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<document id="CAB154A063D9E3A5C05EEE53E7639F00" ID-DOI="10.11646/zootaxa.3980.3.1" ID-GBIF-Dataset="87a07c27-c5f7-490d-9a25-5305095d22de" ID-ISSN="1175-5326" ID-Zenodo-Dep="254013" ID-ZooBank="674D11E9-904C-46E7-A689-013A0B708F3B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1461241738288" checkinUser="plazi" docAuthor="Kaila, Lauri" docDate="2015" docId="C5758781FFA6FFC1FF03BAC29085FF09" docLanguage="en" docName="zt03980p358.pdf" docOrigin="Zootaxa 3980 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Elachista anitella Traugott-Olsen 1985" docType="treatment" docVersion="12" lastPageNumber="323" masterDocId="394CFFF9FFB3FFD7FF94BF26936BFFF1" masterDocTitle="The Elachista dispunctella (Duponchel) complex (Lepidoptera, Elachistidae) revisited, with exceptional level of synonymy" masterLastPageNumber="358" masterPageNumber="301" pageNumber="322" updateTime="1698605267485" updateUser="plazi">
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<mods:title id="537773E2212C59CF748EC80059757786">The Elachista dispunctella (Duponchel) complex (Lepidoptera, Elachistidae) revisited, with exceptional level of synonymy</mods:title>
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<mods:namePart id="76413FA48F4E043EBAEF21F14A78815B">Kaila, Lauri</mods:namePart>
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<treatment id="C5758781FFA6FFC1FF03BAC29085FF09" ID-DOI="http://doi.org/10.5281/zenodo.6102547" ID-GBIF-Taxon="119633122" ID-Zenodo-Dep="6102547" LSID="urn:lsid:plazi:treatment:C5758781FFA6FFC1FF03BAC29085FF09" httpUri="http://treatment.plazi.org/id/C5758781FFA6FFC1FF03BAC29085FF09" lastPageId="22" lastPageNumber="323" pageId="21" pageNumber="322">
<subSubSection id="05C6651CFFA6FFC2FF03BAC29110FA0F" box="[151,635,1508,1534]" pageId="21" pageNumber="322" type="nomenclature">
<paragraph id="4D633697FFA6FFC2FF03BAC29110FA0F" blockId="21.[151,635,1508,1567]" box="[151,635,1508,1534]" pageId="21" pageNumber="322">
<heading id="162B81FBFFA6FFC2FF03BAC29110FA0F" bold="true" box="[151,635,1508,1534]" fontSize="11" level="1" pageId="21" pageNumber="322" reason="1">
<taxonomicName id="8ADC4D14FFA6FFC2FF03BAC29110FA0F" ID-CoL="B6WKQ" authority="Traugott-Olsen, 1985" authorityName="Traugott-Olsen" authorityYear="1985" box="[151,635,1508,1534]" class="Insecta" family="Elachistidae" genus="Elachista" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="anitella">
<emphasis id="7FA8EA85FFA6FFC2FF03BAC29110FA0F" bold="true" box="[151,635,1508,1534]" pageId="21" pageNumber="322">
<emphasis id="7FA8EA85FFA6FFC2FF03BAC29200FA0F" bold="true" box="[151,363,1508,1534]" italics="true" pageId="21" pageNumber="322">Elachista anitella</emphasis>
<bibRefCitation id="294D4B66FFA6FFC2FEE6BAC29110FA0F" author="Traugott-Olsen, E." box="[370,635,1508,1534]" firstAuthor="Traugott-Olsen" journalOrPublisher="SHILAP Revista de lepidopterologia" pageId="21" pageNumber="322" pagination="73 - 76" part="13" refId="ref37094" refString="Traugott-Olsen, E. (1985) Five new Elachista species from Sierra Nevada. SHILAP Revista de lepidopterologia, 13, 73 - 76." title="Five new Elachista species from Sierra Nevada" type="journal article" year="1985">Traugott-Olsen, 1985</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="05C6651CFFA6FFC2FF03B920922EF9EE" box="[151,325,1542,1567]" pageId="21" pageNumber="322" type="description">
<paragraph id="4D633697FFA6FFC2FF03B920922EF9EE" blockId="21.[151,635,1508,1567]" box="[151,325,1542,1567]" pageId="21" pageNumber="322">
<figureCitation id="D5E72A12FFA6FFC2FF03B9209398F9EE" box="[151,243,1542,1567]" captionStart="FIGURES 14 24" captionStartId="9.[151,264,1121,1143]" captionTargetBox="[196,1382,195,1104]" captionTargetId="figure@9.[196,1383,195,1104]" captionTargetPageId="9" captionText="FIGURES 14 24. External appearance of Elachista spp. males. 14. E. dalmatiensis Traugott-Olsen, holotype. 15. E. grandella Traugott-Olsen (Hungary). 16 19. E. parvula Parenti. 16. ♂ (Liguria, from type locality). 17. E. parvula Parenti ♀ (Greece). 18. E. minusculella Traugott-Olsen, holotype. 19. E. blancella Traugott-Olsen, holotype. 20. E. moroccoensis Traugott-Olsen, holotype. 21. E. anitella Traugott-Olsen, holotype. 22 24. Elachista maboulella, all males. 22. E. catalunella Traugott-Olsen, holotype. 23. E. gerdmaritella Traugott-Olsen, holotype. 24. E. gielisi Traugott-Olsen, holotype. Scale bar 2 mm." httpUri="https://zenodo.org/record/254016/files/figure.png" pageId="21" pageNumber="322">Figs. 21</figureCitation>
,
<figureCitation id="D5E72A12FFA6FFC2FF6BB921922EF9EE" box="[255,325,1543,1567]" captionStart="FIGURES 78 79" captionStartId="22.[151,264,1343,1365]" captionTargetBox="[317,1283,317,1321]" captionTargetId="figure@22.[300,1288,301,1322]" captionTargetPageId="22" captionText="FIGURES 78 79. Male genitalia of Elachista anitella Traugott-Olsen. 78. Holotype. 79. Spain (L. Kaila prep. 5354)." httpUri="https://zenodo.org/record/254030/files/figure.png" pageId="21" pageNumber="322">7879</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="05C6651CFFA6FFC2FF03B9689109F995" box="[151,610,1614,1636]" pageId="21" pageNumber="322" type="reference_group">
<paragraph id="4D633697FFA6FFC2FF03B9689109F995" blockId="21.[151,610,1614,1636]" box="[151,610,1614,1636]" pageId="21" pageNumber="322">
<treatmentCitationGroup id="6DCC11B9FFA6FFC2FF03B9689109F995" box="[151,610,1614,1636]" pageId="21" pageNumber="322">
<taxonomicName id="8ADC4D14FFA6FFC2FF03B9689134F995" ID-CoL="B6WKQ" authority="Traugott-Olsen, 1985a: 75" authorityName="Traugott-Olsen" authorityPageNumber="75" authorityYear="1985" box="[151,607,1614,1636]" class="Insecta" family="Elachistidae" genus="Elachista" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="anitella">
<emphasis id="7FA8EA85FFA6FFC2FF03B9689221F995" box="[151,330,1614,1636]" italics="true" pageId="21" pageNumber="322">Elachista anitella</emphasis>
<treatmentCitation id="CC7D1086FFA6FFC2FEC4B9689134F995" author="Traugott-Olsen" box="[336,607,1614,1636]" page="75" pageId="21" pageNumber="322" year="1985">
<bibRefCitation id="294D4B66FFA6FFC2FEC4B9689152F995" author="Traugott-Olsen, E." box="[336,569,1614,1636]" journalOrPublisher="SHILAP Revista de lepidopterologia" pageId="21" pageNumber="322" pagination="73 - 76" part="13" refId="ref37094" refString="Traugott-Olsen, E. (1985) Five new Elachista species from Sierra Nevada. SHILAP Revista de lepidopterologia, 13, 73 - 76." title="Five new Elachista species from Sierra Nevada" type="journal article" year="1985" yearSuffix="a">Traugott-Olsen, 1985a</bibRefCitation>
: 75
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="05C6651CFFA6FFC2FF03B9B7920FF873" pageId="21" pageNumber="322" type="materials_examined">
<paragraph id="4D633697FFA6FFC2FF03B9B7920FF873" blockId="21.[151,1437,1681,2030]" pageId="21" pageNumber="322">
<emphasis id="7FA8EA85FFA6FFC2FF03B9B79148F95B" bold="true" box="[151,547,1681,1706]" pageId="21" pageNumber="322">
Material studied.
<typeStatus id="92678835FFA6FFC2FEE6B9B792C6F95B" box="[370,429,1681,1706]" pageId="21" pageNumber="322">Type</typeStatus>
material.
</emphasis>
<materialsCitation id="FDB43CCAFFA6FFC2FDB9B9B79269F8E7" ID-GBIF-Occurrence="3941583376" collectingDate="1984-08-19" collectionCode="ZMUC" collectorName="E. Traugott-Olsen" country="Spain" elevation="2300" location="Sierra Nevada" municipality="Andalusia" pageId="21" pageNumber="322" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="92678835FFA6FFC2FDB9B9B791FEF95B" box="[557,661,1681,1706]" pageId="21" pageNumber="322" type="holotype">Holotype</typeStatus>
<specimenCount id="5BDAFD1EFFA6FFC2FD34B9B491DEF958" box="[672,693,1682,1705]" pageId="21" pageNumber="322" type="male"></specimenCount>
labelled: Type [rounded with red margin]; Genital praeparat nr. 5800 sex: ♂ E. Traugott-Olsen;
<collectingCountry id="35CB7607FFA6FFC2FD89B99391E8F93F" box="[541,643,1717,1742]" name="Spain" pageId="21" pageNumber="322">Hispania</collectingCountry>
,
<collectingMunicipality id="AD07ACEDFFA6FFC2FD01B9939062F93F" box="[661,777,1717,1742]" pageId="21" pageNumber="322">Andalusia</collectingMunicipality>
,
<location id="4803604CFFA6FFC2FC8EB99390A8F93F" LSID="urn:lsid:plazi:treatment:C5758781FFA6FFC1FF03BAC29085FF09:4803604CFFA6FFC2FC8EB99390A8F93F" box="[794,963,1717,1742]" country="Spain" municipality="Andalusia" name="Sierra Nevada" pageId="21" pageNumber="322">Sierra Nevada</location>
,
<location id="4803604CFFA6FFC2FC40B9909767F93F" LSID="urn:lsid:plazi:treatment:C5758781FFA6FFC1FF03BAC29085FF09:4803604CFFA6FFC2FC40B9909767F93F" box="[980,1036,1718,1742]" country="Spain" municipality="Andalusia" name="Cam" pageId="21" pageNumber="322">Cam</location>
, de
<location id="4803604CFFA6FFC2FBD1B99397E0F93F" LSID="urn:lsid:plazi:treatment:C5758781FFA6FFC1FF03BAC29085FF09:4803604CFFA6FFC2FBD1B99397E0F93F" box="[1093,1163,1717,1742]" country="Spain" municipality="Andalusia" name="Veleta" pageId="21" pageNumber="322">Veleta</location>
<quantity id="8A249B72FFA6FFC2FB0CB990979EF93C" box="[1176,1269,1718,1742]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="21" pageNumber="322" unit="m" value="2300.0">
<elevation id="C6F1D1A4FFA6FFC2FB0CB990979EF93C" box="[1176,1269,1718,1742]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="21" pageNumber="322" unit="m" value="2300.0">2300 m</elevation>
</quantity>
,
<date id="39621057FFA6FFC2FA93B990961CF93F" box="[1287,1399,1718,1742]" pageId="21" pageNumber="322" value="1984-08-19">
<collectingDate id="2926E9BFFFA6FFC2FA93B990961CF93F" box="[1287,1399,1718,1742]" pageId="21" pageNumber="322" value="1984-08-19">19.8.1984</collectingDate>
</date>
<collectorName id="E0295341FFA6FFC2FA10B990922AF903" pageId="21" pageNumber="322">E. Traugott-Olsen</collectorName>
leg.;
<taxonomicName id="8ADC4D14FFA6FFC2FE1BB9FF9137F900" box="[399,604,1753,1777]" class="Insecta" family="Elachistidae" genus="Elachista" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="anitella" status="sp. nov.">
<emphasis id="7FA8EA85FFA6FFC2FE1BB9FF9137F900" box="[399,604,1753,1777]" italics="true" pageId="21" pageNumber="322">Elachista anitella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="649B57FEFFA6FFC2FDFFB9FC91C4F900" box="[619,687,1754,1777]" pageId="21" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
det. E. Traugott-Olsen; DNA sample MM15484 Lepid. Phyl. (
<collectionCode id="2BCDAE52FFA6FFC2FF0BB9D8939DF8E7" box="[159,246,1790,1814]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="21" pageNumber="322">ZMUC</collectionCode>
).
</materialsCitation>
<emphasis id="7FA8EA85FFA6FFC2FE9EB9DB92ADF8E7" bold="true" box="[266,454,1789,1814]" pageId="21" pageNumber="322">Other material.</emphasis>
<materialsCitation id="FDB43CCAFFA6FFC2FE5BB9DB920FF873" ID-GBIF-Occurrence="3941583425" pageId="21" pageNumber="322">
<emphasis id="7FA8EA85FFA6FFC2FE5BB9DB9178F8E7" bold="true" box="[463,531,1789,1814]" pageId="21" pageNumber="322">
<collectingCountry id="35CB7607FFA6FFC2FE5BB9DB9178F8E7" box="[463,531,1789,1814]" name="Spain" pageId="21" pageNumber="322">Spain</collectingCountry>
</emphasis>
:
<collectingCountry id="35CB7607FFA6FFC2FDB0B9DB91EDF8E7" box="[548,646,1789,1814]" name="Grenada" pageId="21" pageNumber="322">Granada</collectingCountry>
, Sierra Nevada,
<quantity id="8A249B72FFA6FFC2FCD0B9D890E6F8E7" box="[836,909,1789,1814]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="322" unit="km" value="20.0">20 km</quantity>
SE.
<collectingCountry id="35CB7607FFA6FFC2FC52B9DB9743F8E7" box="[966,1064,1789,1814]" name="Grenada" pageId="21" pageNumber="322">Granada</collectingCountry>
, 27.
<date id="39621057FFA6FFC2FBC1B9D897B0F8E7" box="[1109,1243,1790,1814]" pageId="21" pageNumber="322" value="2003-07-01">
VII.2003,
<specimenCount id="5BDAFD1EFFA6FFC2FB59B9D897B0F8E7" box="[1229,1243,1790,1814]" pageId="21" pageNumber="322" type="generic">1</specimenCount>
</date>
♂, T. Nupponen leg., L. Kaila prep. 5354,
<collectionCode id="2BCDAE52FFA6FFC2FE28B8049292F8CB" LSID="urn:lsid:biocol.org:col:15536" box="[444,505,1826,1850]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="21" pageNumber="322">DNA</collectionCode>
sample 11423 Lepid. Phyl. (Coll. Nupponen);
<collectingCountry id="35CB7607FFA6FFC2FB9AB807971BF8CB" box="[1038,1136,1825,1850]" name="Grenada" pageId="21" pageNumber="322">Granada</collectingCountry>
, Sierra Nevada,
<quantity id="8A249B72FFA6FFC2FAB8B804961FF8CB" box="[1324,1396,1825,1850]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="322" unit="km" value="20.0">20 km</quantity>
SE
<collectingCountry id="35CB7607FFA6FFC2FF03B8639392F8AF" box="[151,249,1861,1886]" name="Grenada" pageId="21" pageNumber="322">Granada</collectingCountry>
,
<quantity id="8A249B72FFA6FFC2FE97B86092CBF8AC" box="[259,416,1862,1886]" metricMagnitude="3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="21" pageNumber="322" unit="m" value="2800.0" valueMax="2900.0" valueMin="2700.0">27002900 m</quantity>
, 11.
<date id="39621057FFA6FFC2FE5CB8609125F8AF" box="[456,590,1862,1886]" pageId="21" pageNumber="322" value="2010-07-01">
VII.2010,
<specimenCount id="5BDAFD1EFFA6FFC2FDD4B8609125F8AF" box="[576,590,1862,1886]" pageId="21" pageNumber="322" type="generic">1</specimenCount>
</date>
♂, T. Nupponen leg., L. Kaila prep. 5355,
<collectionCode id="2BCDAE52FFA6FFC2FBB9B8609701F8AF" LSID="urn:lsid:biocol.org:col:15536" box="[1069,1130,1862,1886]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="21" pageNumber="322">DNA</collectionCode>
sample 15484 Lepid. Phyl. (Coll. Nupponen).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="05C6651CFFA6FFC2FF53B8AB907AF81F" pageId="21" pageNumber="322" type="diagnosis">
<paragraph id="4D633697FFA6FFC2FF53B8AB907AF81F" blockId="21.[151,1437,1681,2030]" pageId="21" pageNumber="322">
<emphasis id="7FA8EA85FFA6FFC2FF53B8AB9251F857" bold="true" box="[199,314,1933,1958]" pageId="21" pageNumber="322">Diagnosis</emphasis>
.
<taxonomicName id="8ADC4D14FFA6FFC2FED8B8A892AEF854" box="[332,453,1933,1957]" class="Insecta" family="Elachistidae" genus="Elachista" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="anitella">
<emphasis id="7FA8EA85FFA6FFC2FED8B8A892AEF854" box="[332,453,1933,1957]" italics="true" pageId="21" pageNumber="322">E. anitella</emphasis>
</taxonomicName>
is a small species. It closely resembles
<taxonomicName id="8ADC4D14FFA6FFC2FC37B8A89775F854" box="[931,1054,1933,1957]" class="Insecta" family="Elachistidae" genus="Elachista" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="7FA8EA85FFA6FFC2FC37B8A89775F854" box="[931,1054,1933,1957]" italics="true" pageId="21" pageNumber="322">E. parvula</emphasis>
</taxonomicName>
, from which the shorter digitate process of
<taxonomicName id="8ADC4D14FFA6FFC2FE86B89492E3F838" box="[274,392,1969,1993]" class="Insecta" family="Elachistidae" genus="Elachista" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="anitella">
<emphasis id="7FA8EA85FFA6FFC2FE86B89492E3F838" box="[274,392,1969,1993]" italics="true" pageId="21" pageNumber="322">E. anitella</emphasis>
</taxonomicName>
readily separates it. Its phallus is shorter than the valva, narrow, and hardly tapered towards apex. Its uncus lobes are rounded, as long as their width.
</paragraph>
</subSubSection>
<subSubSection id="05C6651CFFA5FFC1FF53BFB192C8FF41" box="[199,419,151,176]" pageId="22" pageNumber="323" type="biology_ecology">
<paragraph id="4D633697FFA5FFC1FF53BFB192C8FF41" blockId="22.[199,1006,151,248]" box="[199,419,151,176]" pageId="22" pageNumber="323">
<emphasis id="7FA8EA85FFA5FFC1FF53BFB19274FF41" bold="true" box="[199,287,151,176]" pageId="22" pageNumber="323">Biology</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="05C6651CFFA5FFC1FF53BF9A92C5FF24" box="[199,430,188,213]" pageId="22" pageNumber="323" type="distribution">
<paragraph id="4D633697FFA5FFC1FF53BF9A92C5FF24" blockId="22.[199,1006,151,248]" box="[199,430,188,213]" pageId="22" pageNumber="323">
<emphasis id="7FA8EA85FFA5FFC1FF53BF9A9232FF24" bold="true" box="[199,345,188,213]" pageId="22" pageNumber="323">Distribution</emphasis>
.
<collectingCountry id="35CB7607FFA5FFC1FEF3BF9A92C1FF24" box="[359,426,188,213]" name="Spain" pageId="22" pageNumber="323">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="05C6651CFFA5FFC1FF53BFF99085FF09" box="[199,1006,223,248]" pageId="22" pageNumber="323" type="discussion">
<paragraph id="4D633697FFA5FFC1FF53BFF99085FF09" blockId="22.[199,1006,151,248]" box="[199,1006,223,248]" pageId="22" pageNumber="323">
<emphasis id="7FA8EA85FFA5FFC1FF53BFF9925FFF09" bold="true" box="[199,308,223,248]" pageId="22" pageNumber="323">Remarks</emphasis>
.
<taxonomicName id="8ADC4D14FFA5FFC1FED6BFC792DCFF09" box="[322,439,224,248]" class="Insecta" family="Elachistidae" genus="Elachista" kingdom="Animalia" order="Lepidoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="anitella">
<emphasis id="7FA8EA85FFA5FFC1FED6BFC792DCFF09" box="[322,439,224,248]" italics="true" pageId="22" pageNumber="323">E. anitella</emphasis>
</taxonomicName>
is referred to as OTU
<quantity id="8A249B72FFA5FFC1FD26BFC69180FF09" box="[690,747,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="22" pageNumber="323" unit="in" value="17.0">17 in</quantity>
<bibRefCitation id="294D4B66FFA5FFC1FD66BFC69082FF09" author="Mutanen, M. &amp; Kekkonen, M. &amp; Prosser, S. W. J. &amp; Hebert, P. D. N. &amp; Kaila, L." box="[754,1001,223,248]" etAl="et al." firstAuthor="Mutanen" journalOrPublisher="Molecular Ecology Resources" pageId="22" pageNumber="323" pagination="967 - 984" part="15" refId="ref36215" refString="Mutanen, M., Kekkonen, M., Prosser, S. W. J., Hebert, P. D. N. &amp; Kaila, L. (2015) One species in eight: DNA barcodes from type specimens resolve a taxonomic quagmire. Molecular Ecology Resources, 15, 967 - 984. http: // dx. doi. org / 10.1111 / 1755 - 0998.12361" title="One species in eight: DNA barcodes from type specimens resolve a taxonomic quagmire" type="journal article" year="2015">
Mutanen
<emphasis id="7FA8EA85FFA5FFC1FCC8BFC790E5FF09" box="[860,910,224,248]" italics="true" pageId="22" pageNumber="323">et al</emphasis>
. (2015)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>