213 lines
20 KiB
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213 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.690.13751" ID-GBIF-Dataset="356a4c29-470b-4ca5-8503-76191b4fef6f" ID-PMC="PMC5672587" ID-Pensoft-Pub="1313-2970-690-1" ID-PubMed="29118595" ID-ZBK="C1B8D7C059E54C3A944F69F4FDE96B20" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-690-1" ModsDocOrigin="ZooKeys 690" ModsDocTitle="A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography" checkinTime="1502920743207" checkinUser="pensoft" docAuthor="Shpeley, Danny, Hunting, Wesley & Ball, George E." docDate="2017" docId="C6959DE428FAE57A04A2D3694A9749FA" docLanguage="en" docName="ZooKeys 690: 1-195" docOrigin="ZooKeys 690" docSource="http://dx.doi.org/10.3897/zookeys.690.13751" docTitle="Selenophorus striatopunctatus Putzeys" docType="treatment" docVersion="5" lastPageNumber="83" masterDocId="FE026465FF86843AE44A7B6EFF8DC422" masterDocTitle="A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography" masterLastPageNumber="195" masterPageNumber="1" pageNumber="82" updateTime="1668164690231" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography</mods:title>
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Shpeley, Danny</mods:namePart>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hunting, Wesley</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ball, George E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>690</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>195</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.690.13751</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.690.13751</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-690-1</mods:identifier>
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<mods:identifier type="ZBK">C1B8D7C059E54C3A944F69F4FDE96B20</mods:identifier>
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<mods:identifier type="ZooBank">C1B8D7C059E54C3A944F69F4FDE96B20</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="132785163" LSID="urn:lsid:plazi:treatment:C6959DE428FAE57A04A2D3694A9749FA" httpUri="http://treatment.plazi.org/id/C6959DE428FAE57A04A2D3694A9749FA" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82">
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<subSubSection pageId="81" pageNumber="82" type="nomenclature">
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<paragraph pageId="81" pageNumber="82">
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<taxonomicName authority="Putzeys" class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus striatopunctatus" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="striatopunctatus">Selenophorus striatopunctatus Putzeys</taxonomicName>
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Figs 58, 59
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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, 60, 61
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</paragraph>
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</subSubSection>
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<subSubSection pageId="81" pageNumber="82" type="reference_group">
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<paragraph pageId="81" pageNumber="82">
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus striatopunctatus" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="striatopunctatus">Selenophorus striatopunctatus</taxonomicName>
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Putzeys, 1878a: 33. SYNTYPES (5) in the Putzeys Collection (IRSB), and (5) in the
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<normalizedToken originalValue="Chaudoir-Oberthür">Chaudoir-Oberthuer</normalizedToken>
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Collection (MNHP). IRSB specimens as follows. 1, male [[indecipherable writing] VII.44 [green paper] // Putzeys Collection label// Type//; 2, male, Chiapas/ 5.7.58 Putzeys Collection label// Type//; 3, Costarico [green paper]// Putzeys Collection label// Type//; 4, male, St. Doming [green paper]// Putzeys Collection label// Type//.-
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<taxonomicName class="Insecta" family="Carabidae" genus="Amblygnathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amblygnathus puncticollis" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="puncticollis">Amblygnathus puncticollis</taxonomicName>
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Putz./Emd. det, 1937//.- 5, male, Mex / 3.7.44// Putzeys Collection label// Type// [specimen a
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<taxonomicName class="Insecta" family="Carabidae" genus="Pelmatellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pelmatellus" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">Pelmatellus</taxonomicName>
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sp.]; LECTOTYPE (here selected), specimen #2, above. MNHP specimens as follows. Box label
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophori" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selenophori striatopunctatus" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="striatopunctatus">striatopunctatus</taxonomicName>
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/ Chaud./ Antilles. 1, male, labelled Rep. Dominginie/
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<normalizedToken originalValue="Sallé">Salle</normalizedToken>
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// 293// 2, 402//; 3, male, labelled Mexique// A. Deyrolle//; 4, female, unlabelled; 5, female, labelled Mexique/.-
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<bibRefCitation pageId="81" pageNumber="82">Csiki 1932</bibRefCitation>
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: 1201.-
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<bibRefCitation author="Darlington, PJ Jr" journalOrPublisher="Psyche" pageId="103" pageNumber="104" pagination="66 - 131" title="New West Indian Carabidae with a list of the Cuban species." url="https://doi.org/10.1155/1934/10161" volume="41" year="1934">Darlington 1934</bibRefCitation>
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: 104.-
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<bibRefCitation pageId="81" pageNumber="82">Blackwelder 1944</bibRefCitation>
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: 50.-
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<bibRefCitation author="Erwin, TL" journalOrPublisher="Quaestiones Entomologicae" pageId="104" pageNumber="105" pagination="350 - 466" title="Carabid beetles of the West Indies (Insects: Coleoptera): a synopsis of the genera and checklists of tribes of Caraboidea, and of the West Indian species." volume="20" year="1984">Erwin and Sims 1984</bibRefCitation>
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: 441.-
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<bibRefCitation author="Bennett, FD" journalOrPublisher="Caribbean Agricultural Research and Development Institute, Wildey" pageId="102" pageNumber="103" title="An annotated check-list of the insects and allied terrestrial arthropods of Barbados." year="1985">Bennett and Alam 1985</bibRefCitation>
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: 20.-
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<bibRefCitation pageId="81" pageNumber="82">Ball 1992</bibRefCitation>
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: 85.-
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<bibRefCitation author="Lorenz, W" journalOrPublisher="Lorenz, Tutzing" pageId="105" pageNumber="106" title="Systematic list of ground beetles of the World (InsectaColeoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). W." year="1998">Lorenz 1998</bibRefCitation>
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: 356.-
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<bibRefCitation author="Peck, SB" journalOrPublisher="Florida Department of Agriculture and Consumer Services, Gainesville" pageId="106" pageNumber="107" title="A distributional checklist of the Beetles (Coleoptera) of Florida. Arthropods of Florida and neighboring land areas, volume 16." year="1998">Peck and Thomas 1998</bibRefCitation>
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: 22.-
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<bibRefCitation pageId="81" pageNumber="82">Lorenz 2005</bibRefCitation>
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: 377.-
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<bibRefCitation author="Peck, SB" journalOrPublisher="Florida Department of Agriculture and Consumer Services, Gainesville" pageId="106" pageNumber="107" title="A checklist of the beetles of Cuba, with data on distribution and bionomics (Insecta: Coleoptera). Arthropods of Florida and neighboring land areas, Volume 18." year="2005">Peck 2005</bibRefCitation>
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: 33.-
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<bibRefCitation author="Perez-Gelabert, DE" journalOrPublisher="Zootaxa" pageId="106" pageNumber="107" pagination="1 - 530" title="Arthropods of Hispaniola (Dominican Republic and Haiti): A checklist and bibliography." volume="1831" year="2008">Perez-Gelabert 2008</bibRefCitation>
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: 80.-
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<bibRefCitation author="Turnbow, RH" journalOrPublisher="Insecta Mundi" pageId="106" pageNumber="107" pagination="1 - 66" title="An annotated checklist of the Coleoptera (Insecta) of the Bahamas." volume="0034" year="2008">Turnbow and Thomas 2008</bibRefCitation>
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: 15.-
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<bibRefCitation author="Peck, SB" journalOrPublisher="Insecta Mundi" pageId="106" pageNumber="107" pagination="1 - 51" title="The Beetles of Barbados, West Indies (Insecta: Coleoptera): diversity, distribution and faunal structure." volume="0073" year="2009">Peck 2009</bibRefCitation>
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: 13.-
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<bibRefCitation author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="102" pageNumber="103" pagination="1 - 1722" title="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico." url="https://doi.org/10.3897/zookeys.245.3416" volume="245" year="2012">Bousquet 2012</bibRefCitation>
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: 1147.
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</paragraph>
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<paragraph pageId="81" pageNumber="82">
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<taxonomicName class="Insecta" family="Carabidae" genus="Hemisopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemisopalus vigilans" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="vigilans">Hemisopalus vigilans</taxonomicName>
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Casey, 1914: 137. LECTOTYPE (here selected) male, labelled Fla// CASEY/ bequest/ 1925// TYPE USNM/ 47869 [red paper]//
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophori" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selenophori vigilans" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="vigilans">vigilans</taxonomicName>
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/ Csy// (USNM).-
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<bibRefCitation author="Peck, SB" journalOrPublisher="Florida Department of Agriculture and Consumer Services, Gainesville" pageId="106" pageNumber="107" title="A distributional checklist of the Beetles (Coleoptera) of Florida. Arthropods of Florida and neighboring land areas, volume 16." year="1998">Peck and Thomas (1998</bibRefCitation>
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: 22).-
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<bibRefCitation author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="102" pageNumber="103" pagination="1 - 1722" title="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico." url="https://doi.org/10.3897/zookeys.245.3416" volume="245" year="2012">Bousquet 2012</bibRefCitation>
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: 1147.
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</paragraph>
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<paragraph pageId="81" pageNumber="82">
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<taxonomicName class="Insecta" family="Carabidae" genus="Hemisopalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemisopalus depressulus" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="depressulus">Hemisopalus depressulus</taxonomicName>
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Casey, 1914: 137. LECTOTYPE selected by
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<bibRefCitation author="Lindroth, CH" journalOrPublisher="The Coleopterists Bulletin" pageId="105" pageNumber="106" pagination="109 - 147" title="Designation of holotypes and lectotypes among ground beetles (Coleoptera, Carabidae) described by Thomas L. Casey." volume="29" year="1975">Lindroth (1975</bibRefCitation>
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: 141) male, labelled Fla// Casey/ bequest/ 1925// TYPE USNM/ 47867 [red paper]//
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophori" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selenophori depressulus" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="depressulus">depressulus</taxonomicName>
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/ Csy [handwritten] (USNM); (synonymy established by
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<bibRefCitation author="Peck, SB" journalOrPublisher="Florida Department of Agriculture and Consumer Services, Gainesville" pageId="106" pageNumber="107" title="A distributional checklist of the Beetles (Coleoptera) of Florida. Arthropods of Florida and neighboring land areas, volume 16." year="1998">Peck and Thomas (1998</bibRefCitation>
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: 22).-
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<bibRefCitation pageId="81" pageNumber="82">Csiki 1932</bibRefCitation>
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: 1199.- Ball and Maddsion 1987: 206.-
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<bibRefCitation author="Lorenz, W" journalOrPublisher="Lorenz, Tutzing" pageId="105" pageNumber="106" title="Systematic list of ground beetles of the World (InsectaColeoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). W." year="1998">Lorenz 1998</bibRefCitation>
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: 355.-
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<bibRefCitation pageId="81" pageNumber="82">Lorenz 2005</bibRefCitation>
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: 377.-
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<bibRefCitation author="Bousquet, Y" journalOrPublisher="ZooKeys" pageId="102" pageNumber="103" pagination="1 - 1722" title="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico." url="https://doi.org/10.3897/zookeys.245.3416" volume="245" year="2012">Bousquet 2012</bibRefCitation>
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: 1147.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="81" pageNumber="82" type="type area">
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<paragraph pageId="81" pageNumber="82">Type area.</paragraph>
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<paragraph pageId="81" pageNumber="82">State of Chiapas, Mexico.</paragraph>
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</subSubSection>
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<subSubSection pageId="81" pageNumber="82" type="diagnosis">
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<paragraph pageId="81" pageNumber="82">Diagnosis.</paragraph>
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<paragraph pageId="81" pageNumber="82">
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This species is readily separated from the other
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">Selenophorus</taxonomicName>
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species by the punctate elytral striae.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82" type="descriptive notes">
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<paragraph pageId="81" pageNumber="82">Descriptive notes.</paragraph>
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<paragraph lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82">
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Data for SBL in Table 1. Habitus as in Fig. 58. Clypeus and labrum with anterior margin of each shallowly concave. Antennae with antennomere 1 testaceous to rufo-testaceous, antennomeres 2-11 darker; mouthparts infuscated or not, testaceous to rufo-testaceous; legs testaceous to dark rufo-testaceous. Head and pronotum brunneous to brunneo-piceous; elytra brunneo-piceous to nearly piceous, suture and apical margin diffusely paler. Ventral surface rufo-brunneous to brunneo-piceous; elytral epipleuron paler than disc. Pronotum with faint bluish metallic luster; elytra with faint to moderate iridescence; ventral surface faintly iridescent. Head shiny, with mesh pattern isodiametric, microlines very fine; pronotum shiny, with mesh pattern slightly
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<pageBreakToken pageId="82" pageNumber="83" start="start">transverse</pageBreakToken>
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, sculpticells about 1.5
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<normalizedToken originalValue="–2×">-2x</normalizedToken>
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wide as long, microlines very fine; elytra very shiny, microlines not visible at 100
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<normalizedToken originalValue="×">x</normalizedToken>
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. Pronotum with posteriolateral impressions punctate; hind angles rounded. Elytral striae punctate, in addition to the standard setigerous punctures in striae 2, 5 and 7. Elytral intervals finely punctate. Males with two terminal setae and females with four terminal setae near the posterior margin on sternum VII.
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</paragraph>
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<paragraph pageId="82" pageNumber="83">
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Male genitalia. Fig. 59
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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. Apical portion of phallic median lobe short, broad, apex symmetrically rounded in dorsal and ventral aspects; endophallus with 17 spines with large bases scattered throughout entire length; without lamina. Ventral surface of shaft smooth.
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</paragraph>
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<paragraph pageId="82" pageNumber="83">Ovipositor and female reproductive tract. Fig. 60. Gonocoxite 2 (gc2) moderately thick, slightly falcate. Bursa copulatrix (bc) markedly long; spermatheca (sp) moderately long, coiled, sausage-like, originating near base of common oviduct (co); markedly long spermathecal gland duct (spgd) originating above base of spermatheca. Spermathecal gland (spg) somewhat dumbbell-like, narrowed in the middle.</paragraph>
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</subSubSection>
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<subSubSection pageId="82" pageNumber="83" type="distribution">
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<paragraph pageId="82" pageNumber="83">Geographical distribution.</paragraph>
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<paragraph pageId="82" pageNumber="83">Fig. 61. This wide-ranging species is found on most of the island groups in the West Indies, with the exception of the islands located between the Greater Antillean Puerto Rico and Lesser Antillean Guadeloupe.</paragraph>
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</subSubSection>
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<subSubSection pageId="82" pageNumber="83" type="chorological affinities and relationships">
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<paragraph pageId="82" pageNumber="83">Chorological affinities and relationships.</paragraph>
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<paragraph pageId="82" pageNumber="83">
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The range of this species overlaps the ranges of most
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">Selenophorus</taxonomicName>
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species. Relationships of
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<taxonomicName lsidName="S. striatopunctatus" pageId="82" pageNumber="83" rank="species" species="striatopunctatus">S. striatopunctatus</taxonomicName>
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are not postulated beyond species group membership.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="82" pageNumber="83" type="materials_examined">
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<paragraph pageId="82" pageNumber="83">Material examined.</paragraph>
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<paragraph pageId="82" pageNumber="83">In addition to type material, we have seen a total of 803 specimens (398 males, 405 females). See Appendix for details.</paragraph>
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<caption pageId="82" pageNumber="83">
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<paragraph pageId="82" pageNumber="83">
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Figure 58. Habitus digital image of
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus striatopunctatus" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="striatopunctatus">Selenophorus striatopunctatus</taxonomicName>
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species group, dorsal aspect,
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<taxonomicName lsidName="S. striatopunctatus" pageId="82" pageNumber="83" rank="species" species="striatopunctatus">S. striatopunctatus</taxonomicName>
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Putzeys. Scale bar 5 mm.
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</paragraph>
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</caption>
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<caption pageId="82" pageNumber="83">
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<paragraph pageId="82" pageNumber="83">
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Figure 59. Digital images of male genitalia of
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus striatopunctatus" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="striatopunctatus">Selenophorus striatopunctatus</taxonomicName>
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species group,
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<taxonomicName lsidName="S. striatopunctatus" pageId="82" pageNumber="83" rank="species" species="striatopunctatus">S. striatopunctatus</taxonomicName>
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Putzeys. A right lateral aspect B dorsal aspect C left lateral aspect. Scale bar 1 mm.
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</paragraph>
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</caption>
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<caption pageId="82" pageNumber="83">
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<paragraph pageId="82" pageNumber="83">
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Figure 60. Line drawing of female reproductive tract of
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus striatopunctatus" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="striatopunctatus">Selenophorus striatopunctatus</taxonomicName>
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species group, ventral aspect,
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<taxonomicName lsidName="S. striatopunctatus" pageId="82" pageNumber="83" rank="species" species="striatopunctatus">S. striatopunctatus</taxonomicName>
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Putzeys. Legend: bc bursa copulatrix co common oviduct gc1 gonocoxite 1 gc2 gonocoxite 2 sp spermatheca spg spermathecal gland spgd spermathecal gland duct v valvifer. Scale bars 1 mm.
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</paragraph>
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</caption>
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<caption pageId="82" pageNumber="83">
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<paragraph pageId="82" pageNumber="83">
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Figure 61. Map of West Indies showing known localities for species of
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<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus striatopunctatus" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="striatopunctatus">Selenophorus striatopunctatus</taxonomicName>
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species group.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |