treatments-xml/data/C5/9E/C9/C59EC9CABCE55F8BB6297DB467323F88.xml

675 lines
57 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="86A35C30D8C7A5B34601FDE90E7F19DD" ID-DOI="10.3897/zookeys.1217.131730" ID-ZooBank="921FBCB0-D6B1-4E61-80F2-DE30167984B0" ID-publisher-id="131730" URI-arpha="233F9E1C-B9DF-5FBD-B718-96D1731CD085" URI-zoobank="http://zoobank.org/921FBCB0-D6B1-4E61-80F2-DE30167984B0" article-type="research-article" checkinTime="1730397883241" checkinUser="pensoft" docAuthor="Pešić, Vladimir, Zawal, Andrzej, Ferreira, Sónia, Benitez-Bosco, Laura, Cruz-Oliveira, Ana, Girão, Dinis, Padilha, Adriana, Turaccio, Paolo, Rossini, Samantha, Ballini, Lorenzo, Staffoni, Giorgia, Fratini, Sara, Ciofi, Claudio, Iannucci, Alessio, Ekrem, Torbjørn &amp; Stur, Elisabeth" docDate="2024" docId="C59EC9CABCE55F8BB6297DB467323F88" docLanguage="en" docName="ZooKeys 1217: 119-171" docOrigin="ZooKeys 1217" docSource="https://zookeys.pensoft.net/article/131730/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Torrenticola (Torrenticola) soniae Pešić &amp; Zawal &amp; Ferreira &amp; Benitez-Bosco &amp; Cruz-Oliveira &amp; Girão &amp; Padilha &amp; Turaccio &amp; Rossini &amp; Ballini &amp; Staffoni &amp; Fratini &amp; Ciofi &amp; Iannucci &amp; Ekrem &amp; Stur 2024, sp. nov." docType="treatment" docUuid="E0A41268-B54C-4366-9886-E2954F1BF00B" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="171" masterDocId="233F9E1CB9DF5FBDB71896D1731CD085" masterDocTitle="DNA barcode library of Portuguese water mites, with the descriptions of two new species (Acari, Hydrachnidia)" masterLastPageNumber="171" masterPageNumber="119" pageNumber="119" updateTime="1730398830545" updateUser="ExternalLinkService">
<mods:mods id="18DA4852BFF2ABF82DE814FA6A902C8D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="741DA7F71DD40A0E472B6CCBB187C7EC">
<mods:title id="B4C2EAEFB2B69B221D3AAD59C9D55C36">DNA barcode library of Portuguese water mites, with the descriptions of two new species (Acari, Hydrachnidia)</mods:title>
</mods:titleInfo>
<mods:name id="F14C624FEE4D5614961C0FFD4DEE6AFD" type="personal">
<mods:role id="D70E939454E66B734355B68072DDA477">
<mods:roleTerm id="BC089DF55D024BFDD93EFEF24026FF47">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A91E821CCB2777FA55B3F1A3359BAE06">Pešić, Vladimir</mods:namePart>
<mods:nameIdentifier id="1D78DE4682B70E4E99853958A99C4480" type="ORCID">0000-0002-9724-345X</mods:nameIdentifier>
<mods:affiliation id="DB162F0903F25A4B7B5C38263F8FEACD">Department of Biology, University of Montenegro, Cetinjski put b. b., 81000 Podgorica, Montenegro</mods:affiliation>
</mods:name>
<mods:name id="43D7AB49F86EAF9AC5F1F58E536A50CD" type="personal">
<mods:role id="AD8C601340B8BBC626C5D4BD1CAE309E">
<mods:roleTerm id="9AD6476EEED61549CDCA3C18982B931B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F5E1B3557AE43819619E996BFD5DB576">Zawal, Andrzej</mods:namePart>
<mods:nameIdentifier id="41378C6B4812501DF749786BA159637F" type="ORCID">0000-0002-5838-6060</mods:nameIdentifier>
<mods:affiliation id="3252921E41A72A08EF9EB362AE78D637">Institute of Marine and Environmental Sciences, Center of Molecular Biology and Biotechnology, University of Szczecin, Wąska 13, 71 415 Szczecin, Poland</mods:affiliation>
</mods:name>
<mods:name id="50920042F0D9637BF7B82026054AAA9C" type="personal">
<mods:role id="02BFBC67EDA17B79ED337DCCF07D50F6">
<mods:roleTerm id="EC547367F5DDB47D1205D1A612BB6904">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9B1DDDEF4758D1C9CF36D4771ABADBBF">Ferreira, Sónia</mods:namePart>
<mods:nameIdentifier id="4CF5B90E129599C140C98B0756ED27AB" type="ORCID">0000-0002-6884-3966</mods:nameIdentifier>
<mods:affiliation id="04ABDDF8FD82AF5BE88DA763379C40E4">CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Vila do Conde, Portugal &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661 Vairão, Vila do Conde, Portugal</mods:affiliation>
</mods:name>
<mods:name id="3C5B4A9BB64FC50130C3DA862B599A53" type="personal">
<mods:role id="42C9DAA8882B6373DAB9D034070969CE">
<mods:roleTerm id="B875E734BC9C7F7010C9294157FDFD00">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B093011055A7E15396686A8B93D70156">Benitez-Bosco, Laura</mods:namePart>
<mods:nameIdentifier id="B10B807C5611F1D47C2A81641731F56E" type="ORCID">0000-0002-8863-8353</mods:nameIdentifier>
<mods:affiliation id="2C29B2D75E5CD79CD0ED303E494A969A">CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Vila do Conde, Portugal &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661 Vairão, Vila do Conde, Portugal &amp; Departamento de Biologia, Faculdade de Ciências, Universidade do Porto, 4099 - 002 Porto, Portugal</mods:affiliation>
</mods:name>
<mods:name id="15BC056CDE56F16C048670B8081C018F" type="personal">
<mods:role id="E4A6AA857C5BA0F013B29A7DB5B5695A">
<mods:roleTerm id="A939DC569C910A942017E88DDCB3BC0C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DFD70F7A23D6E6DDCA9FC8CE672648E1">Cruz-Oliveira, Ana</mods:namePart>
<mods:affiliation id="4EAF1D07AB1DC23281753C15D0567FC3">CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Vila do Conde, Portugal &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661 Vairão, Vila do Conde, Portugal</mods:affiliation>
</mods:name>
<mods:name id="8C9AAA7F59631009F6736EF98F242E62" type="personal">
<mods:role id="C661DC11839B46E41DC59E1E33F93352">
<mods:roleTerm id="F6F9AC030314D2A2680F4AD25A8B7BE5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0E7E1832EC540AF26661BF279CE345A5">Girão, Dinis</mods:namePart>
<mods:nameIdentifier id="72F6B4F8F1638412A9BAE2E42A6F08B7" type="ORCID">https://orcid.org/0009-0008-4842-0721</mods:nameIdentifier>
<mods:affiliation id="DD271AEE43F200047AF56CEC83052481">CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Vila do Conde, Portugal &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661 Vairão, Vila do Conde, Portugal &amp; EBM, Estação Biológica de Mértola, Praça Luís de Camões, Mértola, Portugal</mods:affiliation>
</mods:name>
<mods:name id="D30B13573CF5FEF9FB558F0D1B226457" type="personal">
<mods:role id="E8A95D3668866D9D590E54435BDCB618">
<mods:roleTerm id="7F5F6343D62E386B06850BEA863CF8EF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F2EA414DB6EC9B5A4E1D46677EF59EA6">Padilha, Adriana</mods:namePart>
<mods:affiliation id="02AD05D36399DD6C2FD01A3C6CC9F64C">CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Vila do Conde, Portugal &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661 Vairão, Vila do Conde, Portugal &amp; EBM, Estação Biológica de Mértola, Praça Luís de Camões, Mértola, Portugal</mods:affiliation>
</mods:name>
<mods:name id="CD0D32BBDF7BC7F630ECA2A660F65B06" type="personal">
<mods:role id="E1E11E86CC2791B7A9ADEB20F6802E5B">
<mods:roleTerm id="C848FD708D0C260B278DC574F2447953">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D369084B0A3DA8C9AE8C5A37730F4C2D">Turaccio, Paolo</mods:namePart>
<mods:nameIdentifier id="48ED6D091B55D7A6FEB99DDA2E3B7890" type="ORCID">https://orcid.org/0009-0000-8627-6718</mods:nameIdentifier>
<mods:affiliation id="AB3773432499722A86CD36A98A800E7A">CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Vila do Conde, Portugal &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661 Vairão, Vila do Conde, Portugal &amp; Department of Biology, University of Florence, Sesto Fiorentino, Italy</mods:affiliation>
</mods:name>
<mods:name id="70AA8538D99A218B579ECA5727FC184B" type="personal">
<mods:role id="907E9C0F6A0E71FD3FAE1C1289AC6EAC">
<mods:roleTerm id="14FBE44D96AA4F4F312EDE1621C92147">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="52194DD74D0F1EDA382E9C52F7C8DD6D">Rossini, Samantha</mods:namePart>
<mods:nameIdentifier id="76D020EB733D012B91D6AF947BF9F033" type="ORCID">https://orcid.org/0009-0009-6881-2527</mods:nameIdentifier>
<mods:affiliation id="EEAC9FE87C0DFDE0E8DFBB3FE1FDFDB4">Department of Biology, University of Florence, Sesto Fiorentino, Italy</mods:affiliation>
</mods:name>
<mods:name id="706533D7379ADC9A56ACD1A088854B57" type="personal">
<mods:role id="A57F9FFC13B036168BB2FD3C8A228764">
<mods:roleTerm id="0E553EF4A998D8B36D1797CA901C4C4C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="24FC5A2107DEF40988B389B2657B3A5A">Ballini, Lorenzo</mods:namePart>
<mods:affiliation id="C8DA425E9FEF9F4FFBE1472D95D966BB">Department of Biology, University of Florence, Sesto Fiorentino, Italy</mods:affiliation>
</mods:name>
<mods:name id="ABF4EF4C209936B3CCBC227598970B16" type="personal">
<mods:role id="D24190A9CFE1BBED40C6549FA686D51C">
<mods:roleTerm id="3289AB8427ADDEA16F16F1015D39F1D2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2F991344923391602774D149C06FF474">Staffoni, Giorgia</mods:namePart>
<mods:nameIdentifier id="EFD84CC1243718614B196AA4FF60052F" type="ORCID">https://orcid.org/0009-0003-5916-1757</mods:nameIdentifier>
<mods:affiliation id="908CB0051C7D4F3ADEF9669BA0D55824">Department of Biology, University of Florence, Sesto Fiorentino, Italy</mods:affiliation>
</mods:name>
<mods:name id="D14DEB5EC0FF7B3372B4CC875E59C95C" type="personal">
<mods:role id="1B7860D3EBCC6000F2B082B103E89EB1">
<mods:roleTerm id="B8B1353B3AA074FB9EEFF0A01EE8DCC4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A2AECE735D980923AB9A960D245D5093">Fratini, Sara</mods:namePart>
<mods:affiliation id="95E9EC7E7290DC7F03647C84FB8B6B3F">Department of Biology, University of Florence, Sesto Fiorentino, Italy</mods:affiliation>
</mods:name>
<mods:name id="F7F07F83B41235B4BF2E83BD8796520B" type="personal">
<mods:role id="B16DDAFC03EDC2DC29A2AB1BDFC410C5">
<mods:roleTerm id="5322EC8B02C1A4EC9099A4EA635FB0F7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BC6FAB3D772DDD4DB9903242BB8514AF">Ciofi, Claudio</mods:namePart>
<mods:affiliation id="01260A3CC071F564897C00BC2A3BD7B2">Department of Biology, University of Florence, Sesto Fiorentino, Italy</mods:affiliation>
</mods:name>
<mods:name id="7BB55830F195202C53BE05413BA7266C" type="personal">
<mods:role id="CFF71AD3FB8146264F1199B30B486DEE">
<mods:roleTerm id="222D247AFD8F643DE2A5F537D47B04C9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ECF3D7966D7A666ED8CCF15872BB709E">Iannucci, Alessio</mods:namePart>
<mods:affiliation id="B0304A6C48C82F3807E67058CB66B8B2">Department of Biology, University of Florence, Sesto Fiorentino, Italy</mods:affiliation>
</mods:name>
<mods:name id="2A564384EAFBCD3FAD712356D7E749D9" type="personal">
<mods:role id="EC77D59F7531F89B8A74E1222C9D4C80">
<mods:roleTerm id="E0F2B81CED02D1B5CB9938F965DEF1C6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CBCF572F59C583C0EAB86E65D9EA28C3">Ekrem, Torbjørn</mods:namePart>
<mods:nameIdentifier id="E30EFD4D307FDD1902AC960D4BA8618D" type="ORCID">0000-0003-3469-9211</mods:nameIdentifier>
<mods:affiliation id="5C7065A1502E283081A31D351F8D7F92">Department of Natural History, NTNU University Museum, Norwegian University of Science and Technology, Trondheim, Norway</mods:affiliation>
</mods:name>
<mods:name id="35877D8DA93D8FBAC153A025C105E14A" type="personal">
<mods:role id="8A1819676C382148CFBB95CA7C2C6F68">
<mods:roleTerm id="841AE70F452711939E5B01430CCEFA06">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4FD2F707BA1B3E13E1150C3088DCC676">Stur, Elisabeth</mods:namePart>
<mods:affiliation id="85E9CB203CD407A6210B26E519E314AF">Department of Natural History, NTNU University Museum, Norwegian University of Science and Technology, Trondheim, Norway</mods:affiliation>
</mods:name>
<mods:typeOfResource id="2C71F7F0E0948A6FEB480DEE2E348F84">text</mods:typeOfResource>
<mods:relatedItem id="C7310CA6EE73E7268DF0DE7667AF0F1A" type="host">
<mods:titleInfo id="EEC0A2E5C3DCCA7B8A5235859382D151">
<mods:title id="8D312EC480019F61BD1B3024633E5008">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="2E6665190668E13D417204D9750A297B">
<mods:date id="C94CD4A6C5DFD36E3ED7D91775EB2329">2024</mods:date>
<mods:detail id="06849CE384847B9624F5677CCE5A2BE8" type="pubDate">
<mods:number id="505A03652AAB1CA153CF061156A0662D">2024-10-31</mods:number>
</mods:detail>
<mods:detail id="E6F8072327C09876D1F59DDB05B7AFAE" type="volume">
<mods:number id="CB30F72095810EC07DA72F6D6AA7CFCF">1217</mods:number>
</mods:detail>
<mods:extent id="9165803DAF2DC753191192540599728F" unit="page">
<mods:start id="15569C298B7DF6FFAD0F3785431759EC">119</mods:start>
<mods:end id="96E249C4497935EA0118D92AF13DBBE6">171</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="FEFAFD63099F8EBE52472C7983CA9798">journal article</mods:classification>
<mods:identifier id="EB6745C41DD1E63B06A1DC64614CE822" type="DOI">10.3897/zookeys.1217.131730</mods:identifier>
<mods:identifier id="A71038B7D3FF1B09D50F4E5FC0EF003C" type="ZooBank">921FBCB0-D6B1-4E61-80F2-DE30167984B0</mods:identifier>
</mods:mods>
<treatment id="C59EC9CABCE55F8BB6297DB467323F88" ID-DOI="http://doi.org/10.5281/zenodo.14019467" ID-Zenodo-Dep="14019467" ID-arpha="C59EC9CA-BCE5-5F8B-B629-7DB467323F88" ID-zoobank="https://zoobank.org/E0A41268-B54C-4366-9886-E2954F1BF00B" LSID="urn:lsid:zoobank.org:act:E0A41268-B54C-4366-9886-E2954F1BF00B" LSID-ZBK="urn:lsid:zoobank.org:act:E0A41268-B54C-4366-9886-E2954F1BF00B" httpUri="http://treatment.plazi.org/id/C59EC9CABCE55F8BB6297DB467323F88">
<subSubSection id="AEE8E555941FB940889F4046468E0F14" type="nomenclature">
<paragraph id="E324AFAC0A8FC83E8980113BDE8E64B0">
<taxonomicName id="C51249D709CF6B1FD3F2C604D7135402" authority="Pesic" authorityName="Pešić &amp; Zawal &amp; Ferreira &amp; Benitez-Bosco &amp; Cruz-Oliveira &amp; Girão &amp; Padilha &amp; Turaccio &amp; Rossini &amp; Ballini &amp; Staffoni &amp; Fratini &amp; Ciofi &amp; Iannucci &amp; Ekrem &amp; Stur" authorityYear="2024" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" infraOrder="Hydrachnidia" kingdom="Animalia" order="Trombidiformes" phylum="Arthropoda" rank="species" species="soniae" status="sp. nov." subGenus="Torrenticola">
<emphasis id="3DD07FE889E526AB79ACAD8A90B60151" italics="true">Torrenticola</emphasis>
(
<emphasis id="1F60C2B6A3B43715D9E125E63BEDB626" italics="true">Torrenticola</emphasis>
)
<emphasis id="72753470914F6AB91C7421A76B81EC37" italics="true">soniae</emphasis>
Pešić
</taxonomicName>
<taxonomicNameLabel id="A6F03B68E58E03BD746D09446DE043ED" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="550F04580BCBAC3CC5EF2E4623045D93" type="description">
<paragraph id="4F6B04BA1728DB5D31613EB8E843D05C">
<figureCitation id="FD7071CE33F9E917FA2FC090079B66D9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Torrenticola soniae sp. nov., ♂ holotype A dorsal shield B ventral shield C palp, medial view (inset: ventrodistal projections of P- 2 and P- 3, enlarged 2 ×) D palp, lateral view E gnathosoma and chelicera F ejaculatory complex. Scale bars: 100 μm." figureDoi="10.3897/zookeys.1217.131730.figure1" httpUri="https://binary.pensoft.net/fig/1164865">Figs 1</figureCitation>
,
<figureCitation id="61DDA76CB6DF0512398C096E03D2387D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Torrenticola soniae sp. nov., ♀ paratype A dorsal shield B photograph of dorsal shield C ventral shield D palp, medial view (P- 1 lacking). Scale bars: 100 μm." figureDoi="10.3897/zookeys.1217.131730.figure2" httpUri="https://binary.pensoft.net/fig/1164866">2</figureCitation>
,
<figureCitation id="8F6938D73857AB53E8FF9611B26B037B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A C Photographs of dorsal shield A Torrenticola soniae sp. nov., ♂ holotype B T. elisabethae sp. nov., ♂ holotype C T. tenuipalpis, ♀ (BGE _ 00227 _ F 03) D E Photographs of selected sampling sites D Praia Fluvial de Sabugueiro, locus typicus of Torrenticola soniae sp. nov. E Poço do Inferno, type locality of T. elisabethae sp. nov. Photographs by JC (5 D) and SF (5 E)." figureDoi="10.3897/zookeys.1217.131730.figure5" httpUri="https://binary.pensoft.net/fig/1164869">5 A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5HGK" type="type material">
<paragraph id="8989112C2DEBFD3D69A4A731FBABDFC6">
<heading id="7BB571125ABC440913D90198E005129D" reason="title">Type material examined.</heading>
</paragraph>
<paragraph id="024A334FAFBA44F1B0DDD4B2552FD6CD">
<materialsCitation id="766E8B00FFE0E3F600F4162E51A40992" collectingDate="2023-08-24" collectionCode="RMNH, BOLD" collectorName="Ferreira &amp; Benitez-Bosco &amp; Padilha" country="Portugal" county="Seia" elevation="1021" latitude="40.401" location="Praia Fluvial de Sabugueiro" longLatPrecision="427" longitude="-7.64" municipality="Rio Alva" specimenCode="BSNTN 961-23" specimenCount="1" specimenCount-male="1" stateProvince="Guarda" typeStatus="holotype">
<emphasis id="B608360E09DB28D1638389149C01FA21" bold="true" italics="true">
<typeStatus id="2EA9F08A90C3E5616FFC65186E5C14A3" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="294BA5EF30B329A1BD2C6F4DB5CC3F6C" count="1" type="male"></specimenCount>
, dissected and slide mounted (
<named-content id="E02A04D6E8F42F001522A7EC0EC3ED87" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturalis-biodiversity-center" xlink_title="Naturalis Biodiversity Center in Leiden">
<collectionCode id="76C0B2E50B6EAC76628172AC173874C2" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
</named-content>
),
<collectingCountry id="B5409400436E63174E26B35AE57FF7D3" name="Portugal">Portugal</collectingCountry>
,
<emphasis id="7CAC6EA80233A4A4652E894DD112DA6D" bold="true">
<collectingRegion id="3F224B5316B6717EAE104D6E1F925F89" country="Portugal" name="Guarda">Guarda</collectingRegion>
</emphasis>
,
<collectingCounty id="B19F03DE1D9D8E47F09FE05C8543C779">Seia</collectingCounty>
,
<collectingMunicipality id="F0B8DF23B6FB590E020897218041FDDE">Rio Alva</collectingMunicipality>
,
<location id="C5ED884EF761492562C9F7870D8BAF07" LSID="urn:lsid:plazi:treatment:C59EC9CABCE55F8BB6297DB467323F88:C5ED884EF761492562C9F7870D8BAF07" country="Portugal" county="Seia" latitude="40.401" longLatPrecision="427" longitude="-7.64" municipality="Rio Alva" name="Praia Fluvial de Sabugueiro" stateProvince="Guarda">Praia Fluvial de Sabugueiro</location>
(Fig.
<figureCitation id="4E99231B85931BDA8D25DB327A034638" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A C Photographs of dorsal shield A Torrenticola soniae sp. nov., ♂ holotype B T. elisabethae sp. nov., ♂ holotype C T. tenuipalpis, ♀ (BGE _ 00227 _ F 03) D E Photographs of selected sampling sites D Praia Fluvial de Sabugueiro, locus typicus of Torrenticola soniae sp. nov. E Poço do Inferno, type locality of T. elisabethae sp. nov. Photographs by JC (5 D) and SF (5 E)." figureDoi="10.3897/zookeys.1217.131730.figure5" httpUri="https://binary.pensoft.net/fig/1164869">5 D</figureCitation>
), river,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWIGK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-7.640000,40.401000]}">
<geoCoordinate id="F697CAA8AA0D4D721FCF116CAE4F5150" degrees="40.401" direction="north" orientation="latitude" precision="55" value="40.401">40.401 ° N</geoCoordinate>
,
<geoCoordinate id="823DC853C87394E63AFFCF2E507CB4BB" degrees="7.64" direction="west" orientation="longitude" precision="555" value="-7.64">7.64 ° W</geoCoordinate>
</named-content>
,
<elevation id="2BDA11CE354241DB07F064387377E0F3" metricMagnitude="3" metricUnit="m" metricValue="1.021" unit="m" value="1021.0">
<quantity id="9C94EE72E56869EC21B83E9AF9D974B3" metricMagnitude="3" metricUnit="m" metricValue="1.021" unit="m" value="1021.0">1021 m</quantity>
a. s. l.
</elevation>
,
<date id="E9805C43F39F6D6D4ED2AECBC38BC13A" value="2023-08-24">
<collectingDate id="DB4121011D112D7FD80E9F777581304A" value="2023-08-24">24 Aug. 2023</collectingDate>
</date>
, leg.
<collectorName id="B613C2BCB838FE6AAB15BA8019051601">Ferreira</collectorName>
,
<collectorName id="8ECB685FB31B06A3C131DAF7B63BDCA0">Benitez-Bosco</collectorName>
&amp;
<collectorName id="274A41F4C909A38D63B2A8223C4F83C6">Padilha</collectorName>
, sequenced (
<abbrev id="ABBRID0E2IGK" xlink_title="Barcode of Life Data Systems">
<collectionCode id="0941AF3A867D36C01947536927007673">BOLD</collectionCode>
</abbrev>
ID:
<ext-link id="0408039A26DB1416821D7AB76CC78C5A" ext-link-type="uri" xlink_href="http://www.boldsystems.org/index.php/Public_RecordView?processid=BSNTN961-23" xlink_type="simple">
<specimenCode id="DBD2B561C48A813966FB4CCC6CBAB3C2" collectionCode="BSNTN">BSNTN 961-23</specimenCode>
</ext-link>
)
</materialsCitation>
.
<materialsCitation id="CFC617FBCEA62AA2FFF3A50A61C58EAF" collectingDate="2023-08-23" collectorName="Ferreira &amp; Benitez-Bosco &amp; Padilha" country="Portugal" county="Seia" elevation="312" latitude="40.379" location="Praia" longLatPrecision="69" longitude="-7.736" municipality="Rio Alva" specimenCount="2" specimenCount-male="2" stateProvince="Guarda" typeStatus="paratype">
<emphasis id="A4FC72A36B8CC1308FDC43A2E357FE5E" bold="true" italics="true">
<typeStatus id="50EE7C6B14A75B6CA1DC910986F3F97F" type="paratype">Paratypes</typeStatus>
</emphasis>
: •
<specimenCount id="F37A572F7F7D080C826C2639585357C5" count="2" type="male">2 ♂</specimenCount>
(sequenced),
<collectingCountry id="BC423D180C89A22E0303E5F0CA1D0B4D" name="Portugal">Portugal</collectingCountry>
,
<emphasis id="E70D0E8215B96258FFD6CEFD22787A31" bold="true">
<collectingRegion id="E5DCD9277B0FF23792D54858B9326A1C" country="Portugal" name="Guarda">Guarda</collectingRegion>
</emphasis>
,
<collectingCounty id="7D15C14A5849EF9185BA09C429685574">Seia</collectingCounty>
,
<collectingMunicipality id="B3ACAD726B8C77C733CDC09C026CE8F2">Rio Alva</collectingMunicipality>
,
<location id="EE2FC0813F6FFD59B0CD93F6C725FD8E" LSID="urn:lsid:plazi:treatment:C59EC9CABCE55F8BB6297DB467323F88:EE2FC0813F6FFD59B0CD93F6C725FD8E" country="Portugal" county="Seia" latitude="40.379" longLatPrecision="69" longitude="-7.736" municipality="Rio Alva" name="Praia" stateProvince="Guarda">Praia</location>
fluvial de
<location id="57AA3D930D96DAFC73033E5CCC2F4361" LSID="urn:lsid:plazi:treatment:C59EC9CABCE55F8BB6297DB467323F88:57AA3D930D96DAFC73033E5CCC2F4361" country="Portugal" county="Seia" latitude="40.379" longLatPrecision="69" longitude="-7.736" municipality="Rio Alva" name="Vila Cova" stateProvince="Guarda">Vila Cova</location>
a
<location id="7FBA77629835A3613D0279AF65F641A9" LSID="urn:lsid:plazi:treatment:C59EC9CABCE55F8BB6297DB467323F88:7FBA77629835A3613D0279AF65F641A9" country="Portugal" county="Seia" latitude="40.379" longLatPrecision="69" longitude="-7.736" municipality="Rio Alva" name="Coelheira" stateProvince="Guarda">Coelheira</location>
, river,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMJGK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-7.736000,40.379000]}">
<geoCoordinate id="9B0EBD1CF738D2A58DC2CCA7BB959C13" degrees="40.379" direction="north" orientation="latitude" precision="55" value="40.379">40.379 ° N</geoCoordinate>
,
<geoCoordinate id="5AEC90205CF3DC6842C661304ACABB5D" degrees="7.736" direction="west" orientation="longitude" precision="55" value="-7.736">7.736 ° W</geoCoordinate>
</named-content>
,
<elevation id="E1EAA4B06AC92868C1657FA4941D7F0F" metricMagnitude="2" metricUnit="m" metricValue="3.12" unit="m" value="312.0">
<quantity id="CA312E709D8A0C339F05A1A1FFE8C1C0" metricMagnitude="2" metricUnit="m" metricValue="3.12" unit="m" value="312.0">312 m</quantity>
a. s. l.
</elevation>
,
<date id="BF1AA41EFE31265089B653DB830539A6" value="2023-08-23">
<collectingDate id="47A5E7424B5F64F2317AB1A073996B0A" value="2023-08-23">23 Aug. 2023</collectingDate>
</date>
, leg.
<collectorName id="F0413A4410F3AD6805197DEEF8F24AFD">Ferreira</collectorName>
,
<collectorName id="F01CB37D1A0C20B12A94AB76273F01ED">Benitez-Bosco</collectorName>
&amp;
<collectorName id="C2B4B7E4FE8DC4DE3BCD4BFCA54352EE">Padilha</collectorName>
</materialsCitation>
; •
<materialsCitation id="CDD0DBF9DEFA4A34B32748B3F5068951" collectingDate="2023-08-22" collectorName="Ferreira &amp; Benitez-Bosco &amp; Padilha &amp; Andrade &amp; Stur" country="Portugal" county="Manteigas" elevation="672" latitude="40.403" location="Ponte" longLatPrecision="69" longitude="-7.526" municipality="Zezere" specimenCount="1" specimenCount-male="1" stateProvince="Guarda" typeStatus="paratype">
<specimenCount id="D98C27E9E4639C85514A18BDDD433A94" count="1" type="male">1 ♂</specimenCount>
(sequenced);
<collectingCounty id="FAEC74622857DE41B65F4B4A5ED60180">Manteigas</collectingCounty>
,
<collectingMunicipality id="FCBE60C629498DCB0518B09EC7F44549">Zêzere</collectingMunicipality>
,
<location id="DBE9D97E97440BA5577DD37DC2BBD622" LSID="urn:lsid:plazi:treatment:C59EC9CABCE55F8BB6297DB467323F88:DBE9D97E97440BA5577DD37DC2BBD622" country="Portugal" county="Manteigas" latitude="40.403" longLatPrecision="69" longitude="-7.526" municipality="Zezere" name="Ponte" stateProvince="Guarda">Ponte</location>
dos
<location id="1EB1B4403CF685CC099518ADD828909F" LSID="urn:lsid:plazi:treatment:C59EC9CABCE55F8BB6297DB467323F88:1EB1B4403CF685CC099518ADD828909F" country="Portugal" county="Manteigas" latitude="40.403" longLatPrecision="69" longitude="-7.526" municipality="Zezere" name="Frades" stateProvince="Guarda">Frades</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EUJGK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-7.526000,40.403000]}">
<geoCoordinate id="5B8935454C5D77C348D039620087EB8D" degrees="40.403" direction="north" orientation="latitude" precision="55" value="40.403">40.403 ° N</geoCoordinate>
,
<geoCoordinate id="6763E1CEDB21740DE49138FCA9D302C4" degrees="7.526" direction="west" orientation="longitude" precision="55" value="-7.526">7.526 ° W</geoCoordinate>
</named-content>
,
<elevation id="F6E9C88DC24CA4C5D2DA645F2B6A0F5F" metricMagnitude="2" metricUnit="m" metricValue="6.72" unit="m" value="672.0">
<quantity id="016F89CD4DBAA4A8F19FA646CBCC0C54" metricMagnitude="2" metricUnit="m" metricValue="6.72" unit="m" value="672.0">672 m</quantity>
a. s. l.
</elevation>
,
<date id="2C00EB86F0221C59592B7E70D76433A6" value="2023-08-22">
<collectingDate id="E07A52A2E6DFC03B4D28DD964B43377A" value="2023-08-22">22 Aug. 2023</collectingDate>
</date>
, leg.
<collectorName id="8607FA80DF73EAEB3D1A0B6CDFC9DA49">Ferreira</collectorName>
,
<collectorName id="B8B83648A131C578793CAD2880DEC142">Benitez-Bosco</collectorName>
,
<collectorName id="F98F1F8BDDED7F95308120897BE74B42">Padilha</collectorName>
,
<collectorName id="199AF2F7F8B015C6024290D7A99ECA82">Andrade</collectorName>
&amp;
<collectorName id="E7AD390992930E1F2834E0C7EC9C8C67">Stur</collectorName>
</materialsCitation>
; •
<materialsCitation id="2973E6B36D1F6638C64101025548DA5B" collectingDate="2023-08-24" collectionCode="RMNH" collectorName="Ferreira &amp; Benitez-Bosco &amp; Padilha" country="Portugal" county="Gouveia" elevation="976" latitude="40.454" location="Casais de Folgosinho" longLatPrecision="69" longitude="-7.493" municipality="Rio Mondego" specimenCount="1" specimenCount-female="1" stateProvince="Guarda" typeStatus="paratype">
<specimenCount id="EC011C5327280E50EB0889A207644656" count="1" type="female">1 ♀</specimenCount>
(sequenced), dissected and slide mounted (
<named-content id="51B28EF1CB8B16CC4D4EAE3EF8363F88" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/naturalis-biodiversity-center" xlink_title="Naturalis Biodiversity Center in Leiden">
<collectionCode id="6FDEFE9B5295F4A7B955DAD32603DCF4" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
</named-content>
),
<collectingRegion id="712FAF8BBE43CBA9FAD83818550B2973" country="Portugal" name="Guarda">Guarda</collectingRegion>
,
<collectingCounty id="3B3B3571BFEA64FCD8FA49651236BBB5">Gouveia</collectingCounty>
,
<collectingMunicipality id="FB1A39E287283032033324C2570DBA46">Rio Mondego</collectingMunicipality>
,
<location id="85E958232FBF99B5FC37A895D8EF4B7D" LSID="urn:lsid:plazi:treatment:C59EC9CABCE55F8BB6297DB467323F88:85E958232FBF99B5FC37A895D8EF4B7D" country="Portugal" county="Gouveia" latitude="40.454" longLatPrecision="69" longitude="-7.493" municipality="Rio Mondego" name="Casais de Folgosinho" stateProvince="Guarda">Casais de Folgosinho</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EAKGK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-7.493000,40.454000]}">
<geoCoordinate id="27D4D5DA8ED1BD4DCD407BD98DD6EED9" degrees="40.454" direction="north" orientation="latitude" precision="55" value="40.454">40.454 ° N</geoCoordinate>
,
<geoCoordinate id="3F236CDA0DC8B648357D99597081F2B1" degrees="7.493" direction="west" orientation="longitude" precision="55" value="-7.493">7.493 ° W</geoCoordinate>
</named-content>
,
<elevation id="D91AD5E2A8134731F5B5C9E687CA4346" metricMagnitude="2" metricUnit="m" metricValue="9.76" unit="m" value="976.0">
<quantity id="8FC95B0E44C15DEF921A41AE66ACBD43" metricMagnitude="2" metricUnit="m" metricValue="9.76" unit="m" value="976.0">976 m</quantity>
a. s. l.
</elevation>
,
<date id="F1C4FF714810A1AC0DC8FD5529B7BAB6" value="2023-08-24">
<collectingDate id="0C22ECE6DFF0A5F191E7EA3D6394CCF0" value="2023-08-24">24 Aug. 2023</collectingDate>
</date>
, leg.
<collectorName id="FBEF558C93393F6AC4B26CC5CD77AFDC">Ferreira</collectorName>
,
<collectorName id="BD4CA58270628700E505D53BC2683498">Benitez-Bosco</collectorName>
&amp;
<collectorName id="9AD0E5F989AFB36760916027DB752BD7">Padilha</collectorName>
</materialsCitation>
; •
<materialsCitation id="4C95E37EE36E4A76E45F83F70128A114" collectingDate="2023-07-13" collectorName="Ferreira &amp; Padilha." country="Portugal" elevation="256" latitude="41.665" location="Torre de Dona Chama" longLatPrecision="69" longitude="-7.146" municipality="Mirandela" specimenCount="1" specimenCount-female="1" stateProvince="Braganca" typeStatus="paratype">
<specimenCount id="92A600DD55E94627B844CC887E295C24" count="1" type="female">1 ♀</specimenCount>
(juv.; sequenced),
<emphasis id="DF917A72F8C8F9EB869014F81049D219" bold="true">
<collectingRegion id="574D8599983550280E90FF7290D5EC4D" country="Portugal" name="Braganca">Bragança</collectingRegion>
</emphasis>
,
<collectingMunicipality id="B8978D51367C8137A33F7FD216EE655A">Mirandela</collectingMunicipality>
,
<location id="35B132160D8E6E4259FBCE19F3EC67D1" LSID="urn:lsid:plazi:treatment:C59EC9CABCE55F8BB6297DB467323F88:35B132160D8E6E4259FBCE19F3EC67D1" country="Portugal" latitude="41.665" longLatPrecision="69" longitude="-7.146" municipality="Mirandela" name="Torre de Dona Chama" stateProvince="Braganca">Torre de Dona Chama</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKKGK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-7.146000,41.665000]}">
<geoCoordinate id="B1FEB8B93054D6A2A86C32A20FB0E2C6" degrees="41.665" direction="north" orientation="latitude" precision="55" value="41.665">41.665 ° N</geoCoordinate>
,
<geoCoordinate id="B3FBD062DB11CD501067674629E647BC" degrees="7.146" direction="west" orientation="longitude" precision="55" value="-7.146">7.146 ° W</geoCoordinate>
</named-content>
,
<elevation id="B3D6C0A627BE551A93025DC4D7C99D73" metricMagnitude="2" metricUnit="m" metricValue="2.56" unit="m" value="256.0">
<quantity id="5FFD0218ECB76E3F42DDAA8CCAB91310" metricMagnitude="2" metricUnit="m" metricValue="2.56" unit="m" value="256.0">256 m</quantity>
a. s. l.
</elevation>
,
<date id="A6C8F858F1375B69388FF4A51B4F5972" value="2023-07-13">
<collectingDate id="C6BF0890F8FBCA582505633E085CE4B3" value="2023-07-13">13 Jul. 2023</collectingDate>
</date>
, leg.
<collectorName id="3F0A12FB6210D43673521302932CA714">Ferreira</collectorName>
&amp;
<collectorName id="917895088F9D9A0D27468105739F1D96">Padilha.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPKGK" type="diagnosis">
<paragraph id="10B29F910F6226561E1C52A9799AFFBD">
<heading id="7B461E52B50B28EA11E5130A9E801B55" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="1375F506C88333733F6FB7CD9932C820">
<emphasis id="B91EABD94B2A9F722A831AA7E3FF0CC2" bold="true" italics="true">Morphological</emphasis>
:
<abbrev id="ABBRID0E1KGK" xlink_title="first coxae">Cx-I</abbrev>
relatively short, anteriorly broad; suture lines of Cx-IV prominent, starting at right angle from genital field; ejaculatory complex with well-developed anterior keel and proximal arms; gnathosomal rostrum short, less than width of gnathosoma; P- 3 with a subrectangular, apically serrated ventrodistal projection.
<emphasis id="C9764D5C52164AD8A51581916F590EBD" bold="true" italics="true">Molecular</emphasis>
: this lineage represented by a unique
<abbrev id="ABBRID0EBLGK" xlink_title="Barcode Index Numbers">
<collectionCode id="960B3B0E11D6479B1CFD255713DF1DD5">BIN</collectionCode>
</abbrev>
(
<ext-link id="D223411AD01F0EC51857CEA7C44C8BAC" ext-link-type="uri" xlink_href="http://boldsystems.org/index.php/Public_BarcodeCluster?clusteruri=BOLD:AFW5337" xlink_type="simple">
<collectionCode id="74F7B578810F09ABE54E10080521468D">BOLD</collectionCode>
: AFW 5337
</ext-link>
) differs from
<taxonomicName id="BB06932E7D50269F398FB3F7F9C3993C" baseAuthorityName="Halbert" baseAuthorityYear="1911" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="487FD3D5999A25E861EE21A1A9EF72F8" italics="true">T. brevirostris</emphasis>
</taxonomicName>
clade by 12.27 %
<abbrev id="ABBRID0EVLGK" xlink_title="Kimura 2-parameter model">K 2 P</abbrev>
for
<abbrev id="ABBRID0EZLGK" xlink_title="mitochondrial cytochrome c oxidase subunit I">COI</abbrev>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4LGK" type="description">
<paragraph id="A44C767D854C46BDA7B36EFCB07692BD">
<heading id="F7D96C512F1EBC34A5EF744A8BA31A86" reason="title">Description.</heading>
</paragraph>
<paragraph id="BA02A3E730E0EC6473FA2043C93757DD">
<emphasis id="488C88206772E033AC1B979D2F3C973E" bold="true" italics="true">General features.</emphasis>
Idiosoma roundish; dorsal shield without a color pattern as photographed in Figs
<figureCitation id="F6946E8DB9D0A1FAD51F71F302406B03" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Torrenticola soniae sp. nov., ♀ paratype A dorsal shield B photograph of dorsal shield C ventral shield D palp, medial view (P- 1 lacking). Scale bars: 100 μm." figureDoi="10.3897/zookeys.1217.131730.figure2" httpUri="https://binary.pensoft.net/fig/1164866">2 B</figureCitation>
,
<figureCitation id="AD46460BB1F8A490B1DD5CBC650D0B37" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A C Photographs of dorsal shield A Torrenticola soniae sp. nov., ♂ holotype B T. elisabethae sp. nov., ♂ holotype C T. tenuipalpis, ♀ (BGE _ 00227 _ F 03) D E Photographs of selected sampling sites D Praia Fluvial de Sabugueiro, locus typicus of Torrenticola soniae sp. nov. E Poço do Inferno, type locality of T. elisabethae sp. nov. Photographs by JC (5 D) and SF (5 E)." figureDoi="10.3897/zookeys.1217.131730.figure5" httpUri="https://binary.pensoft.net/fig/1164869">5 A</figureCitation>
; area of primary sclerotization of the dorsal plate with two dorsoglandularia (Fig.
<figureCitation id="3D545225555062EFB6540922AA9EC226" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Torrenticola soniae sp. nov., ♂ holotype A dorsal shield B ventral shield C palp, medial view (inset: ventrodistal projections of P- 2 and P- 3, enlarged 2 ×) D palp, lateral view E gnathosoma and chelicera F ejaculatory complex. Scale bars: 100 μm." figureDoi="10.3897/zookeys.1217.131730.figure1" httpUri="https://binary.pensoft.net/fig/1164865">1 A</figureCitation>
); frontal platelets broad, relatively short;
<abbrev id="ABBRID0EUMGK" xlink_title="first coxae">Cx-I</abbrev>
relatively short, anteriorly broad; gnathosomal bay U-shaped, proximally rounded;
<abbrev id="ABBRID0EYMGK" xlink_title="coxoglandularia 4">Cxgl- 4</abbrev>
subapical; medial suture line of Cx-II + III relatively short; postgenital area extended; excretory pore and Vgl- 2 away from the line of primary sclerotization, excretory pore on the level of Vgl- 2; gnathosomal rostrum short, less than depth of gnathosoma (Fig.
<figureCitation id="1138141192E276739FF7A637CAC22FCD" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Torrenticola soniae sp. nov., ♂ holotype A dorsal shield B ventral shield C palp, medial view (inset: ventrodistal projections of P- 2 and P- 3, enlarged 2 ×) D palp, lateral view E gnathosoma and chelicera F ejaculatory complex. Scale bars: 100 μm." figureDoi="10.3897/zookeys.1217.131730.figure1" httpUri="https://binary.pensoft.net/fig/1164865">1 E</figureCitation>
); P- 2 ventral margin nearly straight or slightly convex, P- 2 ventrodistal protrusion bluntly pointed, apically serrated, P- 3 with a subrectangular, apically serrated ventrodistal projection, P- 4 with a ventral tubercle bearing one long and three shorter setae (Fig.
<figureCitation id="F4F8C49C4687CF419B81C1489C92797D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Torrenticola soniae sp. nov., ♂ holotype A dorsal shield B ventral shield C palp, medial view (inset: ventrodistal projections of P- 2 and P- 3, enlarged 2 ×) D palp, lateral view E gnathosoma and chelicera F ejaculatory complex. Scale bars: 100 μm." figureDoi="10.3897/zookeys.1217.131730.figure1" httpUri="https://binary.pensoft.net/fig/1164865">1 C, D</figureCitation>
).
<emphasis id="E0A4F6A893E83BD6DE0FB35A446392D8" bold="true">Male.</emphasis>
Suture line of Cx-IV evident, medially starting from posterior margin of genital field in a right angle to the main idiosoma axis; genital field large, subrectangular; ejaculatory complex conventional in shape, anterior keel, proximal and distal arms well developed (Fig.
<figureCitation id="9DAEBEFDAEFCDDF30B5C300A71F047FC" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Torrenticola soniae sp. nov., ♂ holotype A dorsal shield B ventral shield C palp, medial view (inset: ventrodistal projections of P- 2 and P- 3, enlarged 2 ×) D palp, lateral view E gnathosoma and chelicera F ejaculatory complex. Scale bars: 100 μm." figureDoi="10.3897/zookeys.1217.131730.figure1" httpUri="https://binary.pensoft.net/fig/1164865">1 F</figureCitation>
).
<emphasis id="35A6AFB6D49F2A22BC70651E3DA7311D" bold="true">Female.</emphasis>
Genital field large and pentagonal in shape, suture lines of Cx-IV extending posteriorly beyond posterior margin of genital field, laterally curved.
</paragraph>
<caption id="B76289EC955DB8DF77DFC65516AF0A8F" ID-DOI="10.3897/zookeys.1217.131730.figure1" ID-arpha="6545011D-8DC9-5EDE-8E15-262718D388EB" httpUri="https://binary.pensoft.net/fig/1164865" startId="F1">
<paragraph id="4B13D5A930A28378302C3E7A82DE1BDC">
<label id="550EAAEB963BF799C85153A0B65EE21F">Figure 1.</label>
</paragraph>
<paragraph id="99D001811A83DE8E13C9514ABE0166E5">
<taxonomicName id="EF14E1909265D492E28013B452286D98" authorityName="Pešić &amp; Zawal &amp; Ferreira &amp; Benitez-Bosco &amp; Cruz-Oliveira &amp; Girão &amp; Padilha &amp; Turaccio &amp; Rossini &amp; Ballini &amp; Staffoni &amp; Fratini &amp; Ciofi &amp; Iannucci &amp; Ekrem &amp; Stur" authorityYear="2024" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" phylum="Arthropoda" rank="species" species="soniae" status="sp. nov.">
<emphasis id="A34EEFFCA1307AD83F0336095C72C553" italics="true">Torrenticola soniae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36F46545E123AAD380377DA6333EB5A0" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ holotype
<emphasis id="B72A7C1553A030759F85B6442F837E92" bold="true">A</emphasis>
dorsal shield
<emphasis id="E73E97CFF5705141102A3F609189A440" bold="true">B</emphasis>
ventral shield
<emphasis id="D568982FE6ADB9932881D1A51125416B" bold="true">C</emphasis>
palp, medial view (inset: ventrodistal projections of P- 2 and P- 3, enlarged 2 ×)
<emphasis id="F1A7F24C692A67AE08716E4D2F2D663F" bold="true">D</emphasis>
palp, lateral view
<emphasis id="1E41A5723779114881F50A65354384C7" bold="true">E</emphasis>
gnathosoma and chelicera
<emphasis id="6567C33F8071799787ABB65424FD1872" bold="true">F</emphasis>
ejaculatory complex. Scale bars: 100 μm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EQOGK" type="description">
<paragraph id="84BC6B131018963EB6558644C5702272">
<heading id="6BC0445E1F72B487760B823F03167ADD" reason="title">Measurements.</heading>
</paragraph>
<paragraph id="DDE74CAB03E8BB29DA160023CAE59D82">
<emphasis id="40CE40ED4C844863EAC3E66C48AA86E1" bold="true">Male</emphasis>
(
<typeStatus id="479F9A442600A3D447965B8B5665678D" type="holotype">holotype</typeStatus>
). Idiosoma (ventral view: Fig.
<figureCitation id="5BE084E2CCD1D86B88B8B4337A8775D4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Torrenticola soniae sp. nov., ♂ holotype A dorsal shield B ventral shield C palp, medial view (inset: ventrodistal projections of P- 2 and P- 3, enlarged 2 ×) D palp, lateral view E gnathosoma and chelicera F ejaculatory complex. Scale bars: 100 μm." figureDoi="10.3897/zookeys.1217.131730.figure1" httpUri="https://binary.pensoft.net/fig/1164865">1 B</figureCitation>
) L 912, W 669; dorsal shield (Fig.
<figureCitation id="35EA4684CD2ECD3A89A9C8C8152A641C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Torrenticola soniae sp. nov., ♂ holotype A dorsal shield B ventral shield C palp, medial view (inset: ventrodistal projections of P- 2 and P- 3, enlarged 2 ×) D palp, lateral view E gnathosoma and chelicera F ejaculatory complex. Scale bars: 100 μm." figureDoi="10.3897/zookeys.1217.131730.figure1" httpUri="https://binary.pensoft.net/fig/1164865">1 A</figureCitation>
) L 756, W 581, L / W ratio 1.3; dorsal plate L 700; shoulder plate L 220222, W 97, L / W ratio 2.272.29; frontal plate L 150156, W 7888, L / W ratio 1.81.9; shoulder / frontal plate L 1.421.47. Gnathosomal bay L 171,
<abbrev id="ABBRID0EAPGK" xlink_title="first coxae">Cx-I</abbrev>
total L 359,
<abbrev id="ABBRID0EEPGK" xlink_title="first coxae">Cx-I</abbrev>
<abbrev id="ABBRID0EIPGK" xlink_title="medial length">mL</abbrev>
188, Cx-II + III
<abbrev id="ABBRID0EMPGK" xlink_title="medial length">mL</abbrev>
137; ratio
<abbrev id="ABBRID0EQPGK" xlink_title="first coxae">Cx-I</abbrev>
L / Cx-II + III
<abbrev id="ABBRID0EUPGK" xlink_title="medial length">mL</abbrev>
2.63;
<abbrev id="ABBRID0EYPGK" xlink_title="first coxae">Cx-I</abbrev>
<abbrev id="ABBRID0E3PGK" xlink_title="medial length">mL</abbrev>
/ Cx-II + III
<abbrev id="ABBRID0EAQGK" xlink_title="medial length">mL</abbrev>
1.37. Genital field L / W 191 / 163, ratio 1.17; distance genital field-excretory pore 116, genital field-caudal idiosoma margin 209. Ejaculatory complex L 291.
</paragraph>
<paragraph id="C5EE22F3B2250ACA8434F6D6FC609FAF">
Gnathosoma vL 331, chelicera L 375; palp total L 398,
<abbrev id="ABBRID0EGQGK" xlink_title="dorsal length">dL</abbrev>
/ H,
<abbrev id="ABBRID0EKQGK" xlink_title="dorsal length">dL</abbrev>
/ H ratio:
<abbrev id="ABBRID0EOQGK" xlink_title="palp segments 15">P- 1</abbrev>
, 39 / 39, 1.0; P- 2, 117 / 73, 1.59; P- 3, 84 / 63, 1.35; P- 4, 120 / 42, 2.86;
<abbrev id="ABBRID0ESQGK" xlink_title="palp segments 15">P- 5</abbrev>
, 38 / 17, 2.18; L ratio P- 2 / P- 4, 0.98.
<abbrev id="ABBRID0EWQGK" xlink_title="dorsal length">dL</abbrev>
of
<abbrev id="ABBRID0E1QGK" xlink_title="fourth-sixth segments of first leg">I-L- 46</abbrev>
: 134, 150, 136; I-L- 6 H 100;
<abbrev id="ABBRID0E5QGK" xlink_title="dorsal length">dL</abbrev>
/ H I-L- 6 ratio 1.36.
</paragraph>
<paragraph id="96E18C16A35BDD87A240FD1C5AD593CE">
<emphasis id="E65F3E04847A5362175643EC5DF78B84" bold="true">Female</emphasis>
(
<typeStatus id="B354E0F691F4331E14173EEA2073F8E0" type="paratype">paratype</typeStatus>
from Casais de Folgosinho, BGE _ 00227 _ F 02). Idiosoma (ventral view: Fig.
<figureCitation id="DA1F5A10D1BE851754F7FB15801DC817" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Torrenticola soniae sp. nov., ♀ paratype A dorsal shield B photograph of dorsal shield C ventral shield D palp, medial view (P- 1 lacking). Scale bars: 100 μm." figureDoi="10.3897/zookeys.1217.131730.figure2" httpUri="https://binary.pensoft.net/fig/1164866">2 C</figureCitation>
) L 1033, W 828; dorsal shield (Fig.
<figureCitation id="6FDE030FDB3DE3DBB447BBD969B36586" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Torrenticola soniae sp. nov., ♀ paratype A dorsal shield B photograph of dorsal shield C ventral shield D palp, medial view (P- 1 lacking). Scale bars: 100 μm." figureDoi="10.3897/zookeys.1217.131730.figure2" httpUri="https://binary.pensoft.net/fig/1164866">2 A, B</figureCitation>
) L 844, W 725, L / W ratio 1.16; dorsal plate L 781; shoulder plate L 222235, W 100102, L / W ratio 2.22.3; frontal plate L 173175, W 9798, L / W ratio 1.79; shoulder / frontal plate L 1.281.34. Gnathosomal bay L 200,
<abbrev id="ABBRID0EORGK" xlink_title="first coxae">Cx-I</abbrev>
total L 384,
<abbrev id="ABBRID0ESRGK" xlink_title="first coxae">Cx-I</abbrev>
<abbrev id="ABBRID0EWRGK" xlink_title="medial length">mL</abbrev>
184, Cx-II + III
<abbrev id="ABBRID0E1RGK" xlink_title="medial length">mL</abbrev>
18; ratio
<abbrev id="ABBRID0E5RGK" xlink_title="first coxae">Cx-I</abbrev>
L / Cx-II + III
<abbrev id="ABBRID0ECSGK" xlink_title="medial length">mL</abbrev>
21.3;
<abbrev id="ABBRID0EGSGK" xlink_title="first coxae">Cx-I</abbrev>
<abbrev id="ABBRID0EKSGK" xlink_title="medial length">mL</abbrev>
/ Cx-II + III
<abbrev id="ABBRID0EOSGK" xlink_title="medial length">mL</abbrev>
10.2. Genital field L / W 221 / 198, ratio 1.12; distance genital field-excretory pore 250, genital field-caudal idiosoma margin 391.
</paragraph>
<caption id="E03C83EA79914D84867065D0461B2B0A" ID-DOI="10.3897/zookeys.1217.131730.figure2" ID-arpha="A3FEA330-9B52-50EB-888A-825DDB520F6B" httpUri="https://binary.pensoft.net/fig/1164866" startId="F2">
<paragraph id="9A055CFE98D5AFF0132652FCC8D9EC4F">
<label id="11897E80EA700324A67AAFC836702559">Figure 2.</label>
</paragraph>
<paragraph id="472D5B0A6A65640C97261CD3D4A27378">
<taxonomicName id="4330FAE121C68CC9665F52BB7084EE97" authorityName="Pešić &amp; Zawal &amp; Ferreira &amp; Benitez-Bosco &amp; Cruz-Oliveira &amp; Girão &amp; Padilha &amp; Turaccio &amp; Rossini &amp; Ballini &amp; Staffoni &amp; Fratini &amp; Ciofi &amp; Iannucci &amp; Ekrem &amp; Stur" authorityYear="2024" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" phylum="Arthropoda" rank="species" species="soniae" status="sp. nov.">
<emphasis id="3063E9D1DCE5270F0D0A8D2FC589C535" italics="true">Torrenticola soniae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="096E7AA756B7CBFD0D1959E8F01F02E1" rank="species">sp. nov.</taxonomicNameLabel>
, ♀ paratype
<emphasis id="E8C94B163DEE78BC17FD2D0D4AE9A970" bold="true">A</emphasis>
dorsal shield
<emphasis id="1E67BAE952FCFCD93A1B0BC0E5980BD8" bold="true">B</emphasis>
photograph of dorsal shield
<emphasis id="2001344A35C985D6EDF381D14BE9BB39" bold="true">C</emphasis>
ventral shield
<emphasis id="BDC380624D982EECBB5AA151FA217058" bold="true">D</emphasis>
palp, medial view (
<abbrev id="ABBRID0ENTGK" xlink_title="palp segments 15">P- 1</abbrev>
lacking). Scale bars: 100 μm.
</paragraph>
</caption>
<paragraph id="448FCDCBC41E42951A2B93F34CB97C4A">
Gnathosoma vL 350, chelicera L 409; palp total L 435,
<abbrev id="ABBRID0EYTGK" xlink_title="dorsal length">dL</abbrev>
/ H,
<abbrev id="ABBRID0E3TGK" xlink_title="dorsal length">dL</abbrev>
/ H ratio:
<abbrev id="ABBRID0EAUGK" xlink_title="palp segments 15">P- 1</abbrev>
, 44 / 41, 1.08; P- 2, 127 / 76, 1.67; P- 3, 93 / 65, 1.44; P- 4, 134 / 42, 3.18;
<abbrev id="ABBRID0EEUGK" xlink_title="palp segments 15">P- 5</abbrev>
, 37 / 18, 2.0; L ratio P- 2 / P- 4, 0.94.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIUGK" type="etymology">
<paragraph id="05D0ACE84E05C1E22DFBBC14A48D4515">
<heading id="67C833CD2B5B1FF6C1A4758B1CBCBC77" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="A04BCA78BF736C108CD649B0F794BE0A">
The species is dedicated to Sónia Ferreira (
<collectionCode id="E5EBE1D1519E3FBD1905B9CD90BEFCDE">CIBIO</collectionCode>
,
<collectingCountry id="FFA3267B1DC28B2A2E8A4DAD571F914F" name="Portugal">Portugal</collectingCountry>
) for collecting numerous specimens used in this study and her enthusiastic support in the study of Portuguese water mites.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENUGK" type="Species delimitation using DNA barcodes">
<paragraph id="CFD65A57705E1A7ECC8AE90044B0F1FB">
<heading id="72D98120481FA91A99268C3A50C13173" reason="title">Species delimitation using DNA barcodes.</heading>
</paragraph>
<paragraph id="BB0F61F505969E909227064B49DFFA0D">
The final alignment for species delimitation using
<abbrev id="ABBRID0ETUGK" xlink_title="mitochondrial cytochrome c oxidase subunit I">COI</abbrev>
sequence data comprised 669 nucleotide positions (nps) of the 130
<taxonomicName id="CB448F5A4FB3A4F53EE743DDB1A2A02C" authorityName="Piersig" authorityYear="1896" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" phylum="Arthropoda" rank="genus">
<emphasis id="4FF4BC7517C50231CFF47EBE74C37D81" italics="true">Torrenticola</emphasis>
</taxonomicName>
specimens listed in Suppl. material
<xref id="B95F6C308D3776983221C27228BA4653" ref-type="supplementary-material" rid="S2">2</xref>
and one outgroup,
<taxonomicName id="B73C90302A8D6176970A6C70B44CF0DE" baseAuthorityName="K. Viets" baseAuthorityYear="1930" class="Arachnida" family="Torrenticolidae" genus="Monatractides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" phylum="Arthropoda" rank="species" species="madritensis">
<emphasis id="E7A709DF27DEAAF9D4BBDDCC5B56F449" italics="true">Monatractides madritensis</emphasis>
</taxonomicName>
from
<collectingCountry id="E1BE50C364B2069F63E72C80C666765E" name="Portugal">Portugal</collectingCountry>
to root the tree. The
<abbrev id="ABBRID0ENVGK" xlink_title="Neighbor-Joining">NJ</abbrev>
tree is presented in Fig.
<figureCitation id="3FFE7D4A5CCAE185B336CCA9A069B8FB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Neighbor-Joining tree of the genus Torrenticola, obtained from 130 nucleotide COI sequences. Bootstrap values &gt; 50 % from 1000 bootstrap replicates on branches." figureDoi="10.3897/zookeys.1217.131730.figure6" httpUri="https://binary.pensoft.net/fig/1164870">6</figureCitation>
. The
<abbrev id="ABBRID0EVVGK" xlink_title="mitochondrial cytochrome c oxidase subunit I">COI</abbrev>
tree sequences retrieved from
<taxonomicName id="C4C4B01FDF9D752D227D1585EE1FA19B" authorityName="Piersig" authorityYear="1896" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" phylum="Arthropoda" rank="genus">
<emphasis id="3C687E6BB820FC0D6992C28A49DF2C35" italics="true">Torrenticola</emphasis>
</taxonomicName>
specimens from
<collectingCountry id="60ED228BB90CDE175C9A9B1A5FC1AF31" name="Portugal">Portugal</collectingCountry>
, here described as
<taxonomicName id="BAC5262B0F90E95C293A22D5289FC635" authorityName="Pešić &amp; Zawal &amp; Ferreira &amp; Benitez-Bosco &amp; Cruz-Oliveira &amp; Girão &amp; Padilha &amp; Turaccio &amp; Rossini &amp; Ballini &amp; Staffoni &amp; Fratini &amp; Ciofi &amp; Iannucci &amp; Ekrem &amp; Stur" authorityYear="2024" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" phylum="Arthropoda" rank="species" species="soniae" status="sp. nov.">
<emphasis id="BE57CB7BEB68A862598BAB1492EE3488" italics="true">T. soniae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7084A271F1B9165CA4675AE9D8F7AAA5" rank="species">sp. nov.</taxonomicNameLabel>
, appeared as a sister group to the cluster of sequences belonging to
<taxonomicName id="9320849D5F8D6D2D3B029C5228913829" authority="(Halbert, 1911)" baseAuthorityName="Halbert" baseAuthorityYear="1911" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="F79B09AC2E9FAA2B4C7C08D014BD01D5" italics="true">T. brevirostris</emphasis>
(Halbert, 1911)
</taxonomicName>
, a rhitrobiontic species widely distributed in Europe. The mean genetic distance between
<abbrev id="ABBRID0EWWGK" xlink_title="mitochondrial cytochrome c oxidase subunit I">COI</abbrev>
sequences of these two clusters was estimated at 12.27 ± 1.42 %
<abbrev id="ABBRID0E1WGK" xlink_title="Kimura 2-parameter model">
K
<specimenCount id="6B4247739E0563406BE6509E4046232E" count="2" type="generic">2 P.</specimenCount>
</abbrev>
The genetic distance was considerably higher than the estimated barcode gap found by
<collectionCode id="7C7D0F84D4D0C7AA98F7284802B73784">ASAP</collectionCode>
analyses (35 %) of all studied
<taxonomicName id="3A94675FCC4A0C56DC6484967AD322A5" authorityName="Piersig" authorityYear="1896" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" phylum="Arthropoda" rank="genus">
<emphasis id="150F3B773BB5B39F8BF8AA5951AC282D" italics="true">Torrenticola</emphasis>
</taxonomicName>
, supporting the species-status of the new taxon. The mean intraspecific
<abbrev id="ABBRID0EFXGK" xlink_title="Kimura 2-parameter model">K 2 P</abbrev>
- divergence within the cluster of the new species was 0.63 ± 0.19 %.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJXGK" type="discussion">
<paragraph id="8F3080248312EE3D45EA08217BF07631">
<heading id="D39F36BD6B711288CBEFDE0887A942FD" reason="title">Discussion.</heading>
</paragraph>
<paragraph id="549D9D3F2119A2179C1A3A30A835F92D">
With regards to the presence of an anteriorly broad and short
<abbrev id="ABBRID0EPXGK" xlink_title="first coxae">Cx-I</abbrev>
, a robust and compact palp, and a deep gnathosoma with a short rostrum, the new species resembles
<taxonomicName id="244BC44146147E864FBCB74218F26781" baseAuthorityName="Halbert" baseAuthorityYear="1911" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="DD6484E98E6241DF42D133035CFE3D38" italics="true">T. brevirostris</emphasis>
</taxonomicName>
. The latter species can be separated from
<taxonomicName id="F7570C1BE28E0A9A55590BB3CB1A0DC2" authorityName="Pešić &amp; Zawal &amp; Ferreira &amp; Benitez-Bosco &amp; Cruz-Oliveira &amp; Girão &amp; Padilha &amp; Turaccio &amp; Rossini &amp; Ballini &amp; Staffoni &amp; Fratini &amp; Ciofi &amp; Iannucci &amp; Ekrem &amp; Stur" authorityYear="2024" class="Arachnida" family="Torrenticolidae" genus="Torrenticola" kingdom="Animalia" order="Trombidiformes" phylum="Arthropoda" rank="species" species="soniae" status="sp. nov.">
<emphasis id="FAFA9662D7363BC8956CE82CE2D1C09B" italics="true">T. soniae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB78D725550D20B22B081FE5AB8A950D" rank="species">sp. nov.</taxonomicNameLabel>
by only slightly protruding ventrodistal projections of P- 2 and particularly of P- 3.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJYGK" type="distribution">
<paragraph id="1FA8C2B6EDF37E27D42039CBAB981E19">
<heading id="7E945ED0A2A666CB4C12763763CCD036" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="07F965C7865A64C020DE394777FDDD5B">
<collectingCountry id="35D548B1AE085213B4ADF81DEBBC7241" name="Portugal">Portugal</collectingCountry>
(this study).
</paragraph>
</subSubSection>
</treatment>
</document>