFC9EED90" author="Rivero-Blanco" box="[531,782,1984,2006]" page="92" pageId="6" pageNumber="7" year="1979">
<bibRefCitation id="2B304B6BFFEA9945FDF1F85EFD7BED90" author="Rivero-Blanco" box="[531,747,1984,2006]" pageId="6" pageNumber="7" refString="Rivero-Blanco, C. (1979) The Neotropical lizard genus Gonatodes Fitzinger (Sauria: Sphaerodactylinae). PhD Thesis, Texas A &amp; M University. 233 pp." type="book" year="1979">Rivero-Blanco, 1979</bibRefCitation>
: 92
</treatmentCitation>
(part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="1BDE6612FFEB9944FF75F8DDFDF8EDB5" httpUri="https://zenodo.org/record/277516/files/figure.png" pageId="7" pageNumber="8" targetBox="[239,1345,203,1826]" targetPageId="7">
<paragraph id="4F1E369AFFEB9944FF75F8DDFDF8EDB5" blockId="7.[151,1436,1859,2035]" pageId="7" pageNumber="8">
<emphasis id="7DD5EA88FFEB9944FF75F8DDFE8AED1F" bold="true" box="[151,282,1859,1882]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
(A) Geographic distribution of
<taxonomicName id="88A14D19FFEB9944FD61F8DAFCE9ED1C" box="[643,889,1860,1882]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="concinnatus">
<emphasis id="7DD5EA88FFEB9944FD61F8DAFCE9ED1C" box="[643,889,1860,1882]" italics="true" pageId="7" pageNumber="8">Gonatodes concinnatus</emphasis>
</taxonomicName>
complex. (B) Detailed geographic distribution of
<taxonomicName id="88A14D19FFEB9944FF75F8FCFE15ED3E" box="[151,389,1890,1912]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="concinnatus">
<emphasis id="7DD5EA88FFEB9944FF75F8FCFE15ED3E" box="[151,389,1890,1912]" italics="true" pageId="7" pageNumber="8">Gonatodes concinnatus</emphasis>
</taxonomicName>
. Red =
<taxonomicName id="88A14D19FFEB9944FE35F8FCFD56ED3E" box="[471,710,1890,1912]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="concinnatus">
<emphasis id="7DD5EA88FFEB9944FE35F8FCFD56ED3E" box="[471,710,1890,1912]" italics="true" pageId="7" pageNumber="8">Gonatodes concinnatus</emphasis>
</taxonomicName>
(Pattern A). Cyan-green =
<taxonomicName id="88A14D19FFEB9944FC3CF8FCFB1DED3E" box="[990,1165,1890,1912]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="ligiae">
<emphasis id="7DD5EA88FFEB9944FC3CF8FCFB1DED3E" box="[990,1165,1890,1912]" italics="true" pageId="7" pageNumber="8">Gonatodes ligiae</emphasis>
</taxonomicName>
(Pattern B). Lime-green =
<taxonomicName id="88A14D19FFEB9944FF75F81FFE12EDD1" box="[151,386,1921,1943]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="tapajonicus">
<emphasis id="7DD5EA88FFEB9944FF75F81FFE12EDD1" box="[151,386,1921,1943]" italics="true" pageId="7" pageNumber="8">Gonatodes tapajonicus</emphasis>
</taxonomicName>
(Pattern C). Yellow =
<taxonomicName id="88A14D19FFEB9944FD8EF81FFCCCEDD1" box="[620,860,1921,1943]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="nascimentoi" status="sp. nov.">
<emphasis id="7DD5EA88FFEB9944FD8EF81FFCCCEDD1" box="[620,860,1921,1943]" italics="true" pageId="7" pageNumber="8">Gonatodes nascimentoi</emphasis>
</taxonomicName>
<emphasis id="7DD5EA88FFEB9944FC81F81CFC25EDD1" bold="true" box="[867,949,1922,1943]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="66E657F3FFEB9944FC81F81CFC25EDD1" box="[867,949,1922,1943]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Pattern D). Blue =
<taxonomicName id="88A14D19FFEB9944FB6BF81FFAD2EDD1" box="[1161,1346,1921,1943]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="riveroi" status="sp. nov.">
<emphasis id="7DD5EA88FFEB9944FB6BF81FFAD2EDD1" box="[1161,1346,1921,1943]" italics="true" pageId="7" pageNumber="8">Gonatodes riveroi</emphasis>
</taxonomicName>
<emphasis id="7DD5EA88FFEB9944FAA8F81CFA0BEDD1" bold="true" box="[1354,1435,1922,1943]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="66E657F3FFEB9944FAA8F81CFA0BEDD1" box="[1354,1435,1922,1943]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Pattern E). Open symbols = type-localities; half-black symbol = literature record; question mark = Pallatanga, Ecuador, probably
<emphasis id="7DD5EA88FFEB9944FF1AF85EFED7ED93" box="[248,327,1984,2005]" italics="true" pageId="7" pageNumber="8">in error</emphasis>
; triangle = locality referred to only as Rio Napo. Numbers refer to localities cited under each species. One number may represent more than one locality.
</paragraph>
</caption>
<subSubSection id="07BB6511FFE4994BFF75FF09FB0CEFEA" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="4F1E369AFFE4994BFF75FF09FB0CEFEA" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="9">
<emphasis id="7DD5EA88FFE4994BFF75FF09FEE5EAF6" bold="true" box="[151,373,151,176]" pageId="8" pageNumber="9">Material examined</emphasis>
(numbers in bold between parentheses refer to
<figureCitation id="D79A2A1FFFE4994BFC63FF09FC54EAE9" box="[897,964,151,175]" captionStart="FIGURE 3" captionStartId="7.[151,250,1859,1882]" captionTargetBox="[239,1345,203,1826]" captionTargetId="figure@7.[235,1351,195,1833]" captionTargetPageId="7" captionText="FIGURE 3. (A) Geographic distribution of Gonatodes concinnatus complex. (B) Detailed geographic distribution of Gonatodes concinnatus. Red = Gonatodes concinnatus (Pattern A). Cyan-green = Gonatodes ligiae (Pattern B). Lime-green = Gonatodes tapajonicus (Pattern C). Yellow = Gonatodes nascimentoi sp. nov. (Pattern D). Blue = Gonatodes riveroi sp. nov. (Pattern E). Open symbols = type-localities; half-black symbol = literature record; question mark = Pallatanga, Ecuador, probably in error; triangle = locality referred to only as Rio Napo. Numbers refer to localities cited under each species. One number may represent more than one locality." httpUri="https://zenodo.org/record/277516/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
):
<emphasis id="7DD5EA88FFE4994BFC3BFF09FBFCEAF6" bold="true" box="[985,1132,151,176]" pageId="8" pageNumber="9">
<collectingCountry id="37B6760AFFE4994BFC3BFF09FBF7EAF6" box="[985,1127,151,176]" name="Ecuador" pageId="8" pageNumber="9">ECUADOR</collectingCountry>
.
</emphasis>
“Pallatanga” (
<emphasis id="7DD5EA88FFE4994BFAEFFF09FA8BEAF6" bold="true" box="[1293,1307,151,176]" pageId="8" pageNumber="9">1</emphasis>
): Photos of
<collectionCode id="29B0AE5FFFE4994BFF75FF22FF7AEA95" box="[151,234,188,211]" pageId="8" pageNumber="9">BMNH</collectionCode>
1946.9.7.15 (formerly
<collectionCode id="29B0AE5FFFE4994BFE09FF22FDAEEA95" box="[491,574,188,211]" pageId="8" pageNumber="9">BMNH</collectionCode>
80.12.8.34,
<typeStatus id="901A8838FFE4994BFD2AFF22FC8DEA92" box="[712,797,188,212]" pageId="8" pageNumber="9" type="syntype">syntype</typeStatus>
of
<taxonomicName id="88A14D19FFE4994BFCA1FF25FBD6EA95" box="[835,1094,187,211]" class="Squamata" family="Sphaerodactylidae" genus="Goniodactylus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="buckleyi">
<emphasis id="7DD5EA88FFE4994BFCA1FF25FBD6EA95" box="[835,1094,187,211]" italics="true" pageId="8" pageNumber="9">Goniodactylus buckleyi</emphasis>
</taxonomicName>
), leg. Buckley. SUCUMBÍOS. Shushufindi, Rio Aguarico (
<emphasis id="7DD5EA88FFE4994BFE39FF40FE79EAB1" bold="true" box="[475,489,222,247]" pageId="8" pageNumber="9">2</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFDE6FF41FDC4EAB1" LSID="urn:lsid:biocol.org:col:34871" box="[516,596,223,247]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">USNM</collectionCode>
234497, female. Santa Cecilia (
<emphasis id="7DD5EA88FFE4994BFC2BFF40FC47EAB1" bold="true" box="[969,983,222,247]" pageId="8" pageNumber="9">3</emphasis>
): KU 10529091, KU 10957273, KU 146643, KU 146645, KU 14758889, KU 152496,
<specimenCount id="59A7FD13FFE4994BFD32FE9CFCB5EB5C" box="[720,805,258,282]" pageId="8" pageNumber="9" type="male">4 males</specimenCount>
and
<specimenCount id="59A7FD13FFE4994BFCB9FE9CFC59EB5C" box="[859,969,258,282]" pageId="8" pageNumber="9" type="female">5 females</specimenCount>
. Lago Agrio (
<emphasis id="7DD5EA88FFE4994BFB86FE9CFBE2EB5D" bold="true" box="[1124,1138,258,283]" pageId="8" pageNumber="9">4</emphasis>
): KU 126683, 12668586, 126688,
<specimenCount id="59A7FD13FFE4994BFF1AFEB8FEDEEB78" box="[248,334,294,318]" pageId="8" pageNumber="9" type="male">2 males</specimenCount>
and
<specimenCount id="59A7FD13FFE4994BFE64FEB8FE65EB78" box="[390,501,294,318]" pageId="8" pageNumber="9" type="female">2 females</specimenCount>
; KU 297992, female, leg. J. Lee. Limoncocha (
<emphasis id="7DD5EA88FFE4994BFBEBFEB8FB87EB78" bold="true" box="[1033,1047,294,318]" pageId="8" pageNumber="9">5</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFBD2FEB8FB10EB78" LSID="urn:lsid:biocol.org:col:34802" box="[1072,1152,294,318]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="8" pageNumber="9">LACM</collectionCode>
73293300, a juvenile,
<specimenCount id="59A7FD13FFE4994BFA6CFEB8FF47EB27" pageId="8" pageNumber="9" type="male">5 males</specimenCount>
and
<specimenCount id="59A7FD13FFE4994BFEE9FED7FEE7EB27" box="[267,375,329,353]" pageId="8" pageNumber="9" type="female">2 females</specimenCount>
, leg. W.R. Heyer;
<collectionCode id="29B0AE5FFFE4994BFDDDFED7FD0BEB27" box="[575,667,329,353]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
54655, male, leg. K. Miyata; KU 144378,
<collectionCode id="29B0AE5FFFE4994BFB8DFED7FB3BEB27" LSID="urn:lsid:biocol.org:col:33791" box="[1135,1195,329,353]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="8" pageNumber="9">MCZ</collectionCode>
86414, a juvenile and a male. Reserva de Producción Faunística de Cuyabeno (RPF Cuyabeno), Estación Biológica de la Pontificia Universidad Catolica del
<collectingCountry id="37B6760AFFE4994BFE57FE0EFD81EBEE" box="[437,529,400,424]" name="Ecuador" pageId="8" pageNumber="9">Ecuador</collectingCountry>
(
<emphasis id="7DD5EA88FFE4994BFDC1FE11FDA1EBEE" bold="true" box="[547,561,399,424]" pageId="8" pageNumber="9">6</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFDA8FE0EFD0EEBEE" box="[586,670,400,424]" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Sam Noble Oklahoma Museum of Natural History" pageId="8" pageNumber="9">OMNH</collectionCode>
3640611, 36424,
<specimenCount id="59A7FD13FFE4994BFC9BFE0EFC5FEBEE" box="[889,975,400,424]" pageId="8" pageNumber="9" type="male">3 males</specimenCount>
and
<specimenCount id="59A7FD13FFE4994BFBEBFE0EFBE8EBEE" box="[1033,1144,400,424]" pageId="8" pageNumber="9" type="female">4 females</specimenCount>
, leg. L.J. Vitt. Reserva de Producción Faunística de Cuyabeno (RPF Cuyabeno), Saladero de Dantas (
<emphasis id="7DD5EA88FFE4994BFC39FE2DFC79EB8A" bold="true" box="[987,1001,435,460]" pageId="8" pageNumber="9">6</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFBE2FE2DFBC5EB8D" box="[1024,1109,435,459]" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Sam Noble Oklahoma Museum of Natural History" pageId="8" pageNumber="9">OMNH</collectionCode>
3641214, a juvenile, a male and a female, leg. L.J. Vitt. Corporación Estatal Petrolera Ecuatoriana (
<collectionCode id="29B0AE5FFFE4994BFC21FE49FB9AEBA9" box="[963,1034,471,495]" pageId="8" pageNumber="9">CEPE</collectionCode>
) oil field (near Tarapoa, near Lago Agrio), Cuyabeno (
<emphasis id="7DD5EA88FFE4994BFE8FFE65FEEBE855" bold="true" box="[365,379,507,531]" pageId="8" pageNumber="9">7</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFE70FE64FE5EE854" LSID="urn:lsid:biocol.org:col:33791" box="[402,462,506,530]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="8" pageNumber="9">MCZ</collectionCode>
163233, male. SUCUMBIOSORELLANA. Rio Napo (undefined locality
<emphasis id="7DD5EA88FFE4994BFAF3FE65FABFE855" bold="true" box="[1297,1327,506,531]" pageId="8" pageNumber="9">- 8</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFAA4FE65FA0CE854" box="[1350,1436,507,530]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="8" pageNumber="9">UMMZ</collectionCode>
90813, female, leg. W.M. Clark.
<collectionCode id="29B0AE5FFFE4994BFDE6FD80FDDFE870" box="[516,591,542,566]" pageId="8" pageNumber="9">NAPO</collectionCode>
. George Kiederle Hacienda, southern bank Rio Napo,
<quantity id="88599B7FFFE4994BFB4DFD80FB77E870" box="[1199,1255,542,566]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="9" unit="km" value="2.0">2 km</quantity>
W Puerto Napo (
<emphasis id="7DD5EA88FFE4994BFF42FDDFFF3EE81C" bold="true" box="[160,174,577,602]" pageId="8" pageNumber="9">9</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFF24FDDFFE86E81F" LSID="urn:lsid:biocol.org:col:34871" box="[198,278,577,601]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">USNM</collectionCode>
16613537,
<specimenCount id="59A7FD13FFE4994BFE45FDDFFE6CE81F" box="[423,508,577,601]" pageId="8" pageNumber="9" type="male">2 males</specimenCount>
and a female, leg. J.A. Peters. Rio Misahualli (
<emphasis id="7DD5EA88FFE4994BFBEAFDDFFBB4E81C" bold="true" box="[1032,1060,577,602]" pageId="8" pageNumber="9">10</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFBDEFDDFFB1CE81F" LSID="urn:lsid:biocol.org:col:34871" box="[1084,1164,577,601]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">USNM</collectionCode>
234490, female, leg. M. Olalla.
<quantity id="88599B7FFFE4994BFF0AFDFBFEB1E83B" box="[232,289,613,637]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="9" unit="km" value="1.0">1 km</quantity>
E Jatun Sacha (
<emphasis id="7DD5EA88FFE4994BFE37FDFAFE7DE83B" bold="true" box="[469,493,612,637]" pageId="8" pageNumber="9">11</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFDE5FDFBFDCBE83B" box="[519,603,613,637]" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Sam Noble Oklahoma Museum of Natural History" pageId="8" pageNumber="9">OMNH</collectionCode>
3637778, a male and a female. San José de Sumaco (
<emphasis id="7DD5EA88FFE4994BFB22FDFAFB4CE83B" bold="true" box="[1216,1244,612,637]" pageId="8" pageNumber="9">12</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFB16FDFBFAD8E83B" LSID="urn:lsid:biocol.org:col:34925" box="[1268,1352,613,637]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="9">AMNH</collectionCode>
28879, female, leg. C. Ollalo. ORELLANA. Between Loreto and Concepcion (
<emphasis id="7DD5EA88FFE4994BFC44FD16FC52E8E7" bold="true" box="[934,962,648,673]" pageId="8" pageNumber="9">13</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFC3BFD16FBABE8E6" box="[985,1083,648,672]" httpUri="http://grbio.org/cool/fxd1-0prj" name="California Academy of Sciences, Stanford University Collection" pageId="8" pageNumber="9">CAS-SU</collectionCode>
1652930, a male and a female, leg. J. Olalla and C.A. Olalla. Loreto (
<emphasis id="7DD5EA88FFE4994BFDA0FD35FDCEE882" bold="true" box="[578,606,683,708]" pageId="8" pageNumber="9">14</emphasis>
).
<collectionCode id="29B0AE5FFFE4994BFD97FD32FD41E882" box="[629,721,684,708]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
338283,
<specimenCount id="59A7FD13FFE4994BFCAAFD32FC26E882" box="[840,950,684,708]" pageId="8" pageNumber="9" type="female">2 females</specimenCount>
, leg. J. Olalla;
<collectionCode id="29B0AE5FFFE4994BFBBFFD32FB3DE882" LSID="urn:lsid:biocol.org:col:34871" box="[1117,1197,684,708]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">USNM</collectionCode>
234494, male, leg. J. Ollala. Rio Punino, affluent of Rio Payamino above Coca (
<emphasis id="7DD5EA88FFE4994BFCCBFD51FCD5E8AE" bold="true" box="[809,837,719,744]" pageId="8" pageNumber="9">15</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFCBFFD51FC09E8A1" LSID="urn:lsid:biocol.org:col:33791" box="[861,921,719,743]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="8" pageNumber="9">MCZ</collectionCode>
<date id="3B1F105AFFE4994BFC43FD51FC67E8A1" box="[929,1015,719,743]" pageId="8" pageNumber="9" value="46-10-13">131046</date>
, juvenile. Rio Payamino (
<emphasis id="7DD5EA88FFE4994BFAFAFD51FAA4E8AE" bold="true" box="[1304,1332,719,744]" pageId="8" pageNumber="9">16</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFAAEFD51FA0CE8A1" LSID="urn:lsid:biocol.org:col:34871" box="[1356,1436,719,743]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">USNM</collectionCode>
23449193, a male and
<specimenCount id="59A7FD13FFE4994BFE49FD6DFD8BE94D" box="[427,539,755,779]" pageId="8" pageNumber="9" type="female">2 females</specimenCount>
, leg. M. Olalla. Mouth of Rio Coca (
<emphasis id="7DD5EA88FFE4994BFC28FD6CFC76E94D" bold="true" box="[970,998,754,779]" pageId="8" pageNumber="9">17</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFBE3FD6DFBC1E94D" LSID="urn:lsid:biocol.org:col:34871" box="[1025,1105,755,779]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">USNM</collectionCode>
234495, male, leg. J. Olalla. PASTAZA. Rio Pindo (
<emphasis id="7DD5EA88FFE4994BFE40FC8BFE2EE968" bold="true" box="[418,446,789,814]" pageId="8" pageNumber="9">18</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFE35FC88FDB7E968" LSID="urn:lsid:biocol.org:col:34871" box="[471,551,790,814]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">USNM</collectionCode>
234500, male, leg. R. Olalla. Canelos (
<emphasis id="7DD5EA88FFE4994BFC08FC8BFB96E968" bold="true" box="[1002,1030,789,814]" pageId="8" pageNumber="9">19</emphasis>
): Photos of
<collectionCode id="29B0AE5FFFE4994BFB76FC89FB77E968" box="[1172,1255,791,814]" pageId="8" pageNumber="9">BMNH</collectionCode>
1046.9.7.1012 (
<typeStatus id="901A8838FFE4994BFF7CFCA7FE90E917" box="[158,256,825,849]" pageId="8" pageNumber="9" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="88A14D19FFE4994BFED2FCA4FDF2E914" box="[304,610,826,850]" class="Squamata" family="Sphaerodactylidae" genus="Goniodactylus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="concinnatus">
<emphasis id="7DD5EA88FFE4994BFED2FCA4FDF2E914" box="[304,610,826,850]" italics="true" pageId="8" pageNumber="9">Goniodactylus concinnatus</emphasis>
</taxonomicName>
, formerly
<collectionCode id="29B0AE5FFFE4994BFD00FCA4FCA5E917" box="[738,821,826,849]" pageId="8" pageNumber="9">BMNH</collectionCode>
80.12.8.2931),
<specimenCount id="59A7FD13FFE4994BFC1EFCA7FBC6E917" box="[1020,1110,825,849]" pageId="8" pageNumber="9" type="male">3 males</specimenCount>
(
<figureCitation id="D79A2A1FFFE4994BFB89FCA7FB24E917" box="[1131,1204,825,849]" captionStart="FIGURE 4" captionStartId="10.[151,250,1973,1996]" captionTargetBox="[265,1321,480,1959]" captionTargetId="figure@10.[265,1321,480,1959]" captionTargetPageId="10" captionText="FIGURE 4. Gonatodes concinnatus (syntypes, BMNH 1946.9.7.10 12). (A) Dorsal, (B) ventral views. Scale bar = 50 mm. (Photo by Colin McCarthy)." httpUri="https://zenodo.org/record/277517/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
), Photos of
<collectionCode id="29B0AE5FFFE4994BFAABFCA4FA0CE917" box="[1353,1436,826,849]" pageId="8" pageNumber="9">BMNH</collectionCode>
1046.9.7.1314 (
<typeStatus id="901A8838FFE4994BFEB4FCC3FE28E933" box="[342,440,861,885]" pageId="8" pageNumber="9" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="88A14D19FFE4994BFE04FCC3FD7DE933" box="[486,749,861,885]" class="Squamata" family="Sphaerodactylidae" genus="Goniodactylus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="buckleyi">
<emphasis id="7DD5EA88FFE4994BFE04FCC3FD7DE933" box="[486,749,861,885]" italics="true" pageId="8" pageNumber="9">Goniodactylus buckleyi</emphasis>
</taxonomicName>
, formerly
<collectionCode id="29B0AE5FFFE4994BFC8FFCC0FC50E933" box="[877,960,862,885]" pageId="8" pageNumber="9">BMNH</collectionCode>
80.12.8.3233),
<specimenCount id="59A7FD13FFE4994BFB67FCC3FB67E933" box="[1157,1271,861,885]" pageId="8" pageNumber="9" type="female">2 females</specimenCount>
, leg. Buckley.
<collectionCode id="29B0AE5FFFE4994BFF75FC1EFF69E9DE" box="[151,249,896,920]" httpUri="http://grbio.org/cool/fxd1-0prj" name="California Academy of Sciences, Stanford University Collection" pageId="8" pageNumber="9">CAS-SU</collectionCode>
1581415,
<collectionCode id="29B0AE5FFFE4994BFE60FC1EFE74E9DE" box="[386,484,896,920]" httpUri="http://grbio.org/cool/fxd1-0prj" name="California Academy of Sciences, Stanford University Collection" pageId="8" pageNumber="9">CAS-SU</collectionCode>
16531, a male and
<specimenCount id="59A7FD13FFE4994BFD28FC1EFCA9E9DE" box="[714,825,896,920]" pageId="8" pageNumber="9" type="female">2 females</specimenCount>
, leg. A. Proana. Sarayacu, (
<emphasis id="7DD5EA88FFE4994BFB9BFC1EFB05E9DF" bold="true" box="[1145,1173,896,921]" pageId="8" pageNumber="9">20</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFB4DFC1EFB7BE9DE" LSID="urn:lsid:biocol.org:col:33791" box="[1199,1259,896,920]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="8" pageNumber="9">MCZ</collectionCode>
3770304, KU
<date id="3B1F105AFFE4994BFF75FC3AFF7CE9FA" box="[151,236,932,956]" pageId="8" pageNumber="9" value="88-12-10">121088</date>
89, a juvenile and
<specimenCount id="59A7FD13FFE4994BFE27FC3AFDA3E9FA" box="[453,563,932,956]" pageId="8" pageNumber="9" type="female">3 females</specimenCount>
. Rio Villano (
<emphasis id="7DD5EA88FFE4994BFD2DFC3AFD7BE9FB" bold="true" box="[719,747,932,957]" pageId="8" pageNumber="9">21</emphasis>
),
<collectionCode id="29B0AE5FFFE4994BFCE1FC3AFCC3E9FA" LSID="urn:lsid:biocol.org:col:34871" box="[771,851,932,956]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">USNM</collectionCode>
234499, female, leg. C. Estrella. Nuevo Golandrina, on trail W toward Rio Curaray,
<quantity id="88599B7FFFE4994BFE1EFC59FDC1E999" box="[508,593,967,991]" metricMagnitude="5" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="9" unit="km" value="130.0">130 km</quantity>
S of Coca (
<emphasis id="7DD5EA88FFE4994BFD38FC59FD66E9A6" bold="true" box="[730,758,967,992]" pageId="8" pageNumber="9">22</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFCEDFC59FCCFE999" LSID="urn:lsid:biocol.org:col:34871" box="[783,863,967,991]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">USNM</collectionCode>
321064, female, leg. R.P. Reynolds and W. Lamar. Tiguino (UNOCAL Base Camp),
<quantity id="88599B7FFFE4994BFDEEFC75FDF0EE45" box="[524,608,1003,1027]" metricMagnitude="5" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="9" unit="km" value="130.0">130 km</quantity>
S of Coca (
<emphasis id="7DD5EA88FFE4994BFD00FC74FD6EEE45" bold="true" box="[738,766,1002,1027]" pageId="8" pageNumber="9">22</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFCF6FC75FCF4EE45" LSID="urn:lsid:biocol.org:col:34871" box="[788,868,1003,1027]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">USNM</collectionCode>
321057, female, leg. R. P. Reynolds and W. Lamar. Rio Conambo (
<emphasis id="7DD5EA88FFE4994BFEA0FB90FECEEE61" bold="true" box="[322,350,1038,1063]" pageId="8" pageNumber="9">23</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFE97FB90FE55EE60" LSID="urn:lsid:biocol.org:col:34871" box="[373,453,1038,1062]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">USNM</collectionCode>
234502, female, leg. R. Olalla. MORONA-SANTIAGO. Cusuime, Rio Cusuime,
<quantity id="88599B7FFFE4994BFAB4FB90FA0CEE60" box="[1366,1436,1038,1062]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="9" unit="km" value="60.0">60 km</quantity>
airline SE Macas (
<emphasis id="7DD5EA88FFE4994BFE84FBAFFE12EE0C" bold="true" box="[358,386,1073,1098]" pageId="8" pageNumber="9">24</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFE78FBACFE7EEE0C" LSID="urn:lsid:biocol.org:col:34925" box="[410,494,1074,1098]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="9">AMNH</collectionCode>
11366667,
<specimenCount id="59A7FD13FFE4994BFD62FBACFD48EE0C" box="[640,728,1074,1098]" pageId="8" pageNumber="9" type="male">2 males</specimenCount>
, leg. B. Malkin. Taisha, Macas (
<emphasis id="7DD5EA88FFE4994BFBA1FBAFFBCFEE0C" bold="true" box="[1091,1119,1073,1098]" pageId="8" pageNumber="9">25</emphasis>
).
<collectionCode id="29B0AE5FFFE4994BFB94FBACFB56EE0C" LSID="urn:lsid:biocol.org:col:34871" box="[1142,1222,1074,1098]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">USNM</collectionCode>
234498, male, leg. M. Ollala;
<collectionCode id="29B0AE5FFFE4994BFEEFFBCBFECDEE2B" LSID="urn:lsid:biocol.org:col:34871" box="[269,349,1109,1133]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">USNM</collectionCode>
283922,
<collectionCode id="29B0AE5FFFE4994BFE21FBCBFD83EE2B" LSID="urn:lsid:biocol.org:col:34871" box="[451,531,1109,1133]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="9">USNM</collectionCode>
283924, a juvenile and a female, leg. C. M. Fugler.
<emphasis id="7DD5EA88FFE4994BFBA8FBC8FB0FEE28" bold="true" box="[1098,1183,1110,1134]" pageId="8" pageNumber="9">
<collectingCountry id="37B6760AFFE4994BFBA8FBC8FB09EE28" box="[1098,1177,1110,1134]" name="Peru" pageId="8" pageNumber="9">PERU</collectingCountry>
:
</emphasis>
LORETO. San Jacinto (
<emphasis id="7DD5EA88FFE4994BFF42FBE6FF2CEED7" bold="true" box="[160,188,1144,1169]" pageId="8" pageNumber="9">26</emphasis>
): KU 222138, female, leg. R. A. Leschen. Moropón (
<emphasis id="7DD5EA88FFE4994BFCFAFBE6FCA4EED7" bold="true" box="[792,820,1144,1169]" pageId="8" pageNumber="9">27</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFCAFFBE7FC0DEED7" box="[845,925,1145,1169]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="8" pageNumber="9">TCWC</collectionCode>
3666571,
<collectionCode id="29B0AE5FFFE4994BFBC0FBE7FBE2EED7" box="[1058,1138,1145,1169]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="8" pageNumber="9">TCWC</collectionCode>
3674346,
<collectionCode id="29B0AE5FFFE4994BFB1AFBE7FAD8EED7" box="[1272,1352,1145,1169]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="8" pageNumber="9">TCWC</collectionCode>
38994,
<collectionCode id="29B0AE5FFFE4994BFF75FB02FF77EEF2" box="[151,231,1180,1204]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="8" pageNumber="9">TCWC</collectionCode>
4121320,
<collectionCode id="29B0AE5FFFE4994BFE8CFB02FE2EEEF2" box="[366,446,1180,1204]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="8" pageNumber="9">TCWC</collectionCode>
4190204,
<collectionCode id="29B0AE5FFFE4994BFDA6FB02FD04EEF2" box="[580,660,1180,1204]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="8" pageNumber="9">TCWC</collectionCode>
4190608,
<collectionCode id="29B0AE5FFFE4994BFCF9FB02FCFBEEF2" box="[795,875,1180,1204]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="8" pageNumber="9">TCWC</collectionCode>
4270613,
<collectionCode id="29B0AE5FFFE4994BFC10FB02FBD2EEF2" box="[1010,1090,1180,1204]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="8" pageNumber="9">TCWC</collectionCode>
42828,
<collectionCode id="29B0AE5FFFE4994BFB42FB02FB60EEF2" box="[1184,1264,1180,1204]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="8" pageNumber="9">TCWC</collectionCode>
43332,
<collectionCode id="29B0AE5FFFE4994BFAAFFB02FA0DEEF2" box="[1357,1437,1180,1204]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="8" pageNumber="9">TCWC</collectionCode>
44230,
<collectionCode id="29B0AE5FFFE4994BFF09FB21FEABEE91" box="[235,315,1215,1239]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="8" pageNumber="9">TCWC</collectionCode>
44283, a juvenile,
<specimenCount id="59A7FD13FFE4994BFDF4FB21FDEAEE91" box="[534,634,1215,1239]" pageId="8" pageNumber="9" type="male">25 males</specimenCount>
and
<specimenCount id="59A7FD13FFE4994BFD56FB21FCA1EE91" box="[692,817,1215,1239]" pageId="8" pageNumber="9" type="female">12 females</specimenCount>
; leg. P. Soini;
<collectionCode id="29B0AE5FFFE4994BFC37FB21FBB5EE91" box="[981,1061,1215,1239]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="8" pageNumber="9">TCWC</collectionCode>
41763, juvenile, leg. J.R. Dixon;
<collectionCode id="29B0AE5FFFE4994BFF75FB7DFF63EEBD" box="[151,243,1251,1275]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
2824849,
<collectionCode id="29B0AE5FFFE4994BFE9EFB7DFE48EEBD" box="[380,472,1251,1275]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
2826063,
<collectionCode id="29B0AE5FFFE4994BFD81FB7DFD2FEEBD" box="[611,703,1251,1275]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
2827379,
<collectionCode id="29B0AE5FFFE4994BFCAAFB7DFC34EEBD" box="[840,932,1251,1275]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
2831113,
<collectionCode id="29B0AE5FFFE4994BFBCFFB7DFB19EEBD" box="[1069,1161,1251,1275]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
28319,
<collectionCode id="29B0AE5FFFE4994BFB08FB7DFAD6EEBD" box="[1258,1350,1251,1275]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
28375,
<collectionCode id="29B0AE5FFFE4994BFF75FA99FF63EF59" box="[151,243,1287,1311]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
39218,
<specimenCount id="59A7FD13FFE4994BFEA9FA99FE55EF59" box="[331,453,1287,1311]" pageId="8" pageNumber="9" type="juvenile">3 juveniles</specimenCount>
,
<specimenCount id="59A7FD13FFE4994BFE2DFA99FDA0EF59" box="[463,560,1287,1311]" pageId="8" pageNumber="9" type="male">12 males</specimenCount>
and
<specimenCount id="59A7FD13FFE4994BFD87FA99FD41EF59" box="[613,721,1287,1311]" pageId="8" pageNumber="9" type="female">4 females</specimenCount>
, leg. P. Soini. Explorama Lodge, junction Rio Yanamono and Rio Amazonas (
<emphasis id="7DD5EA88FFE4994BFEF9FAB4FEA7EF05" bold="true" box="[283,311,1322,1347]" pageId="8" pageNumber="9">28</emphasis>
):
<accessionNumber id="50F2AB79FFE4994BFEADFAB5FE44EF04" box="[335,468,1322,1346]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU220370" pageId="8" pageNumber="9" type="EnaNcbi">
KU
<date id="3B1F105AFFE4994BFE9CFAB4FE44EF04" box="[382,468,1322,1346]" pageId="8" pageNumber="9" value="70-03-22">220370</date>
</accessionNumber>
, juvenile, leg. W. E. Duellman. Iquitos (
<emphasis id="7DD5EA88FFE4994BFC6DFAB4FC3BEF05" bold="true" box="[911,939,1322,1347]" pageId="8" pageNumber="9">29</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFC21FAB4FB83EF04" box="[963,1043,1322,1346]" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="8" pageNumber="9">TCWC</collectionCode>
4113738, a juvenile and a female, leg. P. Soini.
<collectionCode id="29B0AE5FFFE4994BFEC4FAD0FE11EF20" box="[294,385,1358,1382]" pageId="8" pageNumber="9">ACEER</collectionCode>
on Quebrada Grande, close to junction Rio Sucusari and Rio Napo (
<emphasis id="7DD5EA88FFE4994BFB92FAD3FB1CEF20" bold="true" box="[1136,1164,1357,1382]" pageId="8" pageNumber="9">30</emphasis>
):
<accessionNumber id="50F2AB79FFE4994BFB41FAD1FAB6EF20" box="[1187,1318,1358,1382]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU220371" pageId="8" pageNumber="9" type="EnaNcbi">
KU
<date id="3B1F105AFFE4994BFB33FAD0FAB6EF20" box="[1233,1318,1358,1382]" pageId="8" pageNumber="9" value="71-03-22">220371</date>
</accessionNumber>
, male, leg. W. E. Duellman. Yanamono (
<emphasis id="7DD5EA88FFE4994BFE3EFAEFFE68EFCC" bold="true" box="[476,504,1393,1418]" pageId="8" pageNumber="9">31</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFDEDFAEFFDFBEFCF" box="[527,619,1393,1417]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
2835455, a juvenile and a female, leg. P. Soini. Rio Orosa (
<emphasis id="7DD5EA88FFE4994BFAEFFAEFFAB9EFCC" bold="true" box="[1293,1321,1393,1418]" pageId="8" pageNumber="9">32</emphasis>
).
<collectionCode id="29B0AE5FFFE4994BFAA2FAEFFA0CEFCF" box="[1344,1436,1393,1417]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
56657, male, leg. P. Soini. Estirón, Rio Ampiyacu (
<emphasis id="7DD5EA88FFE4994BFD29FA0AFD77EFEB" bold="true" box="[715,743,1428,1453]" pageId="8" pageNumber="9">33</emphasis>
):
<collectionCode id="29B0AE5FFFE4994BFD1CFA0AFCCAEFEA" box="[766,858,1428,1452]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
13458, male, leg. B. Malkin.
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFE4994BFF25FA29FEBEED49" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="4F1E369AFFE4994BFF25FA29FEBEED49" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="9">
<emphasis id="7DD5EA88FFE4994BFF25FA29FEAFEF96" bold="true" box="[199,319,1463,1488]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
A relatively large
<taxonomicName id="88A14D19FFE4994BFDF5FA26FD1EEF96" box="[535,654,1464,1488]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">
<emphasis id="7DD5EA88FFE4994BFDF5FA26FD1EEF96" box="[535,654,1464,1488]" italics="true" pageId="8" pageNumber="9">Gonatodes</emphasis>
</taxonomicName>
, with maximum SLV of
<quantity id="88599B7FFFE4994BFC4CFA26FB87EF96" box="[942,1047,1464,1488]" metricMagnitude="-2" metricUnit="m" metricValue="5.26" pageId="8" pageNumber="9" unit="mm" value="52.6">52.6 mm</quantity>
. Scales around midbody 109131. Ventral scales in a longitudinal row 4961. Proximal subdigital lamellae as wide as digits, in total 1722 under fourth finger, 2127 under fourth toe. Three or four lateral rows of scales on distal portion of fingers and toes. Tail with midventral scales distinctly wider than long, forming a repetitive sequence of two single midventrals (one after the other), each in contact with one laterodistal scale per side, followed by a divided (only on proximal portion of tail, when present) or single midventral in contact with two laterodistal scales per side (respectively 112” and 111”). A white suprahumeral bar, bordered by black, present both in males and females, although more conspicuous in males; it reaches dorsally between the dorsolateral and middorsal regions. Males with head dorsally without vermiculations; back and limbs with a vermiculated pattern of dark and light spots; no dark streaks on gular region (at least in preservative).
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFE4994EFF25F884FBA5EBEA" lastPageId="13" lastPageNumber="14" pageId="8" pageNumber="9" type="description">
<paragraph id="4F1E369AFFE4994BFF25F884FC55EDA2" blockId="8.[151,1437,151,2020]" pageId="8" pageNumber="9">
<emphasis id="7DD5EA88FFE4994BFF25F884FEC6ED75" bold="true" box="[199,342,1818,1843]" pageId="8" pageNumber="9">Description.</emphasis>
Maximum SVL in males of
<quantity id="88599B7FFFE4994BFD79F885FD6FED75" box="[667,767,1819,1843]" metricMagnitude="-2" metricUnit="m" metricValue="5.26" pageId="8" pageNumber="9" unit="mm" value="52.6">52.6 mm</quantity>
(TCWC 36669), in females of
<quantity id="88599B7FFFE4994BFB83F885FB55ED75" box="[1121,1221,1819,1843]" metricMagnitude="-2" metricUnit="m" metricValue="4.92" pageId="8" pageNumber="9" unit="mm" value="49.2">49.2 mm</quantity>
(KU 146645). Tail round in cross section, tapering toward tip, 1.01.4 (1.15 ± 0.09,
<emphasis id="7DD5EA88FFE4994BFC96F8A1FC16ED10" box="[884,902,1855,1878]" italics="true" pageId="8" pageNumber="9">N</emphasis>
=37) times SLV. Head length 0.210.27 (0.24 ± 0.01,
<emphasis id="7DD5EA88FFE4994BFF37F8FDFF77ED3C" box="[213,231,1891,1914]" italics="true" pageId="8" pageNumber="9">N</emphasis>
=137) times SLV, 1.31.6 (1.45 ± 0.06,
<emphasis id="7DD5EA88FFE4994BFD7DF8FDFD22ED3C" box="[671,690,1891,1914]" italics="true" pageId="8" pageNumber="9">N</emphasis>
=136) times as long as wide, 1.21.7 (1.42 ± 0.08,
<emphasis id="7DD5EA88FFE4994BFB04F8FDFB69ED3C" box="[1254,1273,1891,1914]" italics="true" pageId="8" pageNumber="9">N</emphasis>
=135) times as wide as high. Snout round, moderately elongate (
<figureCitation id="D79A2A1FFFE4994BFD51F81BFD64EDDB" box="[691,756,1925,1949]" captionStart="FIGURE 5" captionStartId="11.[151,250,1964,1987]" captionTargetBox="[153,1420,238,1963]" captionTargetId="figure@11.[151,1436,167,2057]" captionTargetPageId="11" captionText="FIGURE 5. Gonatodes concinnatus (MZUSP 54655). (A) Dorsal and (B) ventral views of head; ventral views of (C) right" httpUri="https://zenodo.org/record/277518/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
A), gently sloping toward top of head. Neck slightly narrower than head and anterior portion of body. Body cylindrical. Limbs well developed, lower arm 0.110.15 (0.14 ± 0.01,
<emphasis id="7DD5EA88FFE4994BFF75F853FF3AEDA2" box="[151,170,1997,2020]" italics="true" pageId="8" pageNumber="9">N</emphasis>
=139) times SVL, lower leg 0.130.17 (0.15 ± 0.01,
<emphasis id="7DD5EA88FFE4994BFD0EF853FD6EEDA2" box="[748,766,1997,2020]" italics="true" pageId="8" pageNumber="9">N</emphasis>
=139) times SVL.
</paragraph>
<paragraph id="4F1E369AFFE5994AFF27FF09FD2FE882" blockId="9.[151,1437,151,2020]" pageId="9" pageNumber="10">
Rostral convex, 1.62.9 (2.07 ± 0.16,
<emphasis id="7DD5EA88FFE5994AFD75FF07FD3AEAF6" box="[663,682,153,176]" italics="true" pageId="9" pageNumber="10">N</emphasis>
=139) times as wide as high; posterior margin with a shallow depression medially and slightly indented by 13 medial postrostrals, with or without a median cleft extending anteriorly. Postrostrals 3 (72.0%), 4 (19.4%), or 5 (8.6%), laterals ones (supranasals) distinctly larger than medial one(s) (
<emphasis id="7DD5EA88FFE5994AFF0FFE9DFE90EB5C" box="[237,256,259,282]" italics="true" pageId="9" pageNumber="10">N</emphasis>
=139) (
<figureCitation id="D79A2A1FFFE5994AFEB0FE9CFE1FEB5C" box="[338,399,258,282]" captionStart="FIGURE 5" captionStartId="11.[151,250,1964,1987]" captionTargetBox="[153,1420,238,1963]" captionTargetId="figure@11.[151,1436,167,2057]" captionTargetPageId="11" captionText="FIGURE 5. Gonatodes concinnatus (MZUSP 54655). (A) Dorsal and (B) ventral views of head; ventral views of (C) right" httpUri="https://zenodo.org/record/277518/files/figure.png" pageId="9" pageNumber="10">Fig 5</figureCitation>
A). Nasal bordered by rostral, first supralabial (only a narrow contact in some specimens), 3 (92.1%) or 4 (7.9%) postnasals (
<emphasis id="7DD5EA88FFE5994AFDF5FEB9FDBAEB78" box="[535,554,295,318]" italics="true" pageId="9" pageNumber="10">N</emphasis>
=139), and supranasal; internostril distance 0.200.28 (0.24 ± 0.02,
<emphasis id="7DD5EA88FFE5994AFAA4FEB9FAC9EB78" box="[1350,1369,295,318]" italics="true" pageId="9" pageNumber="10">N</emphasis>
=136) times head width. Supranasal roughly oval or semicircular, 0.81.5 (1.16 ± 0.13;
<emphasis id="7DD5EA88FFE5994AFBDFFED4FBC0EB27" box="[1085,1104,330,353]" italics="true" pageId="9" pageNumber="10">N</emphasis>
=139) times as wide as long. Postnasals slightly larger than, or similar in size to, adjacent loreals. Scales on snout convex, hexagonal to round, juxtaposed, relatively uniform in size. Canthus rostralis rounded. Loreal region with scales slightly more elongate than those on snout, largest on row adjacent to supralabials, 913 (10.4 ± 0.8,
<emphasis id="7DD5EA88FFE5994AFBFCFE2BFBA1EB8A" box="[1054,1073,437,460]" italics="true" pageId="9" pageNumber="10">N</emphasis>
=138) loreals on a line between postnasals and anterior corner of eye. Top and posterior portion of head, as well as supraorbital region, with granular scales. A short supraciliary flap present, anteriorly with 612 (8.4 ± 1.1,
<emphasis id="7DD5EA88FFE5994AFBA3FE62FBC4E855" box="[1089,1108,508,531]" italics="true" pageId="9" pageNumber="10">N</emphasis>
=133) enlarged and flattened scales, among which 07 (2.0 ± 1.3;
<emphasis id="7DD5EA88FFE5994AFDA0FD81FDC5E870" box="[578,597,543,566]" italics="true" pageId="9" pageNumber="10">N</emphasis>
=133) small, conical spines. Pupil round, eye diameter 0.190.26 (0.21 ± 0.01,
<emphasis id="7DD5EA88FFE5994AFF3BFDDCFF7CE81F" box="[217,236,578,601]" italics="true" pageId="9" pageNumber="10">N</emphasis>
=129) times head length. Scales on temporal region similar to those on top of head. Ear-opening much smaller than eye, oval, posterior to, and at same level of, commissure of mouth. Supralabials 57 (6.2 ± 0.5;
<emphasis id="7DD5EA88FFE5994AFF75FD17FF3AE8E6" box="[151,170,649,672]" italics="true" pageId="9" pageNumber="10">N</emphasis>
=139), distinctly enlarged anteriorly and decreasing in size posteriorly, one or two of them posterior to centre of eye, followed to corner of mouth by small scales.
</paragraph>
<paragraph id="4F1E369AFFE5994AFF27FD51FD2FE933" blockId="9.[151,1437,151,2020]" pageId="9" pageNumber="10">
Mental large, distinctly wider anteriorly than posteriorly, with posterior margin forming a wide angle, 0.91.4 (1.1 ± 0.1;
<emphasis id="7DD5EA88FFE5994AFEF3FD6AFEB4E94D" box="[273,292,756,779]" italics="true" pageId="9" pageNumber="10">N</emphasis>
=137) times as wide as long; 1 (1.4%), 2 (82.7%), 3 (12.9%) or 4 (2.9%) postmentals (
<figureCitation id="D79A2A1FFFE5994AFB0AFD6DFABCE94D" box="[1256,1324,755,779]" captionStart="FIGURE 5" captionStartId="11.[151,250,1964,1987]" captionTargetBox="[153,1420,238,1963]" captionTargetId="figure@11.[151,1436,167,2057]" captionTargetPageId="11" captionText="FIGURE 5. Gonatodes concinnatus (MZUSP 54655). (A) Dorsal and (B) ventral views of head; ventral views of (C) right" httpUri="https://zenodo.org/record/277518/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
B). Scales on chin flat, smooth, polygonal, juxtaposed, larger anteriorly, decreasing in size posteriorly. Infralabials 47 (5.5 ± 0.7;
<emphasis id="7DD5EA88FFE5994AFF29FCA5FF4EE914" box="[203,222,827,850]" italics="true" pageId="9" pageNumber="10">N</emphasis>
=138), distinctly enlarged anteriorly and decreasing in size posteriorly; 03 of them posterior to centre of eye, followed to corner of mouth by small scales.
</paragraph>
<paragraph id="4F1E369AFFE5994AFF27FC1EFC8BE9A6" blockId="9.[151,1437,151,2020]" pageId="9" pageNumber="10">Scales on nape small and granular, becoming slightly larger on sides of neck. Scales on throat anteriorly granular, posteriorly flat, smooth, hexagonal or round, increasing in size toward the posterior region, with a short transitional zone between the anterior and posterior parts.</paragraph>
<paragraph id="4F1E369AFFE5994AFF27FC75FEC9EE9E" blockId="9.[151,1437,151,2020]" pageId="9" pageNumber="10">
Dorsals granular, increasing in size toward the flanks. Ventrals larger than dorsals, roughly hexagonal, flat, smooth, imbricate, in oblique rows; 4961 (56.1 ± 2.1;
<emphasis id="7DD5EA88FFE5994AFCF2FB8EFCB3EE61" box="[784,803,1040,1063]" italics="true" pageId="9" pageNumber="10">N</emphasis>
=119) scales along the midventral line between anterior margin of forelimbs and vent; 1720 (18.5 ± 0.9;
<emphasis id="7DD5EA88FFE5994AFD07FBADFD68EE0C" box="[741,760,1075,1098]" italics="true" pageId="9" pageNumber="10">N</emphasis>
=126) scales in a transverse line at midbody, with a short transitional zone between ventrals and scales on flanks. Scales around midbody 109131 (120.7 ± 4.6;
<emphasis id="7DD5EA88FFE5994AFAA2FBC9FAC3EE28" box="[1344,1363,1111,1134]" italics="true" pageId="9" pageNumber="10">N</emphasis>
=110). Scales on preanal plate similar to ventrals, except for those bordering vent, which are very small. Escutcheon present in males on posterior portion of belly and on 35 (mostly four) rows (body-knee direction) on ventral surface of thighs.
</paragraph>
<paragraph id="4F1E369AFFE5994AFF27FB7DFD98EFCF" blockId="9.[151,1437,151,2020]" pageId="9" pageNumber="10">Scales on anterodorsal surface of forelimbs flat, smooth, roundish or rhomboidal, imbricate, largest close to the wrist; on posterodorsal and ventral surface of forelimbs convex, smooth, rhomboidal, juxtaposed, relatively small. Scales on anterodorsal surface of thighs and ventral surface of hind limbs flat, smooth, roundish or rhomboidal, imbricate and relatively larger; on posterodorsal surface of thighs and dorsal surface of lower legs smaller, convex, smooth, rhomboidal, juxtaposed.</paragraph>
<paragraph id="4F1E369AFFE5994AFF27FA0AFE44EC18" blockId="9.[151,1437,151,2020]" pageId="9" pageNumber="10">
Lamellae under second (
<emphasis id="7DD5EA88FFE5994AFE3DFA0BFE65EFEB" bold="true" box="[479,501,1429,1453]" pageId="9" pageNumber="10">II</emphasis>
) through fourth (
<emphasis id="7DD5EA88FFE5994AFD5CFA0BFD4DEFEB" bold="true" box="[702,733,1429,1453]" pageId="9" pageNumber="10">IV</emphasis>
) fingers (proximal lamellae in parentheses):
<emphasis id="7DD5EA88FFE5994AFB00FA0BFB68EFEB" bold="true" box="[1250,1272,1429,1453]" pageId="9" pageNumber="10">II</emphasis>
: 1418 (57),
<emphasis id="7DD5EA88FFE5994AFF75FA26FF28EF96" bold="true" box="[151,184,1464,1488]" pageId="9" pageNumber="10">III</emphasis>
: 1621 (57), and
<emphasis id="7DD5EA88FFE5994AFE6CFA26FE3BEF96" bold="true" box="[398,427,1464,1488]" pageId="9" pageNumber="10">IV</emphasis>
: 1722 (59) (
<figureCitation id="D79A2A1FFFE5994AFDB6FA26FD06EF96" box="[596,662,1464,1488]" captionStart="FIGURE 5" captionStartId="11.[151,250,1964,1987]" captionTargetBox="[153,1420,238,1963]" captionTargetId="figure@11.[151,1436,167,2057]" captionTargetPageId="11" captionText="FIGURE 5. Gonatodes concinnatus (MZUSP 54655). (A) Dorsal and (B) ventral views of head; ventral views of (C) right" httpUri="https://zenodo.org/record/277518/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
C). Lamellae under second through fourth toes (proximal lamellae in parentheses):
<emphasis id="7DD5EA88FFE5994AFED6FA42FEDAEFB2" bold="true" box="[308,330,1500,1524]" pageId="9" pageNumber="10">II</emphasis>
: 1418 (57),
<emphasis id="7DD5EA88FFE5994AFE15FA42FD88EFB2" bold="true" box="[503,536,1500,1524]" pageId="9" pageNumber="10">III</emphasis>
: 1622 (58), and
<emphasis id="7DD5EA88FFE5994AFD14FA42FC83EFB2" bold="true" box="[758,787,1500,1524]" pageId="9" pageNumber="10">IV</emphasis>
: 2127 (913) (
<figureCitation id="D79A2A1FFFE5994AFC32FA45FB85EFB5" box="[976,1045,1499,1523]" captionStart="FIGURE 5" captionStartId="11.[151,250,1964,1987]" captionTargetBox="[153,1420,238,1963]" captionTargetId="figure@11.[151,1436,167,2057]" captionTargetPageId="11" captionText="FIGURE 5. Gonatodes concinnatus (MZUSP 54655). (A) Dorsal and (B) ventral views of head; ventral views of (C) right" httpUri="https://zenodo.org/record/277518/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
D). Claws exposed, non-retractile, between two basal scales. Fingers and toes with three, occasionally four or two (USNM 234497; USNM 234494 and TCWC 42706), lateral scales distally, between the fourth (counted from the claw towards the hand) subdigital lamella and the dorsal scale.
</paragraph>
<paragraph id="4F1E369AFFE5994AFF27F9F7FC26ED75" blockId="9.[151,1437,151,2020]" pageId="9" pageNumber="10">
Scales on tail dorsally and laterally relatively small, rhomboidal, flat, smooth, imbricate. On ventral surface of tail scales smooth, flat, imbricate, increasing in size toward midventral line. Midventral scales, except close to the base of the tail, distinctly wider than long, forming a repetitive sequence of two single midventrals (one after the other), each in contact laterodistally with one scale per side, followed by a divided (only on proximal portion of tail, when present) or single midventral scale in contact laterodistally with two scales per side respectively 112” and 111” in the codification of
<bibRefCitation id="2B304B6BFFE5994AFDE5F885FCD3ED75" author="Avila-Pires" box="[519,835,1819,1843]" pageId="9" pageNumber="10" refString="Avila-Pires, T. C. S. (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen, 299, 1 - 706." type="journal article" year="1995">
Avila-Pires (1995:
<figureCitation id="D79A2A1FFFE5994AFD3EF885FCABED75" box="[732,827,1819,1843]" captionStart="FIGURE 2" captionStartId="6.[151,250,1440,1463]" captionTargetBox="[190,1402,193,1415]" captionTargetId="figure@6.[185,1402,193,1419]" captionTargetPageId="6" captionText="FIGURE 2. Bivariate scattergrams from stepwise discriminant function analysis using scale counts and measurements. Red = Pattern A. Cyan-green = Pattern B. Lime-green = Pattern C. Yellow = Pattern D. Blue = Pattern E." httpUri="https://zenodo.org/record/277515/files/figure.png" pageId="9" pageNumber="10">Figure 2</figureCitation>
)
</bibRefCitation>
(
<figureCitation id="D79A2A1FFFE5994AFCB0F885FC05ED75" box="[850,917,1819,1843]" captionStart="FIGURE 5" captionStartId="11.[151,250,1964,1987]" captionTargetBox="[153,1420,238,1963]" captionTargetId="figure@11.[151,1436,167,2057]" captionTargetPageId="11" captionText="FIGURE 5. Gonatodes concinnatus (MZUSP 54655). (A) Dorsal and (B) ventral views of head; ventral views of (C) right" httpUri="https://zenodo.org/record/277518/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
E).
</paragraph>
<paragraph id="4F1E369AFFE59949FF27F8A0FB4DEA92" blockId="9.[151,1437,151,2020]" lastBlockId="10.[151,1436,151,426]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="7DD5EA88FFE5994AFF27F8A0FE42ED11" bold="true" box="[197,466,1854,1879]" pageId="9" pageNumber="10">Color in preservative.</emphasis>
In males (
<figureCitation id="D79A2A1FFFE5994AFDB6F8A0FD32ED10" box="[596,674,1854,1878]" captionStart="FIGURE 1" captionStartId="5.[151,250,1952,1975]" captionTargetBox="[228,1363,223,1918]" captionTargetId="figure@5.[211,1387,197,1926]" captionTargetPageId="5" captionText="FIGURE 1. Dorsal and ventral color patterns in preservative in the Gonatodes concinnatus complex. (A) pattern A (MZUSP 54655); (B) pattern B (TCWC 57237); (C) pattern C (MZUSP 53674, holotype); (D) pattern D (MPEG 25596, holotype); and (E) pattern E (MZUSP 49153, holotype). Scale bar = 5 mm." httpUri="https://zenodo.org/record/277514/files/figure.png" pageId="9" pageNumber="10">Figs. 1</figureCitation>
A, 4), dorsal surface of head beige, without markings. Back and flanks, base of tail and hind limbs with relatively large, beige and brown vermiculations. A large, conspicuous, white suprahumeral bar, bordered by black, extending dorsally at least up to the dorsolateral region, in some cases almost reaching the middorsal region; never in the form of an ocellus, but
<bibRefCitation id="2B304B6BFFE5994AFC3BF837FB25ED87" author="Vitt" box="[985,1205,1961,1985]" pageId="9" pageNumber="10" refString="Vitt, L. J. &amp; de la Torre, S. 1996. Guia para la Investigacion de las Lagartijas de Cuyabeno - A Research Guide to the Lizards of Cuyabeno. Museo de Zoologia (QCAZ), Centro de Biodiversidad y Ambiente, Pontificia Universidad Catolica del Ecuador, Monografia 1, Quito, 165 p." type="book" year="1996">Vitt &amp; Torre (1996)</bibRefCitation>
present the photo of a specimen with the white bar divided into two spots by a transversal black band. Ventral surface of head, gular region and chest beige or reddish-brown, without oblique streaks; belly gray; underside of limbs beige. Tail brown and/or black dorsally, white and/or brown ventrally. Escutcheon area (belly and thighs) light gray.
</paragraph>
<paragraph id="4F1E369AFFE69949FF27FF7EFCEBEBEC" blockId="10.[151,1436,151,426]" pageId="10" pageNumber="11">In females, dorsal surface of head and limbs with brown and black irregular spots; back gray with dorsolateral pairs of black spots and, in some specimens, pairs of beige spots; flanks gray with black and brown spots. Suprahumeral bar conspicuous, white with black margins, similar in extension but thinner than that of males. Ventral surface of head and gular region white with dark oblique streaks, in contact or not at midventral line; belly and underside of limbs light gray. Tail brown and/or black dorsally, white and/or brown ventrally; original tail distally with white bands that form complete rings around the tail.</paragraph>
<caption id="1BDE6612FFE69949FF75F82BFE25EDAC" httpUri="https://zenodo.org/record/277517/files/figure.png" pageId="10" pageNumber="11" targetBox="[265,1321,480,1959]" targetPageId="10">
<paragraph id="4F1E369AFFE69949FF75F82BFE25EDAC" blockId="10.[151,1437,1973,2026]" pageId="10" pageNumber="11">
<emphasis id="7DD5EA88FFE69949FF75F82BFE84ED8D" bold="true" box="[151,276,1973,1996]" pageId="10" pageNumber="11">FIGURE 4.</emphasis>
<taxonomicName id="88A14D19FFE69949FEFFF828FD9DED8A" box="[285,525,1974,1996]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="concinnatus">
<emphasis id="7DD5EA88FFE69949FEFFF828FD9DED8A" box="[285,525,1974,1996]" italics="true" pageId="10" pageNumber="11">Gonatodes concinnatus</emphasis>
</taxonomicName>
(syntypes, BMNH 1946.9.7.1012). (A) Dorsal, (B) ventral views. Scale bar = 50 mm. (Photo by Colin McCarthy).
</paragraph>
</caption>
<caption id="1BDE6612FFE79948FF75F832FA0CED84" box="[151,1436,1964,1987]" httpUri="https://zenodo.org/record/277518/files/figure.png" pageId="11" pageNumber="12" targetBox="[153,1420,238,1963]" targetPageId="11">
<paragraph id="4F1E369AFFE79948FF75F832FA0CED84" blockId="11.[151,1436,1964,1987]" box="[151,1436,1964,1987]" pageId="11" pageNumber="12">
<emphasis id="7DD5EA88FFE79948FF75F832FE85ED84" bold="true" box="[151,277,1964,1987]" pageId="11" pageNumber="12">FIGURE 5.</emphasis>
<taxonomicName id="88A14D19FFE79948FEFFF832FD9FED84" box="[285,527,1964,1986]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="concinnatus">
<emphasis id="7DD5EA88FFE79948FEFFF832FD9FED84" box="[285,527,1964,1986]" italics="true" pageId="11" pageNumber="12">Gonatodes concinnatus</emphasis>
</taxonomicName>
(MZUSP 54655). (A) Dorsal and (B) ventral views of head; ventral views of (C) right
</paragraph>
</caption>
<caption id="1BDE6612FFE0994FFF75F80EFD34ED83" httpUri="https://zenodo.org/record/277519/files/figure.png" pageId="12" pageNumber="13" targetBox="[151,1436,193,1899]" targetPageId="12">
<paragraph id="4F1E369AFFE0994FFF75F80EFD34ED83" blockId="12.[151,1436,1936,1989]" pageId="12" pageNumber="13">
<emphasis id="7DD5EA88FFE0994FFF75F80EFE84EDE0" bold="true" box="[151,276,1936,1959]" pageId="12" pageNumber="13">FIGURE 6.</emphasis>
<taxonomicName id="88A14D19FFE0994FFEFEF80FFD9BEDE1" box="[284,523,1937,1959]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="concinnatus">
<emphasis id="7DD5EA88FFE0994FFEFEF80FFD9BEDE1" box="[284,523,1937,1959]" italics="true" pageId="12" pageNumber="13">Gonatodes concinnatus</emphasis>
</taxonomicName>
from Cuyabeno, Sucumbios, Ecuador. (A) Adult male (LJV 5014) and (B) adult female (OMNH 36409; LJV 5046) (Photos by Laurie Vitt).
</paragraph>
</caption>
<paragraph id="4F1E369AFFE1994EFF25FF09FF49EB79" blockId="13.[151,1437,151,536]" pageId="13" pageNumber="14">
<emphasis id="7DD5EA88FFE1994EFF25FF09FECBEAF6" bold="true" box="[199,347,151,176]" pageId="13" pageNumber="14">Color in life.</emphasis>
Color in life has been described by
<bibRefCitation id="2B304B6BFFE1994EFD09FF09FC37EAE9" author="Duellman" box="[747,935,151,175]" pageId="13" pageNumber="14" refString="Duellman, W. E. (1978) The biology of an Equatorial Herpetofauna in Amazonian Ecuador. Miscellaneous Publications, Museum of Natural History, University of Kansas, 65, 1 - 352." type="journal article" year="1978">Duellman (1978)</bibRefCitation>
and Vitt &amp; de la
<bibRefCitation id="2B304B6BFFE1994EFB8AFF06FB66EAE9" author="Torre" box="[1128,1270,151,175]" pageId="13" pageNumber="14" refString="Vitt, L. J. &amp; de la Torre, S. 1996. Guia para la Investigacion de las Lagartijas de Cuyabeno - A Research Guide to the Lizards of Cuyabeno. Museo de Zoologia (QCAZ), Centro de Biodiversidad y Ambiente, Pontificia Universidad Catolica del Ecuador, Monografia 1, Quito, 165 p." type="book" year="1996">Torre (1996)</bibRefCitation>
. In males (
<figureCitation id="D79A2A1FFFE1994EFA92FF09FF35EA92" captionStart="FIGURE 6" captionStartId="12.[151,250,1936,1959]" captionTargetBox="[151,1436,193,1899]" captionTargetId="figure@12.[151,1436,193,1899]" captionTargetPageId="12" captionText="FIGURE 6. Gonatodes concinnatus from Cuyabeno, Sucumbios, Ecuador. (A) Adult male (LJV 5014) and (B) adult female (OMNH 36409; LJV 5046) (Photos by Laurie Vitt)." httpUri="https://zenodo.org/record/277519/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
A) head, neck and forelegs orange or reddish brown with cream spots; suprahumeral bar white bordered by black; body olive-green or brown, with reddish brown or yellowish white vermiculations that may present a black margin. Throat orange with cream streaks, belly yellowish gray, and tail black with white rings, or the entire belly and tail black.
</paragraph>
<paragraph id="4F1E369AFFE1994EFF27FED3FBA5EBEA" blockId="13.[151,1437,151,536]" pageId="13" pageNumber="14">
In females (
<figureCitation id="D79A2A1FFFE1994EFEACFED2FE03EB22" box="[334,403,332,356]" captionStart="FIGURE 6" captionStartId="12.[151,250,1936,1959]" captionTargetBox="[151,1436,193,1899]" captionTargetId="figure@12.[151,1436,193,1899]" captionTargetPageId="12" captionText="FIGURE 6. Gonatodes concinnatus from Cuyabeno, Sucumbios, Ecuador. (A) Adult male (LJV 5014) and (B) adult female (OMNH 36409; LJV 5046) (Photos by Laurie Vitt)." httpUri="https://zenodo.org/record/277519/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
B), head, body and tail grayish tan or drab gray with irregular crossbands, brown or black anteriorly and white posteriorly; distally the tail becomes black and white banded. Throat cream with dark streaks; venter creamy tan or yellow. Iris reddish brown with a cream circle around the pupil.
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFE1994EFF25FE29FB86EB89" box="[199,1046,439,464]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="4F1E369AFFE1994EFF25FE29FB86EB89" blockId="13.[151,1437,151,536]" box="[199,1046,439,464]" pageId="13" pageNumber="14">
<emphasis id="7DD5EA88FFE1994EFF25FE29FECDEB96" bold="true" box="[199,349,439,464]" pageId="13" pageNumber="14">Distribution.</emphasis>
Western Amazonia, in
<collectingCountry id="37B6760AFFE1994EFDBFFE29FD28EB89" box="[605,696,439,463]" name="Ecuador" pageId="13" pageNumber="14">Ecuador</collectingCountry>
and northern
<collectingCountry id="37B6760AFFE1994EFCB2FE26FC12EB89" box="[848,898,440,463]" name="Peru" pageId="13" pageNumber="14">Peru</collectingCountry>
(
<figureCitation id="D79A2A1FFFE1994EFC73FE29FC41EB89" box="[913,977,439,463]" captionStart="FIGURE 3" captionStartId="7.[151,250,1859,1882]" captionTargetBox="[239,1345,203,1826]" captionTargetId="figure@7.[235,1351,195,1833]" captionTargetPageId="7" captionText="FIGURE 3. (A) Geographic distribution of Gonatodes concinnatus complex. (B) Detailed geographic distribution of Gonatodes concinnatus. Red = Gonatodes concinnatus (Pattern A). Cyan-green = Gonatodes ligiae (Pattern B). Lime-green = Gonatodes tapajonicus (Pattern C). Yellow = Gonatodes nascimentoi sp. nov. (Pattern D). Blue = Gonatodes riveroi sp. nov. (Pattern E). Open symbols = type-localities; half-black symbol = literature record; question mark = Pallatanga, Ecuador, probably in error; triangle = locality referred to only as Rio Napo. Numbers refer to localities cited under each species. One number may represent more than one locality." httpUri="https://zenodo.org/record/277516/files/figure.png" pageId="13" pageNumber="14">Fig. 3</figureCitation>
AB).
</paragraph>
</subSubSection>
<subSubSection id="07BB6511FFE1994EFF25FE42FAD6E851" pageId="13" pageNumber="14" type="discussion">
<paragraph id="4F1E369AFFE1994EFF25FE42FAD6E851" blockId="13.[151,1437,151,536]" pageId="13" pageNumber="14">
<emphasis id="7DD5EA88FFE1994EFF25FE42FEA9EBB2" bold="true" box="[199,313,476,500]" pageId="13" pageNumber="14">Remarks.</emphasis>
<taxonomicName id="88A14D19FFE1994EFEDDFE42FD7FEBB2" authority="Donoso-Barros, 1967" authorityName="Donoso-Barros" authorityYear="1967" box="[319,751,476,500]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ligiae">
<emphasis id="7DD5EA88FFE1994EFEDDFE42FE6BEBB2" box="[319,507,476,500]" italics="true" pageId="13" pageNumber="14">Gonatodes ligiae</emphasis>
<bibRefCitation id="2B304B6BFFE1994EFDE0FE43FD7FEBB2" author="Donoso-Barros" box="[514,751,476,500]" pageId="13" pageNumber="14" refString="Donoso-Barros, R. (1967) Diagnosis de dos nuevas especies del genero Gonatodes de Venezuela. Noticiario Mensual Del Museo Nacional de Historia Natural de Santiago, 129, no pagination." type="journal volume" year="1967">Donoso-Barros, 1967</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="37B6760AFFE1994EFCD5FE42FC3AEBB2" box="[823,938,476,500]" name="Venezuela" pageId="13" pageNumber="14">Venezuela</collectingCountry>
, is considered in this paper as a valid species. Material from
<collectingCountry id="37B6760AFFE1994EFED4FE61FE32E851" box="[310,418,511,535]" name="Colombia" pageId="13" pageNumber="14">Colombia</collectingCountry>
previously identified as
<taxonomicName id="88A14D19FFE1994EFD52FD9EFCC1E85E" box="[688,849,512,536]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="concinnatus">
<emphasis id="7DD5EA88FFE1994EFD52FD9EFCC1E85E" box="[688,849,512,536]" italics="true" pageId="13" pageNumber="14">G. concinnatus</emphasis>
</taxonomicName>
is here described as a new species,
<taxonomicName id="88A14D19FFE1994EFB3AFD9EFAAFE85E" box="[1240,1343,512,536]" class="Reptilia" family="Sphaerodactylidae" genus="Gonatodes" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="riveroi">
<emphasis id="7DD5EA88FFE1994EFB3AFD9EFAAFE85E" box="[1240,1343,512,536]" italics="true" pageId="13" pageNumber="14">G. riveroi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>